156 lines
18 KiB
XML
156 lines
18 KiB
XML
<document id="FDA136E12AF67E08DB82DF478D0B51EE" ID-CLB-Dataset="51205" ID-DOI="10.11646/zootaxa.4061.5.3" ID-GBIF-Dataset="8e4c370b-2a1d-4bc4-a714-16f9f3a1bd65" ID-ISSN="1175-5326" ID-Zenodo-Dep="264294" ID-ZooBank="6EEE4C42-D292-490A-9549-CB3428430920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459680617424" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2016" docId="038F87EE015E3016FF0BD028FD2CFEF6" docLanguage="en" docName="zootaxa.4061.5.3.pdf" docOrigin="Zootaxa 4061 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus ruvumanus Häckel, 2016, sp. n." docType="treatment" docVersion="10" lastPageNumber="520" masterDocId="FFB6FF9601513006FF9CD433FFDDFFEB" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)" masterLastPageNumber="526" masterPageNumber="504" pageNumber="519" updateTime="1698207076058" updateUser="ExternalLinkService">
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<mods:mods id="2E56B5EA346550471CDDE2858D7AED54" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4D50485FE405C7500AF00FE288124E64">
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<mods:title id="45CCBD733804D4530F3CB47262924CA4">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<mods:name id="0A386E8EF6C40D5C16FA936CF865EA1A" type="personal">
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<mods:roleTerm id="6ED431BC31D1BDEFE92A079B8323D51F">Author</mods:roleTerm>
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<mods:namePart id="21904B7EDF84ABFAC77664EEC323DBCC">Häckel, Martin</mods:namePart>
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<mods:typeOfResource id="4965833319600F2EF90AFBEA10A813E9">text</mods:typeOfResource>
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<mods:title id="50519C5B77082178415EAB3EB11BA7C7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1E011862E8875793BBC271680BF3655F">
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<mods:date id="59383CEFEF2F97A35EB881A13AB9D8E9">2016</mods:date>
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<mods:detail id="45B6856364376B5D9C53219D429DC440" type="volume">
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<mods:number id="D96CE1539A4E69351711C1D2D851ED14">4061</mods:number>
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<mods:detail id="A6D5DF4BBAA1F763FAA057FEE5EA17D5" type="issue">
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<mods:number id="4DF30B022A3FA7BB6D3E973B2F25ACFE">5</mods:number>
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<mods:classification id="2DDC190E2B80F643AD77077456AA51EE">journal article</mods:classification>
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<treatment id="038F87EE015E3016FF0BD028FD2CFEF6" ID-DOI="http://doi.org/10.5281/zenodo.6088143" ID-GBIF-Taxon="119332996" ID-Zenodo-Dep="6088143" LSID="urn:lsid:plazi:treatment:038F87EE015E3016FF0BD028FD2CFEF6" httpUri="http://treatment.plazi.org/id/038F87EE015E3016FF0BD028FD2CFEF6" lastPageId="16" lastPageNumber="520" pageId="15" pageNumber="519">
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<subSubSection id="C33C6573015E3009FF0BD028FDBEFBDE" box="[151,611,1051,1077]" pageId="15" pageNumber="519" type="nomenclature">
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<paragraph id="8B9936F8015E3009FF0BD028FDBEFBDE" blockId="15.[151,712,1051,1110]" box="[151,611,1051,1077]" pageId="15" pageNumber="519">
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<heading id="D0D18194015E3009FF0BD028FDBEFBDE" bold="true" box="[151,611,1051,1077]" fontSize="11" level="1" pageId="15" pageNumber="519" reason="1">
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<emphasis id="B952EAEA015E3009FF0BD028FDBEFBDE" bold="true" box="[151,611,1051,1077]" pageId="15" pageNumber="519">
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11.
