331 lines
61 KiB
XML
331 lines
61 KiB
XML
<document id="9F40BE6CFA942F3C5B6F9CAA54A6976C" ID-DOI="10.37520/aemnp.2023.008" ID-ISSN="1804-6487" ID-Zenodo-Dep="8005824" ID-ZooBank="E1CBAEEC-2D68-4526-8DF4-11BF6DA66B2A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685971164724" checkinUser="felipe" docAuthor="Zhang, Wen-Xuan & Yin, Zi-Wei" docDate="2023" docId="038F87F22C1AFFE66100531F6DD65C20" docLanguage="en" docName="ActaEntMusNatPra.63.1.165-175.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2023.008" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Feabatrus Zhang & Yin 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="168" masterDocId="FFB6FF8A2C1BFFE2605A57726C4C5D75" masterDocTitle="Feabatrus gen. nov., a conspicuous new genus of Batrisitae from Myanmar and China (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="175" masterPageNumber="165" pageNumber="166" updateTime="1685971580993" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="01DF70AC2E8874592769CE6752EFB5AD">Feabatrus gen. nov., a conspicuous new genus of Batrisitae from Myanmar and China (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="DDC4203B41B8CBE38373EC1E07860AED" type="personal">
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<mods:namePart id="8E62C96B34D19ED5835E5EEA8345D015">Zhang, Wen-Xuan</mods:namePart>
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<mods:affiliation id="A1A357E19484CD3C743B275DB9D5BB15">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, Shanghai 200234, China</mods:affiliation>
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<mods:roleTerm id="8CE67973DD2772C5CAA5476A048B5DA8">Author</mods:roleTerm>
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<mods:namePart id="F7D985BF0C9E5F5586E215A5A6BAC37D">Yin, Zi-Wei</mods:namePart>
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<mods:affiliation id="F873E73435E05F5772D606B34FE9CF97">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, Shanghai 200234, China & corresponding author, e-mail: pselaphinae @ gmail. com</mods:affiliation>
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<mods:typeOfResource id="D92CDDBC9556D58978A5734F347E6600">text</mods:typeOfResource>
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<mods:title id="691624757A1C3C88C2BAF052D98BFC9E">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="9023DF3E1E256E7CF7BE9CB4AD3F8E73">
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<mods:date id="30CF0174ACA03351CAA38E3E24777B5A">2023</mods:date>
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<mods:title id="084850DD3407087BC12339F1DCEC272A">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="9864689DEDA59D5874CB97B31AB4484A">2023-06-04</mods:number>
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<mods:number id="ACAFC2AFB694338D307713088F94CB16">63</mods:number>
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<mods:url id="082FC7C828C0B19E69A47729A5E386AB">http://dx.doi.org/10.37520/aemnp.2023.008</mods:url>
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<mods:classification id="F365F9EF45C3443872FB45109433F506">journal article</mods:classification>
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<mods:identifier id="AD2C34D7BBDFF0CE8F6DC85256090575" type="DOI">10.37520/aemnp.2023.008</mods:identifier>
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<treatment id="038F87F22C1AFFE66100531F6DD65C20" LSID="urn:lsid:plazi:treatment:038F87F22C1AFFE66100531F6DD65C20" httpUri="http://treatment.plazi.org/id/038F87F22C1AFFE66100531F6DD65C20" lastPageId="4" lastPageNumber="168" pageId="1" pageNumber="166">
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<subSubSection id="C33C656F2C1AFFE36100531F6E1059EA" pageId="1" pageNumber="166" type="nomenclature">
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<paragraph id="8B9936E42C1AFFE36100531F6E7959F0" blockId="1.[346,565,1133,1157]" box="[346,565,1133,1157]" pageId="1" pageNumber="166">
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<heading id="D0D181882C1AFFE36100531F6E7959F0" box="[346,565,1133,1157]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="166" reason="3">
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<emphasis id="B952EAF62C1AFFE36100531F6E7959F0" bold="true" box="[346,565,1133,1157]" pageId="1" pageNumber="166">
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<taxonomicName id="4C264D672C1AFFE36100531F6D8759F0" authority="Zhang & Yin, 2023" authorityName="Zhang & Yin" authorityYear="2023" box="[346,459,1133,1157]" class="Insecta" family="Staphylinidae" genus="Feabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B952EAF62C1AFFE36100531F6D8759F0" bold="true" box="[346,459,1133,1157]" italics="true" pageId="1" pageNumber="166">Feabatrus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A261578D2C1AFFE3618B531C6E7959F0" box="[465,565,1134,1157]" pageId="1" pageNumber="166" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9936E42C1AFFE3616953FE6E1059EA" blockId="1.[307,604,1162,1183]" box="[307,604,1162,1183]" pageId="1" pageNumber="166">
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Chinese common name:
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<emphasis id="B952EAF62C1AFFE3625D53F86E1059EA" bold="true" box="[519,604,1162,1183]" pageId="1" pageNumber="166">flümDz</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C656F2C1AFFE360CD53CA6D5D599A" pageId="1" pageNumber="166" type="type_taxon">
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<paragraph id="8B9936E42C1AFFE360CD53CA6D5D599A" blockId="1.[151,761,1208,2063]" pageId="1" pageNumber="166">
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<emphasis id="B952EAF62C1AFFE360CD53CA6D7E59BA" bold="true" box="[151,306,1208,1231]" pageId="1" pageNumber="166">
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<typeStatus id="549D88462C1AFFE360CD53CA6C9C59BA" box="[151,208,1208,1231]" pageId="1" pageNumber="166">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C264D672C1AFFE3616753CA6E1959BA" authority="Zhang & Yin, 2023" authorityName="Zhang & Yin" authorityYear="2023" box="[317,597,1208,1231]" class="Insecta" family="Staphylinidae" genus="Feabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="myanmarensis" status="sp. nov.">
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<emphasis id="B952EAF62C1AFFE3616753CA6E1959BA" box="[317,597,1208,1231]" italics="true" pageId="1" pageNumber="166">Feabatrus myanmarensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A261578D2C1AFFE3620553CA6EF659BA" box="[607,698,1208,1231]" pageId="1" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C656F2C1AFFE360CD538A6EF25B9A" pageId="1" pageNumber="166" type="diagnosis">
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<paragraph id="8B9936E42C1AFFE360CD538A6EF25B9A" blockId="1.[151,761,1208,2063]" pageId="1" pageNumber="166">
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<emphasis id="B952EAF62C1AFFE360CD538A6D40587A" bold="true" box="[151,268,1272,1295]" pageId="1" pageNumber="166">Diagnosis.</emphasis>
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Body large-sized, ranging from
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<quantity id="4CDE9B012C1AFFE36235538A6EB8587A" box="[623,756,1272,1295]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.3" metricValueMin="3.2" pageId="1" pageNumber="166" unit="mm" value="3.75" valueMax="4.3" valueMin="3.2">3.2–4.3 mm</quantity>
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, with smooth, shiny elytra and abdomen. Head slightly wi- der than long, sub-rounded at base, with U-shaped sulcus connecting vertexal foveae, with long mediobasal carina, eyes prominent; antenna elongate, lacking modification. Pronotum with broad, laterally carinate median longitudinal sulcus and short, curved discal carinae, with three pairs of large antebasal, discal and marginal spines, with distinct median antebasal and inner and outer pair of basolateral foveae. Each elytron with three basal foveae, discal stria shallow and short, lacking subhumeral fovea. Abdomen elongate, constricted at base and narrowing apically; tergite 1 (IV) longest, with thin inner and thick and short outer marginal carinae; sternites 3–5 (V–VII) each with one pair of basolateral foveae. Aedeagus stout, endophallus armature composed of elongate and curved sclerite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C656F2C1AFFE160CD518A6D3A5C00" lastPageId="3" lastPageNumber="168" pageId="1" pageNumber="166" type="description">
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<paragraph id="8B9936E42C1AFFE360CD518A6EDD5AFA" blockId="1.[151,761,1208,2063]" pageId="1" pageNumber="166">
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<emphasis id="B952EAF62C1AFFE360CD518A6D505A7A" bold="true" box="[151,284,1784,1807]" pageId="1" pageNumber="166">Description</emphasis>
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[figures based on a completely articulated male of
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<taxonomicName id="4C264D672C1AFFE360AB506A6D3B5A5A" authorityName="Zhang & Yin" authorityYear="2023" box="[241,375,1816,1839]" class="Insecta" family="Staphylinidae" genus="Feabatrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="166" phylum="Arthropoda" rank="species" species="leonardoi">
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<emphasis id="B952EAF62C1AFFE360AB506A6D4D5A5A" box="[241,257,1816,1839]" italics="true" pageId="1" pageNumber="166">F</emphasis>
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.
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<emphasis id="B952EAF62C1AFFE3614A506A6D3B5A5A" box="[272,375,1816,1839]" italics="true" pageId="1" pageNumber="166">leonardoi</emphasis>
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</taxonomicName>
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]. Body length
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<quantity id="4CDE9B012C1AFFE36240506A6ED35A5A" box="[538,671,1816,1839]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.3" metricValueMin="3.2" pageId="1" pageNumber="166" unit="mm" value="3.75" valueMax="4.3" valueMin="3.2">3.2–4.3 mm</quantity>
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; habitus elongate, dorsal surface of body covered with long, suberect setae; antenna relatively elongate, extending to more than half of elytral length when bent backward.
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</paragraph>
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<paragraph id="8B9936E42C1AFFE360EF50EA69D15F60" blockId="1.[151,761,1208,2063]" lastBlockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">
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Head (
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<figureCitation id="131D2A612C1AFFE360A350EA6D215ADA" box="[249,365,1944,1967]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Figs 1A–C</figureCitation>
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) roundly rectangular, wider than long; lacking frontal rostrum, antennal tubercles weakly raised, posterior margin with distinct lateral postantennal pit (
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<figureCitation id="131D2A612C1AFFE3629750AA6CF6557A" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1A</figureCitation>
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;
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<emphasis id="B952EAF62C1AFFE36099508A6CA8557A" box="[195,228,2040,2063]" italics="true" pageId="1" pageNumber="166">lpp</emphasis>
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); vertexal foveae (dorsal tentorial pits) (
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<figureCitation id="131D2A612C1AFFE362DB508A6E9F557A" box="[641,723,2040,2063]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1A</figureCitation>
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;
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<emphasis id="B952EAF62C1AFFE36286508A6EA3557A" box="[732,751,2040,2063]" italics="true" pageId="1" pageNumber="166">vf</emphasis>
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) relatively small, asetose, connected by reversed U-shaped sulcus, with long mediobasal (
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<figureCitation id="131D2A612C1AFFE364D557AC68AB5D80" box="[1167,1255,222,245]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1A</figureCitation>
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;
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<emphasis id="B952EAF62C1AFFE364AC57AC696F5D80" box="[1270,1315,222,245]" italics="true" pageId="1" pageNumber="166">mbc</emphasis>
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) and thick lateral carinae; eyes prominent, ocular-mandibular carinae (
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<figureCitation id="131D2A612C1AFFE3631E566C6FD45C40" box="[836,920,286,309]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1B</figureCitation>
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;
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<emphasis id="B952EAF62C1AFFE363FD566C6F9F5C40" box="[935,979,286,309]" italics="true" pageId="1" pageNumber="166">omc</emphasis>
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) complete, extending to mandible. Venter with small gular foveae (posterior tentorial pits) (
|
||
<figureCitation id="131D2A612C1AFFE3651F564C69DB5C20" box="[1349,1431,318,341]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1C</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36366562C6F1D5C00" box="[828,849,350,373]" italics="true" pageId="1" pageNumber="166">gf</emphasis>
|
||
) originating from shared opening, thin gular carina (
|
||
<figureCitation id="131D2A612C1AFFE3652B562C6F115CE0" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1C</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36332560C6FCD5CE0" box="[872,897,382,405]" italics="true" pageId="1" pageNumber="166">gc</emphasis>
|
||
) extending from opening anteriorly to mouthparts; antenna with 11 antennomeres, club indistinct, loosely formed by apical three antennomeres; maxillary palpus with small palpomere 1, 2 basally pedunculate and broad at apex, 3 subtriangular, 4 fusiform, with small cone at apex.
