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<mods:title id="E74E3461A83EE502134A91C1D68314DB">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<treatment id="038F87EE01533005FF0BD762FDB0FF13" ID-DOI="http://doi.org/10.5281/zenodo.6088119" ID-GBIF-Taxon="119333003" ID-Zenodo-Dep="6088119" LSID="urn:lsid:plazi:treatment:038F87EE01533005FF0BD762FDB0FF13" httpUri="http://treatment.plazi.org/id/038F87EE01533005FF0BD762FDB0FF13" lastPageId="3" lastPageNumber="506" pageId="2" pageNumber="506">
<subSubSection id="C33C657301533004FF0BD762FD08FC66" pageId="2" pageNumber="506" type="nomenclature">
<paragraph id="8B9936F801533004FF0BD762FCB9FC80" blockId="2.[151,868,849,909]" box="[151,868,849,875]" pageId="2" pageNumber="506">
<heading id="D0D1819401533004FF0BD762FCB9FC80" bold="true" box="[151,868,849,875]" fontSize="11" level="1" pageId="2" pageNumber="506" reason="1">
<emphasis id="B952EAEA01533004FF0BD762FCB9FC80" bold="true" box="[151,868,849,875]" pageId="2" pageNumber="506">
<taxonomicName id="4C264D7B01533004FF0BD762FE1BFC80" ID-CoL="Z5YR" box="[151,454,849,875]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FF0BD762FE1BFC80" bold="true" box="[151,454,849,875]" italics="true" pageId="2" pageNumber="506">Craspedophorus reflexus</emphasis>
</taxonomicName>
species group (hereby redefined)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936F801533004FF0BD746FD08FC66" blockId="2.[151,868,849,909]" box="[151,725,884,909]" pageId="2" pageNumber="506">
(=see
<bibRefCitation id="EFB74B0901533004FF47D747FE5BFC66" author="Chaudoir" box="[219,390,884,909]" pageId="2" pageNumber="526" refString="Chaudoir, M. de (1879) Essai monographie sur les Panageides (part II). Annales de la Societe entomologique de Belgique, 21, 85 - 186." type="journal article" year="1879">Chaudoir 1879</bibRefCitation>
: 97;
<bibRefCitation id="EFB74B0901533004FE20D747FD5DFC66" author="Basilewsky" box="[444,640,884,909]" pageId="2" pageNumber="525" refString="Basilewsky, P. (1987) Descriptions de quelques especes africaines nouvelles du genre Craspedophorus Hope et notes synonymiques (Coleoptera Carabidae Panagaeini). Bulletin et Annales de La Societe Royale Belge d'entomologie, 123 (7 - 9), 185 - 201." type="journal article" year="1987">Basilewsky 1987</bibRefCitation>
: 196-7)
</paragraph>
</subSubSection>
<subSubSection id="C33C657301533004FF0BD78FFE1CF84B" pageId="2" pageNumber="506" type="description">
<paragraph id="8B9936F801533004FF0BD78FFD52F9B6" blockId="2.[151,1437,956,2025]" pageId="2" pageNumber="506">
<bibRefCitation id="EFB74B0901533004FF0BD78FFE5DFC3E" author="Chaudoir" box="[151,384,956,981]" pageId="2" pageNumber="526" refString="Chaudoir, M. de (1879) Essai monographie sur les Panageides (part II). Annales de la Societe entomologique de Belgique, 21, 85 - 186." type="journal article" year="1879">Chaudoir (1879: 97)</bibRefCitation>
placed species of here established group with paraglossae surpassing glossa and increasing slenderness and length of ligula, protarsi identical in both sexes and the terminal palpomere very dilated, kidneyshaped. He did not use a grouping system and never designated any group in the sense as presently understood. Despite that, the here established
<taxonomicName id="4C264D7B01533004FD90D01AFD5EFBAA" box="[524,643,1064,1089]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FD90D01AFD5EFBAA" box="[524,643,1064,1089]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
</taxonomicName>
group (and other species groups which will be established in following articles) well agree with Chaudoir's concept of the genus
<taxonomicName id="4C264D7B01533004FCD5D07FFBD9FB8E" box="[841,1028,1100,1125]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B952EAEA01533004FCD5D07FFBD9FB8E" box="[841,1028,1100,1125]" italics="true" pageId="2" pageNumber="506">Craspedophorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101533004FB87D07EFB6FFB8E" box="[1051,1202,1100,1125]" pageId="2" pageNumber="506" sensu="stricto">sensu stricto</taxonomicNameLabel>
“. The other species recently placed in this genus, but in other groups, were placed by Chaudoir in the genera
<taxonomicName id="4C264D7B01533004FB37D042FF12FB46" authority="Chaudoir, 1846" authorityName="Chaudoir" authorityYear="1846" class="Insecta" family="Carabidae" genus="Epicosmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B952EAEA01533004FB37D042FAF9FB63" box="[1195,1316,1137,1160]" italics="true" pageId="2" pageNumber="506">Epicosmus</emphasis>
<bibRefCitation id="EFB74B0901533004FAB2D043FF12FB46" author="Chaudoir" pageId="2" pageNumber="526" refString="Chaudoir, M. de (1846) Note sur le groupe des Stomides et description d'un nouveau genre de celuis des Somoplatides. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 19 (2), 511 - 542." type="journal article" year="1846">Chaudoir, 1846</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C264D7B01533004FE9AD0A7FE1DFB47" authority="Laporte" authorityName="Laporte" box="[262,448,1172,1197]" class="Magnoliopsida" family="Brassicaceae" genus="Eudema" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="506" phylum="Tracheophyta" rank="genus">
<emphasis id="B952EAEA01533004FE9AD0A7FEBFFB46" box="[262,354,1172,1197]" italics="true" pageId="2" pageNumber="506">Eudema</emphasis>
Laporte
</taxonomicName>
de
<bibRefCitation id="EFB74B0901533004FE75D0A7FD7CFB46" author="Castelnau" box="[489,673,1172,1197]" pageId="2" pageNumber="526" refString="Laporte de Castelnau, F. L. N. (1840) Histoire Naturelle des Insectes, Animaux articules. Coleopteres. Vol. 1. Dumenil, Paris, cxxv + 24 pp." type="book" year="1840">Castelnau, 1840</bibRefCitation>
, both later synonymized with
<taxonomicName id="4C264D7B01533004FC68D0A7FB72FB46" box="[1012,1199,1172,1197]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B952EAEA01533004FC68D0A7FB72FB46" box="[1012,1199,1172,1197]" italics="true" pageId="2" pageNumber="506">Craspedophorus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB74B0901533004FB46D0A7FF17FB3A" author="Andrewes" pageId="2" pageNumber="525" refString="Andrewes, H. E. (1919) On the types of Oriental Carabidae in the British Museum, and in the Hope Department of the Oxford University Museum. The Transactions of the Royal Entomological Society of London, 1 - 2, 67, 119 - 217. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1919. tb 00006. x" type="journal article" year="1919">Andrewes (1919: 126)</bibRefCitation>
and followed by
<bibRefCitation id="EFB74B0901533004FE3BD08BFD1FFB3A" author="Basilewsky" box="[423,706,1208,1233]" pageId="2" pageNumber="525" refString="Basilewsky, P. (1953) Carabidae (Coleoptera, Adephaga). In: Exploration du Parc National de l´Upemba, Mission G. F. de Witte en collaboration avec W. Adam, A. Janssens, L. van Meel, et R. Verheyen (1946 - 1949), fascicules 10. Bruxelles (Institut de Parcs Nationaux du Congo-Belge), x + 252 pp." type="book chapter" year="1953">Basilewsky (1953: 169)</bibRefCitation>
. Within the
<taxonomicName id="4C264D7B01533004FCC0D08BFBA6FB3A" box="[860,1147,1208,1233]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FCC0D08BFBA6FB3A" box="[860,1147,1208,1233]" italics="true" pageId="2" pageNumber="506">Craspedophorus reflexus</emphasis>
</taxonomicName>
group there are species differing from those of the above listed groups by ventrites not crenulated anteriorly, antennae more slender, head longer, constricted behind eyes, and pronotum not too transverse. Chaudoir classified species here placed in the
<taxonomicName id="4C264D7B01533004FA1ED132FF32FAD6" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FA1ED132FF32FAD6" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
</taxonomicName>
group at the beginning of its system.