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<taxonomicName id="4C264D7B015E3009FF5DD028FDC4FBDE" ID-CoL="Z5YY" box="[193,537,1051,1077]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="ruvumanus" status="sp. nov.">
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<emphasis id="B952EAEA015E3009FF5DD028FDC4FBDE" bold="true" box="[193,537,1051,1077]" italics="true" pageId="15" pageNumber="519">Craspedophorus ruvumanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2615791015E3009FDBCD028FDBEFBDE" box="[544,611,1051,1077]" pageId="15" pageNumber="519" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6573015E3009FF0BD00EFD15FBBD" box="[151,712,1085,1110]" pageId="15" pageNumber="519" type="description">
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<paragraph id="8B9936F8015E3009FF0BD00EFD15FBBD" blockId="15.[151,712,1051,1110]" box="[151,712,1085,1110]" pageId="15" pageNumber="519">
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(Plate 2, Fig. 11,
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<figureCitation id="131D2A7D015E3009FECBD00EFE76FBBD" box="[343,427,1085,1110]" captionStart="FIGURE 26. C" captionStartId="14.[151,250,971,993]" captionTargetBox="[348,1246,223,909]" captionTargetId="figure@14.[340,1247,211,918]" captionTargetPageId="14" captionText="FIGURE 26. C. ruvumanus sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. NMWC labels; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264296/files/figure.png" pageId="15" pageNumber="519">Figs 26</figureCitation>
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a–f, 27d, Plate 4, Fig. 36)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6573015E3009FF0BD0B6FAD9FB29" pageId="15" pageNumber="519" type="materials_examined">
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<paragraph id="8B9936F8015E3009FF0BD0B6FAD9FB29" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">
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<emphasis id="B952EAEA015E3009FF0BD0B6FE9AFB75" bold="true" box="[151,327,1157,1182]" pageId="15" pageNumber="519">
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<typeStatus id="549D885A015E3009FF0BD0B6FF0FFB75" box="[151,210,1157,1182]" pageId="15" pageNumber="519">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="549D885A015E3009FECCD0B6FE65FB75" box="[336,440,1157,1182]" pageId="15" pageNumber="519" type="holotype">Holotype</typeStatus>
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(♂) labelled: “Ugano /
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<quantity id="4CDE9B1D015E3009FD53D0B6FC8DFB75" box="[719,848,1157,1182]" metricMagnitude="2" metricUnit="m" metricValue="8.575" metricValueMax="17.0" metricValueMin="0.15" pageId="15" pageNumber="519" unit="m" value="857.5" valueMax="1700.0" valueMin="15.0">15–1700 m</quantity>
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// Tanganyika-Terr., / Matengo-Hochland / wsw. v. Songea, /
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<date id="FF981038015E3009FE9AD099FEBCFB29" box="[262,353,1194,1218]" pageId="15" pageNumber="519">21.-31.I</date>
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, '036. Zerny [=
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<collectingCountry id="F3317668015E3009FD95D099FDA9FB28" box="[521,628,1194,1219]" name="Tanzania" pageId="15" pageNumber="519">Tanzania</collectingCountry>
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, Ruvuma Region]” (Plate 2, Fig. 11,
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<figureCitation id="131D2A7D015E3009FB88D099FBBAFB28" box="[1044,1127,1194,1219]" captionStart="FIGURE 26. C" captionStartId="14.[151,250,971,993]" captionTargetBox="[348,1246,223,909]" captionTargetId="figure@14.[340,1247,211,918]" captionTargetPageId="14" captionText="FIGURE 26. C. ruvumanus sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. NMWC labels; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264296/files/figure.png" pageId="15" pageNumber="519">Figs 26</figureCitation>
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a–f,
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<collectionCode id="ED37AE3D015E3009FB05D099FB24FB29" box="[1177,1273,1194,1218]" httpUri="http://grbio.org/cool/e7dr-nxad" name="National Museum of Wales" pageId="15" pageNumber="519">NMWC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6573015E3016FF5BD0FEFE54FF3E" lastPageId="16" lastPageNumber="520" pageId="15" pageNumber="519" type="description">
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<paragraph id="8B9936F8015E3009FF5BD0FEFD84FAE1" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">
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<emphasis id="B952EAEA015E3009FF5BD0FEFE84FB0D" bold="true" box="[199,345,1229,1254]" pageId="15" pageNumber="519">Description.</emphasis>
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Length
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<quantity id="4CDE9B1D015E3009FE2AD0FDFDC1FB0D" box="[438,540,1230,1254]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="15" pageNumber="519" unit="mm" value="23.2">23.2 mm</quantity>
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, width
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<quantity id="4CDE9B1D015E3009FDF3D0FDFD15FB0D" box="[623,712,1230,1254]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="15" pageNumber="519" unit="mm" value="9.1">9.1 mm</quantity>
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. Proportions: Pronotum 1.42x wider than long, 1.89x wider than head, elytra 1,44x wider than pronotum.
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</paragraph>
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<paragraph id="8B9936F8015E3009FF5BD126FB4EFAB8" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">Coloration. Body piceous black, opaque, densely covered by long, dark yellow setae; underside piceous black, opaque, antennae and palps brownish. Head wide and short, eyes convex, neck constricted.</paragraph>
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<paragraph id="8B9936F8015E3009FF5BD16CFC1DF9C0" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">
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Pronotum transverse, 1.42x wider than long, almost hexagonal; maximum width immediately behind midlength; anterior margin flat, front angles not projecting anteriorly and strongly rounded; lateral margins evenly rounded; widening gradually to maximum width, then narrowing in weakly sinuate line toward hind angles; each hind angle with a small indentation preceded by an indistinct incision; base medially parallel with elytral base; laterally base slanted forward, causing hind angle to be wide open; sagittal line indistinct, basal pits weakly impressed; surface strongly coarsely and unevenly punctured (
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<figureCitation id="131D2A7D015E3009FCCED221FC7BF9C0" box="[850,934,1554,1579]" captionStart="FIGURE 26. C" captionStartId="14.[151,250,971,993]" captionTargetBox="[348,1246,223,909]" captionTargetId="figure@14.[340,1247,211,918]" captionTargetPageId="14" captionText="FIGURE 26. C. ruvumanus sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. NMWC labels; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264296/files/figure.png" pageId="15" pageNumber="519">Fig. 26</figureCitation>
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a).
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</paragraph>
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<paragraph id="8B9936F8015E3009FF5BD206FDE7F973" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">Elytra fused, almost parallel, weekly convex, flattened on disc, wings reduced. Striae interrupted by punctuation, 1st stria strongly oblique to 2nd stria, joining it near base; intervals fairly convex, finely and densely punctured, sparsely covered by setae.</paragraph>
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<paragraph id="8B9936F8015E3009FF5BD296FB74F90B" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">
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Underside covered by short setae, finely punctured medially, punctured near margins; metepisterna longer than wide, wider anteriorly than posteriorly (
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<figureCitation id="131D2A7D015E3009FDCAD2FBFD75F90A" box="[598,680,1736,1761]" captionStart="FIGURE 26. C" captionStartId="14.[151,250,971,993]" captionTargetBox="[348,1246,223,909]" captionTargetId="figure@14.[340,1247,211,918]" captionTargetPageId="14" captionText="FIGURE 26. C. ruvumanus sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. NMWC labels; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264296/files/figure.png" pageId="15" pageNumber="519">Fig. 26</figureCitation>
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e, f). Legs moderately covered by black setae.
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</paragraph>
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<paragraph id="8B9936F8015E3009FF5BD2DEFD9BF8C2" blockId="15.[151,1436,1157,2014]" pageId="15" pageNumber="519">
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Aedeagus (Plate 4, Fig. 36) similar to those of
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<taxonomicName id="4C264D7B015E3009FD43D2DEFCBFF8ED" box="[735,866,1773,1798]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="impictus">C. impictus</taxonomicName>
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and
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<taxonomicName id="4C264D7B015E3009FC07D2DEFBEEF8ED" box="[923,1075,1773,1798]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="ethmoides">C. ethmoides</taxonomicName>
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, but in dorsal view differing by curvature (Plate 4, cf. Figs 34 and 35).
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</paragraph>
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<paragraph id="8B9936F8015E3016FF5BD306FE54FF3E" blockId="15.[151,1436,1157,2014]" lastBlockId="16.[151,1436,151,285]" lastPageId="16" lastPageNumber="520" pageId="15" pageNumber="519">
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<emphasis id="B952EAEA015E3009FF5BD306FE10F8A5" bold="true" box="[199,461,1845,1870]" pageId="15" pageNumber="519">Differential diagnosis.</emphasis>
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<taxonomicName id="4C264D7B015E3009FE48D306FCCFF8A6" box="[468,786,1845,1870]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="ruvumanus" status="sp. nov.">
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<emphasis id="B952EAEA015E3009FE48D306FCCFF8A6" box="[468,786,1845,1870]" italics="true" pageId="15" pageNumber="519">Craspedophorus ruvumanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2615791015E3009FC85D306FC84F8A5" box="[793,857,1845,1870]" pageId="15" pageNumber="519" rank="species">
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<emphasis id="B952EAEA015E3009FC85D306FC84F8A5" bold="true" box="[793,857,1845,1870]" pageId="15" pageNumber="519">sp. n.</emphasis>
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</taxonomicNameLabel>
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mostly resembles
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<taxonomicName id="4C264D7B015E3009FBB6D305FB60F8A5" box="[1066,1213,1845,1870]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="ethmoides">
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<emphasis id="B952EAEA015E3009FBB6D305FB60F8A5" box="[1066,1213,1845,1870]" italics="true" pageId="15" pageNumber="519">C. ethmoides</emphasis>
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</taxonomicName>
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differing from it by elytral sculpture with punctures much smaller, more rounded, arranged in a more orderly fashion, and spaces between them deeper than centers of intervals. From similarly colored
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<taxonomicName id="4C264D7B015E3009FC46D34DFB80F87E" box="[986,1117,1918,1941]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="impictus">
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<emphasis id="B952EAEA015E3009FC46D34DFB80F87E" box="[986,1117,1918,1941]" italics="true" pageId="15" pageNumber="519">C. impictus</emphasis>
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</taxonomicName>
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the new species differs by smaller size and the same characters as those given by Alluaud for separation of
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<taxonomicName id="4C264D7B015E3009FBABD392FB11F852" box="[1079,1228,1952,1977]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="ethmoides">
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<emphasis id="B952EAEA015E3009FBABD392FB11F852" box="[1079,1228,1952,1977]" italics="true" pageId="15" pageNumber="519">C. ethmoides</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C264D7B015E3009FA88D392FA48F852" box="[1300,1429,1953,1977]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="impictus">
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<emphasis id="B952EAEA015E3009FA88D392FA48F852" box="[1300,1429,1953,1977]" italics="true" pageId="15" pageNumber="519">C. impictus</emphasis>
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</taxonomicName>
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. The aedeagus presents in lateral view the same curvature as in
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<taxonomicName id="4C264D7B015E3009FCF9D3F5FC24F835" box="[869,1017,1989,2014]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="ethmoides">
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<emphasis id="B952EAEA015E3009FCF9D3F5FC24F835" box="[869,1017,1989,2014]" italics="true" pageId="15" pageNumber="519">C. ethmoides</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C264D7B015E3009FBAED3F5FB6EF836" box="[1074,1203,1990,2013]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="519" phylum="Arthropoda" rank="species" species="impictus">
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<emphasis id="B952EAEA015E3009FBAED3F5FB6EF836" box="[1074,1203,1990,2013]" italics="true" pageId="15" pageNumber="519">C. impictus</emphasis>
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</taxonomicName>
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, but in craniocaudal view the margins of the endophallus aperture are near base parallel, whereas in the other two species they converge (Plate 4, Figs 34–36).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301413016FF5BD4ECFD2CFEF6" pageId="16" pageNumber="520" type="etymology">
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<paragraph id="8B9936F801413016FF5BD4ECFD2CFEF6" blockId="16.[151,1436,151,285]" pageId="16" pageNumber="520">
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<emphasis id="B952EAEA01413016FF5BD4ECFE93FF13" bold="true" box="[199,334,223,248]" pageId="16" pageNumber="520">Etymology.</emphasis>
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Named after the Ruvuma Region in
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<collectingCountry id="F331766801413016FD76D4ECFC92FF13" box="[746,847,223,248]" name="Tanzania" pageId="16" pageNumber="520">Tanzania</collectingCountry>
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whre the specimen was found.
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<emphasis id="B952EAEA01413016FF5BD537FEBDFEF6" bold="true" box="[199,352,260,285]" pageId="16" pageNumber="520">Distribution.</emphasis>
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Known only from the
|
||
<typeStatus id="549D885A01413016FDFCD536FD4DFEF6" box="[608,656,261,285]" pageId="16" pageNumber="520">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |