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<document id="CCE2A3CD76C3BB9F9DC8452A23542AA5" ID="10.11646/zootaxa.4281.1.24SLASH11460" ID-CLB-Dataset="28710" ID-DOI="10.11646/zootaxa.4281.1.24" ID-GBIF-Dataset="c164a746-877a-4fcd-a3b7-4134bc96a606" ID-ISSN="1175-5326" ID-Zenodo-Dep="816095" ID-ZooBank="9ABC0A87-E43E-4DE6-83ED-FF3B89D21412" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1498128355953" checkinUser="plazi" docAuthor="Paladini, Andressa &amp; Cavichioli, Rodney Ramiro" docDate="2017" docId="038F87FFD13E8431D58B54B239B04195" docLanguage="en" docName="zootaxa.4281.1.24SLASH11460.pdf" docOrigin="Zootaxa 4281 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tiodus sakakibarai Paladini &amp; Cavichioli 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="244" masterDocId="FFB6FF87D13F8433D51C57123B004655" masterDocTitle="A new species of Tiodus (Hemiptera, Cercopidae) with a key to species of the genus" masterLastPageNumber="245" masterPageNumber="242" pageNumber="243" updateTime="1698448347529" updateUser="ExternalLinkService">
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<mods:title id="998B6EDFDAAA57E3BC9DDE6B1E9108F7">A new species of Tiodus (Hemiptera, Cercopidae) with a key to species of the genus</mods:title>
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<mods:namePart id="483C9F8E855159A55D3C52D8A6B85C5C">Paladini, Andressa</mods:namePart>
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<mods:namePart id="53C04AB8E33CBA8013E3254FE9C94D0A">Cavichioli, Rodney Ramiro</mods:namePart>
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<mods:title id="B9D42D9B0F2236B4A379319E3C5E93C8">Zootaxa</mods:title>
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<mods:date id="1312ECE9284381573EFA8D11DA4ADB84">2017</mods:date>
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<mods:identifier id="E195FD8BA03D3F39235330A822A75708" type="DOI">10.11646/zootaxa.4281.1.24</mods:identifier>
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<treatment id="038F87FFD13E8431D58B54B239B04195" ID-DOI="http://doi.org/10.5281/zenodo.6052437" ID-GBIF-Taxon="150837798" ID-Zenodo-Dep="6052437" LSID="urn:lsid:plazi:treatment:038F87FFD13E8431D58B54B239B04195" httpUri="http://treatment.plazi.org/id/038F87FFD13E8431D58B54B239B04195" lastPageId="2" lastPageNumber="244" pageId="1" pageNumber="243">
<subSubSection id="C33C6562D13E8432D58B54B23A2E458E" pageId="1" pageNumber="243" type="nomenclature">
<paragraph id="8B9936E9D13E8432D58B54B239F845EF" blockId="1.[151,760,928,987]" box="[151,760,928,954]" pageId="1" pageNumber="243">
<heading id="D0D18185D13E8432D58B54B239F845EF" bold="true" box="[151,760,928,954]" fontSize="11" level="1" pageId="1" pageNumber="243" reason="1">
<emphasis id="B952EAFBD13E8432D58B54B239F845EF" bold="true" box="[151,760,928,954]" pageId="1" pageNumber="243">
<taxonomicName id="4C264D6AD13E8432D58B54B2399345EF" ID-CoL="7CCWJ" authority="Paladini &amp; Cavichioli" authorityName="Paladini &amp; Cavichioli" authorityYear="2017" box="[151,659,928,954]" class="Insecta" family="Cercopidae" genus="Tiodus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="sakakibarai" status="sp. nov.">
<emphasis id="B952EAFBD13E8432D58B54B23A7E45EF" bold="true" box="[151,382,928,954]" italics="true" pageId="1" pageNumber="243">Tiodus sakakibarai</emphasis>
Paladini &amp; Cavichioli
</taxonomicName>
<taxonomicNameLabel id="A2615780D13E8432D78654B239F845EF" box="[666,760,928,954]" pageId="1" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936E9D13E8432D58B54D03A2E458E" blockId="1.[151,760,928,987]" box="[151,302,962,987]" pageId="1" pageNumber="243">
(
<figureCitation id="131D2A6CD13E8432D58254D03A25458E" box="[158,293,962,987]" captionStart="FIGURE 1 8" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[159,1407,278,1703]" captionTargetId="figure@2.[150,1436,265,1703]" captionTargetPageId="2" captionText="FIGURE 1 8. Tiodus sakakibarai n. sp. 1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="1" pageNumber="243">Figures 18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33C6562D13E8432D58B5318398E4213" pageId="1" pageNumber="243" type="description">
<paragraph id="8B9936E9D13E8432D58B5318398E4213" blockId="1.[151,1436,1034,2031]" pageId="1" pageNumber="243">
<emphasis id="B952EAFBD13E8432D58B53183A954276" bold="true" box="[151,405,1034,1059]" pageId="1" pageNumber="243">Measurements (mm).</emphasis>
Total length,
<quantity id="4CDE9B0CD13E8432D72E531839984277" box="[562,664,1034,1059]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="243" unit="mm" value="11.6">11.6 mm</quantity>
; length of pronotum,
<quantity id="4CDE9B0CD13E8432D690531938E54277" box="[908,997,1035,1059]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="243" unit="mm" value="2.3">2.3 mm</quantity>
; width of pronotum, 4.0 mm; length of tegmina,
<quantity id="4CDE9B0CD13E8432D5E3533D3A584213" box="[255,344,1071,1095]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="1" pageNumber="243" unit="mm" value="8.3">8.3 mm</quantity>
; width of tegmina,
<quantity id="4CDE9B0CD13E8432D72D533D398A4213" box="[561,650,1071,1095]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="243" unit="mm" value="2.2">2.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C6562D13E8432D5DB53403F0142DA" pageId="1" pageNumber="243" type="diagnosis">
<paragraph id="8B9936E9D13E8432D5DB53403F0142DA" blockId="1.[151,1436,1034,2031]" pageId="1" pageNumber="243">
<emphasis id="B952EAFBD13E8432D5DB53403A41423E" bold="true" box="[199,321,1106,1131]" pageId="1" pageNumber="243">Diagnosis.</emphasis>
Postclypeus orange, moderately inflated, slightly angulated in profile; tegmina black with two reddish orange transverse transcommissural bands; aedeagus robust, not bifid.
</paragraph>
</subSubSection>
<subSubSection id="C33C6562D13E8432D5DB53883AA241D6" pageId="1" pageNumber="243" type="description">
<paragraph id="8B9936E9D13E8432D5DB53883FB240D2" blockId="1.[151,1436,1034,2031]" pageId="1" pageNumber="243">
<emphasis id="B952EAFBD13E8432D5DB53883A5942E6" bold="true" box="[199,345,1178,1203]" pageId="1" pageNumber="243">Description.</emphasis>
Head: black (
<figureCitation id="131D2A6CD13E8432D71B5388397242E6" box="[519,626,1178,1203]" captionStart="FIGURE 1 8" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[159,1407,278,1703]" captionTargetId="figure@2.[150,1436,265,1703]" captionTargetPageId="2" captionText="FIGURE 1 8. Tiodus sakakibarai n. sp. 1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="1" pageNumber="243">Figs 12</figureCitation>
); compound eyes brown, rounded and arranged transversely; vertex smooth and rectangular, lacking median carina; ocelli brown, closer to each other than to compound eyes; tylus smooth and rectangular, lacking median carina; antennae black with scape orange, pedicel setose, basal body of flagellum subcylindrical with one arista as long as flagellum; postclypeus orange, inflated, slightly angulated (forming an obtuse angle) in profile, with one longitudinal carina strongly marked and produced; rostrum extending to mesocoxae. Thorax: reddish orange with basisternum black; pronotum black with anterolateral margins reddish orange, hexagonal, lacking median carina, anterior margin straight, anterolateral margins straight, posterolateral margins slightly sinuous, posterior margin grooved; scutellum black with a light central concavity, lacking horizontal grooves; tegmina black, each with two reddish orange transverse transcommissural bands, one between basal and median third and another on apical third, apical reticulation well developed; hindwings hyaline with orange brown venation, vein Cu1 not thickened at base; legs orange with tibiae and tarsi black; metathoracic tibiae each with two lateral spines (basal spine equal in size to spines of apical crown; apical spine larger than spines of apical crown); apical crown of spines on tibia consisting of two rows; basitarsi each with one row of spines covered by long setae; subungueal process absent. Abdomen: pale to brownish-orange.
</paragraph>
<paragraph id="8B9936E9D13E8432D5DB51803FB1410A" blockId="1.[151,1436,1034,2031]" pageId="1" pageNumber="243">
<emphasis id="B952EAFBD13E8432D5DB51803A7A40FE" bold="true" box="[199,378,1682,1707]" pageId="1" pageNumber="243">Male genitalia.</emphasis>
Genital chamber with finger-like process on posterior margin of pygofer (
<figureCitation id="131D2A6CD13E8432D1D751803E1140FE" box="[1227,1297,1682,1707]" captionStart="FIGURE 1 8" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[159,1407,278,1703]" captionTargetId="figure@2.[150,1436,265,1703]" captionTargetPageId="2" captionText="FIGURE 1 8. Tiodus sakakibarai n. sp. 1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="1" pageNumber="243">Fig. 3</figureCitation>
); subgenital plates long and wide with rounded apex, free for almost entire length (
<figureCitation id="131D2A6CD13E8432D6D851A43F0B409A" box="[964,1035,1718,1743]" captionStart="FIGURE 1 8" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[159,1407,278,1703]" captionTargetId="figure@2.[150,1436,265,1703]" captionTargetPageId="2" captionText="FIGURE 1 8. Tiodus sakakibarai n. sp. 1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="1" pageNumber="243">Fig. 4</figureCitation>
); parameres long and slender, with finger-like apex and rounded process at apical third, without spine or spine-like processes, dorsal margin rounded (
<figureCitation id="131D2A6CD13E8432D58351EC3A054142" box="[159,261,1790,1815]" captionStart="FIGURE 1 8" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[159,1407,278,1703]" captionTargetId="figure@2.[150,1436,265,1703]" captionTargetPageId="2" captionText="FIGURE 1 8. Tiodus sakakibarai n. sp. 1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="1" pageNumber="243">Figs 56</figureCitation>
); aedeagus robust with shaft straight, slightly enlarged at median third, shaft with processes: single dorsal process located on the beginning of apical third; bifid process on apex of shaft, finger-like in lateral view and Cshaped in dorsal view; and two flap-like lateral processes located on median third. (
<figureCitation id="131D2A6CD13E8432D12250543FA5410A" box="[1086,1189,1862,1887]" captionStart="FIGURE 1 8" captionStartId="2.[151,250,1724,1746]" captionTargetBox="[159,1407,278,1703]" captionTargetId="figure@2.[150,1436,265,1703]" captionTargetPageId="2" captionText="FIGURE 1 8. Tiodus sakakibarai n. sp. 1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="1" pageNumber="243">Figs 78</figureCitation>
).
</paragraph>
<paragraph id="8B9936E9D13E8432D5DB50783AA241D6" blockId="1.[151,1436,1034,2031]" box="[199,418,1898,1923]" pageId="1" pageNumber="243">
<emphasis id="B952EAFBD13E8432D5DB50783A2541D6" bold="true" box="[199,293,1898,1923]" pageId="1" pageNumber="243">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33C6562D13E8431D5DB509C3A314681" lastPageId="2" lastPageNumber="244" pageId="1" pageNumber="243" type="discussion">
<paragraph id="8B9936E9D13E8431D5DB509C3A314681" blockId="1.[151,1436,1034,2031]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="244" pageId="1" pageNumber="243">
<emphasis id="B952EAFBD13E8432D5DB509C3A3B41F2" bold="true" box="[199,315,1934,1959]" pageId="1" pageNumber="243">Remarks.</emphasis>
The color pattern of this species is unique in the genus. The male genitalia differ from the other species by the absence of a paramere spine and the entire (not bifid) aedeagus shaft. The postclypeus shape and angle; the general shape of the aedeagus, principally the single process on the apical third and the two processes on the apex; and the elongated parameres with a thinner apex are characters that justify placement of this species in the genus
<taxonomicName id="4C264D6AD13D8431D5C357AE3A2A4680" authorityName="Nast" authorityYear="1951" box="[223,298,188,213]" class="Insecta" family="Cercopidae" genus="Tiodus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B952EAFBD13D8431D5C357AE3A2A4680" box="[223,298,188,213]" italics="true" pageId="2" pageNumber="244">Tiodus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF596661D13D8431D58B51AE391D415A" httpUri="https://zenodo.org/record/816097/files/figure.png" pageId="2" pageNumber="244" targetBox="[159,1407,278,1703]" targetPageId="2">
<paragraph id="8B9936E9D13D8431D58B51AE391D415A" blockId="2.[151,1436,1724,1807]" pageId="2" pageNumber="244">
<emphasis id="B952EAFBD13D8431D58B51AE3A2E4084" bold="true" box="[151,302,1724,1746]" pageId="2" pageNumber="244">FIGURE 18.</emphasis>
<taxonomicName id="4C264D6AD13D8431D42A51AE3AF94087" authority="Paladini &amp; Cavichioli, 2017" authorityName="Paladini &amp; Cavichioli" authorityYear="2017" box="[310,505,1724,1746]" class="Insecta" family="Cercopidae" genus="Tiodus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="244" phylum="Arthropoda" rank="species" species="sakakibarai" status="sp. nov.">
<emphasis id="B952EAFBD13D8431D42A51AE3AF94087" box="[310,505,1724,1746]" italics="true" pageId="2" pageNumber="244">Tiodus sakakibarai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2615780D13D8431D71D51AE393C4084" box="[513,572,1724,1745]" pageId="2" pageNumber="244" rank="species">
<emphasis id="B952EAFBD13D8431D71D51AE393C4084" bold="true" box="[513,572,1724,1745]" pageId="2" pageNumber="244">n. sp.</emphasis>
</taxonomicNameLabel>
1. Holotype habitus, dorsal view. 2. Holotype habitus, lateral view. 3. Male genital chamber, lateral view. 4. Subgenital plates, ventral view. 5. Paramere, lateral view. 6. Paramere, dorsal view. 7. Aedeagus, lateral view. 8. Aedeagus, dorsal view.
</paragraph>
</caption>
<subSubSection id="C33C6562D13D8431D5DB502E395A412C" pageId="2" pageNumber="244" type="etymology">
<paragraph id="8B9936E9D13D8431D5DB502E395A412C" blockId="2.[151,1436,1852,1985]" pageId="2" pageNumber="244">
<emphasis id="B952EAFBD13D8431D5DB502E3A4E4100" bold="true" box="[199,334,1852,1877]" pageId="2" pageNumber="244">Etymology.</emphasis>
The species is named in honor of Dr. Albino Morimasa Sakakibara for his great contributions to the knowledge of the Auchenorrhyncha.
</paragraph>
</subSubSection>
<subSubSection id="C33C6562D13D8431D5DB509639B04195" pageId="2" pageNumber="244" type="materials_examined">
<paragraph id="8B9936E9D13D8431D5DB509639B04195" blockId="2.[151,1436,1852,1985]" pageId="2" pageNumber="244">
<materialsCitation id="3B4E3CB4D13D8431D5DB509639B04195" ID-GBIF-Occurrence="1960124709" collectingDate="2012-08-21" collectionCode="MUSM" collectorName="Km SW &amp; Cavichioli &amp; Melo &amp; Rosa &amp; Santos" country="Peru" elevation="1221" latitude="-13.612" location="Examined" longLatPrecision="76" longitude="-70.446" pageId="2" pageNumber="244" specimenCount="1" stateProvince="Puno" typeStatus="holotype">
<emphasis id="B952EAFBD13D8431D5DB50963AAC41C8" bold="true" box="[199,428,1924,1949]" pageId="2" pageNumber="244">
<location id="8EF96032D13D8431D5DB50963A4041C8" LSID="urn:lsid:plazi:treatment:038F87FFD13E8431D58B54B239B04195:8EF96032D13D8431D5DB50963A4041C8" box="[199,320,1924,1949]" country="Peru" latitude="-13.612" longLatPrecision="76" longitude="-70.446" name="Examined" pageId="2" pageNumber="244" stateProvince="Puno">Examined</location>
material
</emphasis>
:
<typeStatus id="549D884BD13D8431D4A65096392741C8" box="[442,551,1924,1949]" pageId="2" pageNumber="244" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3317679D13D8431D72D5097396741C9" box="[561,615,1925,1948]" name="Peru" pageId="2" pageNumber="244">Peru</collectingCountry>
,
<collectingRegion id="49E2F80BD13D8431D76E509739AB41C9" box="[626,683,1925,1948]" country="Peru" name="Puno" pageId="2" pageNumber="244">Puno</collectingRegion>
20
<collectorName id="26D3533FD13D8431D7D25097382941C8" box="[718,809,1925,1949]" pageId="2" pageNumber="244">Km SW</collectorName>
de San Gaban
<geoCoordinate id="EE12502ED13D8431D6CD50963F2F41C8" box="[977,1071,1924,1949]" direction="south" orientation="latitude" pageId="2" pageNumber="244" precision="55" value="-13.612">13.612S</geoCoordinate>
<geoCoordinate id="EE12502ED13D8431D12950963F9D41C8" box="[1077,1181,1924,1949]" direction="west" orientation="longitude" pageId="2" pageNumber="244" precision="55" value="-70.446">70.446W</geoCoordinate>
,
<elevation id="000BD1DAD13D8431D1B450973FF841C8" box="[1192,1272,1925,1949]" metricMagnitude="3" metricUnit="m" metricValue="1.2209999999999999" pageId="2" pageNumber="244" unit="m" value="1221.0">
<quantity id="4CDE9B0CD13D8431D1B450973FF841C8" box="[1192,1272,1925,1949]" metricMagnitude="3" metricUnit="m" metricValue="1.2209999999999999" pageId="2" pageNumber="244" unit="m" value="1221.0">1221m</quantity>
</elevation>
,
<collectingDate id="EFDCE9C1D13D8431D01F50973E9841C8" box="[1283,1432,1925,1949]" pageId="2" pageNumber="244" value="2012-08-21">
<date id="FF981029D13D8431D01F50973E9841C8" box="[1283,1432,1925,1949]" pageId="2" pageNumber="244" value="2012-08-21">21.VIII.2012</date>
</collectingDate>
,
<collectorName id="26D3533FD13D8431D58B50BA3A0F4194" box="[151,271,1960,1985]" pageId="2" pageNumber="244">Cavichioli</collectorName>
,
<collectorName id="26D3533FD13D8431D40650BA3A584194" box="[282,344,1960,1985]" pageId="2" pageNumber="244">Melo</collectorName>
,
<collectorName id="26D3533FD13D8431D47F50BB3A9B4195" box="[355,411,1961,1984]" pageId="2" pageNumber="244">Rosa</collectorName>
&amp;
<collectorName id="26D3533FD13D8431D4A250BA39094195" box="[446,521,1960,1984]" pageId="2" pageNumber="244">Santos</collectorName>
leg. [
<collectionCode id="ED37AE2CD13D8431D75B50BA39A54195" box="[583,677,1960,1984]" pageId="2" pageNumber="244">MUSM</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>