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<document id="ED223010AA15D9334836E46D12C2B2EC" ID-DOI="10.5281/zenodo.5041813" ID-ISSN="1942-1354" ID-Zenodo-Dep="5041813" ID-ZooBank="DF505A18-63A1-44BB-BF5D-13887FAE0DAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624979060254" checkinUser="carolina" docAuthor="Larochelle &amp; Larivière, Marie-Claude" docDate="2021" docId="038F87CF8F31487BDFED16A8A0973EDF" docLanguage="en" docName="InsectaMundi.0864.1-96.pdf" docOrigin="Insecta Mundi 2021 (864)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Ctenognathus urewera Larochelle and Lariviere E 2021, new species" docType="treatment" docVersion="3" lastPageNumber="38" masterDocId="FFB6FFB78F17485CDF2D100BA17B386D" masterDocTitle="Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="37" updateTime="1698978488161" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="AEB8C7B79015E6739E9DE8590BD173B8">Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="F10F60BD2D39AA6005D1366C951059BE">Larochelle</mods:namePart>
<mods:affiliation id="FB091C5DAFC7BC9DE04ACAAE2A7864C3">Collection, Manaaki Whenua-Landcare Prosphodrus waimana Larochelle</mods:affiliation>
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<mods:namePart id="A1393186E78E646BBEDBB545C6E9C310">Larivière, Marie-Claude</mods:namePart>
<mods:affiliation id="2834C8B88839D247A8C2735FCF440E26">- Landcare Research, Private Bag 92170, Auckland Larochelle and Larivière, new species</mods:affiliation>
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<mods:title id="A1930B6D94EEF3299840FE43818B6E2D">Insecta Mundi</mods:title>
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<mods:date id="707A67B19E1698E6D5402726C78634B1">2021</mods:date>
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<treatment id="038F87CF8F31487BDFED16A8A0973EDF" ID-DOI="http://doi.org/10.5281/zenodo.5041917" ID-Zenodo-Dep="5041917" LSID="urn:lsid:plazi:treatment:038F87CF8F31487BDFED16A8A0973EDF" httpUri="http://treatment.plazi.org/id/038F87CF8F31487BDFED16A8A0973EDF" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37">
<subSubSection id="C33C65528F31487ADFED16A8A2C53EAD" box="[192,958,1699,1728]" pageId="38" pageNumber="37" type="nomenclature">
<paragraph id="8B9936D98F31487ADFED16A8A2C53EAD" blockId="38.[192,958,1699,1763]" box="[192,958,1699,1728]" pageId="38" pageNumber="37">
<heading id="D0D181B58F31487ADFED16A8A2C53EAD" box="[192,958,1699,1728]" fontSize="12" level="2" pageId="38" pageNumber="37" reason="2">
<emphasis id="B952EACB8F31487ADFED16A8A2C53EAD" bold="true" box="[192,958,1699,1728]" pageId="38" pageNumber="37">
<taxonomicName id="4C264D5A8F31487ADFED16A8A2663EAD" ID-CoL="8MLCX" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[192,797,1699,1728]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="urewera" status="sp. nov.">
<emphasis id="B952EACB8F31487ADFED16A8A0A73EAD" bold="true" box="[192,476,1699,1728]" italics="true" pageId="38" pageNumber="37">Ctenognathus urewera</emphasis>
Larochelle and Larivière
</taxonomicName>
,
<taxonomicNameLabel id="A26157B08F31487ADC0416AFA2C53EAD" box="[809,958,1699,1728]" pageId="38" pageNumber="37" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F31487BDFED16C3A5463C4F" lastPageId="39" lastPageNumber="38" pageId="38" pageNumber="37" type="description">
<paragraph id="8B9936D98F31487ADFED16C3A00E3E8F" blockId="38.[192,958,1699,1763]" box="[192,373,1736,1763]" pageId="38" pageNumber="37">
<figureCitation id="131D2A5C8F31487ADFED16C3A0763E8E" box="[192,269,1736,1763]" captionStart="Figures 5760" captionStartId="75.[194,281,1866,1892]" captionTargetBox="[198,1431,256,1834]" captionTargetId="figure-41@75.[192,1439,232,1837]" captionTargetPageId="75" captionText="Figures 5760. Dorsal habitus. 57) Ctenognathus garnerae new species. 58) C. tepaki new species. 59) C. urewera new species. 60) C. cardiophorus. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5041845" httpUri="https://zenodo.org/record/5041845/files/figure.png" pageId="38" pageNumber="37">Fig. 59</figureCitation>
,
<figureCitation id="131D2A5C8F31487ADE3516C3A0393E8F" box="[280,322,1736,1762]" captionStart="Figures 101107" captionStartId="84.[192,279,1866,1892]" captionTargetBox="[252,1384,254,1815]" captionTargetId="figure-54@84.[191,1440,208,1841]" captionTargetPageId="84" captionText="Figures 101107. Aedeagus, lateral. 101) Ctenognathus tepaki new species. 102) C. urewera new species. 103) C. cardiophorus. 104) C. adamsi. 105) C. pictonensis. 106) C. tawanui new species. 107) C. perumalae new species." figureDoi="http://doi.org/10.5281/zenodo.5041881" httpUri="https://zenodo.org/record/5041881/files/figure.png" pageId="38" pageNumber="37">102</figureCitation>
,
<figureCitation id="131D2A5C8F31487ADE6016C3A00E3E8F" box="[333,373,1736,1762]" captionStart="Figures 142147" captionStartId="91.[192,279,1866,1892]" captionTargetBox="[206,1426,213,1839]" captionTargetId="figure-46@91.[206,1426,213,1839]" captionTargetPageId="91" captionText="Figures 142147. Species distribution maps. 142) Ctenognathus pictonensis. 143) C. sandageri. 144) C. takahe new species. 145) C. tawanui new species. 146) C. tepaki new species. 147) C. urewera new species." figureDoi="http://doi.org/10.5281/zenodo.5041903" httpUri="https://zenodo.org/record/5041903/files/figure.png" pageId="38" pageNumber="37">147</figureCitation>
</paragraph>
<paragraph id="8B9936D98F31487ADFCD16FAA0E53FEB" blockId="38.[224,1440,1777,1926]" pageId="38" pageNumber="37">
<taxonomicName id="4C264D5A8F31487ADFCD16FAA3B23F64" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere E" authorityYear="2021" box="[224,713,1777,1801]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="urewera" status="sp. nov.">
<emphasis id="B952EACB8F31487ADFCD16FAA0B93F64" box="[224,450,1777,1801]" italics="true" pageId="38" pageNumber="37">Ctenognathus urewera</emphasis>
<materialsCitation id="3B4E3C848F31487ADEE116FAA3B23F64" box="[460,713,1777,1801]" collectingDate="1994-02-14" collectingDateMax="1994-02-18" collectingDateMin="1994-02-14" collectionCode="NZAC, MONZ" country="New Zealand" elevation="923" pageId="38" pageNumber="37" specimenCount="1" specimenCount-female="1" typeStatus="holotype">Larochelle and Larivière</materialsCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A26157B08F31487ADDFB16FAA2283F64" box="[726,851,1777,1801]" pageId="38" pageNumber="37" rank="species">new species</taxonomicNameLabel>
.
<typeStatus id="549D887B8F31487ADC7216FAA2BB3F64" box="[863,960,1777,1801]" pageId="38" pageNumber="37" type="holotype">Holotype</typeStatus>
: male (NZAC) labeled “
<collectingCountry id="F33176498F31487ADB9316FAA40D3F64" box="[1214,1398,1777,1801]" name="New Zealand" pageId="38" pageNumber="37">NEW ZEALAND</collectingCountry>
GB Urewera NP, Huiarau Summit
<quantity id="4CDE9B3C8F31487ADD1A171BA30B3F45" box="[567,624,1808,1832]" metricMagnitude="2" metricUnit="m" metricValue="9.23" pageId="38" pageNumber="37" unit="m" value="923.0">
<elevation id="000BD1EA8F31487ADD1A171BA30B3F45" box="[567,624,1808,1832]" metricMagnitude="2" metricUnit="m" metricValue="9.23" pageId="38" pageNumber="37" unit="m" value="923.0">923m</elevation>
</quantity>
<date id="FF9810198F31487ADD5A171BA2793F45" box="[631,770,1808,1832]" pageId="38" pageNumber="37" value="1994-02-14" valueMax="1994-02-18" valueMin="1994-02-14">
<collectingDate id="EFDCE9F18F31487ADD5A171BA2793F45" box="[631,770,1808,1832]" pageId="38" pageNumber="37" value="1994-02-14" valueMax="1994-02-18" valueMin="1994-02-14">1418.II.1994</collectingDate>
</date>
Larivière, Larochelle (typed) / Nothofagus + Horopito + tree fern for. [= forest], wet slopes. Pittraps. (typed) /
<typeStatus id="549D887B8F31487ADD941724A2473F2A" box="[697,828,1839,1863]" pageId="38" pageNumber="37" type="holotype">HOLOTYPE</typeStatus>
[male symbol]
<taxonomicName id="4C264D5A8F31487ADCFB1724A04B3F0A" authority="Larochelle &amp; Lariviere, 2021" authorityName="Larochelle &amp; Lariviere" authorityYear="2021" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="urewera">
<emphasis id="B952EACB8F31487ADCFB1724A5CF3F2A" box="[982,1204,1839,1863]" italics="true" pageId="38" pageNumber="37">Ctenognathus urewera</emphasis>
Larochelle &amp; Larivière, 2021
</taxonomicName>
(red label; typed).”
<typeStatus id="549D887B8F31487ADEDB1744A3213F0A" box="[502,602,1871,1895]" pageId="38" pageNumber="37" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D20FD508F31487ADD4E1744A3A03F0A" box="[611,731,1871,1895]" pageId="38" pageNumber="37" type="female">two females</specimenCount>
(MONZ, NZAC) from the same locality as the holotype, bearing blue
<typeStatus id="549D887B8F31487ADE2D1765A0233FEB" box="[256,344,1902,1926]" pageId="38" pageNumber="37" type="paratype">paratype</typeStatus>
labels.
</paragraph>
<paragraph id="8B9936D98F30487BDFED10DEA5463C4F" blockId="39.[191,1441,213,1058]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED10DEA0293882" bold="true" box="[192,338,213,239]" pageId="39" pageNumber="38">Description.</emphasis>
Body length
<quantity id="4CDE9B3C8F30487BDEC710DDA3F4389D" box="[490,655,214,240]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.38" metricValueMin="1.27" pageId="39" pageNumber="38" unit="mm" value="13.25" valueMax="13.8" valueMin="12.7">12.713.8 mm</quantity>
. Head, pronotum, and elytra coal black; abdomen blackish; antennae, palpi, and femora rufopiceous; tibiae and tarsi reddish. Microsculpture very transverse (with microlines) on head, pronotum, and elytra. Iridescence present on head and pronotum. Very shiny. Forebody narrow in comparison to elytra.
<emphasis id="B952EACB8F30487BDE261137A02B393B" bold="true" box="[267,336,316,342]" pageId="39" pageNumber="38">Head.</emphasis>
Narrow. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, moderately emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth truncate apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically.
<emphasis id="B952EACB8F30487BDC8C11A9A28739D1" bold="true" box="[929,1020,418,444]" pageId="39" pageNumber="38">Thorax.</emphasis>
Pronotum slightly convex, obsoletely punctate across apex and base, slightly wrinkled on disc, narrow, strongly cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, angulate; anterior bead complete, less impressed medially; sides strongly rounded and sinuate posteriorly; lateral beads complete; lateral depressions narrow; a single setiferous puncture on each side (anteriorly); posterolateral angles rectangular; laterobasal foveae moderately deep, rather narrow, prolonged forward; posterior bead complete, less impressed medially; base emarginate.
<emphasis id="B952EACB8F30487BDFED1264A1873AE4" bold="true" box="[192,252,623,649]" pageId="39" pageNumber="38">Legs.</emphasis>
Very long. Metafemora with two posteroventral setae. Meso- and metatarsomeres 14 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 glabrous ventrally.
<emphasis id="B952EACB8F30487BDD4712B8A3C23AA0" bold="true" box="[618,697,691,717]" pageId="39" pageNumber="38">Elytra.</emphasis>
Slightly convex, not sloping down toward apex, subovate, widest about middle. Shoulders moderately rounded. Sides strongly rounded. Scutellar setiferous pore invisible. Scutellar striole short or moderately long, impunctate. Striae obsolete, impunctate. Intervals slightly convex; interval 3 with a single shallow, minute subapical setiferous puncture. Umbilicate series with 1718 setiferous punctures. Subapical sinuations feeble. Apices rounded.
<emphasis id="B952EACB8F30487BDD941330A24F3B38" bold="true" box="[697,820,827,853]" pageId="39" pageNumber="38">Abdomen.</emphasis>
Sterna IVVI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with ten long apical ambulatory setae.
<emphasis id="B952EACB8F30487BDEF91374A3363BF4" bold="true" box="[468,589,895,921]" pageId="39" pageNumber="38">Aedeagus.</emphasis>
Lateral view (
<figureCitation id="131D2A5C8F30487BDDC71374A2333BF4" box="[746,840,895,921]" captionStart="Figures 101107" captionStartId="84.[192,279,1866,1892]" captionTargetBox="[252,1384,254,1815]" captionTargetId="figure-54@84.[191,1440,208,1841]" captionTargetPageId="84" captionText="Figures 101107. Aedeagus, lateral. 101) Ctenognathus tepaki new species. 102) C. urewera new species. 103) C. cardiophorus. 104) C. adamsi. 105) C. pictonensis. 106) C. tawanui new species. 107) C. perumalae new species." figureDoi="http://doi.org/10.5281/zenodo.5041881" httpUri="https://zenodo.org/record/5041881/files/figure.png" pageId="39" pageNumber="38">Fig. 102</figureCitation>
): strongly arcuate; base strongly convex dorsally, with basal lobe very wide; middle moderately convex in basal half and straight in apical half dorsally, slightly concave ventrally, with dorsal membranous area very wide and very long; apex subtriangular, straight dorsally and ventrally, with extreme tip narrow and short. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area.
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F30487BDFED1432A2063CEB" pageId="39" pageNumber="38" type="materials_examined">
<paragraph id="8B9936D98F30487BDFED1432A2C53C3E" blockId="39.[192,958,1081,1108]" box="[192,958,1081,1108]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED1432A0D83C3E" bold="true" box="[192,419,1081,1107]" pageId="39" pageNumber="38">Material examined.</emphasis>
<specimenCount id="9D20FD508F30487BDE841432A3373C3E" box="[425,588,1081,1107]" pageId="39" pageNumber="38" type="generic">811 specimens</specimenCount>
(
<collectionCode id="ED37AE1C8F30487BDD771432A3D93C3E" box="[602,674,1081,1107]" pageId="39" pageNumber="38">JNNZ</collectionCode>
,
<collectionCode id="ED37AE1C8F30487BDD801431A3833C39" box="[685,760,1082,1108]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="39" pageNumber="38">LUNZ</collectionCode>
,
<collectionCode id="ED37AE1C8F30487BDC2F1432A22C3C3E" box="[770,855,1081,1107]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="39" pageNumber="38" type="Museum">MONZ</collectionCode>
,
<collectionCode id="ED37AE1C8F30487BDC4F1432A2C93C3E" box="[866,946,1081,1107]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="39" pageNumber="38" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="8B9936D98F30487BDFED1460A2063CEB" blockId="39.[192,893,1131,1158]" box="[192,893,1131,1158]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED1460A0A73CE8" bold="true" box="[192,476,1131,1157]" pageId="39" pageNumber="38">Geographic distribution</emphasis>
(
<figureCitation id="131D2A5C8F30487BDEC61460A3323CE8" box="[491,585,1131,1157]" captionStart="Figures 142147" captionStartId="91.[192,279,1866,1892]" captionTargetBox="[206,1426,213,1839]" captionTargetId="figure-46@91.[206,1426,213,1839]" captionTargetPageId="91" captionText="Figures 142147. Species distribution maps. 142) Ctenognathus pictonensis. 143) C. sandageri. 144) C. takahe new species. 145) C. tawanui new species. 146) C. tepaki new species. 147) C. urewera new species." figureDoi="http://doi.org/10.5281/zenodo.5041903" httpUri="https://zenodo.org/record/5041903/files/figure.png" pageId="39" pageNumber="38">Fig. 147</figureCitation>
). North Island: BP,
<collectingRegion id="49E2F83B8F30487BDC0D1460A23D3CE8" box="[800,838,1131,1157]" country="Guinea-Bissau" name="Gabu" pageId="39" pageNumber="38">GB</collectingRegion>
, HB.
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F30487BDFED1496A5333DD9" pageId="39" pageNumber="38" type="biology_ecology">
<paragraph id="8B9936D98F30487BDFED1496A2FB3C96" blockId="39.[192,1440,1181,1276]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED1496A0583CDA" bold="true" box="[192,291,1181,1207]" pageId="39" pageNumber="38">Ecology.</emphasis>
Lowland, montane. Epigean, fossorial, arboreal. Wet forests (beech, broadleaf) and tree plantations (pine). Associated with stream, seepages, and mud flats. Shaded ground; wet, soaked soil. Nocturnal; hides during the day under logs, fallen branches, and stones. Gregarious.
</paragraph>
<paragraph id="8B9936D98F30487BDFED1518A5FE3D3C" blockId="39.[192,1440,1299,1361]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED1518A05B3D40" bold="true" box="[192,288,1299,1325]" pageId="39" pageNumber="38">Biology.</emphasis>
Seasonality: SeptemberJanuary, MarchApril. Tenerals: DecemberFebruary. Predacious (based on mouthpart morphology). Occasionally infested with mites and fungi (Laboulbeniales).
</paragraph>
<paragraph id="8B9936D98F30487BDFED156CA5293DEF" blockId="39.[192,1106,1383,1411]" box="[192,1106,1383,1411]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED156CA0F83DEF" bold="true" box="[192,387,1383,1410]" pageId="39" pageNumber="38">Dispersal power.</emphasis>
Subapterous (incapable of flight). Occasional climber (on trees).
</paragraph>
<paragraph id="8B9936D98F30487BDFED1592A5333DD9" blockId="39.[192,1096,1433,1460]" box="[192,1096,1433,1460]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED1592A0BD3DDE" bold="true" box="[192,454,1433,1459]" pageId="39" pageNumber="38">Collecting techniques.</emphasis>
Pitfall trapping; turning logs, fallen branches, and stones.
</paragraph>
</subSubSection>
<subSubSection id="C33C65528F30487BDFED15C7A0973EDF" pageId="39" pageNumber="38" type="discussion">
<paragraph id="8B9936D98F30487BDFED15C7A0973EDF" blockId="39.[190,1442,1484,1715]" pageId="39" pageNumber="38">
<emphasis id="B952EACB8F30487BDFED15C7A0543D8B" bold="true" box="[192,303,1484,1510]" pageId="39" pageNumber="38">Remarks.</emphasis>
The species is named after the Urewera National Park (
<collectingRegion id="49E2F83B8F30487BDC8C15C7A2B23D8B" box="[929,969,1484,1510]" country="Guinea-Bissau" name="Gabu" pageId="39" pageNumber="38">GB</collectingRegion>
) which includes the
<typeStatus id="549D887B8F30487BDB9E15C7A5983D8B" box="[1203,1251,1484,1510]" pageId="39" pageNumber="38">type</typeStatus>
locality Huiarau Summit, and is applied as a noun in apposition.
<taxonomicName id="4C264D5A8F30487BDDF615E5A2A93E65" authorityName="Larochelle &amp; Lariviere" authorityYear="2021" box="[731,978,1518,1544]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="urewera">
<emphasis id="B952EACB8F30487BDDF615E5A2A93E65" box="[731,978,1518,1544]" italics="true" pageId="39" pageNumber="38">Ctenognathus urewera</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="4C264D5A8F30487BDA2615E5A1983E47" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F30487BDA2615E5A1983E47" italics="true" pageId="39" pageNumber="38">C. cardiophorus</emphasis>
</taxonomicName>
. In addition to diagnostic characters of the male genitalia,
<taxonomicName id="4C264D5A8F30487BDC4C161BA2AC3E47" authorityName="Larochelle &amp; Lariviere" authorityYear="2021" box="[865,983,1552,1578]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="urewera">
<emphasis id="B952EACB8F30487BDC4C161BA2AC3E47" box="[865,983,1552,1578]" italics="true" pageId="39" pageNumber="38">C. urewera</emphasis>
</taxonomicName>
has the following distinguishing features: pronotum widest before middle, slightly wrinkled on disc, sides strongly sinuate posteriorly, posterolateral angles rectangular; interval 3 of elytra with a minute subapical setiferous puncture. The two species are allopatric:
<taxonomicName id="4C264D5A8F30487BDAA7165FA0623EFD" authorityName="Larochelle &amp; Lariviere" authorityYear="2021" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="urewera">
<emphasis id="B952EACB8F30487BDAA7165FA0623EFD" italics="true" pageId="39" pageNumber="38">C. urewera</emphasis>
</taxonomicName>
occurs in eastern central areas of the North Island (BP,
<collectingRegion id="49E2F83B8F30487BDCAE167DA2D23EFD" box="[899,937,1654,1680]" country="Guinea-Bissau" name="Gabu" pageId="39" pageNumber="38">GB</collectingRegion>
, HB), while
<taxonomicName id="4C264D5A8F30487BDB1E167DA59A3EFD" baseAuthorityName="Chaudoir" baseAuthorityYear="1878" box="[1075,1249,1654,1680]" class="Insecta" family="Carabidae" genus="Ctenognathus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="cardiophorus">
<emphasis id="B952EACB8F30487BDB1E167DA59A3EFD" box="[1075,1249,1654,1680]" italics="true" pageId="39" pageNumber="38">C. cardiophorus</emphasis>
</taxonomicName>
is found in more northern areas (NDWO).
</paragraph>
</subSubSection>
</treatment>
</document>