|
||
</paragraph>
|
||
<paragraph id="8B9936E42C1AFFE36300556C68C55EA0" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">
|
||
Pronotum (
|
||
<figureCitation id="131D2A612C1AFFE3638E556C68075F40" box="[980,1099,542,565]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Figs 1D–F</figureCitation>
|
||
) moderately transverse, lateral margin convergent anteriorly and constricted at basal 1/2, anterolateral margins with small denticles in front of middle, anterior and posterior margin slightly carinate; lateral sides with row of dense pubescence, with broad laterally carinate median and lateral longitudinal sulci (
|
||
<figureCitation id="131D2A612C1AFFE3657955CC69D45FA0" box="[1315,1432,702,725]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Figs 1D, E</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE3636655AC6F2D5F80" box="[828,865,734,757]" italics="true" pageId="1" pageNumber="166">mls</emphasis>
|
||
,
|
||
<emphasis id="B952EAF62C1AFFE3632A55AC6FC55F80" box="[880,905,734,757]" italics="true" pageId="1" pageNumber="166">lls</emphasis>
|
||
) and short mediobasal and curved discal carinae (
|
||
<figureCitation id="131D2A612C1AFFE3631E558C6FD65E60" box="[836,922,766,789]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1D</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE363FD558C6F9F5E60" box="[935,979,766,789]" italics="true" pageId="1" pageNumber="166">mbc</emphasis>
|
||
,
|
||
<emphasis id="B952EAF62C1AFFE363B8558C6FB75E60" box="[994,1019,766,789]" italics="true" pageId="1" pageNumber="166">dc</emphasis>
|
||
), lateral antebasal foveae (
|
||
<figureCitation id="131D2A612C1AFFE36544558C69D45E60" box="[1310,1432,766,789]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Figs 1D, E</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36366546C6F145E40" box="[828,856,798,821]" italics="true" pageId="1" pageNumber="166">laf</emphasis>
|
||
) asetose, with one median antebasal (
|
||
<figureCitation id="131D2A612C1AFFE36559546C69105E40" box="[1283,1372,798,821]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1D</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36531546C69DF5E40" box="[1387,1427,798,821]" italics="true" pageId="1" pageNumber="166">maf</emphasis>
|
||
) and two pairs of basolateral foveae (
|
||
<figureCitation id="131D2A612C1AFFE36491544C696F5E20" box="[1227,1315,830,853]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1D</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE3656A544C69C15E20" box="[1328,1421,830,853]" italics="true" pageId="1" pageNumber="166">oblf, iblf</emphasis>
|
||
), with large marginal, discal and antebasal spines, lacking transverse antebasal sulcus; with thin hypomeral carinae (
|
||
<figureCitation id="131D2A612C1AFFE3631E54EC6FDA5EC0" box="[836,918,926,949]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1E</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE363FB54EC6FF65EC0" box="[929,954,926,949]" italics="true" pageId="1" pageNumber="166">hc</emphasis>
|
||
) and small pit; prosternum with small lateral procoxal foveae (
|
||
<figureCitation id="131D2A612C1AFFE363AF54CC680A5EA0" box="[1013,1094,958,981]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1F</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE3640B54CC68355EA0" box="[1105,1145,958,981]" italics="true" pageId="1" pageNumber="166">lpcf</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936E42C1AFFE3630054AC69D759E0" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">
|
||
Elytra constricted and truncate at bases, each elytron with three basal foveae (
|
||
<figureCitation id="131D2A612C1AFFE36461548C68C35960" box="[1083,1167,1022,1045]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1G</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE364C2548C68F55960" box="[1176,1209,1022,1045]" italics="true" pageId="1" pageNumber="166">bef</emphasis>
|
||
), inner two close,lacking subbasal fovea; with shallow and short discal (
|
||
<figureCitation id="131D2A612C1AFFE3651C536C69DB5940" box="[1350,1431,1054,1077]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1G</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36366534C6F185920" box="[828,852,1086,1109]" italics="true" pageId="1" pageNumber="166">ds</emphasis>
|
||
) and complete sutural striae, lacking subhumeral fovea, slightly carinate marginal striae (
|
||
<figureCitation id="131D2A612C1AFFE364FC532C68B25900" box="[1190,1278,1118,1141]" captionStart="Fig" captionStartId="2.[151,182,1943,1962]" captionTargetBox="[158,1432,377,1901]" captionTargetId="figure-13@2.[151,1436,377,1909]" captionTargetPageId="2" captionText="Fig. 1. Morphology of Feabatrus gen. nov.A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H." figureDoi="http://doi.org/10.5281/zenodo.8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="1" pageNumber="166">Fig. 1H</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36556532C69665900" box="[1292,1322,1118,1141]" italics="true" pageId="1" pageNumber="166">ms</emphasis>
|
||
) extended from approximately middle to posterior margin of elytra.
|
||
</paragraph>
|
||
<paragraph id="8B9936E42C1AFFE3630053EC68865800" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">
|
||
Mesoventrite with median foveae (
|
||
<figureCitation id="131D2A612C1AFFE3649553EC696A59C0" box="[1231,1318,1182,1205]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2A</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE3656B53EC692559C0" box="[1329,1385,1182,1205]" italics="true" pageId="1" pageNumber="166">mmsf</emphasis>
|
||
) widely separated, in shared transverse opening, with large lateral mesoventral foveae (
|
||
<figureCitation id="131D2A612C1AFFE3640C53AC68E45980" box="[1110,1192,1246,1269]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2A</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE364EB53AC68905980" box="[1201,1244,1246,1269]" italics="true" pageId="1" pageNumber="166">lmsf</emphasis>
|
||
) forked internally; metaventrite with large, setose lateral coxal foveae (
|
||
<figureCitation id="131D2A612C1AFFE3652B538C6F2C5840" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2A</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36333526C6FDB5840" box="[873,919,1310,1333]" italics="true" pageId="1" pageNumber="166">lmcf</emphasis>
|
||
), lateral metaventral foveae (
|
||
<figureCitation id="131D2A612C1AFFE36497526C696E5840" box="[1229,1314,1310,1333]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2A</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36576526C69195840" box="[1324,1365,1310,1333]" italics="true" pageId="1" pageNumber="166">lmtf</emphasis>
|
||
) separated, posterior margin with deep, narrow split in middle; metaventral coxae broadly separated.
|
||
</paragraph>
|
||
<paragraph id="8B9936E42C1AFFE36300520C69D15BE0" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">
|
||
Abdomen elongate; tergite 1 (IV) longest, with mediobasal and two pairs of basolateral foveae (
|
||
<figureCitation id="131D2A612C1AFFE3656552EC69D458C0" box="[1343,1432,1438,1461]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2C</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE3636652CC6F2858A0" box="[828,868,1470,1493]" italics="true" pageId="1" pageNumber="166">mbf</emphasis>
|
||
,
|
||
<emphasis id="B952EAF62C1AFFE3632A52CC6FC058A0" box="[880,908,1470,1493]" italics="true" pageId="1" pageNumber="166">blf</emphasis>
|
||
), with thin inner marginal carinae, outer marginal carinae thick and short, with short, nodule-like discal carinae; tergites 2–4 (V–VII) each with one pair of basolateral foveae (
|
||
<figureCitation id="131D2A612C1AFFE363D5516C6FAC5B40" box="[911,992,1566,1589]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE363B3516C68495B40" box="[1001,1029,1566,1589]" italics="true" pageId="1" pageNumber="166">blf</emphasis>
|
||
) and thin lateral carinae; sternite 2 (IV) with one pair of mediobasal (
|
||
<figureCitation id="131D2A612C1AFFE36434514C688D5B20" box="[1134,1217,1598,1621]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2D</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36496514C68B85B20" box="[1228,1268,1598,1621]" italics="true" pageId="1" pageNumber="166">mbf</emphasis>
|
||
) and three pairs of basolateral foveae (
|
||
<figureCitation id="131D2A612C1AFFE3647D512C68375B00" box="[1063,1147,1630,1653]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2D</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE364DC512C68EE5B00" box="[1158,1186,1630,1653]" italics="true" pageId="1" pageNumber="166">blf</emphasis>
|
||
), sternites 3–5 (V–VII) each with one pair of small basolateral foveae (
|
||
<figureCitation id="131D2A612C1AFFE36543510C69255BE0" box="[1305,1385,1662,1685]" captionStart="Fig" captionStartId="3.[151,182,2004,2023]" captionTargetBox="[245,1358,632,1982]" captionTargetId="figure-15@3.[229,1358,625,1986]" captionTargetPageId="3" captionText="Fig. 2. Morphology of Feabatrus gen. nov. A – meso- and metaventrite; B–D – abdomen, lateral (B), dorsal (C), and ventral (D). Abbreviations: blf – basolateral fovea, lmcf – lateral mesocoxal fovea, lmsf – lateral mesoventral fovea, lmtf – lateral metaventral fovea, mbf – mediobasal fovea, mmsf – median mesoventral fovea, s1–5 – sternites 1–5 (III–VII), t1–4 – tergites 1–4 (IV–VII). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8005828" httpUri="https://zenodo.org/record/8005828/files/figure.png" pageId="1" pageNumber="166">Fig. 2B</figureCitation>
|
||
;
|
||
<emphasis id="B952EAF62C1AFFE36528510C69C15BE0" box="[1394,1421,1662,1685]" italics="true" pageId="1" pageNumber="166">blf</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936E42C1AFFE3630051EC689A5BA0" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">Legs elongate, tarsomeres 2 and 3 subequal in length, with one major and one setiform claw.</paragraph>
|
||
<paragraph id="8B9936E42C1AFFE3630051AC68BB5560" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="166">Male has relatively longer antennae than female; trochanters and mesofemora with spine or projection on ventral margin; abdomen greatly elongate in at least one species; aedeagus asymmetric, stout, median lobe with large basal capsule, dorsal lobe short, flat, endophallus armature markedly long and curved. Female with long spines on tergites 2–4 (V–VII) in two species; tergite 5 (VIII) trapezoidal, flat or weakly arched, posterior margin weakly emarginate and with small nodule at middle; genital complex asymmetric, weakly sclerotized.</paragraph>
|
||
<paragraph id="8B9936E42C19FFE160CD57CC6D3A5C00" blockId="2.[151,760,190,341]" lastBlockId="3.[150,760,190,597]" lastPageId="3" lastPageNumber="168" pageId="2" pageNumber="167">
|
||
<emphasis id="B952EAF62C19FFE060CD57CC6D305DA0" bold="true" box="[151,380,190,213]" pageId="2" pageNumber="167">Comparative notes.</emphasis>
|
||
<taxonomicName id="4C264D672C19FFE061DC57CC6DB95DA0" authority="Zhang & Yin, 2023" authorityName="Zhang & Yin" authorityYear="2023" box="[390,501,190,213]" class="Insecta" family="Staphylinidae" genus="Feabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B952EAF62C19FFE061DC57CC6DB95DA0" box="[390,501,190,213]" italics="true" pageId="2" pageNumber="167">Feabatrus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261578D2C19FFE061A557CC6E295DA0" box="[511,613,190,213]" pageId="2" pageNumber="167" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
belongs to a group of Oriental genera including
|
||
<taxonomicName id="4C264D672C19FFE0624F57AC6C9C5C60" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C19FFE0624F57AC6ED55D80" box="[533,665,222,245]" italics="true" pageId="2" pageNumber="167">Tribasodites</emphasis>
|
||
Jeannel, 1960
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D672C19FFE06085578C6E675C60" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[223,555,254,277]" class="Insecta" family="Staphylinidae" genus="Coryphomodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C19FFE06085578C6DC85C60" box="[223,388,254,277]" italics="true" pageId="2" pageNumber="167">Coryphomodes</emphasis>
|
||
Jeannel, 1960
|
||
</taxonomicName>
|
||
, and those genera that possess spinose pronotal lateral margins and have three basal foveae on each elytron (=
|
||
<taxonomicName id="4C264D672C19FFE06266564C6EFD5C20" authorityName="Jeannel" authorityYear="1958" box="[572,689,318,341]" class="Insecta" family="Staphylinidae" genus="Tribasodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C19FFE06266564C6EFD5C20" box="[572,689,318,341]" italics="true" pageId="2" pageNumber="167">Tribasodes</emphasis>
|
||
</taxonomicName>
|
||
-group of
|
||
<bibRefCitation id="EFB74B152C19FFE0630757CC682F5DA0" author="NOMURA S. & IDRIS A. B." box="[861,1123,190,213]" pageId="2" pageNumber="167" pagination="55 - 72" refId="ref7101" refString="NOMURA S. & IDRIS A. B. 2003: Faunistic notes on the batrisine species from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 8 (1 - 2): 55 - 72." type="journal article" year="2003">NOMURA & IDRIS 2003</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D672C19FFE0642257CC68A45DA0" authorityName="Zhang & Yin" authorityYear="2023" box="[1144,1256,190,213]" class="Insecta" family="Staphylinidae" genus="Feabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C19FFE0642257CC68A45DA0" box="[1144,1256,190,213]" italics="true" pageId="2" pageNumber="167">Feabatrus</emphasis>
|
||
</taxonomicName>
|
||
differs from all known Asian genera of this group in a combination of 1) large-sized body with long, suberect setae on dorsal surface and antennae; 2) pronotum with large marginal, discal and antebasal spines; 3) elytra each with a smooth disc, inner two basal foveae close, discal stria shallow, lacking subhumeral fovea; 4) sternites 3–5 (V–VII) with only one pair of small basolateral foveae; 5) constriction between elytra and abdomen distinct, and 6) flat, sub-trapezoidal female tergite 5 (VIII) in middle with a small nodule of the posterior margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF59666C2C19FFE060CD50E568AB5560" ID-DOI="http://doi.org/10.5281/zenodo.8005826" ID-Zenodo-Dep="8005826" httpUri="https://zenodo.org/record/8005826/files/figure.png" pageId="2" pageNumber="167" startId="2.[151,182,1943,1962]" targetBox="[158,1432,377,1901]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8B9936E42C19FFE060CD50E568AB5560" blockId="2.[151,1437,1943,2069]" pageId="2" pageNumber="167">
|
||
Fig. 1. Morphology of
|
||
<taxonomicName id="4C264D672C19FFE0610350E56DFD5ADF" authority="Zhang & Yin, 2023" authorityName="Zhang & Yin" authorityYear="2023" box="[345,433,1943,1962]" class="Insecta" family="Staphylinidae" genus="Feabatrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="167" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B952EAF62C19FFE0610350E56DFD5ADF" box="[345,433,1943,1962]" italics="true" pageId="2" pageNumber="167">Feabatrus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A261578D2C19FFE061EF50E56E485ADF" box="[437,516,1943,1962]" pageId="2" pageNumber="167" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
A–C – head, dorsal (A), lateral (B), and ventral (C); D–F – prothorax, dorsal (D), lateral (E), and ventral (F); G – right elytron; H – apical half of elytron, lateral. Abbreviations: bef – basal elytral foveae, dc – discal carina, ds – discal stria, gc – gular carina, gf – gular fovea, hc – hypomeral carina, iblf – inner basolateral fovea, laf – lateral antebasal fovea, lls – lateral longitudinal sulcus, lpcf – lateral procoxal fovea, lpp – lateral postantennal pits, maf – median antebasal fovea, mbc – mediobasal carina, mls – median longitudinal sulcus, ms – marginal stria, oblf – outer basolateral fovea, omc – ocular-mandibular carina, vf – vertexal fovea. Scale bars: 0.2 mm in A–F; 0.25 mm in G, H.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33C656F2C18FFE160CC560C6DA55F60" pageId="3" pageNumber="168" type="etymology">
|
||
<paragraph id="8B9936E42C18FFE160CC560C6DA55F60" blockId="3.[150,760,190,597]" pageId="3" pageNumber="168">
|
||
<emphasis id="B952EAF62C18FFE160CC560C6D545CE0" bold="true" box="[150,280,382,405]" pageId="3" pageNumber="168">Etymology.</emphasis>
|
||
The generic name is a combination of the family name of Leonardo Fea, an Italian naturalist who collected the material of both new species described here, and a partially abbreviated name of
|
||
<taxonomicName id="4C264D672C18FFE1624C56AC6EBF5C80" authority="Aube, 1833" authorityName="Aube" authorityYear="1833" box="[534,755,478,501]" class="Insecta" family="Staphylinidae" genus="Batrisus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C18FFE1624C56AC6E235C80" box="[534,623,478,501]" italics="true" pageId="3" pageNumber="168">Batrisus</emphasis>
|
||
Aubé, 1833
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="549D88462C18FFE160E4568C6CA05F60" box="[190,236,510,533]" pageId="3" pageNumber="168">type</typeStatus>
|
||
genus of the
|
||
<taxonomicName id="4C264D672C18FFE16121568C6DA95F60" authorityName="Reitter" authorityYear="1882" box="[379,485,510,533]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="165" phylum="Arthropoda" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C656F2C18FFE660CC556C6DD65C20" lastPageId="4" lastPageNumber="169" pageId="3" pageNumber="168" type="discussion">
|
||
<paragraph id="8B9936E42C18FFE660CC556C6DD65C20" blockId="3.[150,760,190,597]" lastBlockId="4.[151,760,190,341]" lastPageId="4" lastPageNumber="169" pageId="3" pageNumber="168">
|
||
<emphasis id="B952EAF62C18FFE160CC556C6DD85F40" bold="true" box="[150,404,542,565]" pageId="3" pageNumber="168">Comments on biology.</emphasis>
|
||
The female genital complexes of all three species are slightly asymmetric, with membranous structures pressing genital plate to one side. For
|
||
<taxonomicName id="4C264D672C18FFE1651A57CC69D05DA0" authorityName="Reitter" authorityYear="1882" box="[1344,1436,190,213]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
this character state has not been known by us, and is postulated to be associated with an undocumented, specialized mating behavior of the species. Note that the female of two of the three
|
||
<taxonomicName id="4C264D672C18FFE163B1564C68E15C20" box="[1003,1197,318,341]" pageId="3" pageNumber="168">
|
||
<emphasis id="B952EAF62C18FFE163B1564C68155C20" box="[1003,1113,318,341]" italics="true" pageId="3" pageNumber="168">Feabatrus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(see below) possesses long dorsal spines on tergites 2–4 (V–VII), and a similar condition can be found in a few species of the genera
|
||
<taxonomicName id="4C264D672C18FFE1636656EC68305CC0" authority="Kurbatov, 2007" authorityName="Kurbatov" authorityYear="2007" box="[828,1148,414,437]" class="Insecta" family="Staphylinidae" genus="Intestinarius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C18FFE1636656EC6F8A5CC0" box="[828,966,414,437]" italics="true" pageId="3" pageNumber="168">Intestinarius</emphasis>
|
||
Kurbatov, 2007
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C264D672C18FFE164D456EC68A35CC0" authorityName="Reitter" authorityYear="1882" box="[1166,1263,414,437]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C264D672C18FFE1656956EC6F825CA0" authority="Raffray, 1905" authorityName="Raffray" authorityYear="1905" class="Insecta" family="Staphylinidae" genus="Horniella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAF62C18FFE1656956EC69D15CC0" box="[1331,1437,414,437]" italics="true" pageId="3" pageNumber="168">Horniella</emphasis>
|
||
Raffray, 1905
|
||
</taxonomicName>
|
||
(Pselaphitae:
|
||
<taxonomicName id="4C264D672C18FFE1643056CC68E35CA0" authorityName="Reitter" authorityYear="1882" box="[1130,1199,446,469]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="tribe" tribe="Tyrini">Tyrini</taxonomicName>
|
||
), etc. The function of such spines in
|
||
<taxonomicName id="4C264D672C18FFE1638D56AC681A5C80" authorityName="Latreille" authorityYear="1802" box="[983,1110,478,501]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
|
||
is unknown, but a comparable case has been documented for the water strider
|
||
<taxonomicName id="4C264D672C18FFE16569568C68CD5F40" authority="Drake & Hottes, 1925" authorityName="Drake & Hottes" authorityYear="1925" class="Insecta" family="Gerridae" genus="Gerris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="168" phylum="Arthropoda" rank="species" species="incognitus">
|
||
<emphasis id="B952EAF62C18FFE16569568C6FDA5F40" italics="true" pageId="3" pageNumber="168">Gerris incognitus</emphasis>
|
||
Drake & Hottes, 1925
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFB74B152C18FFE164D2556C69D15F40" author="ARNQVIST G. & ROWE L." box="[1160,1437,542,565]" pageId="3" pageNumber="168" pagination="123 - 127" refId="ref6822" refString="ARNQVIST G. & ROWE L. 1995: Sexual conflict and arms races between the sexes: a morphological adaptation for control of mating in a female insect. Proceedings of the Royal Society B: Biological Sciences 261: 123 - 127." type="journal article" year="1995">ARNQVIST & ROWE (1995)</bibRefCitation>
|
||
provided experimental evidence that female abdominal spines of
|
||
<taxonomicName id="4C264D672C1FFFE660A057CC6DCB5DA0" authorityName="Drake & Hottes" authorityYear="1925" box="[250,391,190,213]" class="Insecta" family="Gerridae" genus="Gerris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="species" species="incognitus">
|
||
<emphasis id="B952EAF62C1FFFE660A057CC6D415DA0" box="[250,269,190,213]" italics="true" pageId="4" pageNumber="169">G</emphasis>
|
||
.
|
||
<emphasis id="B952EAF62C1FFFE6614257CC6DCB5DA0" box="[280,391,190,213]" italics="true" pageId="4" pageNumber="169">incognitus</emphasis>
|
||
</taxonomicName>
|
||
are used to thwart harassing males, while increasing female control over copulation. However, this ‘sexual-conflict’ hypothesis has never been tested for
|
||
<taxonomicName id="4C264D672C1FFFE660CD566C6D555C40" authorityName="Latreille" authorityYear="1802" box="[151,281,286,309]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="169" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
|
||
, and further field observations or laboratory experiments are needed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |