169 lines
21 KiB
XML
169 lines
21 KiB
XML
<document id="A15B757D4CF1DFC9C2EFE22F46A12880" ID-DOI="10.11646/zootaxa.4999.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5119246" ID-ZooBank="3E4F9A7C-CAA5-4B95-8345-836A63AB51B1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626872801607" checkinUser="plazi" docAuthor="Ahrens, Dirk, Liu, Wangang, Fabrizi, Silvia & Bai, Ming" docDate="2021" docId="038F87DF3330FFEEFF00FE6ACA4FF82E" docLanguage="en" docName="zootaxa.4999.4.4.pdf" docOrigin="Zootaxa 4999 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trioserica daweishana Ahrens, Liu & Fabrizi 2021, new species" docType="treatment" docVersion="3" lastPageNumber="351" masterDocId="FFB6FFA73338FFE6FF97FF2FCE6AFFDB" masterDocTitle="Taxonomic review on the Trioserica Moser, 1922 species of China (Coleoptera Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="355" masterPageNumber="343" pageNumber="351" updateTime="1699011907055" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7780559A9E803502A513FA0F3B95C244">Taxonomic review on the Trioserica Moser, 1922 species of China (Coleoptera Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:name id="03CCD934007EA5752F9C1DBD7CA44F4D" type="personal">
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<mods:namePart id="41F45D07C47F6F75F98CAE92CCE083A3">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="36E84280D556C57DD69E1CC80CE57243">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 160, 53113 Bonn, Germany.</mods:affiliation>
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<mods:namePart id="0F0B8EEDC39781118A67120FE4FBC8DE">Liu, Wangang</mods:namePart>
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<mods:affiliation id="076072F2C9457F590F1B1817962A6BEB">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 160, 53113 Bonn, Germany. & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Box 92, No. 1, Beichen West Road, Chaoyang District, Beijing, 100101, P. R. China & Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061 P. R. China</mods:affiliation>
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</mods:name>
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<mods:name id="8BE3751A237326EAA605CD0C39A43778" type="personal">
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<mods:roleTerm id="89A5F6C2E417F34D2F46450E7F0E193C">Author</mods:roleTerm>
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<mods:namePart id="007B858AD28DF2EDF1A7267811758036">Fabrizi, Silvia</mods:namePart>
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<mods:affiliation id="220FEB1F783FD7EE753A5B41989CEE35">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 160, 53113 Bonn, Germany.</mods:affiliation>
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<mods:namePart id="A6F665C00BC8C56C61F21D14CDDA774F">Bai, Ming</mods:namePart>
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<mods:affiliation id="A39FC794053B26A9C8B21D6F4D96B873">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Box 92, No. 1, Beichen West Road, Chaoyang District, Beijing, 100101, P. R. China & Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061 P. R. China</mods:affiliation>
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<mods:title id="B14F5A62CBAAB97BC1CA81120500D77D">Zootaxa</mods:title>
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<mods:part id="73A787839C4BCE5ED39BF0AE21CC337F">
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<mods:date id="8A72665EDCA6D18AD0EDB0ECE9006014">2021</mods:date>
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<mods:number id="18093B3BD017E678C8AA8FD0E9714E29">2021-07-13</mods:number>
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<mods:number id="86D468F2B4F80375A8A63E9B11F73FFB">4999</mods:number>
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<mods:classification id="5315AA8FDB0582BDDC9C211C11762873">journal article</mods:classification>
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<mods:identifier id="A565D959EED05079F63D986BF082089D" type="DOI">10.11646/zootaxa.4999.4.4</mods:identifier>
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<mods:identifier id="C7ED535F9DC476D1074197C458A678BE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038F87DF3330FFEEFF00FE6ACA4FF82E" ID-DOI="http://doi.org/10.5281/zenodo.5119258" ID-Zenodo-Dep="5119258" LSID="urn:lsid:plazi:treatment:038F87DF3330FFEEFF00FE6ACA4FF82E" httpUri="http://treatment.plazi.org/id/038F87DF3330FFEEFF00FE6ACA4FF82E" lastPageNumber="351" pageId="8" pageNumber="351">
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<subSubSection id="C33C65423330FFEEFF00FE6ACD0CFE84" box="[151,870,324,351]" pageId="8" pageNumber="351" type="nomenclature">
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<paragraph id="8B9936C93330FFEEFF00FE6ACD0CFE84" blockId="8.[151,870,324,387]" box="[151,870,324,351]" pageId="8" pageNumber="351">
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<heading id="D0D181A53330FFEEFF00FE6ACD0CFE84" bold="true" box="[151,870,324,351]" fontSize="11" level="1" pageId="8" pageNumber="351" reason="1">
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<emphasis id="B952EADB3330FFEEFF00FE6ACD0CFE84" bold="true" box="[151,870,324,351]" pageId="8" pageNumber="351">
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<taxonomicName id="4C264D4A3330FFEEFF00FE6ACCA0FE84" authority="Ahrens, Liu & Fabrizi" authorityName="Ahrens, Liu & Fabrizi" authorityYear="2021" box="[151,714,324,351]" class="Insecta" family="Melolonthidae" genus="Trioserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="daweishana" status="sp. nov.">
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<emphasis id="B952EADB3330FFEEFF00FE6ACFC2FE84" bold="true" box="[151,424,324,351]" italics="true" pageId="8" pageNumber="351">Trioserica daweishana</emphasis>
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Ahrens, Liu & Fabrizi
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26157A03330FFEEFD41FE6ACD0CFE84" box="[726,870,325,351]" pageId="8" pageNumber="351" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C65423330FFEEFF00FE46CF77FE58" box="[151,285,361,387]" pageId="8" pageNumber="351" type="description">
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<paragraph id="8B9936C93330FFEEFF00FE46CF77FE58" blockId="8.[151,870,324,387]" box="[151,285,361,387]" pageId="8" pageNumber="351">
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<figureCitation id="131D2A4C3330FFEEFF00FE46CF77FE58" box="[151,285,361,387]" captionStart="FIGURE 2" captionStartId="9.[151,250,1907,1931]" captionTargetBox="[211,1371,192,1868]" captionTargetId="figure-19@9.[199,1388,181,1883]" captionTargetPageId="9" captionText="FIGURE 2. A–D: Trioserica shiyiduiensis Ahrens, Liu & Fabrizi, new species (holotype); E–H: T. cangyuanensis Ahrens, Liu & Fabrizi, new species (holotype); I–L: T. daweishana Ahrens, Liu & Fabrizi, new species (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.5119252" httpUri="https://zenodo.org/record/5119252/files/figure.png" pageId="8" pageNumber="351">Figure 2I–L</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C65423330FFEEFF00FE84CCD1FE33" pageId="8" pageNumber="351" type="materials_examined">
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<paragraph id="8B9936C93330FFEEFF00FE84CCD1FE33" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">
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<emphasis id="B952EADB3330FFEEFF00FE84CFAEFE1F" bold="true" box="[151,452,426,453]" pageId="8" pageNumber="351">Type material examined.</emphasis>
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<materialsCitation id="3B4E3C943330FFEEFE5AFE84CCD1FE33" collectingDate="1956-06-20" collectionCode="IZAS" collectorName="Huang Keren" location="Pingbian" municipality="Mt. Daweishan" pageId="8" pageNumber="351" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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<typeStatus id="549D886B3330FFEEFE5AFE84CC52FE1E" box="[461,568,427,453]" pageId="8" pageNumber="351" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="9D20FD403330FFEEFDD1FE84CC34FE1E" box="[582,606,427,453]" pageId="8" pageNumber="351" type="male">♂</specimenCount>
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: “
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<collectingMunicipality id="6BFDACB33330FFEEFDE0FE84CD43FE1E" box="[631,809,427,453]" pageId="8" pageNumber="351">Mt. Daweishan</collectingMunicipality>
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,
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<location id="8EF960123330FFEEFCA1FE84CDF6FE1E" LSID="urn:lsid:plazi:treatment:038F87DF3330FFEEFF00FE6ACA4FF82E:8EF960123330FFEEFCA1FE84CDF6FE1E" box="[822,924,427,453]" municipality="Mt. Daweishan" name="Pingbian" pageId="8" pageNumber="351" stateProvince="Yunnan">Pingbian</location>
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,
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<collectingRegion id="49E2F82B3330FFEEFC3EFE84CA68FE1E" box="[937,1026,427,453]" country="China" name="Yunnan" pageId="8" pageNumber="351">Yunnan</collectingRegion>
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,
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<date id="FF9810093330FFEEFB98FE84CAF3FE1D" box="[1039,1177,427,454]" pageId="8" pageNumber="351" value="1956-06-20">
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<collectingDate id="EFDCE9E13330FFEEFB98FE84CAF3FE1D" box="[1039,1177,427,454]" pageId="8" pageNumber="351" value="1956-06-20">1956.VI.20.</collectingDate>
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</date>
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,
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<collectingMethod id="52674EDE3330FFEEFB31FE84CB7BFE1E" box="[1190,1297,427,453]" pageId="8" pageNumber="351">light trap</collectingMethod>
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, leg.
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<collectorName id="26D3531F3330FFEEFAC5FE84CEB1FE33" pageId="8" pageNumber="351">Huang Keren</collectorName>
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etc. / LW-560 /
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<taxonomicName id="4C264D4A3330FFEEFE06FEE0CC5FFE33" box="[401,565,462,488]" class="Insecta" family="Melolonthidae" genus="Lasioserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B952EADB3330FFEEFE06FEE0CC7EFE33" box="[401,532,463,488]" italics="true" pageId="8" pageNumber="351">Lasioserica</emphasis>
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sp
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</taxonomicName>
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86” (
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<collectionCode id="ED37AE0C3330FFEEFDFCFEE1CCC5FE33" box="[619,687,462,488]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="8" pageNumber="351">IZAS</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C65423330FFEEFF50FEDECD2DF8B3" pageId="8" pageNumber="351" type="description">
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<paragraph id="8B9936C93330FFEEFF50FEDECD96FD8A" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">
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<emphasis id="B952EADB3330FFEEFF50FEDECC37FDD0" bold="true" box="[199,605,497,523]" pageId="8" pageNumber="351">
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Description of the
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<typeStatus id="549D886B3330FFEEFE30FEDECC65FDD0" box="[423,527,497,523]" pageId="8" pageNumber="351" type="holotype">holotype</typeStatus>
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, male.
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</emphasis>
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Length:
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<quantity id="4CDE9B2C3330FFEEFD53FEDECD77FDD7" box="[708,797,497,524]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="351" unit="mm" value="8.5">8.5 mm</quantity>
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, length of elytra:
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<quantity id="4CDE9B2C3330FFEEFC70FEDECA2AFDD7" box="[999,1088,497,524]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="351" unit="mm" value="6.2">6.2 mm</quantity>
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, width:
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<quantity id="4CDE9B2C3330FFEEFB0AFEDECA9DFDD7" box="[1181,1271,497,524]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="351" unit="mm" value="4.9">4.9 mm</quantity>
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. Body oblong, yellowish brown, dorsal surface with small or larger dark spots, frons dark brown with greenish shine, antenna yellow, dorsal surface dull and glabrous except a few short white setae on elytra.
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</paragraph>
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<paragraph id="8B9936C93330FFEEFF50FD74CFD9FC55" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">
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Labroclypeus subtrapezoidal and wide, widest at base, lateral margins convex and convergent to strongly rounded anterior angles; margins weakly reflexed, anterior margin narrowly emarginate medially; lateral margin and ocular canthus producing a blunt angle; surface nearly flat, shiny, finely and densely punctate, distance between punctures less than their diameter, with a few short, erect setae. Frontoclypeal suture feebly incised and medially curved; smooth area in front of eye short and approximately twice as wide as long. Ocular canthus short and triangular, laterally sparsely punctate, with a single, short terminal seta. Frons dull, with fine and evenly dense punctures, with a few short setae beside eyes. Eyes small, ratio of diameter/ interocular width: 0.6. Antenna yellow, composed of 10 antennomeres; club with four lamellae, little longer than remaining antennomeres combined.
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<collectingRegion id="49E2F82B3330FFEEFA94FC7DCB08FCB7" box="[1283,1378,850,876]" country="Myanmar" name="Chin" pageId="8" pageNumber="351">Mentum</collectingRegion>
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convexly elevated anteriorly.
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</paragraph>
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<paragraph id="8B9936C93330FFEEFF50FCB7CD6BFBB9" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">Pronotum moderately wide, widest at base, lateral margins weakly but evenly convex and weakly convergent anteriorly; anterior angles distinctly produced and sharp; posterior angles blunt; anterior margin convexly produced medially, with a distinct and complete marginal line; basal margin without marginal line; surface with even, dense and fine punctures, posterior midline narrowly impunctate, punctures with microscopic setae; lateral margins sparsely setose. Hypomeron distinctly carinate. Scutellum narrow and short, with fine and dense punctures, medially broadly smooth, punctures with microscopic setae.</paragraph>
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<paragraph id="8B9936C93330FFEEFF50FB44CD93FB35" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">Elytra oblong, widest shortly behind middle; striae distinctly impressed, finely and densely punctate; intervals densely finely punctate, flat, odd intervals with a few fine and short setae, otherwise with minute setae in punctures; epipleural margin robust, ending at strongly curved external apical angle of elytra; epipleura sparsely setose; apical border narrowly membraneous, with a very fine rim of short microtrichomes.</paragraph>
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<paragraph id="8B9936C93330FFEEFF50FBD8CF2DFA19" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">Ventral surface dull, with large and dense punctures, sparsely and shortly setose, setae partly adpressed. Mesosternum between mesocoxae as wide as mesofemur, with irregularly scattered robust and long setae. Ratio of length of metepisternum/ metacoxa: 1/ 1.71. Metacoxa glabrous, laterally with numerous robust setae. Each abdominal sternite with indistinct transversal row of coarse punctures bearing each a robust seta between fine and dense punctation. Pygidium weakly convex, finely and densely punctate, without smooth midline, along apical margin with a few short setae.</paragraph>
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<paragraph id="8B9936C93330FFEEFF50FAE4CA81F89E" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">Legs narrow and moderately long, with shiny surface; femur with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur ventrally dull, sharply carinate anteriorly, without a submarginal serrated line; posterior margin straight, ventral posterior margin in distal half only weakly widened and smooth; dorsal margin smooth, with short setae. Metatibia moderately long and wide, widest shortly before middle; ratio width/ length: 1/ 3.5; dorsal margin completely sharply carinate, with two groups of spines; basal group of spines shortly before half of metatibial length, apical one at three quarters of metatibial length; lateral face longitudinally convex, with a few fine and very sparse punctures; ventral margin finely serrated, with four robust, equidistant spines; medial face impunctate, apex interiorly near tarsal articulation sharply truncate. Tarsomeres dorsally glabrous and sparsely finely punctate; pro- and mesotarsomeres ventrally with dense and short setae. Metatarsomeres ventrally glabrous, with a strongly serrated, longitudinal carina; metatarsomere 1 as long as following two tarsomeres combined and little longer than dorsal tibial spur. Protibia moderately long, tridentate; protarsal claws symmetric.</paragraph>
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<paragraph id="8B9936C93330FFEEFF50F861CD2DF8B3" blockId="8.[151,1437,426,2038]" box="[199,839,1870,1896]" pageId="8" pageNumber="351">
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Aedeagus:
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<figureCitation id="131D2A4C3330FFEEFED3F861CFDFF8B3" box="[324,437,1870,1896]" captionStart="FIGURE 2" captionStartId="9.[151,250,1907,1931]" captionTargetBox="[211,1371,192,1868]" captionTargetId="figure-19@9.[199,1388,181,1883]" captionTargetPageId="9" captionText="FIGURE 2. A–D: Trioserica shiyiduiensis Ahrens, Liu & Fabrizi, new species (holotype); E–H: T. cangyuanensis Ahrens, Liu & Fabrizi, new species (holotype); I–L: T. daweishana Ahrens, Liu & Fabrizi, new species (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.5119252" httpUri="https://zenodo.org/record/5119252/files/figure.png" pageId="8" pageNumber="351">Fig. 2I–K</figureCitation>
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. Habitus:
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<figureCitation id="131D2A4C3330FFEEFDB2F861CC19F8B3" box="[549,627,1870,1896]" captionStart="FIGURE 2" captionStartId="9.[151,250,1907,1931]" captionTargetBox="[211,1371,192,1868]" captionTargetId="figure-19@9.[199,1388,181,1883]" captionTargetPageId="9" captionText="FIGURE 2. A–D: Trioserica shiyiduiensis Ahrens, Liu & Fabrizi, new species (holotype); E–H: T. cangyuanensis Ahrens, Liu & Fabrizi, new species (holotype); I–L: T. daweishana Ahrens, Liu & Fabrizi, new species (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.5119252" httpUri="https://zenodo.org/record/5119252/files/figure.png" pageId="8" pageNumber="351">Fig. 2I</figureCitation>
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. Female unknown.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C65423330FFEEFF50F85ECFF3F809" pageId="8" pageNumber="351" type="diagnosis">
|
||
<paragraph id="8B9936C93330FFEEFF50F85ECFF3F809" blockId="8.[151,1437,426,2038]" pageId="8" pageNumber="351">
|
||
<emphasis id="B952EADB3330FFEEFF50F85ECF28F850" bold="true" box="[199,322,1905,1931]" pageId="8" pageNumber="351">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C264D4A3330FFEEFEDBF85DCC3AF850" authority="Ahrens & Liu & Fabrizi & Bai, 2021" authorityName="Ahrens & Liu & Fabrizi & Bai" authorityYear="2021" box="[332,592,1905,1931]" class="Insecta" family="Melolonthidae" genus="Trioserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="daweishana" status="sp. nov.">
|
||
<emphasis id="B952EADB3330FFEEFEDBF85DCC3AF850" box="[332,592,1905,1931]" italics="true" pageId="8" pageNumber="351">Trioserica daweishana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157A03330FFEEFDCCF85DCC8CF850" box="[603,742,1905,1932]" pageId="8" pageNumber="351" rank="species">new species</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C264D4A3330FFEEFC12F85DCA28F850" baseAuthorityName="Miyake & Yamaya" baseAuthorityYear="2001" box="[901,1090,1905,1931]" class="Insecta" family="Melolonthidae" genus="Trioserica" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="maculipennis">
|
||
<emphasis id="B952EADB3330FFEEFC12F85DCA28F850" box="[901,1090,1905,1931]" italics="true" pageId="8" pageNumber="351">Tr. maculipennis</emphasis>
|
||
</taxonomicName>
|
||
by the basally more strongly curved left paramere, which is in apical half nearly straight rather than curved (lateral view), as well as by the more narrow right paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C65423330FFEEFF50F8F3CA4FF82E" box="[199,1061,2011,2038]" pageId="8" pageNumber="351" type="etymology">
|
||
<paragraph id="8B9936C93330FFEEFF50F8F3CA4FF82E" blockId="8.[151,1437,426,2038]" box="[199,1061,2011,2038]" pageId="8" pageNumber="351">
|
||
<emphasis id="B952EADB3330FFEEFF50F8F3CF24F82D" bold="true" box="[199,334,2012,2038]" pageId="8" pageNumber="351">Etymology.</emphasis>
|
||
The new species is named after its
|
||
<typeStatus id="549D886B3330FFEEFD4CF8F3CD61F82D" box="[731,779,2012,2038]" pageId="8" pageNumber="351">type</typeStatus>
|
||
locality, Mt. Daweishan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |