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<document id="BB9A1881CC54C8AC54F54D17A7A8CB56" ID-CLB-Dataset="299020" ID-DOI="10.5252/zoosystema2024v46a14" ID-GBIF-Dataset="b5619e29-fcbd-4ddc-9231-138300251568" ID-ISSN="1638-9387" ID-Zenodo-Dep="12548683" ID-ZooBank="23D972D0-F573-429C-816B-F42AFF02D0B1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1719412818962" checkinUser="felipe" docAuthor="Walter, Angelo Vannozzi &amp; Renda, Walter" docDate="2024" docId="038F87DDFF8CFFE22015FF2C3DF5CCF6" docLanguage="en" docName="zoosystema.2024.46.14.pdf" docOrigin="Zoosystema 46 (14)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Caecum miliarium Walter &amp; Renda 2024, n. sp." docType="treatment" docVersion="6" lastPageNumber="336" masterDocId="FFB6FFA5FF87FFE92128FFFB396CCE26" masterDocTitle="Sampling the depth: New data on the Caecidae (Mollusca, Gastropoda) from northeastern Papua New Guinea" masterLastPageNumber="347" masterPageNumber="327" pageNumber="336" updateTime="1719441926131" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F175CC42736446733F2BD927C46D283A">Sampling the depth: New data on the Caecidae (Mollusca, Gastropoda) from northeastern Papua New Guinea</mods:title>
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<mods:name id="A4D6570DA9038357A111947EDB82FC4F" type="personal">
<mods:role id="031377CC7A711137EB1D64AE004C0C89">
<mods:roleTerm id="8C6832AF5F2BFE1BAB04A99FA458DEF9">Author</mods:roleTerm>
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<mods:namePart id="DD3B5682A97EB624E3CB1F0F46F288D2">Walter, Angelo Vannozzi</mods:namePart>
<mods:affiliation id="DA9B72BD52CA7A087D3CBA253D2CF05E">Via M. L. Longo 8, 00151 Rome (Italy)</mods:affiliation>
<mods:nameIdentifier id="6B120724F2202EE1EF5E94E69DE52470" type="email">ang.vannozzi@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B5839F1C20B87326AF641358720A4B98">Renda, Walter</mods:namePart>
<mods:affiliation id="221C51614FB5A6568A816DB1170E0C9A">Via Bologna 18 / A, 87032 Amantea (CS) (Italy)</mods:affiliation>
<mods:nameIdentifier id="25EE28E9F23ACEB9C1B94D1EE0523EE4" type="email">w.renda1@tin.it</mods:nameIdentifier>
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<mods:title id="BA28BB88F7E8BA3C4CBAF6E2204808AC">Zoosystema</mods:title>
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<mods:part id="8DBC10D06401AD64C44585A403FCB004">
<mods:date id="CDAD564A355B32209D15A27B6B7C4198">2024</mods:date>
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<mods:number id="067630BEC0981848834222B52FD308BB">2024-06-07</mods:number>
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<mods:identifier id="337272E3FE59976D75F8402E2A339047" type="CLB-Dataset">299020</mods:identifier>
<mods:identifier id="EFFB6DE7D5A2B5AF255EB3C85C225E5A" type="DOI">10.5252/zoosystema2024v46a14</mods:identifier>
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<treatment id="038F87DDFF8CFFE22015FF2C3DF5CCF6" ID-DOI="http://doi.org/10.5281/zenodo.12554878" ID-GBIF-Taxon="233799623" ID-Zenodo-Dep="12554878" LSID="urn:lsid:plazi:treatment:038F87DDFF8CFFE22015FF2C3DF5CCF6" httpUri="http://treatment.plazi.org/id/038F87DDFF8CFFE22015FF2C3DF5CCF6" lastPageNumber="336" pageId="11" pageNumber="336">
<subSubSection id="C33C6540FF8CFFE22015FF2C3B22CED4" box="[317,590,215,242]" pageId="11" pageNumber="336" type="nomenclature">
<paragraph id="8B9936CBFF8CFFE22015FF2C3B22CED4" blockId="11.[317,590,215,274]" box="[317,590,215,242]" pageId="11" pageNumber="336">
<heading id="D0D181A7FF8CFFE22015FF2C3B22CED4" box="[317,590,215,242]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="336" reason="2">
<taxonomicName id="4C264D48FF8CFFE22015FF2C3B60CED4" authority="Walter &amp; Renda, 2024" authorityName="Walter &amp; Renda" authorityYear="2024" box="[317,524,215,242]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="miliarium" status="sp. nov.">
<emphasis id="B952EAD9FF8CFFE22015FF2C3B60CED4" bold="true" box="[317,524,215,242]" italics="true" pageId="11" pageNumber="336">Caecum miliarium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157A2FF8CFFE2233BFF233B22CED4" box="[531,590,216,242]" pageId="11" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C6540FF8CFFE220B6FF0C3B9ECF60" pageId="11" pageNumber="336" type="description">
<paragraph id="8B9936CBFF8CFFE220B6FF0C3882CF34" blockId="11.[317,590,215,274]" box="[414,494,247,274]" pageId="11" pageNumber="336">
(
<figureCitation id="131D2A4EFF8CFFE2208DFF0C3888CF34" box="[421,484,247,274]" captionStart="FIG" captionStartId="12.[133,144,1984,2002]" captionTargetBox="[132,1456,215,1895]" captionTargetId="figure-19@12.[132,1456,215,1895]" captionTargetPageId="12" captionText="FIG. 5. — Caecum miliarium n. sp.:A-D, holotype MNHN-IM-2000-38946; E-I, paratypes MNHN-IM-2000-38947; E, growth stage; F, specimen with partially worn temporary septum; H-I, postlarval specimen, different views. Scale bar: 1 mm, details not to scale." figureDoi="http://doi.org/10.5281/zenodo.12548696" httpUri="https://zenodo.org/record/12548696/files/figure.png" pageId="11" pageNumber="336">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="8B9936CBFF8CFFE221B1FECF3B9ECF60" blockId="11.[153,754,308,326]" box="[153,754,308,326]" pageId="11" pageNumber="336">
<uri id="FFB73AC9FF8CFFE221B1FECF3B9ECF60" box="[153,754,308,326]" pageId="11" pageNumber="336">
urn:lsid:zoobank.org:act:
<uuid id="FF800C1EFF8CFFE22076FECF3B9ECF60" box="[350,754,308,326]" pageId="11" pageNumber="336">8FFDAF1B-E8DE-435D-B8AF-4953A5C66CBC</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C33C6540FF8CFFE221ACFE9C3880CC6B" pageId="11" pageNumber="336" type="materials_examined">
<paragraph id="8B9936CBFF8CFFE221ACFE9C3840CFEB" blockId="11.[132,777,358,515]" pageId="11" pageNumber="336">
<materialsCitation id="3B4E3C96FF8CFFE221ACFE9C3840CFEB" ID-GBIF-Occurrence="4899671322" collectingDate="2014-08-28" collectionCode="MNHN-" collectorName="KAVIENG" country="Papua New Guinea" elevation="338" latitude="-2.45" location="New Ireland" longLatPrecision="1307" longitude="150.66667" pageId="11" pageNumber="336" specimenCode="MNHN- IM-2000-38946" specimenCount="1" stateProvince="New Ireland" typeStatus="holotype">
<smallCapsWord id="8D7FA017FF8CFFE221ACFE9C39DACF5A" baselines="376,376" box="[132,182,359,382]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="11" pageNumber="336">TYPE</smallCapsWord>
<smallCapsWord id="8D7FA017FF8CFFE22197FE973873CF5B" baselines="377" box="[191,287,364,381]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="11" pageNumber="336">MATERIAL</smallCapsWord>
. —
<emphasis id="B952EAD9FF8CFFE22067FE9C38DCCF59" bold="true" box="[335,432,359,383]" pageId="11" pageNumber="336">
<typeStatus id="549D8869FF8CFFE22067FE9C38DCCF59" box="[335,432,359,383]" pageId="11" pageNumber="336" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B952EAD9FF8CFFE22096FE9C3BEBCFA6" bold="true" box="[446,647,359,384]" pageId="11" pageNumber="336">
<collectingCountry id="F331765BFF8CFFE22096FE9C3BEBCFA6" box="[446,647,359,384]" name="Papua New Guinea" pageId="11" pageNumber="336">Papua New Guinea</collectingCountry>
</emphasis>
• sh (length
<quantity id="4CDE9B2EFF8CFFE221AEFE7A39BACFBE" box="[134,214,385,409]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="336" unit="mm" value="1.3">1.3 mm</quantity>
,
<figureCitation id="131D2A4EFF8CFFE221C8FE7A387ACFBF" box="[224,278,385,409]" captionStart="FIG" captionStartId="12.[133,144,1984,2002]" captionTargetBox="[132,1456,215,1895]" captionTargetId="figure-19@12.[132,1456,215,1895]" captionTargetPageId="12" captionText="FIG. 5. — Caecum miliarium n. sp.:A-D, holotype MNHN-IM-2000-38946; E-I, paratypes MNHN-IM-2000-38947; E, growth stage; F, specimen with partially worn temporary septum; H-I, postlarval specimen, different views. Scale bar: 1 mm, details not to scale." figureDoi="http://doi.org/10.5281/zenodo.12548696" httpUri="https://zenodo.org/record/12548696/files/figure.png" pageId="11" pageNumber="336">Fig. 5</figureCitation>
A-D);
<collectingRegion id="49E2F829FF8CFFE2207FFE7A38BFCFBF" box="[343,467,385,409]" country="Papua New Guinea" name="New Ireland" pageId="11" pageNumber="336">New Ireland</collectingRegion>
,
<collectorName id="26D3531DFF8CFFE220F5FE7A3B27CFBF" box="[477,587,385,409]" pageId="11" pageNumber="336">KAVIENG</collectorName>
2014 Stn CP4418;
<geoCoordinate id="EE12500CFF8CFFE221ACFE6039A5CF95" box="[132,201,411,435]" degrees="2" direction="south" minutes="27" orientation="latitude" pageId="11" pageNumber="336" precision="925" value="-2.45">2°27S</geoCoordinate>
,
<geoCoordinate id="EE12500CFF8CFFE221FFFE60385BCF95" box="[215,311,411,435]" degrees="150" direction="east" minutes="40" orientation="longitude" pageId="11" pageNumber="336" precision="925" value="150.66667">150°40E</geoCoordinate>
;
<elevation id="000BD1F8FF8CFFE2206FFE603B6ECF95" box="[327,514,411,435]" metricMagnitude="2" metricUnit="m" metricValue="3.375" metricValueMax="3.4" metricValueMin="3.35" pageId="11" pageNumber="336" unit="m" value="337.5" valueMax="340.0" valueMin="335.0">
<quantity id="4CDE9B2EFF8CFFE2206FFE6038D7CF95" box="[327,443,411,435]" metricMagnitude="2" metricUnit="m" metricValue="3.375" metricValueMax="3.4" metricValueMin="3.35" pageId="11" pageNumber="336" unit="m" value="337.5" valueMax="340.0" valueMin="335.0">335-340 m</quantity>
depth
</elevation>
;
<date id="FF98100BFF8CFFE22327FE603BF0CF95" box="[527,668,411,435]" pageId="11" pageNumber="336" value="2014-08-28">
<collectingDate id="EFDCE9E3FF8CFFE22327FE603BF0CF95" box="[527,668,411,435]" pageId="11" pageNumber="336" value="2014-08-28">28.VIII.2014</collectingDate>
</date>
;
<specimenCode id="DB809EB0FF8CFFE22380FE67384ACFE8" pageId="11" pageNumber="336">MNHN-IM-2000-38946</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9936CBFF8CFFE221ACFE2B3B60CC24" blockId="11.[132,777,358,515]" pageId="11" pageNumber="336">
<materialsCitation id="3B4E3C96FF8CFFE221ACFE2B3B6ACC25" ID-GBIF-Occurrence="4899671385" collectingDate="2014-08-28" collectionCode="MNHN-" collectorName="Stn CP" country="Papua New Guinea" elevation="338" latitude="-2.45" location="Papua New Guinea" longLatPrecision="1307" longitude="150.66667" pageId="11" pageNumber="336" specimenCode="MNHN-IM-2000-38947" specimenCount="116" stateProvince="New Ireland" typeStatus="paratype">
<emphasis id="B952EAD9FF8CFFE221ACFE2B38DACFCC" bold="true" box="[132,438,464,490]" pageId="11" pageNumber="336">
<typeStatus id="549D8869FF8CFFE221ACFE2B398ACFCE" box="[132,230,464,488]" pageId="11" pageNumber="336" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F331765BFF8CFFE221DAFE2B38DACFCC" box="[242,438,464,490]" name="Papua New Guinea" pageId="11" pageNumber="336">Papua New Guinea</collectingCountry>
</emphasis>
<specimenCount id="9D20FD42FF8CFFE220E6FE2B3898CFCE" box="[462,500,464,488]" pageId="11" pageNumber="336">106</specimenCount>
sh (
<specimenCount id="9D20FD42FF8CFFE22308FE2B3B56CFCE" box="[544,570,464,488]" pageId="11" pageNumber="336">10</specimenCount>
larval); same data as for holotype;
<specimenCode id="DB809EB0FF8CFFE22021FE103B6ACC25" box="[265,518,491,515]" pageId="11" pageNumber="336">MNHN-IM-2000-38947</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9936CBFF8CFFE221ACFDE03880CC6B" blockId="11.[131,776,538,589]" pageId="11" pageNumber="336">
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<smallCapsWord id="8D7FA017FF8CFFE22191FDDB387FCC17" baselines="557" box="[185,275,544,561]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="11" pageNumber="336">LOCALITY</smallCapsWord>
. — Pacific Ocean,
<collectingCountry id="F331765BFF8CFFE220E0FDE03BEECC14" box="[456,642,538,562]" name="Papua New Guinea" pageId="11" pageNumber="336">Papua New Guinea</collectingCountry>
,
<collectingRegion id="49E2F829FF8CFFE223A3FDE03A68CC14" box="[651,772,538,562]" country="Papua New Guinea" name="New Ireland" pageId="11" pageNumber="336">New Ireland</collectingRegion>
;
<geoCoordinate id="EE12500CFF8CFFE221ABFDCE39A9CC6B" box="[131,197,565,589]" degrees="2" direction="south" minutes="27" orientation="latitude" pageId="11" pageNumber="336" precision="925" value="-2.45">2°27S</geoCoordinate>
,
<geoCoordinate id="EE12500CFF8CFFE221E7FDCE3847CC6B" box="[207,299,565,589]" degrees="150" direction="east" minutes="40" orientation="longitude" pageId="11" pageNumber="336" precision="925" value="150.66667">150°40E</geoCoordinate>
;
<quantity id="4CDE9B2EFF8CFFE22010FDCE38CACC6A" box="[312,422,565,589]" metricMagnitude="2" metricUnit="m" metricValue="3.375" metricValueMax="3.4" metricValueMin="3.35" pageId="11" pageNumber="336" unit="m" value="337.5" valueMax="340.0" valueMin="335.0">335-340 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C33C6540FF8CFFE221ACFD9E3BF9CC5A" box="[132,661,612,637]" pageId="11" pageNumber="336" type="distribution">
<paragraph id="8B9936CBFF8CFFE221ACFD9E3BF9CC5A" blockId="11.[132,661,612,637]" box="[132,661,612,637]" pageId="11" pageNumber="336">
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. — Known only from the
<typeStatus id="549D8869FF8CFFE22331FD9E3B2FCC5B" box="[537,579,613,637]" pageId="11" pageNumber="336">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33C6540FF8CFFE221ACFD6E3990CCDD" pageId="11" pageNumber="336" type="etymology">
<paragraph id="8B9936CBFF8CFFE221ACFD6E3990CCDD" blockId="11.[132,777,660,763]" pageId="11" pageNumber="336">
<smallCapsWord id="8D7FA017FF8CFFE221ACFD6E3992CC8D" baselines="678,679" box="[132,254,661,684]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="11" pageNumber="336">ETYMOLOGY</smallCapsWord>
. — From the Latin adjective
<emphasis id="B952EAD9FF8CFFE22303FD6F3BECCC8A" box="[555,640,660,684]" italics="true" pageId="11" pageNumber="336">miliarius</emphasis>
(meaning: of a thousand, containing a thousand) because the sample collected in the
<typeStatus id="549D8869FF8CFFE221EDFD323983CCC7" box="[197,239,713,737]" pageId="11" pageNumber="336">type</typeStatus>
locality contained a great number of specimens of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C33C6540FF8CFFE221ACFCCD38FBCB88" pageId="11" pageNumber="336" type="description">
<paragraph id="8B9936CBFF8CFFE221ACFCCD3879CD68" blockId="11.[130,778,822,1454]" box="[132,277,822,848]" pageId="11" pageNumber="336">
<smallCapsWord id="8D7FA017FF8CFFE221ACFCCD3879CD68" baselines="841,841" box="[132,277,822,848]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="11" pageNumber="336">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9936CBFF8CFFE221ACFCAE38FBCB88" blockId="11.[130,778,822,1454]" pageId="11" pageNumber="336">
Shell small for the genus, smooth, colourless, semitransparent, somewhat iridescent. Protoconch planorbid, multispiral of
<emphasis id="B952EAD9FF8CFFE221ACFC6D39F9CD89" box="[132,149,918,943]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
2 whorls, smooth and shiny, the last portion detached and straight, ending with a simple aperture with slightly reflected lip. Teleoconch tubular, arched, smooth. Early teleoconch subcylindrical, slightly twisted. Adult shell subcylindrical, slightly dextrally twisted in ventral view. Surface covered by very fine wavy longitudinal and discontinuous threads, evenly spaced but becoming more and more numerous and crowded going from the apex to the aperture. Macula none detected. Aperture somewhat inclined, without a clear swelling and ending with a fine reflected lip. Septum low dome shaped without mucro, slightly recessed below the cutting plane. Temporary septum thick, opaque, hemispherical, persistent, lying on the permanent one. Young stage with flatter and more recessed septum. Periostracum none detected. Operculum and soft parts unknown.
<typeStatus id="549D8869FF8CFFE223BBFAA83A6FCB4B" box="[659,771,1363,1389]" pageId="11" pageNumber="336" type="holotype">Holotype</typeStatus>
: length
<quantity id="4CDE9B2EFF8CFFE221FCFA883843CBA8" box="[212,303,1395,1422]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="336" unit="mm" value="1.3">1.3 mm</quantity>
, max diam
<quantity id="4CDE9B2EFF8CFFE22099FA883B77CBA8" box="[433,539,1395,1422]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="336" unit="mm" value="0.25">0.25 mm</quantity>
, min diam
<quantity id="4CDE9B2EFF8CFFE223B3FA883A6ACBA8" box="[667,774,1395,1422]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="336" unit="mm" value="0.2">0.20 mm</quantity>
. Protoconch size 395 µm.
</paragraph>
</subSubSection>
<subSubSection id="C33C6540FF8CFFE221ACFA283DF5CCF6" pageId="11" pageNumber="336" type="discussion">
<paragraph id="8B9936CBFF8CFFE221ACFA283984CBCD" blockId="11.[130,776,1491,2027]" box="[132,232,1491,1517]" pageId="11" pageNumber="336">
<smallCapsWord id="8D7FA017FF8CFFE221ACFA283984CBCD" baselines="1510,1510" box="[132,232,1491,1517]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="11" pageNumber="336">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8B9936CBFF8CFFE221ACFA0838B6C9AD" blockId="11.[130,776,1491,2027]" pageId="11" pageNumber="336">
Among the characteristics of this species, a rather persistent temporary septum can be highlighted. It is entirely preserved in the
<typeStatus id="549D8869FF8CFFE221EDF9C9384FC86A" box="[197,291,1586,1612]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
and partially preserved in the
<typeStatus id="549D8869FF8CFFE22377F9C83BD7C86B" box="[607,699,1587,1613]" pageId="11" pageNumber="336" type="paratype">paratype</typeStatus>
shown in
<figureCitation id="131D2A4EFF8CFFE2218AF9A93862C84B" box="[162,270,1618,1645]" captionStart="FIG" captionStartId="12.[133,144,1984,2002]" captionTargetBox="[132,1456,215,1895]" captionTargetId="figure-19@12.[132,1456,215,1895]" captionTargetPageId="12" captionText="FIG. 5. — Caecum miliarium n. sp.:A-D, holotype MNHN-IM-2000-38946; E-I, paratypes MNHN-IM-2000-38947; E, growth stage; F, specimen with partially worn temporary septum; H-I, postlarval specimen, different views. Scale bar: 1 mm, details not to scale." figureDoi="http://doi.org/10.5281/zenodo.12548696" httpUri="https://zenodo.org/record/12548696/files/figure.png" pageId="11" pageNumber="336">
Figure
<specimenCount id="9D20FD42FF8CFFE221D9F9A9387EC84B" box="[241,274,1618,1645]" count="5" pageId="11" pageNumber="336" type="generic">5F.</specimenCount>
</figureCitation>
The presence of numerous specimens in the type locality, including the complete growth series, allowed a clear and exhaustive knowledge of shell ontogeny of the new species. In fact, except for
<specimenCount id="9D20FD42FF8CFFE220B2F9493B5BC8EA" box="[410,567,1714,1740]" count="3" pageId="11" pageNumber="336" type="juvenile">three juveniles</specimenCount>
, likely belonging to
<taxonomicName id="4C264D48FF8CFFE221ABF9293812C8CA" authority="Vannozzi, 2019" authorityName="Vannozzi" authorityYear="2019" box="[131,382,1746,1772]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="frugi">
<emphasis id="B952EAD9FF8CFFE221ABF92939BEC8CA" box="[131,210,1746,1772]" italics="true" pageId="11" pageNumber="336">C. frugi</emphasis>
Vannozzi, 2019
</taxonomicName>
, the lot originating from Stn CP4418 contained only specimens
<taxonomicName id="4C264D48FF8CFFE22088F9093B47C92A" authority="Walter &amp; Renda, 2024" authorityName="Walter &amp; Renda" authorityYear="2024" box="[416,555,1778,1804]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="miliarium" status="n. sp.">
<emphasis id="B952EAD9FF8CFFE22088F9093B47C92A" box="[416,555,1778,1804]" italics="true" pageId="11" pageNumber="336">C. miliarium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157A2FF8CFFE2231CF9093B1CC92A" box="[564,624,1778,1804]" pageId="11" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
Furthermore, the presence of all growth stages, all well preserved, suggests that the actual habitat was sampled during the expedition and that this species lives in the aphotic zone, unlike the majority of other members of the family.
</paragraph>
<paragraph id="8B9936CBFF8CFFE221B3F86A3C0BCFF6" blockId="11.[130,776,1491,2027]" lastBlockId="11.[811,1456,215,720]" pageId="11" pageNumber="336">
The new species can be compared with
<taxonomicName id="4C264D48FF8CFFE2231AF8693BD7C98D" authorityName="Vannozzi" authorityYear="2019" box="[562,699,1937,1964]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="frugi">
<emphasis id="B952EAD9FF8CFFE2231AF8693BD7C98D" box="[562,699,1937,1964]" italics="true" pageId="11" pageNumber="336">Caecum frugi</emphasis>
</taxonomicName>
, which shows a similar size and septum. However,
<taxonomicName id="4C264D48FF8CFFE22368F8493BA4C9ED" authority="Walter &amp; Renda, 2024" authorityName="Walter &amp; Renda" authorityYear="2024" box="[576,712,1969,1996]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="miliarium" status="n. sp.">
<emphasis id="B952EAD9FF8CFFE22368F8493BA4C9ED" box="[576,712,1969,1996]" italics="true" pageId="11" pageNumber="336">C. miliarium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157A2FF8CFFE223E7F8493A6BC9EA" box="[719,775,1970,1996]" pageId="11" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
is readily distinguished by the wider tube and the longitudinal microsculpture visible also at low magnification, whereas in
<taxonomicName id="4C264D48FF8CFFE22205FF033A11CF37" authorityName="Vannozzi" authorityYear="2019" box="[813,893,247,274]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="frugi">
<emphasis id="B952EAD9FF8CFFE22205FF033A11CF37" box="[813,893,247,274]" italics="true" pageId="11" pageNumber="336">C. frugi</emphasis>
</taxonomicName>
the microsculpture is denser and visible only at very high magnification. Furthermore,
<taxonomicName id="4C264D48FF8CFFE22588FEEC3D9DCF17" authorityName="Vannozzi" authorityYear="2019" box="[1184,1265,279,305]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="frugi">
<emphasis id="B952EAD9FF8CFFE22588FEEC3D9DCF17" box="[1184,1265,279,305]" italics="true" pageId="11" pageNumber="336">C. frugi</emphasis>
</taxonomicName>
shows a different shape of the tube, more cylindrical and with inflated abapical portion. A similar microsculpture is shown also by
<taxonomicName id="4C264D48FF8CFFE22477FEAC3D39CFB7" authority="de Folin, 1868" authorityName="de Folin" authorityYear="1868" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="neocaledonicum">
<emphasis id="B952EAD9FF8CFFE22477FEAC3AC2CFB7" italics="true" pageId="11" pageNumber="336">C. neocaledonicum</emphasis>
de Folin, 1868
</taxonomicName>
, described from
<collectingCountry id="F331765BFF8CFFE22420FE8C3CC3CFB7" box="[1288,1455,375,401]" name="New Caledonia" pageId="11" pageNumber="336">New Caledonia</collectingCountry>
and occurring in the whole IWP, but this species is larger and otherwise different in several aspects as shown below.
</paragraph>
<paragraph id="8B9936CBFF8CFFE2226BFE2C3DF5CCF6" blockId="11.[811,1456,215,720]" pageId="11" pageNumber="336">
A further comparison can be made with
<taxonomicName id="4C264D48FF8CFFE225DEFE2C3A04CC37" authority="Odhner, 1925" authorityName="Odhner" authorityYear="1925" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="suteri">
<emphasis id="B952EAD9FF8CFFE225DEFE2C3C21CFD6" box="[1270,1357,471,497]" italics="true" pageId="11" pageNumber="336">C. suteri</emphasis>
Odhner, 1925
</taxonomicName>
, a small species described from
<collectingCountry id="F331765BFF8CFFE2259AFE0C3C28CC36" box="[1202,1348,502,529]" name="New Zealand" pageId="11" pageNumber="336">New Zealand</collectingCountry>
, currently in the synonymy of
<taxonomicName id="4C264D48FF8CFFE22528FDEC3C35CC16" authority="Hedley, 1904" authorityName="Hedley" authorityYear="1904" box="[1024,1369,534,561]" class="Gastropoda" family="Caecidae" genus="Caecum" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="digitulum">
<emphasis id="B952EAD9FF8CFFE22528FDEC3DADCC16" box="[1024,1217,534,561]" italics="true" pageId="11" pageNumber="336">Caecum digitulum</emphasis>
Hedley, 1904
</taxonomicName>
. In fact, although this species differs by the small finger-like mucro, absent in the new species, it shares a quite similar protoconch showing about the same number of whorls and a prolonged straight portion before the transition with the teleoconch (
<bibRefCitation id="EFB74B3AFF8CFFE2221FFD4E3A94CCF6" author="ODHNER N. H." box="[823,1016,693,720]" pageId="11" pageNumber="336" pagination="1 - 90" refId="ref17164" refString="ODHNER N. H. 1924. - Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 16. XIX. New Zealand Mollusca. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Kobenhavn 77: 1 - 90, pls 1, 2. https: // www. biodiversitylibrary. org / item / 88784" type="journal article" year="1924">Odhner 1924: 29</bibRefCitation>
; pl. 1; fig. 19).
</paragraph>
</subSubSection>
</treatment>
</document>