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<document id="46245DC4CCF90672D849AC2EA645DDA9" ID-CLB-Dataset="3791" ID-DOI="10.11646/zootaxa.5060.1.1" ID-GBIF-Dataset="40f89bbe-7f7b-4abd-aa02-a45e410da232" ID-ISSN="1175-5326" ID-Zenodo-Dep="5606671" ID-ZooBank="14126B35-AEEA-40F5-A892-FE6A8D64E5D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1635406161957" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2021" docId="038F87D5FF90FFBDFF6D21C5FE989BA7" docLanguage="en" docName="zootaxa.5060.1.1.pdf" docOrigin="Zootaxa 5060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pacifiosoma triangulatum Mikhaljova 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="33" masterDocId="FFB6FFADFF8FFF9DFFFA2217FF8C9A76" masterDocTitle="The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)" masterLastPageNumber="44" masterPageNumber="1" pageNumber="32" updateTime="1699419728057" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4EB5D1B7BA07FF6B75C17199FFA66E1A">The millipede family Diplomaragnidae Attems, 1907 in the Asian part of Russia with the descriptions of two new genera and seven new species (Diplopoda, Chordeumatida)</mods:title>
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<mods:namePart id="FD4390FFEB92C8C8CA1DB1B1CFEE6B18">Mikhaljova, Elena V.</mods:namePart>
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<mods:date id="4CB923DCB8731B6B1730B624C05D85ED">2021</mods:date>
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<treatment id="038F87D5FF90FFBDFF6D21C5FE989BA7" ID-DOI="http://doi.org/10.5281/zenodo.5606683" ID-GBIF-Taxon="189548892" ID-Zenodo-Dep="5606683" LSID="urn:lsid:plazi:treatment:038F87D5FF90FFBDFF6D21C5FE989BA7" httpUri="http://treatment.plazi.org/id/038F87D5FF90FFBDFF6D21C5FE989BA7" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C33C6548FF90FF82FF6D21C5FDB5999B" box="[151,569,978,1005]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B9936C3FF90FF82FF6D21C5FDB5999B" blockId="31.[151,569,978,1041]" box="[151,569,978,1005]" pageId="31" pageNumber="32">
<heading id="D0D181AFFF90FF82FF6D21C5FDB5999B" bold="true" box="[151,569,978,1005]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B952EAD1FF90FF82FF6D21C5FDB5999B" bold="true" box="[151,569,978,1005]" pageId="31" pageNumber="32">
<taxonomicName id="4C264D40FF90FF82FF6D21C5FE58999B" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[151,468,978,1005]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="triangulatum" status="sp. nov.">
<emphasis id="B952EAD1FF90FF82FF6D21C5FE58999B" bold="true" box="[151,468,978,1005]" italics="true" pageId="31" pageNumber="32">Pacifiosoma triangulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157AAFF90FF82FE2621C4FDB5999B" box="[476,569,979,1005]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF90FF82FF6D21E0FE989E67" box="[151,276,1014,1041]" pageId="31" pageNumber="32" type="description">
<paragraph id="8B9936C3FF90FF82FF6D21E0FE989E67" blockId="31.[151,569,978,1041]" box="[151,276,1014,1041]" pageId="31" pageNumber="32">
<figureCitation id="131D2A46FF90FF82FF6D21E0FE989E67" box="[151,276,1014,1041]" captionStart="FIGURES 5657" captionStartId="32.[151,264,1038,1062]" captionTargetBox="[158,1432,489,1012]" captionTargetId="figure-134@32.[151,1436,489,1015]" captionTargetPageId="32" captionText="FIGURES 5657. Pacifiosoma triangulatum sp. nov., male holotype. 56, gonopods, caudal view; 57, gonopods, front view; pp, posterior angicoxal process; c, clawlike outgrowth; n, triangular plate; ap, anterior angiocoxal process; cx, coxosternum. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.5606713" httpUri="https://zenodo.org/record/5606713/files/figure.png" pageId="31" pageNumber="32">Figs 5657</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF90FF82FF6D2629FA989EB3" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B9936C3FF90FF82FF6D2629FA989EB3" blockId="31.[151,1437,1086,2013]" pageId="31" pageNumber="32">
<emphasis id="B952EAD1FF90FF82FF6D2629FE089E2E" bold="true" box="[151,388,1086,1112]" pageId="31" pageNumber="32">Material examined.</emphasis>
<materialsCitation id="3B4E3C9EFF90FF82FE752628FA989EB3" ID-GBIF-Occurrence="3395035345" collectingDate="1972-09" collectionCode="FSCB" collectorName="A. N. Kupyanskaya &amp; A. N. Kupyanskaya." location="Pimorsky Krai" municipality="Dalnegorsky District" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549D8861FF90FF82FE752628FE709E2F" box="[399,508,1087,1113]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="ED37AE06FF90FF82FDA92628FD119E2F" box="[595,669,1087,1113]" pageId="31" pageNumber="32">FSCB</collectionCode>
: 2 permanent micropreparations, N 351352: antenna, leg pairs 1 and 7, gnathochilarium, gonopods; the head and trunk lost; a draft of a description),
<location id="8EF96018FF90FF82FBA22674FA8D9E0B" LSID="urn:lsid:plazi:treatment:038F87D5FF90FFBDFF6D21C5FE989BA7:8EF96018FF90FF82FBA22674FA8D9E0B" box="[1112,1281,1123,1149]" municipality="Dalnegorsky District" name="Pimorsky Krai" pageId="31" pageNumber="32">Pimorsky Krai</location>
,
<collectingMunicipality id="6BFDACB9FF90FF82FAF72674FF629ED7" pageId="31" pageNumber="32">Dalnegorsky District</collectingMunicipality>
,
<taxonomicName id="4C264D40FF90FF82FF012690FEB79ED6" box="[251,315,1159,1184]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
<emphasis id="B952EAD1FF90FF82FF012690FEB79ED6" box="[251,315,1159,1184]" italics="true" pageId="31" pageNumber="32">Picea</emphasis>
</taxonomicName>
forest,
<date id="FF981003FF90FF82FE682690FE7D9ED7" box="[402,497,1159,1185]" pageId="31" pageNumber="32" value="1972-09">
<collectingDate id="EFDCE9EBFF90FF82FE682690FE7D9ED7" box="[402,497,1159,1185]" pageId="31" pageNumber="32" value="1972-09">IX.1972</collectingDate>
</date>
, leg.
<collectorName id="26D35315FF90FF82FDD52690FC869ED7" box="[559,778,1159,1185]" pageId="31" pageNumber="32">A.N. Kupyanskaya</collectorName>
;
<typeStatus id="549D8861FF90FF82FCED2690FCF09ED7" box="[791,892,1159,1185]" pageId="31" pageNumber="32" type="paratype">paratype</typeStatus>
:
<specimenCount id="9D20FD4AFF90FF82FC7D2690FC589ED7" box="[903,980,1159,1185]" pageId="31" pageNumber="32" type="male">1 male</specimenCount>
(
<collectionCode id="ED37AE06FF90FF82FC1F2690FBA39ED7" box="[997,1071,1159,1185]" pageId="31" pageNumber="32">FSCB</collectionCode>
: 1 constant micropreparation, N 353: gonopod; the head and trunk lost), same locality as in
<typeStatus id="549D8861FF90FF82FCD426BCFC1F9EB3" box="[814,915,1195,1221]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
,
<date id="FF981003FF90FF82FC6426BCFC719EB3" box="[926,1021,1195,1221]" pageId="31" pageNumber="32" value="1972-09">
<collectingDate id="EFDCE9EBFF90FF82FC6426BCFC719EB3" box="[926,1021,1195,1221]" pageId="31" pageNumber="32" value="1972-09">IX.1972</collectingDate>
</date>
, leg.
<collectorName id="26D35315FF90FF82FBCD26BCFA989EB3" box="[1079,1300,1195,1221]" pageId="31" pageNumber="32">A.N. Kupyanskaya.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF90FF82FF3D26D9FBA69FB7" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8B9936C3FF90FF82FF3D26D9FEF99F47" blockId="31.[151,1437,1086,2013]" pageId="31" pageNumber="32">
<emphasis id="B952EAD1FF90FF82FF3D26D9FECE9E9E" bold="true" box="[199,322,1230,1256]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
Differs from its congeners mainly by the clawlike process of the posterior gonopod posterior angiocoxal process, as well as in the plate-like mesal margin of the colpocoxite and the flagelliform anterior angiocoxal process.
</paragraph>
<paragraph id="8B9936C3FF90FF82FF3D272CFBA69FB7" blockId="31.[151,1437,1086,2013]" pageId="31" pageNumber="32">
This species seems to be closely related to
<taxonomicName id="4C264D40FF90FF82FD50272CFCAC9F23" authorityName="Mikhaljova" authorityYear="2019" box="[682,800,1339,1365]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="koiensis">
<emphasis id="B952EAD1FF90FF82FD50272CFCAC9F23" box="[682,800,1339,1365]" italics="true" pageId="31" pageNumber="32">P. koiensis</emphasis>
</taxonomicName>
but differs in the configuration of the posterior gonopod posterior angiocoxal process supplied with a strong clawlike outgrowth (
<emphasis id="B952EAD1FF90FF82FC3F2748FC5D9F0F" bold="true" box="[965,977,1375,1401]" pageId="31" pageNumber="32">c</emphasis>
,
<figureCitation id="131D2A46FF90FF82FC252748FBA39F0F" box="[991,1071,1374,1401]" captionStart="FIGURES 5657" captionStartId="32.[151,264,1038,1062]" captionTargetBox="[158,1432,489,1012]" captionTargetId="figure-134@32.[151,1436,489,1015]" captionTargetPageId="32" captionText="FIGURES 5657. Pacifiosoma triangulatum sp. nov., male holotype. 56, gonopods, caudal view; 57, gonopods, front view; pp, posterior angicoxal process; c, clawlike outgrowth; n, triangular plate; ap, anterior angiocoxal process; cx, coxosternum. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.5606713" httpUri="https://zenodo.org/record/5606713/files/figure.png" pageId="31" pageNumber="32">Fig. 56</figureCitation>
) (vs without clawlike outgrowth in
<taxonomicName id="4C264D40FF90FF82FF422794FEBD9FEB" authorityName="Mikhaljova" authorityYear="2019" box="[184,305,1411,1437]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="koiensis">
<emphasis id="B952EAD1FF90FF82FF422794FEBD9FEB" box="[184,305,1411,1437]" italics="true" pageId="31" pageNumber="32">P. koiensis</emphasis>
</taxonomicName>
) as well as in the more slender distal part of the colpocoxite. In addition, the gonopod posterior angiocoxal processes of the new species are similar to those of
<taxonomicName id="4C264D40FF90FF82FCA627B0FC459FB6" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[860,969,1447,1472]" class="Diplopoda" family="Diplomaragnidae" genus="Pacifiosoma" kingdom="Animalia" order="Chordeumatida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="acutum" status="sp. nov.">
<emphasis id="B952EAD1FF90FF82FCA627B0FC459FB6" box="[860,969,1447,1472]" italics="true" pageId="31" pageNumber="32">P. acutum</emphasis>
</taxonomicName>
<emphasis id="B952EAD1FF90FF82FC2A27B0FBA69FB7" bold="true" box="[976,1066,1447,1473]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A26157AAFF90FF82FC2A27B0FBA69FB7" box="[976,1066,1447,1473]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33C6548FF90FFBDFF3D27DDFE1D9BFF" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="description">
<paragraph id="8B9936C3FF90FF82FF3D27DDFCEA9C7F" blockId="31.[151,1437,1086,2013]" pageId="31" pageNumber="32">
<emphasis id="B952EAD1FF90FF82FF3D27DDFED59F92" bold="true" box="[199,345,1482,1508]" pageId="31" pageNumber="32">Description.</emphasis>
Male. Length
<quantity id="4CDE9B26FF90FF82FDFA27DCFDF79F93" box="[512,635,1483,1509]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="31" pageNumber="32" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
, width about 1.1 mm with paraterga. Coloration in alcohol beige. Legs brownish, their distal parts brown. Antennae brown. Eyes black.
</paragraph>
<paragraph id="8B9936C3FF90FF82FF3D2404FEC59CCB" blockId="31.[151,1437,1086,2013]" pageId="31" pageNumber="32">Number of body rings unknown, most likely, body with 32 rings. Head densely setose. Eye patches triangular, each composed of about 30 ocelli. Antennae slender. Paraterga relatively well-developed from ring 2, on rings 527 their caudolateral corners beak-shaped pointed. Metazonital macrochaetae in a transverse row on somites 2830, like an extended triangle on preceding rings. Anterolateral macrochaetae shortest. All macrochaetae pointed, but not very sharply so.</paragraph>
<paragraph id="8B9936C3FF90FF82FF3D24D0FBD59D07" blockId="31.[151,1437,1086,2013]" pageId="31" pageNumber="32">Legs long and slender. Leg pairs 1 and 2 typically reduced in size, with usual tarsal brushes. Claw of the leg pairs 1 and 2 at base with a long setoid outgrowth ventrally and two small additional claws dorsally. Other pregonopodal legs somewhat enlarged. Leg pair 7 with a group of funnelshaped tarsal papillae apically near claw. Claws 7 at base with a long setoid outgrowth ventrally but without additional claws. Information on tarsal papillae and the claw structure of other legs absent. Legs 10 and 11 normal, with coxal glands.</paragraph>
<paragraph id="8B9936C3FF90FFBDFF3D256CFDDB9B13" blockId="31.[151,1437,1086,2013]" lastBlockId="32.[151,1437,150,465]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
Gonopods as in figs 5657. Flagelliform anterior gonopod telopodites pass inside a short sheath groove with elevated edges. Apices of the anterior gonopod telopodites unmodified. Posterior gonopod colpocoxites fused basally, their distal parts slightly curved caudally. Each colpocoxite entire, almost completely covered with dense papillae. Mesal edge of gonopod colpocoxite drawn into a triangular plate (
<emphasis id="B952EAD1FFAFFFBDFC132280FC759AC7" bold="true" box="[1001,1017,151,177]" pageId="32" pageNumber="33">n</emphasis>
,
<figureCitation id="131D2A46FFAFFFBDFBFD2280FBD79AC7" box="[1031,1115,150,177]" captionStart="FIGURES 5657" captionStartId="32.[151,264,1038,1062]" captionTargetBox="[158,1432,489,1012]" captionTargetId="figure-134@32.[151,1436,489,1015]" captionTargetPageId="32" captionText="FIGURES 5657. Pacifiosoma triangulatum sp. nov., male holotype. 56, gonopods, caudal view; 57, gonopods, front view; pp, posterior angicoxal process; c, clawlike outgrowth; n, triangular plate; ap, anterior angiocoxal process; cx, coxosternum. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.5606713" httpUri="https://zenodo.org/record/5606713/files/figure.png" pageId="32" pageNumber="33">Fig. 56</figureCitation>
). Colpocoxite sheath groove without evident processes. Posterior gonopod angiocoxites with moderately developed elongated subglobules in posterior view, depressed centrally in anterior view. Posterior angiocoxal process (
<emphasis id="B952EAD1FFAFFFBDFBB722C8FBE09A8F" bold="true" box="[1101,1132,223,249]" pageId="32" pageNumber="33">pp</emphasis>
,
<figureCitation id="131D2A46FFAFFFBDFB8722C8FB5C9A8F" box="[1149,1232,222,249]" captionStart="FIGURES 5657" captionStartId="32.[151,264,1038,1062]" captionTargetBox="[158,1432,489,1012]" captionTargetId="figure-134@32.[151,1436,489,1015]" captionTargetPageId="32" captionText="FIGURES 5657. Pacifiosoma triangulatum sp. nov., male holotype. 56, gonopods, caudal view; 57, gonopods, front view; pp, posterior angicoxal process; c, clawlike outgrowth; n, triangular plate; ap, anterior angiocoxal process; cx, coxosternum. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.5606713" httpUri="https://zenodo.org/record/5606713/files/figure.png" pageId="32" pageNumber="33">Fig. 56</figureCitation>
) large, with wide rounded apex covered with papillae and clawlike outgrowth (
<emphasis id="B952EAD1FFAFFFBDFC9B2314FCE19B6B" bold="true" box="[865,877,259,285]" pageId="32" pageNumber="33">с</emphasis>
,
<figureCitation id="131D2A46FFAFFFBDFC852314FC5A9B6B" box="[895,982,258,285]" captionStart="FIGURES 5657" captionStartId="32.[151,264,1038,1062]" captionTargetBox="[158,1432,489,1012]" captionTargetId="figure-134@32.[151,1436,489,1015]" captionTargetPageId="32" captionText="FIGURES 5657. Pacifiosoma triangulatum sp. nov., male holotype. 56, gonopods, caudal view; 57, gonopods, front view; pp, posterior angicoxal process; c, clawlike outgrowth; n, triangular plate; ap, anterior angiocoxal process; cx, coxosternum. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.5606713" httpUri="https://zenodo.org/record/5606713/files/figure.png" pageId="32" pageNumber="33">Fig. 56</figureCitation>
). Anterior angiocoxal process (
<emphasis id="B952EAD1FFAFFFBDFABB2314FAD39B6B" bold="true" box="[1345,1375,259,285]" pageId="32" pageNumber="33">ap</emphasis>
,
<figureCitation id="131D2A46FFAFFFBDFA8A2314FF389B37" captionStart="FIGURES 5657" captionStartId="32.[151,264,1038,1062]" captionTargetBox="[158,1432,489,1012]" captionTargetId="figure-134@32.[151,1436,489,1015]" captionTargetPageId="32" captionText="FIGURES 5657. Pacifiosoma triangulatum sp. nov., male holotype. 56, gonopods, caudal view; 57, gonopods, front view; pp, posterior angicoxal process; c, clawlike outgrowth; n, triangular plate; ap, anterior angiocoxal process; cx, coxosternum. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.5606713" httpUri="https://zenodo.org/record/5606713/files/figure.png" pageId="32" pageNumber="33">Fig. 57</figureCitation>
) long, flagelliform, its distal portion placed inside frontal fold of colpocoxite. Posterior gonopod telopodite 2- segmented, setose; distal segment large.
</paragraph>
<paragraph id="8B9936C3FFAFFFBDFF3D2378FE1D9BFF" blockId="32.[151,1437,150,465]" box="[199,401,367,393]" pageId="32" pageNumber="33">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C33C6548FFAFFFBDFF3D2384FE989BA7" pageId="32" pageNumber="33" type="etymology">
<paragraph id="8B9936C3FFAFFFBDFF3D2384FE989BA7" blockId="32.[151,1437,150,465]" pageId="32" pageNumber="33">
<emphasis id="B952EAD1FFAFFFBDFF3D2384FEC29BDB" bold="true" box="[199,334,403,429]" pageId="32" pageNumber="33">Etymology.</emphasis>
The specific epithet refers to the triangular mesal edge of the gonopod colpocoxite, an adjective in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>