187 lines
21 KiB
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187 lines
21 KiB
XML
<document id="756BFC65BCEA0AD5C79F60BAEB4441FD" ID-CLB-Dataset="51205" ID-DOI="10.11646/zootaxa.4061.5.3" ID-GBIF-Dataset="8e4c370b-2a1d-4bc4-a714-16f9f3a1bd65" ID-ISSN="1175-5326" ID-Zenodo-Dep="264294" ID-ZooBank="6EEE4C42-D292-490A-9549-CB3428430920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459680617424" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2016" docId="038F87EE01523002FF0BD2FFFEB3FB2C" docLanguage="en" docName="zootaxa.4061.5.3.pdf" docOrigin="Zootaxa 4061 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus bonnyi Bates 1890" docType="treatment" docVersion="10" lastPageNumber="508" masterDocId="FFB6FF9601513006FF9CD433FFDDFFEB" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)" masterLastPageNumber="526" masterPageNumber="504" pageNumber="507" updateTime="1698207076058" updateUser="ExternalLinkService">
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<mods:title id="564208E6B816C98CCC7889646CDBED61">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="17CBF3426DDF24BF1B552F3CEE7EE503">Häckel, Martin</mods:namePart>
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<mods:typeOfResource id="EDDA89034C79CC7418CB24A6E6BD86E9">text</mods:typeOfResource>
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<mods:title id="55F938D88C1DF95CFAAD0EB9AB6A405B">Zootaxa</mods:title>
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<mods:date id="3458E787FA9E45D9BA47DF64059FC576">2016</mods:date>
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<mods:number id="D4073C6110625087FB46FC9FD0B7EB48">4061</mods:number>
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<treatment id="038F87EE01523002FF0BD2FFFEB3FB2C" ID-DOI="http://doi.org/10.5281/zenodo.6088123" ID-GBIF-Taxon="119333005" ID-Zenodo-Dep="6088123" LSID="urn:lsid:plazi:treatment:038F87EE01523002FF0BD2FFFEB3FB2C" httpUri="http://treatment.plazi.org/id/038F87EE01523002FF0BD2FFFEB3FB2C" lastPageId="4" lastPageNumber="508" pageId="3" pageNumber="507">
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<subSubSection id="C33C657301523005FF0BD2FFFDB7F90D" box="[151,618,1740,1766]" pageId="3" pageNumber="507" type="nomenclature">
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<paragraph id="8B9936F801523005FF0BD2FFFDB7F90D" blockId="3.[151,618,1740,1799]" box="[151,618,1740,1766]" pageId="3" pageNumber="507">
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<heading id="D0D1819401523005FF0BD2FFFDB7F90D" bold="true" box="[151,618,1740,1766]" fontSize="11" level="1" pageId="3" pageNumber="507" reason="1">
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<emphasis id="B952EAEA01523005FF0BD2FFFDB7F90D" bold="true" box="[151,618,1740,1766]" pageId="3" pageNumber="507">
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2.