<bibRefCitation id="EFB74B0901533004FD3CD117FCBAFAD6" author="Andrewes" box="[672,871,1316,1341]" pageId="2" pageNumber="525" refString="Andrewes, H. E. (1919) On the types of Oriental Carabidae in the British Museum, and in the Hope Department of the Oxford University Museum. The Transactions of the Royal Entomological Society of London, 1 - 2, 67, 119 - 217. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1919. tb 00006. x" type="journal article" year="1919">Andrewes (1919)</bibRefCitation>
, designated
<taxonomicName id="4C264D7B01533004FC68D116FAE5FAD6" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[1012,1336,1316,1341]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FC68D116FBB3FAD6" box="[1012,1134,1316,1341]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901533004FBE2D117FAEDFAD6" author="Fabricius" box="[1150,1328,1316,1341]" pageId="2" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
the
<typeStatus id="549D885A01533004FAF0D116FA41FAD6" box="[1388,1436,1317,1341]" pageId="2" pageNumber="506">type</typeStatus>
species of the genus. Therefore, I consider the
<taxonomicName id="4C264D7B01533004FD3AD17AFCC3FA8A" box="[678,798,1352,1377]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FD3AD17AFCC3FA8A" box="[678,798,1352,1377]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
</taxonomicName>
group to be “nominotypical” sensu
<bibRefCitation id="EFB74B0901533004FB2AD17BFA41FA8B" author="Chaudoir" box="[1206,1436,1352,1377]" pageId="2" pageNumber="526" refString="Chaudoir, M. de (1879) Essai monographie sur les Panageides (part II). Annales de la Societe entomologique de Belgique, 21, 85 - 186." type="journal article" year="1879">Chaudoir (1879: 97)</bibRefCitation>
and including firstly two species,
<taxonomicName id="4C264D7B01533004FDBCD15EFCB5FA6E" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[544,872,1388,1413]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FDBCD15EFD41FA6E" box="[544,668,1388,1413]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901533004FD30D15FFCBDFA6E" author="Fabricius" box="[684,864,1388,1413]" pageId="2" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C264D7B01533004FC39D15EFB21FA6E" authority="Boheman, 1848" authorityName="Boheman" authorityYear="1848" box="[933,1276,1388,1413]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="impictus">
<emphasis id="B952EAEA01533004FC39D15EFBF5FA6F" box="[933,1064,1389,1412]" italics="true" pageId="2" pageNumber="506">C. impictus</emphasis>
(
<bibRefCitation id="EFB74B0901533004FBA6D15FFB29FA6E" author="Boheman" box="[1082,1268,1388,1413]" pageId="2" pageNumber="525" refString="Boheman, C. H. (1848) Insecta Caffraria annis 1838 - 1845 a J. A. Wahlberg collecta amici auxillio suffultus. Pars I, Fasc. 1. (Carabici, Hydrocanthari, Gyrinii et Staphylinii). Fritze &amp; Norstedt, Holmiae, viii + 625 pp. [1851]" type="book chapter" year="1848">Boheman, 1848</bibRefCitation>
)
</taxonomicName>
. I also accept Bates', Alluaud's and Basilewsky's conclusions increasing it with the species
<taxonomicName id="4C264D7B01533004FB87D1A2FAFCFA43" authority="Bates, 1890" authorityName="Bates" authorityYear="1890" box="[1051,1313,1424,1449]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="bonnyi">
<emphasis id="B952EAEA01533004FB87D1A2FB51FA42" box="[1051,1164,1424,1449]" italics="true" pageId="2" pageNumber="506">C. bonnyi</emphasis>
<bibRefCitation id="EFB74B0901533004FB0AD1A2FAFCFA43" author="Bates" box="[1174,1313,1424,1448]" pageId="2" pageNumber="525" refString="Bates, H. W. (1890) On some Coleopterous Insects collected by Mr. W. Bonny in the Aruwimi Valley. Proceedings Zoological Society London, 1890, 479 - 578." type="proceedings paper" year="1890">Bates, 1890</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FAACD1A2FE9FFA26" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="bozasi">
<emphasis id="B952EAEA01533004FAACD1A2FA41FA42" box="[1328,1436,1424,1449]" italics="true" pageId="2" pageNumber="506">C. bozasi</emphasis>
<bibRefCitation id="EFB74B0901533004FF0BD187FE9FFA26" author="Alluaud" box="[151,322,1460,1485]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FEC8D186FD5BFA26" authority="Kolbe, 1889" authorityName="Kolbe" authorityYear="1889" box="[340,646,1460,1485]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="buettneri">
<emphasis id="B952EAEA01533004FEC8D186FE3CFA26" box="[340,481,1460,1485]" italics="true" pageId="2" pageNumber="506">C. buettneri</emphasis>
<bibRefCitation id="EFB74B0901533004FE6DD187FD5BFA26" author="Kolbe" box="[497,646,1460,1485]" pageId="2" pageNumber="526" refString="Kolbe, H. J. (1889) Bericht uber die von Herrn Dr. R. Buttner im Gebiete des unteren Quango und Kongo gesammelten Coleopteren. Entomologische Zeitung, 1889 (4 - 6), 113 - 133." type="journal article" year="1889">Kolbe, 1889</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FD04D186FBD4FA26" authority="Harold, 1879" authorityName="Harold" authorityYear="1879" box="[664,1033,1460,1485]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="carbonarius">
<emphasis id="B952EAEA01533004FD04D186FC97FA26" box="[664,842,1460,1485]" italics="true" pageId="2" pageNumber="506">C. carbonarius</emphasis>
(
<bibRefCitation id="EFB74B0901533004FCFCD187FBDCFA26" author="Harold" box="[864,1025,1460,1485]" pageId="2" pageNumber="526" refString="Harold, E. von (1879) Bericht uber die von den Herren A. von Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Kafer. Coleopterologische Hefte, 16, 1 - 221." type="journal article" year="1879">Harold, 1879</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FB80D186FAB2FA26" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[1052,1391,1460,1485]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="ethmoides">
<emphasis id="B952EAEA01533004FB80D186FB68FA26" box="[1052,1205,1460,1485]" italics="true" pageId="2" pageNumber="506">C. ethmoides</emphasis>
<bibRefCitation id="EFB74B0901533004FB58D187FAB2FA26" author="Alluaud" box="[1220,1391,1460,1485]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FA1ED186FE3DFA1B" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="latemaculatus">
<emphasis id="B952EAEA01533004FA1ED186FEE8FA1A" italics="true" pageId="2" pageNumber="506">C. latemaculatus</emphasis>
<bibRefCitation id="EFB74B0901533004FEA0D1EBFE3DFA1B" author="Alluaud" box="[316,480,1496,1521]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FE71D1EAFCC4FA1B" authority="Alluaud, 1932" authorityName="Alluaud" authorityYear="1932" box="[493,793,1496,1521]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="lebaudyi">
<emphasis id="B952EAEA01533004FE71D1EAFDB2FA1A" box="[493,623,1496,1521]" italics="true" pageId="2" pageNumber="506">C. lebaudyi</emphasis>
<bibRefCitation id="EFB74B0901533004FDEBD1EBFCC4FA1B" author="Alluaud" box="[631,793,1496,1521]" pageId="2" pageNumber="525" refString="Alluaud, C. (1932) III. Carabiques d Afrique tropicale. Afra, Cahiers d'Entomologie, 5, 9 - 17." type="journal article" year="1932">Alluaud, 1932</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C264D7B01533004FCCED1EAFBA5FA1B" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[850,1144,1496,1521]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01533004FCCED1EAFC16FA1A" box="[850,971,1496,1521]" italics="true" pageId="2" pageNumber="506">C. stanleyi</emphasis>
<bibRefCitation id="EFB74B0901533004FC4FD1EBFBA5FA1B" author="Alluaud" box="[979,1144,1496,1521]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C264D7B01533004FB18D1EBFEF2F9FE" authority="Hope, 1842" authorityName="Hope" authorityYear="1842" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="savagei">
<emphasis id="B952EAEA01533004FB18D1EBFA46FA1B" box="[1156,1435,1496,1521]" italics="true" pageId="2" pageNumber="506">Craspedophorus savagei</emphasis>
(
<bibRefCitation id="EFB74B0901533004FF03D1CEFEFAF9FE" author="Hope" box="[159,295,1533,1557]" pageId="2" pageNumber="526" refString="Hope, F. W. (1842) On some new Insects from Western Africa. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 10, 91 - 93." type="journal article" year="1842">Hope, 1842</bibRefCitation>
)
</taxonomicName>
is hereby synonymized with
<taxonomicName id="4C264D7B01533004FD15D1CEFC0AF9FE" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[649,983,1532,1557]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FD15D1CEFCD8F9FE" box="[649,773,1532,1557]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901533004FC8AD1CFFC13F9FE" author="Fabricius" box="[790,974,1532,1557]" pageId="2" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C264D7B01533004FC78D1CFFB25F9FF" box="[996,1272,1532,1557]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="arnosti" status="sp. nov.">
<emphasis id="B952EAEA01533004FC78D1CFFB25F9FF" box="[996,1272,1532,1557]" italics="true" pageId="2" pageNumber="506">Craspedophorus arnosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101533004FA9FD1CFFA9BF9FE" box="[1283,1350,1532,1557]" pageId="2" pageNumber="506" rank="species">
<emphasis id="B952EAEA01533004FA9FD1CFFA9BF9FE" bold="true" box="[1283,1350,1532,1557]" pageId="2" pageNumber="506">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C264D7B01533004FA1ED1CEFEC9F9D3" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="ruvumanus" status="sp. nov.">
<emphasis id="B952EAEA01533004FA1ED1CEFEC9F9D3" italics="true" pageId="2" pageNumber="506">C. ruvumanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101533004FE87D213FE86F9D2" box="[283,347,1568,1593]" pageId="2" pageNumber="506" rank="species">
<emphasis id="B952EAEA01533004FE87D213FE86F9D2" bold="true" box="[283,347,1568,1593]" pageId="2" pageNumber="506">sp. n.</emphasis>
</taxonomicNameLabel>
by its features are placed as well in the
<taxonomicName id="4C264D7B01533004FCBDD212FC47F9D2" box="[801,922,1568,1593]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FCBDD212FC47F9D2" box="[801,922,1568,1593]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
</taxonomicName>
group also. Presently the group encompasses 12 species, one of them with two subspecies.
</paragraph>
<paragraph id="8B9936F801533004FF5BD25BFE1CF84B" blockId="2.[151,1437,956,2025]" pageId="2" pageNumber="506">
<emphasis id="B952EAEA01533004FF5BD25BFE89F96A" bold="true" box="[199,340,1640,1665]" pageId="2" pageNumber="506">Characters.</emphasis>
Larger species (
<quantity id="4CDE9B1D01533004FD8ED25BFD4EF96B" box="[530,659,1640,1664]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.9" metricValueMin="1.8" pageId="2" pageNumber="506" unit="mm" value="23.5" valueMax="29.0" valueMin="18.0">1829 mm</quantity>
). Head longer, markedly constricted behind eyes. Antennae slender. Palps with terminal article very dilated, kidney-shaped. Metepisterna of characteristic shape, longer than wide (
<figureCitation id="131D2A7D01533004FF03D283FF28F922" box="[159,245,1712,1737]" captionStart="FIGURE 25. C" captionStartId="13.[151,250,1927,1949]" captionTargetBox="[362,1246,1183,1901]" captionTargetId="figure@13.[340,1247,1183,1906]" captionTargetPageId="13" captionText="FIGURE 25. C. arnosti sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. pronotum, dorsal view; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264295/files/figure.png" pageId="2" pageNumber="506">Figs 25</figureCitation>
e, f, 26e, f, 27c). Ventrites anteriorly not crenulated along posterior margin of preceding segment (
<figureCitation id="131D2A7D01533004FAA9D283FA45F922" box="[1333,1432,1712,1737]" captionStart="FIGURE 25. C" captionStartId="13.[151,250,1927,1949]" captionTargetBox="[362,1246,1183,1901]" captionTargetId="figure@13.[340,1247,1183,1906]" captionTargetPageId="13" captionText="FIGURE 25. C. arnosti sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. pronotum, dorsal view; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264295/files/figure.png" pageId="2" pageNumber="506">Figs 25c</figureCitation>
,
<figureCitation id="131D2A7D01533004FF0BD2E7FF1EF906" box="[151,195,1748,1773]" captionStart="FIGURE 26. C" captionStartId="14.[151,250,971,993]" captionTargetBox="[348,1246,223,909]" captionTargetId="figure@14.[340,1247,211,918]" captionTargetPageId="14" captionText="FIGURE 26. C. ruvumanus sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. NMWC labels; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264296/files/figure.png" pageId="2" pageNumber="506">26c</figureCitation>
). Elytral coloration varies from two relatively large orange fasciae extending from medial intervals to margin as in
<taxonomicName id="4C264D7B01533004FF4AD2CAFD9EF8FB" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[214,579,1784,1809]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="latemaculatus">
<emphasis id="B952EAEA01533004FF4AD2CAFE4BF8FA" box="[214,406,1784,1809]" italics="true" pageId="2" pageNumber="506">C. latemaculatus</emphasis>
<bibRefCitation id="EFB74B0901533004FE02D2CBFD9EF8FB" author="Alluaud" box="[414,579,1784,1809]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FDCDD2CAFC47F8FB" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[593,922,1784,1809]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01533004FDCDD2CAFD16F8FA" box="[593,715,1784,1809]" italics="true" pageId="2" pageNumber="506">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901533004FD47D2CBFC4DF8FB" author="Fabricius" box="[731,912,1784,1809]" pageId="2" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FC3BD2CAFB67F8FB" authority="Bates, 1890" authorityName="Bates" authorityYear="1890" box="[935,1210,1784,1809]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="bonnyi">
<emphasis id="B952EAEA01533004FC3BD2CAFBCBF8FA" box="[935,1046,1784,1809]" italics="true" pageId="2" pageNumber="506">C. bonnyi</emphasis>
(
<bibRefCitation id="EFB74B0901533004FBBBD2CBFB6FF8FB" author="Bates" box="[1063,1202,1784,1808]" pageId="2" pageNumber="525" refString="Bates, H. W. (1890) On some Coleopterous Insects collected by Mr. W. Bonny in the Aruwimi Valley. Proceedings Zoological Society London, 1890, 479 - 578." type="proceedings paper" year="1890">Bates, 1890</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FB5BD2CAFF0FF8DE" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="bozasi">
<emphasis id="B952EAEA01533004FB5BD2CAFAECF8FA" box="[1223,1329,1784,1809]" italics="true" pageId="2" pageNumber="506">C. bozasi</emphasis>
<bibRefCitation id="EFB74B0901533004FAA7D2CBFF0FF8DE" author="Alluaud" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FF41D32EFE2BF8DE" authority="Kolbe, 1889" authorityName="Kolbe" authorityYear="1889" box="[221,502,1820,1845]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="buettneri">
<emphasis id="B952EAEA01533004FF41D32EFEBEF8DE" box="[221,355,1820,1845]" italics="true" pageId="2" pageNumber="506">C. buettneri</emphasis>
<bibRefCitation id="EFB74B0901533004FEF7D32FFE2BF8DE" author="Kolbe" box="[363,502,1820,1845]" pageId="2" pageNumber="526" refString="Kolbe, H. J. (1889) Bericht uber die von Herrn Dr. R. Buttner im Gebiete des unteren Quango und Kongo gesammelten Coleopteren. Entomologische Zeitung, 1889 (4 - 6), 113 - 133." type="journal article" year="1889">Kolbe, 1889</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C264D7B01533004FDB1D32EFC87F8DE" authority="Alluaud, 1932" authorityName="Alluaud" authorityYear="1932" box="[557,858,1820,1845]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="lebaudyi">
<emphasis id="B952EAEA01533004FDB1D32EFD72F8DE" box="[557,687,1820,1845]" italics="true" pageId="2" pageNumber="506">C. lebaudyi</emphasis>
<bibRefCitation id="EFB74B0901533004FD2BD32FFC87F8DE" author="Alluaud" box="[695,858,1820,1845]" pageId="2" pageNumber="525" refString="Alluaud, C. (1932) III. Carabiques d Afrique tropicale. Afra, Cahiers d'Entomologie, 5, 9 - 17." type="journal article" year="1932">Alluaud, 1932</bibRefCitation>
</taxonomicName>
, to strongly reduced elytral coloration with either a single humeral macula (
<taxonomicName id="4C264D7B01533004FE3AD372FDCBF8B3" box="[422,534,1857,1880]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="arnosti" status="sp. nov.">
<emphasis id="B952EAEA01533004FE3AD372FDCBF8B3" box="[422,534,1857,1880]" italics="true" pageId="2" pageNumber="506">C. arnosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101533004FD82D372FDABF8B3" box="[542,630,1856,1880]" pageId="2" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
) or 23 residual macular spots as in
<taxonomicName id="4C264D7B01533004FB88D372FAB1F8B3" authority="Harold, 1879" authorityName="Harold" authorityYear="1879" box="[1044,1388,1856,1881]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="carbonarius">
<emphasis id="B952EAEA01533004FB88D372FB63F8B2" box="[1044,1214,1856,1881]" italics="true" pageId="2" pageNumber="506">C. carbonarius</emphasis>
(
<bibRefCitation id="EFB74B0901533004FB51D373FAB8F8B3" author="Harold" box="[1229,1381,1856,1881]" pageId="2" pageNumber="526" refString="Harold, E. von (1879) Bericht uber die von den Herren A. von Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Kafer. Coleopterologische Hefte, 16, 1 - 221." type="journal article" year="1879">Harold, 1879</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C264D7B01533004FF0BD356FE66F896" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[151,443,1892,1917]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01533004FF0BD356FED2F896" box="[151,271,1892,1917]" italics="true" pageId="2" pageNumber="506">C. stanleyi</emphasis>
<bibRefCitation id="EFB74B0901533004FE8BD357FE66F896" author="Alluaud" box="[279,443,1892,1917]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
, or even entirely black as in
<taxonomicName id="4C264D7B01533004FC9FD356FB9FF896" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[771,1090,1892,1917]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="ethmoides">
<emphasis id="B952EAEA01533004FC9FD356FC4BF896" box="[771,918,1892,1917]" italics="true" pageId="2" pageNumber="506">C. ethmoides</emphasis>
<bibRefCitation id="EFB74B0901533004FC02D357FB9FF896" author="Alluaud" box="[926,1090,1892,1917]" pageId="2" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C264D7B01533004FBD2D356FA41F896" authority="Boheman, 1848" authorityName="Boheman" authorityYear="1848" box="[1102,1436,1892,1917]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="impictus">
<emphasis id="B952EAEA01533004FBD2D356FB12F897" box="[1102,1231,1893,1916]" italics="true" pageId="2" pageNumber="506">C. impictus</emphasis>
(
<bibRefCitation id="EFB74B0901533004FB43D357FA49F896" author="Boheman" box="[1247,1428,1892,1917]" pageId="2" pageNumber="525" refString="Boheman, C. H. (1848) Insecta Caffraria annis 1838 - 1845 a J. A. Wahlberg collecta amici auxillio suffultus. Pars I, Fasc. 1. (Carabici, Hydrocanthari, Gyrinii et Staphylinii). Fritze &amp; Norstedt, Holmiae, viii + 625 pp. [1851]" type="book chapter" year="1848">Boheman, 1848</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C264D7B01533004FF5AD3BAFEBEF84B" box="[198,355,1929,1952]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="506" phylum="Arthropoda" rank="species" species="ruvumanus" status="sp. nov.">
<emphasis id="B952EAEA01533004FF5AD3BAFEBEF84B" box="[198,355,1929,1952]" italics="true" pageId="2" pageNumber="506">C. ruvumanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101533004FEF6D3BAFE1CF84B" box="[362,449,1929,1952]" pageId="2" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C33C657301533005FF5BD39FFDB0FF13" lastPageId="3" lastPageNumber="507" pageId="2" pageNumber="506" type="distribution">
<paragraph id="8B9936F801533005FF5BD39FFDB0FF13" blockId="2.[151,1437,956,2025]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="507" pageId="2" pageNumber="506">
<emphasis id="B952EAEA01533004FF5BD39FFE10F82E" bold="true" box="[199,461,1964,1989]" pageId="2" pageNumber="506">Distribution remarks.</emphasis>
These species occur in a large region covering West African savanna from
<collectingCountry id="F331766801533004FAB9D39FFAA3F82E" box="[1317,1406,1964,1989]" name="Senegal" pageId="2" pageNumber="506">Senegal</collectingCountry>
to
<collectingCountry id="F331766801533004FF0BD3E3FED2F803" box="[151,271,2000,2024]" name="Cameroon" pageId="2" pageNumber="506">Cameroon</collectingCountry>
, continuing southward to the mouth of the
<collectingCountry id="F331766801533004FD6ED3E3FCE0F803" box="[754,829,2000,2024]" name="Republic of the Congo" pageId="2" pageNumber="506">Congo</collectingCountry>
River, and Central Africa from
<collectingCountry id="F331766801533004FB39D3E3FB2DF802" box="[1189,1264,2000,2025]" name="Gabon" pageId="2" pageNumber="506">Gabon</collectingCountry>
to north of
<collectingCountry id="F331766801533005FAE9D3E2FF39FF5B" lastPageId="3" lastPageNumber="507" name="Democratic Republic of the Congo" pageId="2" pageNumber="506">DR Congo</collectingCountry>
, reaching the Great Lakes and eastward
<collectingCountry id="F331766801523005FD28D4A4FCD3FF5B" box="[692,782,151,176]" name="Uganda" pageId="3" pageNumber="507">Uganda</collectingCountry>
,
<collectingCountry id="F331766801523005FC87D4A4FC73FF5B" box="[795,942,151,176]" name="South Sudan" pageId="3" pageNumber="507">South Sudan</collectingCountry>
,
<collectingCountry id="F331766801523005FC26D4A4FBC1FF5B" box="[954,1052,151,176]" name="Ethiopia" pageId="3" pageNumber="507">Ethiopia</collectingCountry>
,
<collectingCountry id="F331766801523005FBB5D4ABFBA8FF5B" box="[1065,1141,152,176]" name="Kenya" pageId="3" pageNumber="507">Kenya</collectingCountry>
, continental
<collectingCountry id="F331766801523005FA9AD4A4FAB6FF5B" box="[1286,1387,151,176]" name="Tanzania" pageId="3" pageNumber="507">Tanzania</collectingCountry>
and
<collectingCountry id="F331766801523005FF0BD48FFF20FF3E" box="[151,253,188,213]" name="Tanzania" pageId="3" pageNumber="507">Zanzibar</collectingCountry>
. Distributions of some species (e.g.
<taxonomicName id="4C264D7B01523005FD3CD48EFCFEFF3F" box="[672,803,189,212]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="impictus">
<emphasis id="B952EAEA01523005FD3CD48EFCFEFF3F" box="[672,803,189,212]" italics="true" pageId="3" pageNumber="507">C. impictus</emphasis>
</taxonomicName>
) extend to
<collectingCountry id="F331766801523005FC35D48FFBE2FF3E" box="[937,1087,188,213]" name="South Africa" pageId="3" pageNumber="507">South Africa</collectingCountry>
,
<collectingCountry id="F331766801523005FBD0D48FFB79FF3E" box="[1100,1188,188,213]" name="Zambia" pageId="3" pageNumber="507">Zambia</collectingCountry>
,
<collectingCountry id="F331766801523005FB2ED48FFAF7FF3E" box="[1202,1322,188,213]" name="Zimbabwe" pageId="3" pageNumber="507">Zimbabwe</collectingCountry>
and
<collectingCountry id="F331766801523005FAFAD48FFA41FF3F" box="[1382,1436,188,212]" name="South Africa" pageId="3" pageNumber="507">RSA</collectingCountry>
(Limpopo, Mpumalanga,
<collectingCountry id="F331766801523005FE2AD4ECFE28FF13" box="[438,501,223,248]" name="South Africa" pageId="3" pageNumber="507">Natal</collectingCountry>
) (Fig, 37).
</paragraph>
</subSubSection>
</treatment>
</document>