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<taxonomicName id="4C264D7B01523005FF28D2FFFDB7F90D" ID-CoL="Z5TG" authority="Bates, 1890" authorityName="Bates" authorityYear="1890" box="[180,618,1740,1766]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="bonnyi">
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<emphasis id="B952EAEA01523005FF28D2FFFE08F90D" bold="true" box="[180,469,1740,1766]" italics="true" pageId="3" pageNumber="507">Craspedophorus bonnyi</emphasis>
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<bibRefCitation id="EFB74B0901523005FE40D2FFFDB7F90D" author="Bates" box="[476,618,1740,1766]" pageId="3" pageNumber="525" refString="Bates, H. W. (1890) On some Coleopterous Insects collected by Mr. W. Bonny in the Aruwimi Valley. Proceedings Zoological Society London, 1890, 479 - 578." type="proceedings paper" year="1890">Bates, 1890</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301523005FF0BD2DDFE5FF8EC" box="[151,386,1774,1799]" pageId="3" pageNumber="507" type="description">
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<paragraph id="8B9936F801523005FF0BD2DDFE5FF8EC" blockId="3.[151,618,1740,1799]" box="[151,386,1774,1799]" pageId="3" pageNumber="507">(Plate 3, Figs 17, 18)</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301523005FF0BD304FAFBF880" pageId="3" pageNumber="507" type="reference_group">
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<paragraph id="8B9936F801523005FF0BD304FAFBF880" blockId="3.[151,1436,1847,1899]" pageId="3" pageNumber="507">
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<treatmentCitationGroup id="AB3611D601523005FF0BD304FAFBF880" pageId="3" pageNumber="507">
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<taxonomicName id="4C264D7B01523005FF0BD304FD92F8A6" ID-CoL="Z5TG" authority="Bates, 1890: 480" authorityName="Bates" authorityPageNumber="480" authorityYear="1890" box="[151,591,1847,1869]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="species" species="bonnyi">
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<emphasis id="B952EAEA01523005FF0BD304FE4DF8A6" box="[151,400,1847,1869]" italics="true" pageId="3" pageNumber="507">Craspedophorus bonnyi</emphasis>
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<treatmentCitation id="0A8710E901523005FE06D30BFD92F8A6" author="Bates" box="[410,591,1847,1869]" page="480" pageId="3" pageNumber="507" year="1890">
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<bibRefCitation id="EFB74B0901523005FE06D30BFDC7F8A6" author="Bates" box="[410,538,1847,1869]" pageId="3" pageNumber="525" refString="Bates, H. W. (1890) On some Coleopterous Insects collected by Mr. W. Bonny in the Aruwimi Valley. Proceedings Zoological Society London, 1890, 479 - 578." type="proceedings paper" year="1890">Bates, 1890</bibRefCitation>
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: 480
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</treatmentCitation>
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</taxonomicName>
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(
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<typeStatus id="549D885A01523005FDFCD304FD50F8A6" box="[608,653,1847,1869]" pageId="3" pageNumber="507">type</typeStatus>
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loc. “R[iver]. Aruwimi” [=Democratic
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<collectingCountry id="F331766801523005FBABD304FBA3F8A6" box="[1079,1150,1847,1869]" name="Republic of the Congo" pageId="3" pageNumber="507">Congo</collectingCountry>
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: Orientale Province: forest region towards Albert Lake]).
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<treatmentCitation id="0A8710E901523005FE67D366FD66F880" author="Alluaud" box="[507,699,1877,1899]" page="154" pageId="3" pageNumber="507" year="1915">
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<bibRefCitation id="EFB74B0901523005FE67D366FD54F880" author="Alluaud" box="[507,649,1877,1899]" pageId="3" pageNumber="525" refString="Alluaud, C. (1915) Contributions a l'etude des carabiques d'Afrique et de Madagascar (Col. Carabidae). V. Les grands Panageides du bassin du Chari. Bulletin de la Societe Entomologique de France, 1915, 152 - 156." type="journal article" year="1915">Alluaud 1915</bibRefCitation>
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: 154
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</treatmentCitation>
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,
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<treatmentCitation id="0A8710E901523005FD5AD365FC56F880" author="Burgeon" box="[710,907,1877,1899]" page="159" pageId="3" pageNumber="507" year="1930">
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<bibRefCitation id="EFB74B0901523005FD5AD365FC84F880" author="Burgeon" box="[710,857,1877,1899]" pageId="3" pageNumber="526" refString="Burgeon, L. (1930) Les Panagaeini du Musee du Congo. Revue de Zoologie et de Botanique Africaines, 19, 151 - 166." type="journal article" year="1930">Burgeon 1930</bibRefCitation>
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: 159
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</treatmentCitation>
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, 1935b: 181.
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<treatmentCitation id="0A8710E901523005FB89D366FAFFF880" author="Hackel" box="[1045,1314,1877,1899]" page="79" pageId="3" pageNumber="507" year="2012">
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<bibRefCitation id="EFB74B0901523005FB89D366FB20F880" author="Hackel" box="[1045,1277,1877,1899]" pageId="3" pageNumber="526" refString="Hackel, M. & Farkac, J. (2012) A check-list of the subfamily Panagaeinae Hope, 1838 of the World (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 8 (1 - 2), 67 - 116." type="journal article" year="2012">Häckel et Farkač 2012</bibRefCitation>
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: 79
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301523002FF0BD3ABFDA4FE8E" lastPageId="4" lastPageNumber="508" pageId="3" pageNumber="507" type="materials_examined">
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<paragraph id="8B9936F801523002FF0BD3ABFEC0FF3E" blockId="3.[151,1436,1944,2005]" lastBlockId="4.[151,1437,151,1223]" lastPageId="4" lastPageNumber="508" pageId="3" pageNumber="507">
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<emphasis id="B952EAEA01523005FF0BD3ABFE97F85A" bold="true" box="[151,330,1944,1969]" pageId="3" pageNumber="507">
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<typeStatus id="549D885A01523005FF0BD3ABFF0FF85A" box="[151,210,1944,1969]" pageId="3" pageNumber="507">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="549D885A01523005FECAD3ABFE63F85A" box="[342,446,1944,1969]" pageId="3" pageNumber="507" type="holotype">Holotype</typeStatus>
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(♀): “R. / Aruwimi [handwritten, white label] //
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<taxonomicName id="4C264D7B01523005FB8FD3ABFB13F85A" box="[1043,1230,1944,1969]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="507" phylum="Arthropoda" rank="genus">
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<emphasis id="B952EAEA01523005FB8FD3ABFB13F85A" box="[1043,1230,1944,1969]" italics="true" pageId="3" pageNumber="507">Craspedophorus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C264D7B01523005FB72D3AAFAE0F85B" box="[1262,1341,1945,1968]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="bonnyi">
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<emphasis id="B952EAEA01523005FB72D3AAFAE0F85B" box="[1262,1341,1945,1968]" italics="true" pageId="3" pageNumber="507">Bonnyi</emphasis>
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</taxonomicName>
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/ Bates [handwritten, white label] // Ex-Musaeo / H. W. Bates / 1892 [printed, white label] //
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<collectionCode id="ED37AE3D01523005FBFDD38EFB7AF83F" box="[1121,1191,1981,2004]" pageId="3" pageNumber="507">
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<typeStatus id="549D885A01523005FBFDD38EFB7AF83F" box="[1121,1191,1981,2004]" pageId="3" pageNumber="507">TYPE</typeStatus>
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</collectionCode>
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[printed, red label] // Muséum Paris // ex Coll. / R. Oberthür / 1952 [printed, blue label]”, (Plate 3, Fig. 18,
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<collectionCode id="ED37AE3D01553002FB16D4ABFB3FFF44" LSID="urn:lsid:biocol.org:col:34988" box="[1162,1250,152,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="508">MNHN</collectionCode>
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, Oberthür/Bates Collection).
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</paragraph>
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<paragraph id="8B9936F801553002FF5BD4ECFA88FEF7" blockId="4.[151,1437,151,1223]" pageId="4" pageNumber="508">
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<emphasis id="B952EAEA01553002FF5BD4ECFD3AFF13" bold="true" box="[199,743,223,248]" pageId="4" pageNumber="508">
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Other material examined. Democratic
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<collectingCountry id="F331766801553002FD0ED4ECFD3FFF13" box="[658,738,223,248]" name="Republic of the Congo" pageId="4" pageNumber="508">Congo</collectingCountry>
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.
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</emphasis>
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<specimenCount id="9D20FD7101553002FD71D4D3FCCCFF13" box="[749,785,223,248]" pageId="4" pageNumber="508" type="generic">
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1
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<emphasis id="B952EAEA01553002FD60D4ECFCCCFF13" bold="true" box="[764,785,223,248]" pageId="4" pageNumber="508">♂</emphasis>
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</specimenCount>
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: “Urwald Beni [=Nord-Kivu Province]”, (Plate 3, Fig. 17, cMH);
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<emphasis id="B952EAEA01553002FF74D537FEEFFEF6" bold="true" box="[232,306,260,285]" pageId="4" pageNumber="508">Error.</emphasis>
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1♀: “S-India, Madras state, Kurumbagaran, ex col. Hlinkel, 2011 [erroneous locality]”, (cIB).
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</paragraph>
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<paragraph id="8B9936F801553002FF5BD514FDA4FE8E" blockId="4.[151,1437,151,1223]" pageId="4" pageNumber="508">
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<emphasis id="B952EAEA01553002FF5BD514FEDBFEAB" bold="true" box="[199,262,295,320]" pageId="4" pageNumber="508">Note.</emphasis>
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The original description of this species is based on a single specimen collected by William Bonny, who was a military man in Stanley's expedition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301553002FF5BD55CFB22FBDC" pageId="4" pageNumber="508" type="description">
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<paragraph id="8B9936F801553002FF5BD55CFB22FBDC" blockId="4.[151,1437,151,1223]" pageId="4" pageNumber="508">
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<emphasis id="B952EAEA01553002FF5BD55CFE8FFE63" bold="true" box="[199,338,367,392]" pageId="4" pageNumber="508">Description</emphasis>
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(in part, see
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<bibRefCitation id="EFB74B0901553002FE7AD543FDB5FE63" author="Bates" box="[486,616,368,392]" pageId="4" pageNumber="525" refString="Bates, H. W. (1890) On some Coleopterous Insects collected by Mr. W. Bonny in the Aruwimi Valley. Proceedings Zoological Society London, 1890, 479 - 578." type="proceedings paper" year="1890">Bates 1890</bibRefCitation>
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: 480). “Larger body, head relatively smaller than pronotum, elytra wider and convex. Black, glossy, head convex, weakly constricted behind eyes; pronotum almost ovoid, with almost rectangular posterior angles, base flat, lateral margins posteriorly sinuate and elevated, quite coarsely punctured and sparsely covered by setae; elytra smooth, striated, punctured, with intervals pubescent quite sparsely and finely for this genus, each with two transverse straw-coloured maculae extending from 4th to 8th interval… ventrites anteriorly not crenulated, [met]episterna square, slightly longer than wide” [from Latin]. “Length
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<quantity id="4CDE9B1D01553002FB64D614FA90FDD4" box="[1272,1357,551,575]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="508" unit="mm" value="23.0">23 mm</quantity>
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. In the proportions of head and thorax to the pterothorax similar to
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<taxonomicName id="4C264D7B01553002FCD3D678FBFFFD88" authority="Laferte" authorityName="Laferte" box="[847,1058,586,611]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="508" phylum="Arthropoda" rank="species" species="eximius">
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<emphasis id="B952EAEA01553002FCD3D678FC1AFD88" box="[847,967,587,611]" italics="true" pageId="4" pageNumber="508">C. eximius</emphasis>
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Laferté
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</taxonomicName>
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[=
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<taxonomicName id="4C264D7B01553002FBD8D678FA4DFD88" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[1092,1424,586,611]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="508" phylum="Arthropoda" rank="species" species="reflexus">
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<emphasis id="B952EAEA01553002FBD8D678FB63FD88" box="[1092,1214,586,611]" italics="true" pageId="4" pageNumber="508">C. reflexus</emphasis>
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(
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<bibRefCitation id="EFB74B0901553002FB4CD679FA5AFD88" author="Fabricius" box="[1232,1415,586,611]" pageId="4" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
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)
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</taxonomicName>
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], with which it also agrees in the moderate strangulation of the neck and in the suboval outline of the thorax, which has, however, more sinuated sides behind the middle, and distinct, almost rectangular, hind angles. The elytra, in the single example which appears not to be abraded, is nearly glabrous, the lateral interstices only having a few hairs; the striae are deep and punctured and the punctuation of the interstices is rather shallow and nowhere very dense: the fasciae are straw-yellow, narrow, and macular, the spot on the fifth interstice of both fasciae being much shorter than the others; on the hinder fascia the spot on the sixth interstice is extended forward and that on the seventh prolonged behind.” The coloration of
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<taxonomicName id="4C264D7B01553002FD32D774FCC3FCB4" box="[686,798,838,863]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="508" phylum="Arthropoda" rank="species" species="bonnyi">
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<emphasis id="B952EAEA01553002FD32D774FCC3FCB4" box="[686,798,838,863]" italics="true" pageId="4" pageNumber="508">C. bonnyi</emphasis>
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</taxonomicName>
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most closely resembles
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<taxonomicName id="4C264D7B01553002FBA0D774FA85FCB4" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[1084,1368,838,863]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="508" phylum="Arthropoda" rank="species" species="bozasi">
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<emphasis id="B952EAEA01553002FBA0D774FB7AFCB4" box="[1084,1191,838,863]" italics="true" pageId="4" pageNumber="508">C. bozasi</emphasis>
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<bibRefCitation id="EFB74B0901553002FB2DD775FA85FCB4" author="Alluaud" box="[1201,1368,838,863]" pageId="4" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
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</taxonomicName>
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, from which it differs by pronotal shape (more transverse, sharper and more projecting hind angles), elytral sculpture (striae more finely punctate), and the length and width of humeral maculae (Plate 3, cf. Figs 17, 18 and 19). From other species of the group near
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<taxonomicName id="4C264D7B01553002FE67D780FDA9FC20" box="[507,628,946,971]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="508" phylum="Arthropoda" rank="species" species="reflexus">
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<emphasis id="B952EAEA01553002FE67D780FDA9FC20" box="[507,628,946,971]" italics="true" pageId="4" pageNumber="508">C. reflexus</emphasis>
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</taxonomicName>
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it differs by shape of pronotal hind angles (sharp, more projecting) and shape of the maculae, namely humeral. The shape of the antennae (not flattened), front margins of ventrites (not crenulate) and the metepisterna place this species in the
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<taxonomicName id="4C264D7B01553002FC9AD7C8FCA3FBF8" box="[774,894,1018,1043]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="508" phylum="Arthropoda" rank="species" species="reflexus">
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<emphasis id="B952EAEA01553002FC9AD7C8FCA3FBF8" box="[774,894,1018,1043]" italics="true" pageId="4" pageNumber="508">C. reflexus</emphasis>
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</taxonomicName>
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group, near species with two orange maculae on each elytron (Plates 1, 3). The differential characters are given in the key to species presented below.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C657301553002FF5BD071FEB3FB2C" pageId="4" pageNumber="508" type="distribution">
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<paragraph id="8B9936F801553002FF5BD071FC35FBB0" blockId="4.[151,1437,151,1223]" box="[199,1000,1090,1115]" pageId="4" pageNumber="508">
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<emphasis id="B952EAEA01553002FF5BD071FEBDFBB0" bold="true" box="[199,352,1090,1115]" pageId="4" pageNumber="508">Distribution.</emphasis>
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Democratic
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|
<collectingCountry id="F331766801553002FE6DD070FD9DFBB0" box="[497,576,1091,1115]" name="Republic of the Congo" pageId="4" pageNumber="508">Congo</collectingCountry>
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: Orientale and Nord- Kivu Provinces.
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</paragraph>
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<paragraph id="8B9936F801553002FF5BD05BFEB3FB2C" blockId="4.[151,1437,151,1223]" pageId="4" pageNumber="508">
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<bibRefCitation id="EFB74B0901553002FF5BD05BFE4CFB94" author="Bates" box="[199,401,1127,1151]" pageId="4" pageNumber="525" refString="Bates, H. W. (1890) On some Coleopterous Insects collected by Mr. W. Bonny in the Aruwimi Valley. Proceedings Zoological Society London, 1890, 479 - 578." type="proceedings paper" year="1890">Bates (1890: 479)</bibRefCitation>
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noted. “Mr. Bonny informs me that they [specimens] were all taken at Yambuya Camp and on the march through the forest-region towards Albert Nyanza, between the months of
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<date id="FF98103801553002FB59D0B9FAB8FB48" box="[1221,1381,1162,1187]" pageId="4" pageNumber="508" value="1887-10">October 1887</date>
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and
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<date id="FF98103801553002FF0BD09DFE92FB2C" box="[151,335,1198,1223]" pageId="4" pageNumber="508" value="1888-11">November 1888</date>
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...”
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |