1376 lines
199 KiB
XML
1376 lines
199 KiB
XML
<document id="B7A88339C6BA7D14EC55B92226D5FD4D" ID-DOI="10.1590/2358-2936e2022008" ID-ISSN="2358-2936" ID-Zenodo-Dep="10904872" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712001447768" checkinUser="felipe" docAuthor="Paramasivam, Purushothaman, Dhinakaran, A., Ajith Kumar, T. T. & Lal, Kuldeep K." docDate="2022" docId="038F87BDFFB6FF9E2619FAF46527F8D3" docLanguage="en" docName="Nauplius.30.e2022008.pdf" docOrigin="Nauplius (e 2022008) 30" docSource="http://dx.doi.org/10.1590/2358-2936e2022008" docStyle="DocumentStyle:64D98C772C5FBF4D3A503C51F9604E62.1:Nauplius.2017-.journal_article" docStyleId="64D98C772C5FBF4D3A503C51F9604E62" docStyleName="Nauplius.2017-.journal_article" docStyleVersion="1" docTitle="Actinimenes koyas Paramasivam & Dhinakaran & Ajith Kumar & Lal 2022, sp. nov." docType="treatment" docUuid="E455A624-F36D-4386-BFF4-8A548568C10C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="14" masterDocId="FFB6FFC5FFB2FF93275EFFF0611FFFE5" masterDocTitle="A new species of the genus Actinimenes Ďuriš and Horká, 2017 (Crustacea: Decapoda: Palaemonidae) from the Arabian Sea, Lakshadweep Islands, India" masterLastPageNumber="18" masterPageNumber="1" pageNumber="5" updateTime="1712690790463" updateUser="carolina" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="5057217118736857BFEE386F21B320A2">A new species of the genus Actinimenes Ďuriš and Horká, 2017 (Crustacea: Decapoda: Palaemonidae) from the Arabian Sea, Lakshadweep Islands, India</mods:title>
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<mods:roleTerm id="E748754F5F2E9D0D41917287C5A4A07E">Author</mods:roleTerm>
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<mods:namePart id="63BD429CE28BA9C7A35F0188EE281828">Paramasivam, Purushothaman</mods:namePart>
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<mods:roleTerm id="4E2619A1B432E930D83B751B6B33B0DC">Author</mods:roleTerm>
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<mods:namePart id="A7799EADBD8C4748D08C24D8A856C4FD">Dhinakaran, A.</mods:namePart>
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<mods:namePart id="4DC32BCCEDB2541D6C015224B7B360BA">Ajith Kumar, T. T.</mods:namePart>
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<mods:name id="ECAF377C2DCF964A3C1F93A93E2DCB06" type="personal">
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<mods:roleTerm id="2468B383D89D24DF37352E4B65586E7F">Author</mods:roleTerm>
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<mods:namePart id="3C0273E9871B04C73950CE4E628DC8FA">Lal, Kuldeep K.</mods:namePart>
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<mods:typeOfResource id="E679DAE8233EECEC7926CC60CFA7E421">text</mods:typeOfResource>
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<mods:title id="49A53395B933072CAD87ED412618F7A1">Nauplius</mods:title>
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</mods:titleInfo>
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<mods:part id="03522AE0E6A5D09688FCE3976CA5BE5E">
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<mods:date id="78C906F22BDBF6671903F890FCDBEDEE">2022</mods:date>
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<mods:title id="F3FEC9D4171C93AFC6AC6CE5DB74C41B">e 2022008</mods:title>
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<mods:number id="0DAD35C03173C37F5F76BC002A3B1204">2022-04-11</mods:number>
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<mods:detail id="86D31614A09A2774336A48FB25265354" type="volume">
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<mods:number id="071DF8EA5D19B75C2838311229BD6DA0">30</mods:number>
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<mods:identifier id="B15E5C067FF46EF4ADDD6C76AEF175CD" type="ZooBank">45058CD0-568E-4CEA-A4EE-CE7490617076</mods:identifier>
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<treatment id="038F87BDFFB6FF9E2619FAF46527F8D3" LSID="urn:lsid:zoobank.org:act:E455A624-F36D-4386-BFF4-8A548568C10C" httpUri="http://treatment.plazi.org/id/038F87BDFFB6FF9E2619FAF46527F8D3" lastPageId="13" lastPageNumber="14" pageId="4" pageNumber="5">
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<subSubSection id="C33C6520FFB6FF972619FAF4638CFAC4" box="[327,659,1284,1313]" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="8B9936ABFFB6FF972619FAF4638CFAC4" blockId="4.[197,789,1284,1430]" box="[327,659,1284,1313]" pageId="4" pageNumber="5">
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<heading id="D0D181C7FFB6FF972619FAF4638CFAC4" bold="true" box="[327,659,1284,1313]" centered="true" fontSize="12" level="2" pageId="4" pageNumber="5" reason="2">
|
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<emphasis id="B952EAB9FFB6FF972619FAF4638CFAC4" bold="true" box="[327,659,1284,1313]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C264D28FFB6FF972619FAF46332FAC4" ID-CoL="9H272" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[327,557,1284,1313]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB6FF972619FAF46332FAC4" bold="true" box="[327,557,1284,1313]" italics="true" pageId="4" pageNumber="5">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB6FF97256DFAF5638CFAC4" box="[563,659,1285,1313]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6520FFB6FF9726F4FADC63DBFA73" pageId="4" pageNumber="5" type="description">
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<paragraph id="8B9936ABFFB6FF9726F4FADC632EFAAC" blockId="4.[197,789,1284,1430]" box="[426,561,1323,1353]" pageId="4" pageNumber="5">
|
||
(
|
||
<figureCitation id="131D2A2EFFB6FF9726EBFADB633BFAAD" box="[437,548,1323,1352]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStartId-0="5.[181,248,1456,1479]" captionStartId-1="6.[181,249,1830,1853]" captionStartId-2="8.[181,250,1837,1860]" captionStartId-3="9.[181,250,1831,1854]" captionStartId-4="10.[181,250,1916,1939]" captionTargetBox-0="[212,1437,151,1416]" captionTargetBox-1="[185,1475,169,1769]" captionTargetBox-2="[210,1421,151,1781]" captionTargetBox-3="[252,1357,907,1793]" captionTargetBox-4="[181,1466,1022,1879]" captionTargetId-0="figure-39@5.[212,1437,151,1416]" captionTargetId-1="figure-40@6.[170,1485,151,1797]" captionTargetId-2="figure-40@8.[195,1452,151,1805]" captionTargetId-3="figure-318@9.[195,1449,888,1822]" captionTargetId-4="figure-365@10.[181,1466,1022,1879]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="Figure 2.Actinimenes koyas sp.nov.from Agattiisland,India.(a) Holotype:ovigerous female(CL 3.5 mm,NBFGR/PALAKOY─01); (b) paratype, male (CL 3.2 mm, NBFGR: DBTLDA62, Ethanol preserved)." captionText-1="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." captionText-2="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." captionText-3="Figure 5.Actinimenes koyas sp. nov. holotype, ovigerous female, CL 3.5 mm, NBFGR/PALAKOY–01. A, Fourth sternal plate with median incision (inner view); B, left dactylus of the thirdpereopod (closer view); C, dorsal view of telson with uropod; D, posterior end of telson (closer view)." captionText-4="Figure 6.Collection site, assemblages of Heteractis magnifica at Agatti Island, Lakshadweep, India with live colorationof Actinimenes koyas sp. nov. (Depth 1–2 m)." figureDoi-0="http://doi.org/10.5281/zenodo.10904876" figureDoi-1="http://doi.org/10.5281/zenodo.10904878" figureDoi-2="http://doi.org/10.5281/zenodo.10904880" figureDoi-3="http://doi.org/10.5281/zenodo.10904882" figureDoi-4="http://doi.org/10.5281/zenodo.10904884" httpUri-0="https://zenodo.org/record/10904876/files/figure.png" httpUri-1="https://zenodo.org/record/10904878/files/figure.png" httpUri-2="https://zenodo.org/record/10904880/files/figure.png" httpUri-3="https://zenodo.org/record/10904882/files/figure.png" httpUri-4="https://zenodo.org/record/10904884/files/figure.png" pageId="4" pageNumber="5">Figs. 2–6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB6FF97279BFAA263DBFA73" blockId="4.[197,789,1284,1430]" pageId="4" pageNumber="5">
|
||
Zoobank:
|
||
<uri id="FFB73AA9FFB6FF97261AFAA263DBFA73" pageId="4" pageNumber="5">
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||
urn:lsid:zoobank.org:pub:
|
||
<uuid id="FF800C7EFFB6FF972520FAA263DBFA73" pageId="4" pageNumber="5">45058CD0- 568E-4CEA-A4EE-CE7490617076</uuid>
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||
</uri>
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6520FFB6FF972785FA4E64A4FBEC" pageId="4" pageNumber="5" type="materials_examined">
|
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<paragraph id="8B9936ABFFB6FF972785FA4E6239F97B" blockId="4.[179,808,1470,1967]" pageId="4" pageNumber="5">
|
||
<emphasis id="B952EAB9FFB6FF972785FA4E60A3FA3E" box="[219,444,1470,1499]" italics="true" pageId="4" pageNumber="5">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B4E3CF6FFB6FF972693FA4E6239F97B" collectingDate="2019-12" collectingMethod="scoop net" collectionCode="NBFGR" country="India" latitude="10.820278" location="Agatti Island" longLatPrecision="21" longitude="72.16861" municipality="Arabian Sea" pageId="4" pageNumber="5" specimenCode="DBTLDA59" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="549D8809FFB6FF972693FA4E6358FA3E" box="[461,583,1470,1499]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
|
||
: ovigerous female (CL3.0 mm; accession
|
||
<superScript id="7C539BE3FFB6FF97269DFA1660C4FA11" attach="left" box="[451,475,1508,1538]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
:
|
||
<collectionCode id="ED37AE6EFFB6FF9726B6FA156354F9E7" box="[488,587,1509,1538]" country="India" httpUri="http://grbio.org/cool/j0qy-cctt" name="National Bureau of Fish Genetic Resources (Indian Council of Agricultural Research)" pageId="4" pageNumber="5">NBFGR</collectionCode>
|
||
/PALAKOY─01), ID
|
||
<superScript id="7C539BE3FFB6FF9727BFF9FD61E5F9FE" attach="left" box="[225,250,1547,1577]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
<specimenCode id="DB809ED0FFB6FF97265EF9FC6084F9CC" box="[256,411,1548,1577]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA59</specimenCode>
|
||
,
|
||
<geoCoordinate id="EE12506CFFB6FF9726F7F9FC6328F9CC" box="[425,567,1548,1577]" degrees="10" direction="north" minutes="49" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="13" value="10.820278">10°49’13”N</geoCoordinate>
|
||
<geoCoordinate id="EE12506CFFB6FF97251FF9FC63D0F9CC" box="[577,719,1548,1577]" degrees="72" direction="east" minutes="10" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="07" value="72.16861">72°10’07”E</geoCoordinate>
|
||
, depth 1.0–2.0 m,
|
||
<collectingMethod id="52674EBCFFB6FF972614F9C460D5F9B5" box="[330,458,1588,1616]" pageId="4" pageNumber="5">scoop net</collectingMethod>
|
||
, associated with
|
||
<taxonomicName id="4C264D28FFB6FF9725F7F9C46368F992" authority="(Quay and Gaimard, 1833)" baseAuthorityName="Quay and Gaimard" baseAuthorityYear="1833" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="5" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFB6FF9725F7F9C46038F993" italics="true" pageId="4" pageNumber="5">Heteractis magnifica</emphasis>
|
||
(Quay and Gaimard, 1833)
|
||
</taxonomicName>
|
||
,
|
||
<location id="8EF96070FFB6FF9725DFF9AA623FF992" LSID="urn:lsid:plazi:treatment:038F87BDFFB6FF9E2619FAF46527F8D3:8EF96070FFB6FF9725DFF9AA623FF992" box="[641,800,1626,1655]" country="India" latitude="10.820278" longLatPrecision="21" longitude="72.16861" municipality="Arabian Sea" name="Agatti Island" pageId="4" pageNumber="5">Agatti Island</location>
|
||
, Lakshadweep,
|
||
<collectingCountry id="F331763BFFB6FF97263CF97160BCF97B" box="[354,419,1665,1694]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
|
||
,
|
||
<collectingMunicipality id="6BFDACD1FFB6FF9726F2F9716324F97B" box="[428,571,1665,1694]" pageId="4" pageNumber="5">Arabian Sea</collectingMunicipality>
|
||
, on
|
||
<date id="FF98106BFFB6FF972538F971623DF97B" box="[614,802,1665,1694]" pageId="4" pageNumber="5" value="2019-12">
|
||
<collectingDate id="EFDCE983FFB6FF972538F971623DF97B" box="[614,802,1665,1694]" pageId="4" pageNumber="5" value="2019-12">December 2019</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB6FF972787F95964A4FBEC" blockId="4.[179,808,1470,1967]" lastBlockId="4.[843,1467,845,1033]" pageId="4" pageNumber="5">
|
||
<materialsCitation id="3B4E3CF6FFB6FF972787F959606BF8F6" collectingDate="2020-03" collectionCode="CL, NBFGR" country="India" latitude="10.822222" location="Island" longLatPrecision="21" longitude="72.17083" municipality="Arabian Sea" pageId="4" pageNumber="5" specimenCode="DBTLDA62" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus id="549D8809FFB6FF972787F9596041F920" box="[217,350,1705,1733]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
|
||
: male (
|
||
<collectionCode id="ED37AE6EFFB6FF97269DF95860F1F920" box="[451,494,1704,1733]" pageId="4" pageNumber="5">CL</collectionCode>
|
||
<quantity id="4CDE9B4EFFB6FF9726A2F9586377F920" box="[508,616,1704,1733]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="5" unit="mm" value="3.2">3.2 mm</quantity>
|
||
; accession
|
||
<superScript id="7C539BE3FFB6FF97245EF9596204F952" attach="right" box="[768,795,1703,1733]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
:
|
||
<collectionCode id="ED37AE6EFFB6FF9727EBF93F6005F909" box="[181,282,1743,1772]" collectionName="NBFGR" country="India" httpUri="http://grbio.org/cool/j0qy-cctt" name="National Bureau of Fish Genetic Resources (Indian Council of Agricultural Research)" pageId="4" pageNumber="5">NBFGR</collectionCode>
|
||
/PALAKOY─02), ID
|
||
<superScript id="7C539BE3FFB6FF972579F9206320F93B" attach="left" box="[551,575,1742,1772]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
<specimenCode id="DB809ED0FFB6FF97251AF93F63C0F909" box="[580,735,1743,1772]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA62</specimenCode>
|
||
, same data as holotype.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4E3CF6FFB6FF972629F906623DF84A" collectingDate="2020-03" collectingMethod="scoop net" collectionCode="CL, ZSI, C" country="India" latitude="10.822222" location="Island" longLatPrecision="21" longitude="72.17083" municipality="Arabian Sea" pageId="4" pageNumber="5" specimenCode="DBTLDA111" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D20FD22FFB6FF972629F90660C7F8F6" box="[375,472,1782,1811]" count="1" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
|
||
and
|
||
<specimenCount id="9D20FD22FFB6FF972555F9066349F8F6" box="[523,598,1782,1811]" count="1" pageId="4" pageNumber="5" type="male">1male</specimenCount>
|
||
(
|
||
<collectionCode id="ED37AE6EFFB6FF97253CF9066395F8F6" box="[610,650,1782,1811]" pageId="4" pageNumber="5">CL</collectionCode>
|
||
2.0–
|
||
<quantity id="4CDE9B4EFFB6FF97259AF9066200F8F7" box="[708,799,1782,1811]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="5" unit="mm" value="2.5">2.5mm</quantity>
|
||
; accession
|
||
<superScript id="7C539BE3FFB6FF972673F8ED6057F8C9" attach="right" box="[301,328,1820,1849]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
:
|
||
<collectionCode id="ED37AE6EFFB6FF972606F8ED6094F8DF" box="[344,395,1821,1850]" collectionName="ZSI" pageId="4" pageNumber="5">ZSI</collectionCode>
|
||
:
|
||
<collectionCode id="ED37AE6EFFB6FF9726C8F8ED60B4F8DF" box="[406,427,1821,1850]" pageId="4" pageNumber="5">C</collectionCode>
|
||
8867/2), ID
|
||
<superScript id="7C539BE3FFB6FF97251AF8ED6342F8C9" attach="left" box="[580,605,1820,1849]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
<specimenCode id="DB809ED0FFB6FF97253AF8ED6213F8DF" box="[612,780,1821,1850]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA111</specimenCode>
|
||
A,
|
||
<specimenCode id="DB809ED0FFB6FF9727E9F8B46078F884" box="[183,359,1860,1889]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA111</specimenCode>
|
||
B,
|
||
<geoCoordinate id="EE12506CFFB6FF9726D0F8B46339F884" box="[398,550,1860,1889]" degrees="10" direction="north" minutes="49" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="20" value="10.822222">10°49’20”N</geoCoordinate>
|
||
<geoCoordinate id="EE12506CFFB6FF97256BF8B363D7F885" box="[565,712,1859,1888]" degrees="72" direction="east" minutes="10" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="15" value="72.17083">72°10’15”E</geoCoordinate>
|
||
, depth 1.0–2.0 m,
|
||
<collectingMethod id="52674EBCFFB6FF972661F89B60A8F862" box="[319,439,1899,1927]" pageId="4" pageNumber="5">scoop net</collectingMethod>
|
||
, associated with
|
||
<taxonomicName id="4C264D28FFB6FF9725D8F89C601EF84B" authority=", Agatti" authorityName="Agatti" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="5" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFB6FF9725D8F89C6201F862" box="[646,798,1899,1928]" italics="true" pageId="4" pageNumber="5">H. magnifica</emphasis>
|
||
, Agatti
|
||
</taxonomicName>
|
||
<location id="8EF96070FFB6FF972657F8616049F84B" LSID="urn:lsid:plazi:treatment:038F87BDFFB6FF9E2619FAF46527F8D3:8EF96070FFB6FF972657F8616049F84B" box="[265,342,1937,1966]" country="India" latitude="10.822222" longLatPrecision="21" longitude="72.17083" municipality="Arabian Sea" name="Island" pageId="4" pageNumber="5">Island</location>
|
||
, Lakshadweep,
|
||
<collectingCountry id="F331763BFFB6FF972546F8616344F84B" box="[536,603,1937,1966]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
|
||
, in
|
||
<date id="FF98106BFFB6FF9725D9F861623DF84A" box="[647,802,1937,1967]" pageId="4" pageNumber="5" value="2020-03">
|
||
<collectingDate id="EFDCE983FFB6FF9725D9F861623DF84A" box="[647,802,1937,1967]" pageId="4" pageNumber="5" value="2020-03">March 2020</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B4E3CF6FFB6FF972415FCA0653CFC76" collectingDate="2019-12" collectionCode="CL" country="India" latitude="10.820278" location="Island" longLatPrecision="21" longitude="72.16861" pageId="4" pageNumber="5" specimenCode="DBTLDA60, DBTLDA66" specimenCount="7" specimenCount-female="7" specimenCount-male="6" typeStatus="paratype">
|
||
<specimenCount id="9D20FD22FFB6FF972415FCA062A4FC89" box="[843,955,847,876]" count="7" pageId="4" pageNumber="5" type="female">7 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED37AE6EFFB6FF972496FCBF62EDFC89" box="[968,1010,847,876]" pageId="4" pageNumber="5">CL</collectionCode>
|
||
<quantity id="4CDE9B4EFFB6FF9724A9FCBF6594FC89" box="[1015,1163,847,876]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="4.2" metricValueMin="1.9" pageId="4" pageNumber="5" unit="mm" value="3.05" valueMax="4.2" valueMin="1.9">1.9–4.2 mm</quantity>
|
||
), ID
|
||
<superScript id="7C539BE3FFB6FF97239BFCA065C2FCBB" attach="left" box="[1221,1245,846,876]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
<specimenCode id="DB809ED0FFB6FF9723BEFCBF6466FC89" box="[1248,1401,847,876]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA60</specimenCode>
|
||
─64,
|
||
<specimenCode id="DB809ED0FFB6FF972415FC8662FBFC76" box="[843,996,886,915]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA66</specimenCode>
|
||
─67;
|
||
</materialsCitation>
|
||
<materialsCitation id="3B4E3CF6FFB6FF972375FC8664A7FBEC" collectingDate="2019-12" collectingMethod="scoop net" collectionCode="DBTLDA" country="India" latitude="10.820278" location="Agatti Island" longLatPrecision="21" longitude="72.16861" pageId="4" pageNumber="5" specimenCode="DBTLDA68" specimenCount="6" specimenCount-female="7" specimenCount-male="6" typeStatus="paratype">
|
||
<specimenCount id="9D20FD22FFB6FF972375FC866596FC76" box="[1067,1161,886,915]" count="6" pageId="4" pageNumber="5" type="male">6 males</specimenCount>
|
||
(CL 1.9–3.0 mm), ID
|
||
<superScript id="7C539BE3FFB6FF9722FCFC8764A5FC60" attach="left" box="[1442,1466,885,915]" fontSize="7" pageId="4" pageNumber="5">no</superScript>
|
||
<specimenCode id="DB809ED0FFB6FF972415FC6D62F5FC5F" box="[843,1002,925,954]" collectionCode="DBTLDA" pageId="4" pageNumber="5">DBTLDA68</specimenCode>
|
||
─73,
|
||
<geoCoordinate id="EE12506CFFB6FF97236DFC6E65DAFC5E" box="[1075,1221,926,955]" degrees="10" direction="north" minutes="49" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="13" value="10.820278">10°49’13”N</geoCoordinate>
|
||
<geoCoordinate id="EE12506CFFB6FF972391FC6E647DFC5E" box="[1231,1378,926,955]" degrees="72" direction="east" minutes="10" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="07" value="72.16861">72°10’07”E</geoCoordinate>
|
||
, depth 1.0–2.0 m,
|
||
<collectingMethod id="52674EBCFFB6FF97248BFC356552FC04" box="[981,1101,965,993]" pageId="4" pageNumber="5">scoop net</collectingMethod>
|
||
, associated with
|
||
<taxonomicName id="4C264D28FFB6FF972242FC36628BFBEC" authority=", Agatti" authorityName="Agatti" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="5" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFB6FF972242FC3664ACFC04" box="[1308,1459,965,994]" italics="true" pageId="4" pageNumber="5">H. magnifica</emphasis>
|
||
,
|
||
<location id="8EF96070FFB6FF972415FC1C62FCFBEC" LSID="urn:lsid:plazi:treatment:038F87BDFFB6FF9E2619FAF46527F8D3:8EF96070FFB6FF972415FC1C62FCFBEC" box="[843,995,1004,1033]" country="India" latitude="10.820278" longLatPrecision="21" longitude="72.16861" name="Agatti Island" pageId="4" pageNumber="5">Agatti Island</location>
|
||
</taxonomicName>
|
||
, Lakshadweep,
|
||
<collectingCountry id="F331763BFFB6FF9723C6FC1C65C6FBEC" box="[1176,1241,1004,1033]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
|
||
, in
|
||
<date id="FF98106BFFB6FF9723A3FC1C64A7FBEC" box="[1277,1464,1004,1033]" pageId="4" pageNumber="5" value="2019-12">
|
||
<collectingDate id="EFDCE983FFB6FF9723A3FC1C64A7FBEC" box="[1277,1464,1004,1033]" pageId="4" pageNumber="5" value="2019-12">December 2019</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6520FFB6FF992431FBC363CBFC59" lastPageId="10" lastPageNumber="11" pageId="4" pageNumber="5" type="description">
|
||
<paragraph id="8B9936ABFFB6FF972431FBC362F6FA32" blockId="4.[841,1469,1074,1967]" pageId="4" pageNumber="5">
|
||
<emphasis id="B952EAB9FFB6FF972431FBC362EDFBAA" box="[879,1010,1075,1103]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
|
||
. Small-sized palaemonid shrimp, body (
|
||
<figureCitation id="131D2A2EFFB6FF972409FBA962C8FB93" box="[855,983,1113,1142]" captionStart="Figure 2" captionStartId="5.[181,248,1456,1479]" captionTargetBox="[212,1437,151,1416]" captionTargetId="figure-39@5.[212,1437,151,1416]" captionTargetPageId="5" captionText="Figure 2.Actinimenes koyas sp.nov.from Agattiisland,India.(a) Holotype:ovigerous female(CL 3.5 mm,NBFGR/PALAKOY─01); (b) paratype, male (CL 3.2 mm, NBFGR: DBTLDA62, Ethanol preserved)." figureDoi="http://doi.org/10.5281/zenodo.10904876" httpUri="https://zenodo.org/record/10904876/files/figure.png" pageId="4" pageNumber="5">Fig. 2A, B</figureCitation>
|
||
) usually glabrous. Rostrum (
|
||
<figureCitation id="131D2A2EFFB6FF972208FBA964A4FB93" box="[1366,1467,1113,1142]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="4" pageNumber="5">Fig. 3A,</figureCitation>
|
||
B) well developed, straight or slightly depressed, about 0.65–0.72 times as long as carapace, reaching distal margin of the third antennular peduncle; dorsal margin bearing 5 to 8 acute teeth, slightly decreasing in size, distally with a row of 7–14 long plumose setae on rostrum underneath of each tooth; first dorsal tooth found posterior to posterior orbital margin; ventral margin convex with small ventral tooth at 0.7 of length.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB6FF97242FFA126494F84A" blockId="4.[841,1469,1074,1967]" pageId="4" pageNumber="5">
|
||
Carapace (
|
||
<figureCitation id="131D2A2EFFB6FF9724ABFA126575FA1B" box="[1013,1130,1506,1535]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="4" pageNumber="5">Fig. 3A, B</figureCitation>
|
||
) smooth, glabrous, without epigastric or supraorbital spines; inferior orbital angle well developed, acute in lateral view; antennal spine moderate in size, marginal, found in bottom of inferior orbital angle, slightly exceeding distal end of orbital angle;hepatic spine prominent, smaller than antennal, situated slightly lower than level of antennal spine; anterolateral angle of carapace region simply rounded. Eyes (
|
||
<figureCitation id="131D2A2EFFB6FF9724E8F8EC6519F8DC" box="[950,1030,1820,1849]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="4" pageNumber="5">Fig.3C</figureCitation>
|
||
)well developed with globular-shaped cornea and distinct accessory pigment spot present posterodorsally; eyestalks almost equal in length to proximal width and slightly swollen proximally.
|
||
</paragraph>
|
||
<caption id="DF596623FFB7FF9627EBFA4062B5FA02" ID-DOI="http://doi.org/10.5281/zenodo.10904876" ID-Zenodo-Dep="10904876" httpUri="https://zenodo.org/record/10904876/files/figure.png" pageId="5" pageNumber="6" startId="5.[181,248,1456,1479]" targetBox="[212,1437,151,1416]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B9936ABFFB7FF9627EBFA4062B5FA02" blockId="5.[181,1467,1454,1512]" pageId="5" pageNumber="6">
|
||
<emphasis id="B952EAB9FFB7FF9627EBFA40600DFA22" bold="true" box="[181,274,1456,1479]" pageId="5" pageNumber="6">Figure 2.</emphasis>
|
||
<taxonomicName id="4C264D28FFB7FF96264CFA4060A5FA22" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[274,442,1455,1479]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB7FF96264CFA4060A5FA22" box="[274,442,1455,1479]" italics="true" pageId="5" pageNumber="6">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB7FF9626E0FA40631AFA22" box="[446,517,1456,1479]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Agatti island, India.(
|
||
<emphasis id="B952EAB9FFB7FF96245AFA40620FFA2D" bold="true" box="[772,784,1456,1480]" pageId="5" pageNumber="6">a</emphasis>
|
||
) Holotype: ovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01);
|
||
<emphasis id="B952EAB9FFB7FF9627EBFA3F61D1FA02" bold="true" box="[181,206,1487,1511]" pageId="5" pageNumber="6">(b</emphasis>
|
||
) paratype, male (CL 3.2 mm, NBFGR: DBTLDA62, Ethanol preserved).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936ABFFB7FF962785F9C1658EF849" blockId="5.[180,807,1585,1964]" lastBlockId="5.[841,1467,1585,1964]" pageId="5" pageNumber="6">
|
||
Antennular peduncle (
|
||
<figureCitation id="131D2A2EFFB7FF9626B0F9C1637FF9A8" box="[494,608,1585,1614]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="5" pageNumber="6">Fig. 3A, E</figureCitation>
|
||
) reaching distal margin of scaphocerite. Proximal segment broad, thin, about 1.9 times longer than central width; lateral margin barely convex, anterolateral margin produced angular lobate with small acute distolateral tooth, row of plumose long setae on proximal margin; ventral margin with small submedian tooth found at 0.4 the length of segment. Stylocerite well developed, slender, acute reaches near middle of proximal segment. Intermediate segment short, about 1.1 times longer than width with small lateral lobe, 0.78 times as long as distal segments, and together equals 0.66 times of proximal segment length. Antennular f lagellum (
|
||
<figureCitation id="131D2A2EFFB7FF9623E8F955641AF926" box="[1206,1285,1701,1731]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="5" pageNumber="6">Fig. 3E</figureCitation>
|
||
) long, tapered; outer f lagellum biramous, proximal three segments of rami fused, shorter free ramus consists of 7 to 8 segments with 12–14 groups of aesthetascs, longer free ramus filiform, with 29–31 segments; inner flagellum slender, filiform, almost equal to outer one with 34–36 segments.
|
||
</paragraph>
|
||
<caption id="DF596623FFB4FF9527EBF8D665BCF87B" ID-DOI="http://doi.org/10.5281/zenodo.10904878" ID-Zenodo-Dep="10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="6" pageNumber="7" startId="6.[181,249,1830,1853]" targetBox="[185,1475,169,1769]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B9936ABFFB4FF9527EBF8D665BCF87B" blockId="6.[180,1467,1828,1950]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAB9FFB4FF9527EBF8D6600CF8D8" bold="true" box="[181,275,1830,1853]" pageId="6" pageNumber="7">Figure 3.</emphasis>
|
||
<taxonomicName id="4C264D28FFB4FF95264DF8D660A4F8DB" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[275,443,1830,1854]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB4FF95264DF8D660A4F8DB" box="[275,443,1830,1854]" italics="true" pageId="6" pageNumber="7">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB4FF9526E1F8D66317F8D8" box="[447,520,1830,1853]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, from Agatti island, India.
|
||
<emphasis id="B952EAB9FFB4FF95245FF8D5620DF8D8" bold="true" box="[769,786,1829,1853]" pageId="6" pageNumber="7">A</emphasis>
|
||
, Carapace of ovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view);
|
||
<emphasis id="B952EAB9FFB4FF952662F8B66052F8BB" bold="true" box="[316,333,1862,1886]" pageId="6" pageNumber="7">B</emphasis>
|
||
, anterior part of male carapace (DBTLDA62, lateral view;
|
||
<emphasis id="B952EAB9FFB4FF9524CEF8B562BDF8B8" bold="true" box="[912,930,1861,1885]" pageId="6" pageNumber="7">C</emphasis>
|
||
, dorsal view of cornea and eyestalk);
|
||
<emphasis id="B952EAB9FFB4FF95224DF8B66439F8BB" bold="true" box="[1299,1318,1862,1886]" pageId="6" pageNumber="7">D</emphasis>
|
||
, dorsal view of right scaphocerite;
|
||
<emphasis id="B952EAB9FFB4FF95262FF896609EF89B" bold="true" box="[369,385,1894,1918]" pageId="6" pageNumber="7">E</emphasis>
|
||
, dorsal view of antennular peduncle and flagellum;
|
||
<emphasis id="B952EAB9FFB4FF95242AF896629CF89B" bold="true" box="[884,899,1894,1918]" pageId="6" pageNumber="7">F</emphasis>
|
||
, left mandible (outer view);
|
||
<emphasis id="B952EAB9FFB4FF9523C6F89565B4F898" bold="true" box="[1176,1195,1893,1917]" pageId="6" pageNumber="7">G</emphasis>
|
||
, left maxillula (outer view);
|
||
<emphasis id="B952EAB9FFB4FF9527EAF87661D7F87B" bold="true" box="[180,200,1926,1950]" pageId="6" pageNumber="7">H</emphasis>
|
||
, left maxilla (outer view);
|
||
<emphasis id="B952EAB9FFB4FF952691F87660C6F87B" bold="true" box="[463,473,1926,1950]" pageId="6" pageNumber="7">I</emphasis>
|
||
, left first maxilliped (outer view);
|
||
<emphasis id="B952EAB9FFB4FF952472F8766229F87B" bold="true" box="[812,822,1926,1950]" pageId="6" pageNumber="7">J</emphasis>
|
||
, left second maxilliped (outer view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936ABFFB5FF942785FF676304FE0E" blockId="7.[179,807,151,1971]" pageId="7" pageNumber="8">
|
||
Antenna: basicerite bearing acute distolateral tooth. Ischiocerite and merocerite short. Carpocerite reaching middle of scaphocerite. Scaphocerite (
|
||
<figureCitation id="131D2A2EFFB5FF9425A6FF1561C5FECC" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="7" pageNumber="8">Fig. 3D</figureCitation>
|
||
) long with lamella reaching just beyond distal margin of distal antennular peduncle; lateral margin straight, with well-developed distolateral tooth; lamella rounded, about 2.1–2.3 times longer than breadth, with long distal plumose setae, extending beyond the distolateral tooth.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF942785FE0560AAFC72" blockId="7.[179,807,151,1971]" pageId="7" pageNumber="8">
|
||
Mandible(
|
||
<figureCitation id="131D2A2EFFB5FF942603FE0560B5FDF6" box="[349,426,501,531]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="7" pageNumber="8">Fig. 3F</figureCitation>
|
||
) without palp; the molar process stout, subcylindrical with 2 blunt teeth distally, numerous rows of small stout setae sub-distally; incisor processes short, slightly tapering distally, obliquely truncate, with 5 teeth distally and lateralmost slightly enlarged. Maxillula (
|
||
<figureCitation id="131D2A2EFFB5FF942500FD4863A7FD30" box="[606,696,696,725]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="7" pageNumber="8">Fig. 3G</figureCitation>
|
||
) normal form, palp feebly bilobed, outer lobe obscure, inner lobe with few small setae; upper lacinia with 10 stout long spines, several spiniform setae and plumose setae; lower lacinia subcylindrical, tapering distally, with numerous long setae.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF942785FC526047FBBF" blockId="7.[179,807,151,1971]" pageId="7" pageNumber="8">
|
||
Maxilla (
|
||
<figureCitation id="131D2A2EFFB5FF94260CFC5260ADFC5A" box="[338,434,930,959]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="7" pageNumber="8">Fig. 3H</figureCitation>
|
||
) with broad scaphognathite, without setae on inner and outer surfaces, anterior lobe wider than posterior with numerous long setae terminally; basal endite bilobed with spine-like long setae distally.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF942785FB9460DEFA5C" blockId="7.[179,807,151,1971]" pageId="7" pageNumber="8">
|
||
First maxilliped (
|
||
<figureCitation id="131D2A2EFFB5FF9426FDFB9460F5FB67" box="[419,490,1124,1154]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="7" pageNumber="8">Fig. 3I</figureCitation>
|
||
)endopodite short, slender, reaching distal margin of basal segment, with single long seta distally. Basal endite broad, rounded with long setae marginally, median margin with slender simple marginal setae. Coxal endite broadly rounded, elevated from basal endite by notch. Flagellum of exopodite well developed with 3 to 4 long plumose distal setae. Caridean lobe well developed; epipod bilobed without setae.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF942785FA33631FF915" blockId="7.[179,807,151,1971]" pageId="7" pageNumber="8">
|
||
Second maxilliped (
|
||
<figureCitation id="131D2A2EFFB5FF942681FA33632EFA05" box="[479,561,1475,1504]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="7" pageNumber="8">Fig. 3J</figureCitation>
|
||
) dactylus segment narrow, densely fringed with numerous serrulate setae. Propodal segment longer than dactylus, distomedial margin produced, with10 long spiniform setae.Carpus and merus short without setae. Ischium fused with basal segment. Coxal segment triangular-shaped medially with single setae. Exopod slender, with few long plumose setae distally.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF942785F90A6596FE20" blockId="7.[179,807,151,1971]" lastBlockId="7.[841,1469,151,1971]" pageId="7" pageNumber="8">
|
||
Third maxilliped (
|
||
<figureCitation id="131D2A2EFFB5FF94268DF90A632EF8F2" box="[467,561,1786,1815]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
|
||
) slender, reaching middle of antennal scaphocerite. Ultimate segment tapering, about 4.0 times longer than proximal width, 0.63 times as long as penultimate segment, with rows of short serrulate setae laterally, numerous longer setae distally. Penultimate segment slender, about 6.5 times longer than maximum width, 0.82 times as long as antepenultimate, with rows of long simple setae laterally. The antepenultimate segment slightly convex, about 4.5 times as long as proximal width. Coxa with small rounded epipod. Exopod slender, reaching distal end of antepenultimate segment, with few plumose setae distally.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF94242FFE3E6519FC00" blockId="7.[841,1469,151,1971]" pageId="7" pageNumber="8">
|
||
First pereopod (
|
||
<figureCitation id="131D2A2EFFB5FF942371FE3E6561FE09" box="[1071,1150,462,492]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="7" pageNumber="8">Fig. 4B</figureCitation>
|
||
) slender, extended beyond the distal end of scaphocerite by chela; fingers slender, sub-equal, concave, dactylus about 1.3–1.4 times longer than palm length, with small hooked terminal tooth, with numerous long setae; fixed finger with 6 to 7 groups serrulate setae on dorsal side. Palm short, sub-cylindrical, proximal region with row of setae. Carpus sub-cylindrical, about 4.6 times long as distal width, 2.9–3.0 times longer than palm, with distoventral cleaning setae. Merus cylindrical, about 0.9–0.95 times as long as carpus, 5.2 times longer than maximum width. Ischium equal to basis, about 0.35 times of merus length. Coxa with small setae on the medial process.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF94242FFC0062FBF92C" blockId="7.[841,1469,151,1971]" pageId="7" pageNumber="8">
|
||
Second pereopod: major and minor chelipeds (
|
||
<figureCitation id="131D2A2EFFB5FF942409FBE662CEFBD6" box="[855,977,1046,1076]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="7" pageNumber="8">Fig. 4C, D</figureCitation>
|
||
) well developed, similar, barely unequal in length, overreaching beyond the distal margin of antennular peduncle by length of chela.Major cheliped (
|
||
<figureCitation id="131D2A2EFFB5FF942409FB7B62C5FB4C" box="[855,986,1163,1193]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="7" pageNumber="8">Fig. 4C, C</figureCitation>
|
||
<figureCitation id="131D2A2EFFB5FF942485FB7C62F9FB4C" box="[987,998,1164,1193]" captionStart="Figure 1" captionStartId="2.[181,250,1847,1870]" captionTargetBox="[184,1459,472,1818]" captionTargetId="figure-146@2.[170,1473,463,1830]" captionTargetPageId="2" captionText="Figure 1. Map showing (a–d) location of Agatti Island, Lakshadweep, India, the type locality of Actinimenes koyas sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10904874" httpUri="https://zenodo.org/record/10904874/files/figure.png" pageId="7" pageNumber="8">1</figureCitation>
|
||
, C
|
||
<figureCitation id="131D2A2EFFB5FF942356FB7C6508FB4C" box="[1032,1047,1164,1193]" captionStart="Figure 2" captionStartId="5.[181,248,1456,1479]" captionTargetBox="[212,1437,151,1416]" captionTargetId="figure-39@5.[212,1437,151,1416]" captionTargetPageId="5" captionText="Figure 2.Actinimenes koyas sp.nov.from Agattiisland,India.(a) Holotype:ovigerous female(CL 3.5 mm,NBFGR/PALAKOY─01); (b) paratype, male (CL 3.2 mm, NBFGR: DBTLDA62, Ethanol preserved)." figureDoi="http://doi.org/10.5281/zenodo.10904876" httpUri="https://zenodo.org/record/10904876/files/figure.png" pageId="7" pageNumber="8">2</figureCitation>
|
||
): fingers slender, less than half of palm length (0.45–0.47times);dactylus slender, about 3.8 times as long as proximal depth, slowly tapering distally with stout terminal tooth, proximal half with 2 acute teeth, distal half of cutting edge concave, with numerous long setae; fixed finger with 4 to 5 teeth proximally, distal half concave with terminal tooth, with numerous long setae. Palm sub-cylindrical, about 2.9–3.0 times longer than maximum width. Carpus short, stout, about 1.1–1.3 times as long as distal width, tapering proximally, unarmed. Merus cylindrical, uniform, unarmed, about 2.1 times as long as carpus, 4.5 times as long as wide. Ischium with 0.7–0.78 times as long as merus, 0.45 times of palm length.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB5FF94242FF9236427F856" blockId="7.[841,1469,151,1971]" pageId="7" pageNumber="8">
|
||
Minor cheliped (
|
||
<figureCitation id="131D2A2EFFB5FF942315F92365B9F914" box="[1099,1190,1747,1777]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="7" pageNumber="8">Fig. 4D</figureCitation>
|
||
): fingers slender, less than half of palm length (0.50–0.51 times); dactylus slender, about 4.5 times as long as proximal depth, slowly tapering distally with stout terminal tooth, sub-proximal half with 2 acute teeth. Carpus, merus, and ischium similar with major cheliped.
|
||
</paragraph>
|
||
<caption id="DF596623FFBAFF9B27EBF8DD62E9F840" ID-DOI="http://doi.org/10.5281/zenodo.10904880" ID-Zenodo-Dep="10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="8" pageNumber="9" startId="8.[181,250,1837,1860]" targetBox="[210,1421,151,1781]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B9936ABFFBAFF9B27EBF8DD62E9F840" blockId="8.[181,1467,1837,1957]" pageId="8" pageNumber="9">
|
||
<emphasis id="B952EAB9FFBAFF9B27EBF8DD6007F8A1" bold="true" box="[181,280,1837,1860]" pageId="8" pageNumber="9">Figure 4.</emphasis>
|
||
<taxonomicName id="4C264D28FFBAFF9B267EF8DD60D1F8A0" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[288,462,1837,1861]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBAFF9B267EF8DD60D1F8A0" box="[288,462,1837,1861]" italics="true" pageId="8" pageNumber="9">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBAFF9B268BF8DD633FF8A1" box="[469,544,1837,1860]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62.
|
||
<emphasis id="B952EAB9FFBAFF9B231CF8DD654CF8A0" bold="true" box="[1090,1107,1837,1861]" pageId="8" pageNumber="9">A</emphasis>
|
||
, Left third maxilliped (outer view);
|
||
<emphasis id="B952EAB9FFBAFF9B27EBF8BD61D9F880" bold="true" box="[181,198,1869,1893]" pageId="8" pageNumber="9">B</emphasis>
|
||
, left first pereopod (lateral view);
|
||
<emphasis id="B952EAB9FFBAFF9B257DF8BD632AF880" bold="true" box="[547,565,1869,1893]" pageId="8" pageNumber="9">C</emphasis>
|
||
, major second pereopod (left, lateral view);
|
||
<emphasis id="B952EAB9FFBAFF9B24A7F8BD6507F880" bold="true" box="[1017,1048,1869,1893]" pageId="8" pageNumber="9">C1</emphasis>
|
||
, lateral view of major chela of the second pereopod;
|
||
<emphasis id="B952EAB9FFBAFF9B267DF89D605DF860" bold="true" box="[291,322,1901,1925]" pageId="8" pageNumber="9">C2</emphasis>
|
||
, dorsal view of major chela of same;
|
||
<emphasis id="B952EAB9FFBAFF9B25EDF89D63D9F860" bold="true" box="[691,710,1901,1925]" pageId="8" pageNumber="9">D</emphasis>
|
||
, minor second pereopod, right (lateral view);
|
||
<emphasis id="B952EAB9FFBAFF9B23CBF89D65BAF860" bold="true" box="[1173,1189,1901,1925]" pageId="8" pageNumber="9">E</emphasis>
|
||
, left third pereopod (lateral view);
|
||
<emphasis id="B952EAB9FFBAFF9B27A4F87D6016F840" bold="true" box="[250,265,1933,1957]" pageId="8" pageNumber="9">F</emphasis>
|
||
, right fourth pereopod (lateral view);
|
||
<emphasis id="B952EAB9FFBAFF9B25DAF87D6388F840" bold="true" box="[644,663,1933,1957]" pageId="8" pageNumber="9">G</emphasis>
|
||
, right fifth pereopod (lateral view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936ABFFBBFF9A2785FF67637CFD19" blockId="9.[179,807,151,842]" pageId="9" pageNumber="10">
|
||
Ambulatory pereopods(
|
||
<figureCitation id="131D2A2EFFBBFF9A26ACFF67637BFF51" box="[498,612,151,180]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="9" pageNumber="10">Fig.4E–G</figureCitation>
|
||
): slender,similar in structure,and moderately long.Third pereopod (
|
||
<figureCitation id="131D2A2EFFBBFF9A25A4FF4E61CBFEE7" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="9" pageNumber="10">Fig. 4E</figureCitation>
|
||
) extending beyond distal margin of scaphocerite by length of dactylus; dactylus acute, slender, about 0.28 times propodus length, 2.1 times as long as proximal width, with curved unguis, terminal end sharp with numerous long setae (
|
||
<figureCitation id="131D2A2EFFBBFF9A269BFE716306FE78" box="[453,537,384,414]" captionStart="Figure 5" captionStartId="9.[181,250,1831,1854]" captionTargetBox="[252,1357,907,1793]" captionTargetId="figure-318@9.[195,1449,888,1822]" captionTargetPageId="9" captionText="Figure 5.Actinimenes koyas sp. nov. holotype, ovigerous female, CL 3.5 mm, NBFGR/PALAKOY–01. A, Fourth sternal plate with median incision (inner view); B, left dactylus of the thirdpereopod (closer view); C, dorsal view of telson with uropod; D, posterior end of telson (closer view)." figureDoi="http://doi.org/10.5281/zenodo.10904882" httpUri="https://zenodo.org/record/10904882/files/figure.png" pageId="9" pageNumber="10">Fig. 5B</figureCitation>
|
||
). Propodus about 5.5 times as long as width, with numerous short and long setae distally. Carpus, merus, and ischium are about 0.60, 1.17,and0.55 times propodus length, respectively. Fourth pereopod(
|
||
<figureCitation id="131D2A2EFFBBFF9A26D6FDEC60CBFDDF" box="[392,468,540,570]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="9" pageNumber="10">Fig.4F</figureCitation>
|
||
)and fifth pereopod(
|
||
<figureCitation id="131D2A2EFFBBFF9A2599FDEC6206FDDF" box="[711,793,540,570]" captionStart="Figure 4" captionStartId="8.[181,250,1837,1860]" captionTargetBox="[210,1421,151,1781]" captionTargetId="figure-40@8.[195,1452,151,1805]" captionTargetPageId="8" captionText="Figure 4. Actinimenes koyas sp. nov. paratype, male, (CL 3.2 mm), NBFGR: DBTLDA62. A, Left third maxilliped (outer view); B, left first pereopod (lateral view); C, major second pereopod (left, lateral view); C1, lateral view of major chela of the second pereopod; C2, dorsal view of major chela of same; D, minor second pereopod, right (lateral view); E, left third pereopod (lateral view); F, right fourth pereopod (lateral view); G, right fifth pereopod (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.10904880" httpUri="https://zenodo.org/record/10904880/files/figure.png" pageId="9" pageNumber="10">Fig.4G</figureCitation>
|
||
) slightly longer than third; dactylus about 0.25 times as long as propodus, 2.1 times longer than proximal width; propodus about 6.4 times as long as wide; carpus, merus, and ischium about 0.52, 1.06, and 0.55 times propodus length, respectively.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBBFF9A2785FCF66209FCAF" blockId="9.[179,807,151,842]" pageId="9" pageNumber="10">
|
||
Fourth thoracic sternite (
|
||
<figureCitation id="131D2A2EFFBBFF9A2546FCF6636EFCC6" box="[536,625,774,803]" captionStart="Figure 5" captionStartId="9.[181,250,1831,1854]" captionTargetBox="[252,1357,907,1793]" captionTargetId="figure-318@9.[195,1449,888,1822]" captionTargetPageId="9" captionText="Figure 5.Actinimenes koyas sp. nov. holotype, ovigerous female, CL 3.5 mm, NBFGR/PALAKOY–01. A, Fourth sternal plate with median incision (inner view); B, left dactylus of the thirdpereopod (closer view); C, dorsal view of telson with uropod; D, posterior end of telson (closer view)." figureDoi="http://doi.org/10.5281/zenodo.10904882" httpUri="https://zenodo.org/record/10904882/files/figure.png" pageId="9" pageNumber="10">Fig. 5A</figureCitation>
|
||
) with angular broad lateral ridges and “
|
||
<collectionCode id="ED37AE6EFFBBFF9A26BEFCDD60E9FCAF" box="[480,502,813,842]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="10" type="Museum">V</collectionCode>
|
||
”-shaped median notch.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBBFF9A242FFF6765E0FE78" blockId="9.[841,1467,151,803]" pageId="9" pageNumber="10">
|
||
First pleopods: exopod broad, outer margin fringed with long setae; endopod short, about 0.38 times as long as exopod, lateral border with row of 9 plumose setae.
|
||
<collectingRegion id="49E2F849FFBBFF9A24CFFEFC62D0FECC" box="[913,975,268,297]" country="Maldives" name="Maale" pageId="9" pageNumber="10">Male</collectingRegion>
|
||
second pleopod with appendix masculina short and stout, reaching middle of exopod, distal outer margin fringed with few long setae; exopod broad, outer margin with long setae.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBBFF9A242FFE5862A6FCC6" blockId="9.[841,1467,151,803]" pageId="9" pageNumber="10">
|
||
Abdominal segments(
|
||
<figureCitation id="131D2A2EFFBBFF9A232AFE5865C1FE21" box="[1140,1246,424,453]" captionStart="Figure 2" captionStartId="5.[181,248,1456,1479]" captionTargetBox="[212,1437,151,1416]" captionTargetId="figure-39@5.[212,1437,151,1416]" captionTargetPageId="5" captionText="Figure 2.Actinimenes koyas sp.nov.from Agattiisland,India.(a) Holotype:ovigerous female(CL 3.5 mm,NBFGR/PALAKOY─01); (b) paratype, male (CL 3.2 mm, NBFGR: DBTLDA62, Ethanol preserved)." figureDoi="http://doi.org/10.5281/zenodo.10904876" httpUri="https://zenodo.org/record/10904876/files/figure.png" pageId="9" pageNumber="10">Fig.2A,B</figureCitation>
|
||
)smooth,glabrous, about 2.8 times as long as carapace; third abdominal tergite not produced in entire posterior margin. Fourth and fifth abdominal segments posterior-dorsally rounded; sixth abdominal segment elongate, about 1.6 times as long as fifth, produced barely pointed posteroventral angle, posterolateral angle acute. Pleura of first to fifth abdominal segments broadly rounded; fourth and fifth slightly produced posteriorly and rounded.
|
||
</paragraph>
|
||
<caption id="DF596623FFBBFF9A27EBF8D760DDF89B" ID-DOI="http://doi.org/10.5281/zenodo.10904882" ID-Zenodo-Dep="10904882" httpUri="https://zenodo.org/record/10904882/files/figure.png" pageId="9" pageNumber="10" startId="9.[181,250,1831,1854]" targetBox="[252,1357,907,1793]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B9936ABFFBBFF9A27EBF8D760DDF89B" blockId="9.[180,1467,1830,1918]" pageId="9" pageNumber="10">
|
||
<emphasis id="B952EAB9FFBBFF9A27EBF8D76009F8DB" bold="true" box="[181,278,1831,1854]" pageId="9" pageNumber="10">Figure 5.</emphasis>
|
||
<taxonomicName id="4C264D28FFBBFF9A2647F8D760DAF8DB" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[281,453,1830,1854]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBBFF9A2647F8D760DAF8DB" box="[281,453,1830,1854]" italics="true" pageId="9" pageNumber="10">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBBFF9A2694F8D7630BF8DB" box="[458,532,1831,1854]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype, ovigerous female, CL 3.5 mm, NBFGR/PALAKOY
|
||
<emphasis id="B952EAB9FFBBFF9A2324F8D66598F8DB" bold="true" box="[1146,1159,1830,1854]" pageId="9" pageNumber="10">–</emphasis>
|
||
01.
|
||
<emphasis id="B952EAB9FFBBFF9A23F5F8D665A3F8DB" bold="true" box="[1195,1212,1830,1854]" pageId="9" pageNumber="10">A</emphasis>
|
||
, Fourth sternal plate with median incision (inner view);
|
||
<emphasis id="B952EAB9FFBBFF9A2680F8B660EAF8BB" bold="true" box="[478,501,1862,1886]" pageId="9" pageNumber="10">B,</emphasis>
|
||
left dactylus of the third pereopod (closer view);
|
||
<emphasis id="B952EAB9FFBBFF9A2488F8B662F7F8BB" bold="true" box="[982,1000,1862,1886]" pageId="9" pageNumber="10">C</emphasis>
|
||
, dorsal view of telson with uropod;
|
||
<emphasis id="B952EAB9FFBBFF9A221AF8B66448F8BB" bold="true" box="[1348,1367,1862,1886]" pageId="9" pageNumber="10">D</emphasis>
|
||
, posterior end of telson (closer view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936ABFFB8FF992785FF67623BFD40" blockId="10.[180,806,151,677]" pageId="10" pageNumber="11">
|
||
Telson (
|
||
<figureCitation id="131D2A2EFFB8FF99261DFF6760DCFF51" box="[323,451,151,180]" captionStart="Figure 5" captionStartId="9.[181,250,1831,1854]" captionTargetBox="[252,1357,907,1793]" captionTargetId="figure-318@9.[195,1449,888,1822]" captionTargetPageId="9" captionText="Figure 5.Actinimenes koyas sp. nov. holotype, ovigerous female, CL 3.5 mm, NBFGR/PALAKOY–01. A, Fourth sternal plate with median incision (inner view); B, left dactylus of the thirdpereopod (closer view); C, dorsal view of telson with uropod; D, posterior end of telson (closer view)." figureDoi="http://doi.org/10.5281/zenodo.10904882" httpUri="https://zenodo.org/record/10904882/files/figure.png" pageId="10" pageNumber="11">Fig. 5C, D</figureCitation>
|
||
) slender, about 1.5 times as long as sixth abdominal segment, 3.5 times longer than anterior width, with 2 pairs of movable, well-developed dorsal spines appearing at 0.25 and 0.6 of telson length. Posterior margin triangular-shaped, 0.30 times anterior width, with 3 pairs of spines; lateral spine short and subequal to dorsal spines; intermediate spines well developed, slender, about 0.16 times telson length, 1.6 times length of submedian spines.Uropods exceeding length of telson; exopod slightly longer than endopod, about 2.25 times as long as width, with small fixed distolateral tooth; endopod with leaf-like structure, outer and inner margin bearing long setae.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB8FF992785FD2063CBFC59" blockId="10.[179,805,720,956]" pageId="10" pageNumber="11">
|
||
<emphasis id="B952EAB9FFB8FF992785FD2060A8FD08" box="[219,439,720,749]" italics="true" pageId="10" pageNumber="11">Coloration in life</emphasis>
|
||
. Body and appendages are transparent or pale brownish (
|
||
<figureCitation id="131D2A2EFFB8FF992553FD096356FCF3" box="[525,585,761,790]" captionStart="Figure 6" captionStartId="10.[181,250,1916,1939]" captionTargetBox="[181,1466,1022,1879]" captionTargetId="figure-365@10.[181,1466,1022,1879]" captionTargetPageId="10" captionText="Figure 6.Collection site, assemblages of Heteractis magnifica at Agatti Island, Lakshadweep, India with live colorationof Actinimenes koyas sp. nov. (Depth 1–2 m)." figureDoi="http://doi.org/10.5281/zenodo.10904884" httpUri="https://zenodo.org/record/10904884/files/figure.png" pageId="10" pageNumber="11">Fig.6</figureCitation>
|
||
). Hepatic pancreas and cardiac regions visible as pale orange through integument. Abdominal segments and telson are almost fully translucent without any spots. Eyes are greenish with a dark mark posterior-dorsally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6520FFB8FF98242FFF676374FB4F" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="8B9936ABFFB8FF98242FFF67604CFCED" blockId="10.[841,1469,151,955]" lastBlockId="11.[180,807,151,776]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="B952EAB9FFB8FF99242FFF67658AFF51" box="[881,1173,151,180]" italics="true" pageId="10" pageNumber="11">Distribution and habitat</emphasis>
|
||
. Presently known from the coral atolls of Agatti Island (near Kalpetti Island),
|
||
<collectingCountry id="F331763BFFB8FF992417FF1962EDFEE3" box="[841,1010,233,262]" name="India" pageId="10" pageNumber="11">Lakshadweep</collectingCountry>
|
||
,
|
||
<collectingCountry id="F331763BFFB8FF9924A2FF19655FFEE3" box="[1020,1088,233,262]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
|
||
. The specimen
|
||
<taxonomicName id="4C264D28FFB8FF9923A4FF196444FEE3" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1274,1371,233,262]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB8FF9923A4FF196444FEE3" box="[1274,1371,233,262]" italics="true" pageId="10" pageNumber="11">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB8FF99223EFF1964A6FEE0" box="[1376,1465,233,261]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found in two locations at the sampling site. This species was seen in association with the magnificent sea anemone
|
||
<taxonomicName id="4C264D28FFB8FF9924E2FE946553FE9A" box="[956,1100,355,384]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFB8FF9924E2FE946553FE9A" box="[956,1100,355,384]" italics="true" pageId="10" pageNumber="11">H. magnifica</emphasis>
|
||
</taxonomicName>
|
||
, assemblages (
|
||
<figureCitation id="131D2A2EFFB8FF9923A8FE93642BFE65" box="[1270,1332,355,384]" captionStart="Figure 6" captionStartId="10.[181,250,1916,1939]" captionTargetBox="[181,1466,1022,1879]" captionTargetId="figure-365@10.[181,1466,1022,1879]" captionTargetPageId="10" captionText="Figure 6.Collection site, assemblages of Heteractis magnifica at Agatti Island, Lakshadweep, India with live colorationof Actinimenes koyas sp. nov. (Depth 1–2 m)." figureDoi="http://doi.org/10.5281/zenodo.10904884" httpUri="https://zenodo.org/record/10904884/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
|
||
)belonging to the order
|
||
<taxonomicName id="4C264D28FFB8FF9924BDFE7C657EFE4C" authorityName="Hertwig" authorityYear="1882" box="[995,1121,396,425]" class="Anthozoa" kingdom="Animalia" order="Actiniaria" pageId="10" pageNumber="11" phylum="Cnidaria" rank="order">Actiniaria</taxonomicName>
|
||
, which is widely distributed in the
|
||
<collectingCountry id="F331763BFFB8FF9924C7FE44655FFE34" box="[921,1088,436,465]" name="India" pageId="10" pageNumber="11">Lakshadweep</collectingCountry>
|
||
islands. It was noticed that the shrimp mostly appeared in between the tentacles of the sea anemone, where they can easily hide from predators. The aggregations of shrimps on the sea anemone were typically observed in the daytime only. We recorded a salinity of 34 ppt and a temperature of 26 °C at the sampling locations. It was observed that several peacock-tail anemone shrimp
|
||
<taxonomicName id="4C264D28FFB8FF992258FD2264A6FD0A" box="[1286,1465,722,751]" class="Malacostraca" family="Palaemonidae" genus="Ancylocaris" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevicarpalis">
|
||
<emphasis id="B952EAB9FFB8FF992258FD2264A6FD0A" box="[1286,1465,722,751]" italics="true" pageId="10" pageNumber="11">A. brevicarpalis</emphasis>
|
||
</taxonomicName>
|
||
and marbled shrimp
|
||
<taxonomicName id="4C264D28FFB8FF992303FD0B628FFCA4" authority="(Olivier, 1811)" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Malacostraca" family="Hippolytidae" genus="Saron" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="marmoratus">
|
||
<emphasis id="B952EAB9FFB8FF992303FD0B645AFCFD" box="[1117,1349,763,792]" italics="true" pageId="10" pageNumber="11">Saron marmoratus</emphasis>
|
||
(Olivier, 1811)
|
||
</taxonomicName>
|
||
were also found in the same collection site. However,
|
||
<taxonomicName id="4C264D28FFB8FF99249DFCBD653CFC8C" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[963,1059,844,873]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB8FF99249DFCBD653CFC8C" box="[963,1059,844,873]" italics="true" pageId="10" pageNumber="11">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB8FF992379FCBD6560FC8C" box="[1063,1151,845,873]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was the dominant (~93 %) species group on the sea anemone assemblage. The faunal assemblage and habitat of
|
||
<taxonomicName id="4C264D28FFB8FF9923B0FC6E644BFC5E" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1262,1364,926,955]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB8FF9923B0FC6E644BFC5E" box="[1262,1364,926,955]" italics="true" pageId="10" pageNumber="11">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB8FF992200FC6F64A2FC5E" box="[1374,1469,927,955]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were documented as a video (https://youtu.be/ jb9DmGSRn_o). Further, ornamental fishes were also noticed in the collection site such as the swallowtail cardinalfish,
|
||
<taxonomicName id="4C264D28FFB9FF98260CFEFE63C0FECE" authority="(Weber, 1909)" baseAuthorityName="Weber" baseAuthorityYear="1909" box="[338,735,270,299]" family="Apogonidae" genus="Verulux" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cypselurus">
|
||
<emphasis id="B952EAB9FFB9FF98260CFEFE6335FECE" box="[338,554,270,299]" italics="true" pageId="11" pageNumber="12">Verulux cypselurus</emphasis>
|
||
(Weber, 1909)
|
||
</taxonomicName>
|
||
; bluespotted cardinalfish,
|
||
<taxonomicName id="4C264D28FFB9FF98269AFEC6601FFE9F" authority="(Bleeker, 1856)" baseAuthorityName="Bleeker" baseAuthorityYear="1856" family="Apogonidae" genus="Apogon" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="apogonides">
|
||
<emphasis id="B952EAB9FFB9FF98269AFEC663B4FEB6" box="[452,683,310,339]" italics="true" pageId="11" pageNumber="12">Apogon apogonides</emphasis>
|
||
(Bleeker, 1856)
|
||
</taxonomicName>
|
||
; Kashmir snapper,
|
||
<taxonomicName id="4C264D28FFB9FF9826BEFEAF601FFE47" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kasmira">
|
||
<emphasis id="B952EAB9FFB9FF9826BEFEAF63B6FE9E" box="[480,681,350,379]" italics="true" pageId="11" pageNumber="12">Lutjanus kasmira</emphasis>
|
||
(Forsskål, 1775)
|
||
</taxonomicName>
|
||
; three spot damsel,
|
||
<taxonomicName id="4C264D28FFB9FF982555FE75606BFE2F" authority="(Ruppell, 1829)" baseAuthorityName="Ruppell" baseAuthorityYear="1829" family="Pomacentridae" genus="Dascyllus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="trimaculatus">
|
||
<emphasis id="B952EAB9FFB9FF982555FE75623BFE47" box="[523,804,389,418]" italics="true" pageId="11" pageNumber="12">Dascyllus trimaculatus</emphasis>
|
||
(Rüppell, 1829)
|
||
</taxonomicName>
|
||
; cleaner wrasse,
|
||
<taxonomicName id="4C264D28FFB9FF98256AFE5D60A7FE17" authority="(Valenciennes 1839)" baseAuthorityName="Valenciennes" baseAuthorityYear="1839" family="Labridae" genus="Labroides" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dimidiatus">
|
||
<emphasis id="B952EAB9FFB9FF98256AFE5D623AFE2F" box="[564,805,429,458]" italics="true" pageId="11" pageNumber="12">Labroides dimidiatus</emphasis>
|
||
(Valenciennes 1839)
|
||
</taxonomicName>
|
||
; blue-green damsel,
|
||
<taxonomicName id="4C264D28FFB9FF9825E0FE2560A7FDFF" authority="(Cuvier, 1830)" baseAuthorityName="Cuvier" baseAuthorityYear="1830" family="Pomacentridae" genus="Chromis" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="viridis">
|
||
<emphasis id="B952EAB9FFB9FF9825E0FE2561E2FDFC" italics="true" pageId="11" pageNumber="12">Chromis viridis</emphasis>
|
||
(Cuvier, 1830)
|
||
</taxonomicName>
|
||
; scarlet soldierfish,
|
||
<taxonomicName id="4C264D28FFB9FF9825FAFE0D60B9FDA4" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" family="Holocentridae" genus="Myripristis" kingdom="Animalia" order="Beryciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pralinia">
|
||
<emphasis id="B952EAB9FFB9FF9825FAFE0D6013FDA4" italics="true" pageId="11" pageNumber="12">Myripristis pralinia</emphasis>
|
||
Cuvier, 1829
|
||
</taxonomicName>
|
||
;twinspot damselfish,
|
||
<taxonomicName id="4C264D28FFB9FF9825FDFDD46369FD8C" authority="(Quoy and Gaimard, 1825)" baseAuthorityName="Quoy and Gaimard" baseAuthorityYear="1825" family="Pomacentridae" genus="Chrysiptera" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="biocellata">
|
||
<emphasis id="B952EAB9FFB9FF9825FDFDD4603BFD8C" italics="true" pageId="11" pageNumber="12">Chrysiptera biocellata</emphasis>
|
||
(Quoy and Gaimard, 1825)
|
||
</taxonomicName>
|
||
; moon wrasse,
|
||
<taxonomicName id="4C264D28FFB9FF9827EBFD84639FFD75" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[181,640,627,657]" family="Labridae" genus="Thalassoma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="lunare">
|
||
<emphasis id="B952EAB9FFB9FF9827EBFD846083FD74" box="[181,412,628,657]" italics="true" pageId="11" pageNumber="12">Thalassoma lunare</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
; black streak surgeon,
|
||
<taxonomicName id="4C264D28FFB9FF98267AFD6B61EAFD05" authority="Duncker and Mohr, 1929" authorityName="Duncker and Mohr" authorityYear="1929" family="Acanthuridae" genus="Acanthurus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="nigricauda">
|
||
<emphasis id="B952EAB9FFB9FF98267AFD6B6330FD5D" box="[292,559,667,696]" italics="true" pageId="11" pageNumber="12">Acanthurus nigricauda</emphasis>
|
||
Duncker and Mohr, 1929
|
||
</taxonomicName>
|
||
; and
|
||
<collectingCountry id="F331763BFFB9FF982671FD33608DFD05" box="[303,402,707,736]" name="Maldives" pageId="11" pageNumber="12">Maldive</collectingCountry>
|
||
anemonefish,
|
||
<taxonomicName id="4C264D28FFB9FF982565FD336050FCED" authority="Regan, 1908" authorityName="Regan" authorityYear="1908" family="Pomacentridae" genus="Amphiprion" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="nigripes">
|
||
<emphasis id="B952EAB9FFB9FF982565FD33623AFD05" box="[571,805,707,736]" italics="true" pageId="11" pageNumber="12">Amphiprion nigripes</emphasis>
|
||
Regan, 1908
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF596623FFB8FF9927EBF88C60C3F856" ID-DOI="http://doi.org/10.5281/zenodo.10904884" ID-Zenodo-Dep="10904884" httpUri="https://zenodo.org/record/10904884/files/figure.png" pageId="10" pageNumber="11" startId="10.[181,250,1916,1939]" targetBox="[181,1466,1022,1879]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8B9936ABFFB8FF9927EBF88C60C3F856" blockId="10.[181,1465,1915,1971]" pageId="10" pageNumber="11">
|
||
<emphasis id="B952EAB9FFB8FF9927EBF88C600BF876" bold="true" box="[181,276,1916,1939]" pageId="10" pageNumber="11">Figure 6.</emphasis>
|
||
Collection site, assemblages of
|
||
<taxonomicName id="4C264D28FFB8FF99251DF88C621CF876" box="[579,771,1915,1939]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="10" pageNumber="11" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFB8FF99251DF88C621CF876" box="[579,771,1915,1939]" italics="true" pageId="10" pageNumber="11">Heteractis magnifica</emphasis>
|
||
</taxonomicName>
|
||
at Agatti Island, Lakshadweep, India with live coloration of
|
||
<taxonomicName id="4C264D28FFB8FF992219F88C61F5F856" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB8FF992219F88C61F5F856" italics="true" pageId="10" pageNumber="11">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB8FF9927AEF86C6025F856" box="[240,314,1948,1971]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Depth 1
|
||
<emphasis id="B952EAB9FFB8FF9926C4F86B60B8F856" bold="true" box="[410,423,1947,1971]" pageId="10" pageNumber="11">–</emphasis>
|
||
2 m).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936ABFFB9FF982785FCC163F1FBF1" blockId="11.[180,805,817,1044]" pageId="11" pageNumber="12">
|
||
<emphasis id="B952EAB9FFB9FF982785FCC16040FCAB" box="[219,351,817,846]" italics="true" pageId="11" pageNumber="12">Etymology</emphasis>
|
||
. Koyas are an important ethnic community of
|
||
<collectingCountry id="F331763BFFB9FF982622FCA86330FC90" box="[380,559,856,885]" name="India" pageId="11" pageNumber="12">Lakshadweep</collectingCountry>
|
||
. Koyas have made a significant contribution to the development and preservation of the heritage of the society at
|
||
<collectingCountry id="F331763BFFB9FF9827EBFC3F6041FC09" box="[181,350,975,1004]" name="India" pageId="11" pageNumber="12">Lakshadweep</collectingCountry>
|
||
. The present species is named
|
||
<taxonomicName id="4C264D28FFB9FF98259AFC206204FC08" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[708,795,976,1005]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="koyas">“Koyas</taxonomicName>
|
||
” to honor the local community at
|
||
<collectingCountry id="F331763BFFB9FF98251EFC0763F5FBF1" box="[576,746,1015,1044]" name="India" pageId="11" pageNumber="12">Lakshadweep</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB9FF982785FBCE6374FB4F" blockId="11.[180,806,1085,1194]" pageId="11" pageNumber="12">
|
||
<emphasis id="B952EAB9FFB9FF982785FBCE6099FBBF" box="[219,390,1085,1114]" italics="true" pageId="11" pageNumber="12">DNA barcodes</emphasis>
|
||
. COI and 16S sequences from the
|
||
<typeStatus id="549D8809FFB9FF9827E8FB956000FB67" box="[182,287,1125,1154]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="549D8809FFB9FF98260DFB9560A2FB64" box="[339,445,1125,1153]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
|
||
, and
|
||
<taxonomicName id="4C264D28FFB9FF9826ABFB956361FB67" box="[501,638,1125,1154]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFB9FF9826ABFB956361FB67" box="[501,638,1125,1154]" italics="true" pageId="11" pageNumber="12">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
were obtained and submitted to GenBank (
|
||
<tableCitation id="C6A40310FFB9FF98254FFB7C6347FB4F" box="[529,600,1164,1194]" captionStart="Table 1" captionStartId="4.[181,244,152,175]" captionTargetPageId="4" captionText="Table 1. Details of DNA sequences of Actinimenes koyas sp. nov., two other Actinimenes species and outgroup species obtained from GenBank." pageId="11" pageNumber="12">Tab. 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6520FFB9FF9E2785FB226527F8D3" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="8B9936ABFFB9FF982785FB2264A6FEB5" blockId="11.[180,806,1234,1939]" lastBlockId="11.[841,1469,151,1938]" pageId="11" pageNumber="12">
|
||
<emphasis id="B952EAB9FFB9FF982785FB226055FB0A" box="[219,330,1234,1263]" italics="true" pageId="11" pageNumber="12">Remarks</emphasis>
|
||
. Generally, shrimps in the genus
|
||
<taxonomicName id="4C264D28FFB9FF9827EBFB0B605CFAF2" authorityName="Duris and Horka" authorityYear="2017" box="[181,323,1275,1303]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAB9FFB9FF9827EBFB0B605CFAF2" box="[181,323,1275,1303]" italics="true" pageId="11" pageNumber="12">Actinimenes</emphasis>
|
||
</taxonomicName>
|
||
are associated with sea anemones and commonly distributed in Indo-West Pacific regions (
|
||
<bibRefCitation id="EFB74B5AFFB9FF9827E0FABA604CFA82" author="Kemp, S. W." box="[190,339,1354,1383]" pageId="11" pageNumber="12" pagination="113 - 288" refId="ref10211" refString="Kemp, S. W. 1922. Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24: 113 - 288." type="journal article" year="1922">Kemp,1922</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF982602FABA60F3FA82" author="Bruce, A. J." box="[348,492,1354,1383]" pageId="11" pageNumber="12" pagination="253 - 278" refId="ref8735" refString="Bruce, A. J. 1969. Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen, 43: 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9826ABFABA632CFA82" author="Bruce, A. J." box="[501,563,1354,1383]" pageId="11" pageNumber="12" pagination="89 - 153" refId="ref8864" refString="Bruce, A. J. 1976. A report on some pontoniinid shrimps collected from the Seychelle Islands by the FRV Manihine, 1972, with a review of the Seychelles pontoniinid shrimp fauna. Zoological Journal of the Linnean Society, 59: 89 - 153." type="journal article" year="1976">1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF982562FABA6364FA82" author="Bruce, A. J." box="[572,635,1354,1383]" pageId="11" pageNumber="12" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9825DDFABA63DDFA82" author="Bruce, A. J." box="[643,706,1354,1383]" pageId="11" pageNumber="12" pagination="1 - 39" refId="ref9008" refString="Bruce, A. J. 1980. On some pontoniine shrimps from Noumea, New Caledonia. Cahiers de l'Indo-Pacifique, 2: 1 - 39." type="journal article" year="1980">1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF982595FABA6064FA6B" author="Miyake, S. & Fujino, T." pageId="11" pageNumber="12" pagination="399 - 431" refId="ref10737" refString="Miyake, S. and Fujino, T. 1968. Pontoniid shrimps from the Palau Islands (Crustacea, Decapoda, Palaemonidae). Journal of the Faculty of Agriculture, Kyushu University, 14: 399 - 431." type="journal article" year="1968">Miyake and Fujino, 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9826DAFA8163ABFA6A" author="Suzuki, K. & Hayashi, K. I." box="[388,692,1393,1423]" pageId="11" pageNumber="12" pagination="193 - 208" refId="ref11233" refString="Suzuki, K. and Hayashi, K. I. 1977. Five caridean shrimps associated with sea anemones in central Japan. Publications of the Seto Marine Biological Laboratory, 24: 193 - 208." type="journal article" year="1977">Suzuki and Hayashi, 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9825E3FA8261EAFA53" author="Fransen, C. H. J. M." pageId="11" pageNumber="12" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF98265DFA6B6330FA53" author="Duris, Z." box="[259,559,1433,1467]" pageId="11" pageNumber="12" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D28FFB9FF98251BFA69623CFA53" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[581,803,1433,1462]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB9FF98251BFA69623CFA53" box="[581,803,1433,1462]" italics="true" pageId="11" pageNumber="12">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB9FF9827EBFA31600CFA38" box="[181,275,1473,1501]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the fourth species in the genus and has the following morphological characters similar to the three existing species in the genus (
|
||
<tableCitation id="C6A40310FFB9FF98259EF9E0620FF9C8" box="[704,784,1552,1581]" captionStart="Table 2" captionStartId="12.[181,242,152,175]" captionTargetPageId="12" captionText="Table 2.Comparison of morphologicalfeatures of Actinimenes koyas sp.nov.and other Actinimenes spp.:Anp–Antennular peduncle and TeL–Telson length." pageId="11" pageNumber="12">Tab. 2</tableCitation>
|
||
): (i) rostrum reaching or exceeding the antennular peduncles; (ii) rostral formula:
|
||
<quantity id="4CDE9B4EFFB9FF982563F9AF6385F998" box="[573,666,1631,1661]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.54" metricValueMin="1.27" pageId="11" pageNumber="12" unit="in" value="7.5" valueMax="10.0" valueMin="5.0">5–10 in</quantity>
|
||
dorsal and 0–2 teeth in ventral; (iii) antennal spine acute or only minutely produced; (iv) shorter free ramus of the outer antennular f lagellum with 7–8 segments and 8–14 groups of aesthetascs; (v) the fourth thoracic sternal plate produced anteriorly with a “U” or “
|
||
<collectionCode id="ED37AE6EFFB9FF9825FAF8D663A5F8A6" box="[676,698,1830,1859]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="12" type="Museum">V</collectionCode>
|
||
”-shaped median, narrow and deep notch; (vi) first pereopod: length ratio of finger and palm is1.0–1.4 times, length ratio of carpus and palm is 1.2 to 3.0 times; (vii) second pereopod merus is 0.66–0.75 times as long as palm; (viii) the dactylus of ambulatory pereopods have minute spinulate or simple structure around the base of unguis (
|
||
<bibRefCitation id="EFB74B5AFFB9FF9824A0FEC3658DFEB5" author="Bruce, A. J." box="[1022,1170,307,336]" pageId="11" pageNumber="12" pagination="20 - 36" refId="ref9236" refString="Bruce, A. J. 2010. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the Coral reefs 2009 Heron Island expedition. Zootaxa, 2604: 20 - 36." type="journal article" year="2010">Bruce, 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9823C5FEC564B2FEB5" author="Duris, Z." box="[1179,1453,307,341]" pageId="11" pageNumber="12" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB9FF982431FEAB6498FCC0" blockId="11.[841,1469,151,1938]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C264D28FFB9FF982431FEAB6525FE92" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[879,1082,346,375]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB9FF982431FEAB6525FE92" box="[879,1082,346,375]" italics="true" pageId="11" pageNumber="12">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB9FF982360FEAB658CFE92" box="[1086,1171,347,375]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
showed some remarkable morphological differences between the sexes: the rostrum of the male is mostly horizontal and reaches the distal end of the antennular peduncle (
|
||
<figureCitation id="131D2A2EFFB9FF98220AFE2064B7FE08" box="[1364,1448,464,493]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="11" pageNumber="12">Fig. 3B</figureCitation>
|
||
), whereas in the female the rostrum is slightly depressed and barely reaches the distal end of the antennular peduncle (
|
||
<figureCitation id="131D2A2EFFB9FF982490FDB56539FD87" box="[974,1062,581,610]" captionStart="Figure 3" captionStartId="6.[181,249,1830,1853]" captionTargetBox="[185,1475,169,1769]" captionTargetId="figure-40@6.[170,1485,151,1797]" captionTargetPageId="6" captionText="Figure 3.Actinimenes koyas sp.nov., from Agatti island,India.A, Carapace ofovigerous female (CL 3.5 mm, NBFGR/PALAKOY─01, lateral view); B, anterior part of male carapace (DBTLDA62, lateral view; C, dorsal view of cornea and eyestalk); D, dorsal view of right scaphocerite; E, dorsal view of antennular peduncle and flagellum; F, left mandible (outer view); G, left maxillula (outer view); H, left maxilla (outer view); I, left first maxilliped (outer view); J, left second maxilliped (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10904878" httpUri="https://zenodo.org/record/10904878/files/figure.png" pageId="11" pageNumber="12">Fig. 3A</figureCitation>
|
||
). The range of the rostral dorsal teeth is five to seven for males and eight for females. The inferior orbital angle of the male is slightly wider than in females; five to six cleaning setae are present on the distoventral region of carpus of the first pereopod for males and six to eight are present in females.
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFB9FF98242FFCDF62FEF877" blockId="11.[841,1469,151,1938]" pageId="11" pageNumber="12">
|
||
Among the three allied species,
|
||
<taxonomicName id="4C264D28FFB9FF98225BFCC06482FCA9" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1285,1437,815,844]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF98225BFCC06482FCA9" box="[1285,1437,815,844]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
is morphologically most similar to
|
||
<taxonomicName id="4C264D28FFB9FF98238BFCA76427FC96" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1237,1336,854,883]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB9FF98238BFCA76427FC96" box="[1237,1336,854,883]" italics="true" pageId="11" pageNumber="12">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB9FF982260FCA76488FC96" box="[1342,1431,855,883]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the shape of the rostrum; general appearance of the hepatic spine on the carapace; and the length ratio and width of the scaphocerite (
|
||
<tableCitation id="C6A40310FFB9FF9823CAFC3B65C0FC0C" box="[1172,1247,971,1001]" captionStart="Table 2" captionStartId="12.[181,242,152,175]" captionTargetPageId="12" captionText="Table 2.Comparison of morphologicalfeatures of Actinimenes koyas sp.nov.and other Actinimenes spp.:Anp–Antennular peduncle and TeL–Telson length." pageId="11" pageNumber="12">Tab. 2</tableCitation>
|
||
). In both species, the stylocerite is slender and reaches near the middle of the first antennular segment; the first pereopods have subspatulate fingers; there is dentition on the chela of the second pereopod; and the telson has two pairs of dorsal and three pairs of posterior spines. However,
|
||
<taxonomicName id="4C264D28FFB9FF982498FB466536FB36" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[966,1065,1206,1235]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFB9FF982498FB466536FB36" box="[966,1065,1206,1235]" italics="true" pageId="11" pageNumber="12">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFB9FF982371FB466596FB37" box="[1071,1161,1206,1234]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C264D28FFB9FF982278FB4664A5FB36" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1318,1466,1206,1235]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF982278FB4664A5FB36" box="[1318,1466,1206,1235]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
(summarized in
|
||
<tableCitation id="C6A40310FFB9FF982344FB2D657AFB1F" box="[1050,1125,1245,1274]" captionStart="Table 2" captionStartId="12.[181,242,152,175]" captionTargetPageId="12" captionText="Table 2.Comparison of morphologicalfeatures of Actinimenes koyas sp.nov.and other Actinimenes spp.:Anp–Antennular peduncle and TeL–Telson length." pageId="11" pageNumber="12">Tab. 2</tableCitation>
|
||
) by having biramous outer antennular flagellum with proximal three segments of rami fused (
|
||
<emphasis id="B952EAB9FFB9FF9824A7FADC6509FAAD" box="[1017,1046,1324,1352]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
|
||
four in
|
||
<taxonomicName id="4C264D28FFB9FF98232CFADB641CFAAD" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1138,1283,1323,1352]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF98232CFADB641CFAAD" box="[1138,1283,1323,1352]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
); a shorter free ramus consisting of seven to eight segments with 12–14 groups of aesthetascs (
|
||
<emphasis id="B952EAB9FFB9FF9823EAFA8A65CEFA73" box="[1204,1233,1402,1430]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
|
||
shorter free ramus with only four segments and eight to nine groups of aesthetascs in
|
||
<taxonomicName id="4C264D28FFB9FF9824A6FA386595FA01" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1016,1162,1479,1508]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF9824A6FA386595FA01" box="[1016,1162,1479,1508]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
); the length of the telson is 3.5 times longer than anterior width (2.4 times for
|
||
<taxonomicName id="4C264D28FFB9FF982414F9E662C5F9D7" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[842,986,1557,1586]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF982414F9E662C5F9D7" box="[842,986,1557,1586]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
); fourth thoracic sternite with V-shaped median incision (
|
||
<emphasis id="B952EAB9FFB9FF982349F9CD652BF9BC" box="[1047,1076,1597,1625]" italics="true" pageId="11" pageNumber="12">vs.</emphasis>
|
||
U-shaped for
|
||
<taxonomicName id="4C264D28FFB9FF982389F9CD647AF9BC" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1239,1381,1596,1625]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF982389F9CD647AF9BC" box="[1239,1381,1596,1625]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
);third maxilliped penultimate segment 0.82 times as long as antepenultimate (0.75 times as long in
|
||
<taxonomicName id="4C264D28FFB9FF98224AF97B64B9F942" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1300,1446,1674,1703]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF98224AF97B64B9F942" box="[1300,1446,1674,1703]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
); and dactylus of ambulatory pereopods are without spinulation on the base of unguis(spinulated minutely in
|
||
<taxonomicName id="4C264D28FFB9FF982439F8F062EDF8F8" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[871,1010,1792,1821]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF982439F8F062EDF8F8" box="[871,1010,1792,1821]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFB74B5AFFB9FF982352F8F06580F8F8" author="Bruce, A. J." box="[1036,1183,1792,1821]" pageId="11" pageNumber="12" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce, 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9823F9F8F065FAF8F8" author="Bruce, A. J." box="[1191,1253,1792,1821]" pageId="11" pageNumber="12" pagination="20 - 36" refId="ref9236" refString="Bruce, A. J. 2010. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the Coral reefs 2009 Heron Island expedition. Zootaxa, 2604: 20 - 36." type="journal article" year="2010">2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9823B0F8F26297F8A1" author="Duris, Z." pageId="11" pageNumber="12" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
). Consequently,
|
||
<taxonomicName id="4C264D28FFB9FF982313F8D765C0F8A1" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1101,1247,1831,1860]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFB9FF982313F8D765C0F8A1" box="[1101,1247,1831,1860]" italics="true" pageId="11" pageNumber="12">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
was reported only from the
|
||
<collectingCountry id="F331763BFFB9FF9824E9F8BE659EF88E" box="[951,1153,1870,1899]" name="Marshall Islands" pageId="11" pageNumber="12">Marshall Islands</collectingCountry>
|
||
and Heron Island (
|
||
<bibRefCitation id="EFB74B5AFFB9FF982239F8BE6294F877" author="Bruce, A. J." pageId="11" pageNumber="12" pagination="39 - 81" refId="ref8913" refString="Bruce, A. J. 1977. Pontoniine shrimps in the collections of the Australian Museum. Records of the Australian Museum, 31: 39 - 81." type="journal article" year="1977">Bruce, 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFB9FF9824CBF88562CAF877" author="Bruce, A. J." box="[917,981,1909,1938]" pageId="11" pageNumber="12" pagination="20 - 36" refId="ref9236" refString="Bruce, A. J. 2010. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the Coral reefs 2009 Heron Island expedition. Zootaxa, 2604: 20 - 36." type="journal article" year="2010">2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF596623FFBEFF9F27EBFF6860BDFF2A" pageId="12" pageNumber="13" startId="12.[181,242,152,175]" targetBox="[189,1454,227,1323]" targetIsTable="true" targetPageId="12" targetType="table">
|
||
<paragraph id="8B9936ABFFBEFF9F27EBFF6860BDFF2A" blockId="12.[180,1467,151,207]" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F27EBFF686012FF4A" bold="true" box="[181,269,152,175]" pageId="12" pageNumber="13">Table 2.</emphasis>
|
||
Comparison of morphological features of
|
||
<taxonomicName id="4C264D28FFBEFF9F25F9FF67624CFF4A" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[679,851,151,175]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBEFF9F25F9FF67624CFF4A" box="[679,851,151,175]" italics="true" pageId="12" pageNumber="13">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBEFF9F2409FF6862BFFF4A" box="[855,928,152,175]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and other
|
||
<taxonomicName id="4C264D28FFBEFF9F2358FF6765B5FF4A" box="[1030,1194,151,175]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B952EAB9FFBEFF9F2358FF676567FF4B" box="[1030,1144,151,174]" italics="true" pageId="12" pageNumber="13">Actinimenes</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
: Anp
|
||
<emphasis id="B952EAB9FFBEFF9F2382FF6765F6FF4A" bold="true" box="[1244,1257,151,175]" pageId="12" pageNumber="13">–</emphasis>
|
||
Antennular peduncle and TeL
|
||
<emphasis id="B952EAB9FFBEFF9F2659FF47600BFF2A" bold="true" box="[263,276,183,207]" pageId="12" pageNumber="13">–</emphasis>
|
||
Telson length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936ABFFBEFF9F27E3FF006476FACE" pageId="12" pageNumber="13">
|
||
<table id="F926C40BFFBE006C27E3FF1364B1FACE" box="[189,1454,227,1323]" gridcols="6" gridrows="16" pageId="12" pageNumber="13">
|
||
<tr id="351634E9FFBE006C27E3FF1364B1FEF5" box="[189,1454,227,272]" gridrow="0" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FF1361F0FEF5" box="[189,239,227,272]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F27E3FF0061F0FEE1" bold="true" box="[189,239,240,260]" pageId="12" pageNumber="13">S. No.</emphasis>
|
||
</th>
|
||
<th id="76C75D95FFBE006C2657FF1360CEFEF5" box="[265,465,227,272]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F2660FF006084FEE6" bold="true" box="[318,411,240,259]" pageId="12" pageNumber="13">Characters</emphasis>
|
||
</th>
|
||
<th id="76C75D95FFBE006C26B9FF1363D6FEF5" box="[487,713,227,272]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F26A3FF14638BFEF5" authority="(Kemp, 1922)" baseAuthorityName="Kemp" baseAuthorityYear="1922" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F26A3FF14638BFEF5" bold="true" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F26A3FF1463ABFF12" bold="true" box="[509,692,228,247]" italics="true" pageId="12" pageNumber="13">Actinimenes inornatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F257FFF0D6391FEF5" author="Kemp, S. W." box="[545,654,253,272]" pageId="12" pageNumber="13" pagination="113 - 288" refId="ref10211" refString="Kemp, S. W. 1922. Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24: 113 - 288." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76C75D95FFBE006C2581FF1362DFFEF5" box="[735,960,227,272]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F25ADFF146294FEF5" authority="(Bruce, 1979)" baseAuthorityName="Bruce" baseAuthorityYear="1979" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBEFF9F25ADFF146294FEF5" bold="true" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F25ADFF1462B3FF13" bold="true" box="[755,940,227,247]" italics="true" pageId="12" pageNumber="13">Actinimenes ornatellus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F2447FF0D6299FEF5" author="Bruce, A. J." box="[793,902,253,272]" pageId="12" pageNumber="13" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce, 1979</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76C75D95FFBE006C2486FF1365A9FEF5" box="[984,1206,227,272]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F24AAFF14659CFEF5" authority="(Bruce, 1969)" baseAuthorityName="Bruce" baseAuthorityYear="1969" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F24AAFF14659CFEF5" bold="true" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F24AAFF146585FF12" bold="true" box="[1012,1178,228,247]" italics="true" pageId="12" pageNumber="13">Actinimenes ornatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F234FFF0D6562FEF5" author="Bruce, A. J." box="[1041,1149,253,272]" pageId="12" pageNumber="13" pagination="253 - 278" refId="ref8735" refString="Bruce, A. J. 1969. Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen, 43: 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76C75D95FFBE006C2391FF1364B1FEF5" box="[1231,1454,227,272]" gridcol="5" gridrow="0" pageId="12" pageNumber="13">
|
||
<emphasis id="B952EAB9FFBEFF9F23AAFF146440FEF5" bold="true" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F23AAFF146495FF13" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1268,1418,227,247]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBEFF9F23AAFF146495FF13" bold="true" box="[1268,1418,227,247]" italics="true" pageId="12" pageNumber="13">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBEFF9F2241FF0D6440FEF5" box="[1311,1375,253,272]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FED164B1FED0" box="[189,1454,289,309]" gridrow="1" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FED161F0FED0" box="[189,239,289,309]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">1</th>
|
||
<td id="76C75D95FFBE006C2657FED160CEFED0" box="[265,465,289,309]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">Rostrum</td>
|
||
<td id="76C75D95FFBE006C26B9FED163D6FED0" box="[487,713,289,309]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">Little beyond Anp</td>
|
||
<td id="76C75D95FFBE006C2581FED162DFFED0" box="[735,960,289,309]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">Exceed Anp</td>
|
||
<td id="76C75D95FFBE006C2486FED165A9FED0" box="[984,1206,289,309]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">Equal to Anp</td>
|
||
<td id="76C75D95FFBE006C2391FED164B1FED0" box="[1231,1454,289,309]" gridcol="5" gridrow="1" pageId="12" pageNumber="13">Equal to Anp</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FEB564B1FE97" box="[189,1454,325,370]" gridrow="2" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FEB561F0FE97" box="[189,239,325,370]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">2</th>
|
||
<td id="76C75D95FFBE006C2657FEB560CEFE97" box="[265,465,325,370]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">Rostral formula (dorsal/ventral)</td>
|
||
<td id="76C75D95FFBE006C26B9FEB563D6FE97" box="[487,713,325,370]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">6─8 0─2</td>
|
||
<td id="76C75D95FFBE006C2581FEB562DFFE97" box="[735,960,325,370]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">7─10 1─2</td>
|
||
<td id="76C75D95FFBE006C2486FEB565A9FE97" box="[984,1206,325,370]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">7 1</td>
|
||
<td id="76C75D95FFBE006C2391FEB564B1FE97" box="[1231,1454,325,370]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">5─8 1</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FE7364B1FE73" box="[189,1454,387,406]" gridrow="3" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FE7361F0FE73" box="[189,239,387,406]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">3</th>
|
||
<td id="76C75D95FFBE006C2657FE7360CEFE73" box="[265,465,387,406]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">Inferior orbital angle</td>
|
||
<td id="76C75D95FFBE006C26B9FE7363D6FE73" box="[487,713,387,406]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">Round lobe, slightly pointed</td>
|
||
<td id="76C75D95FFBE006C2581FE7362DFFE73" box="[735,960,387,406]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">Acute, without lower border</td>
|
||
<td id="76C75D95FFBE006C2486FE7365A9FE73" box="[984,1206,387,406]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">Minutely produced</td>
|
||
<td id="76C75D95FFBE006C2391FE7364B1FE73" box="[1231,1454,387,406]" gridcol="5" gridrow="3" pageId="12" pageNumber="13">Produced, angular</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FE5764B1FE5F" box="[189,1454,423,442]" gridrow="4" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FE5761F0FE5F" box="[189,239,423,442]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">4</th>
|
||
<td id="76C75D95FFBE006C2657FE5760CEFE5F" box="[265,465,423,442]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">Antennal spine</td>
|
||
<td id="76C75D95FFBE006C26B9FE5763D6FE5F" box="[487,713,423,442]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">Slender, marginal</td>
|
||
<td id="76C75D95FFBE006C2581FE5762DFFE5F" box="[735,960,423,442]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">Acute and sub-marginal</td>
|
||
<td id="76C75D95FFBE006C2486FE5765A9FE5F" box="[984,1206,423,442]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">Slender, marginal</td>
|
||
<td id="76C75D95FFBE006C2391FE5764B1FE5F" box="[1231,1454,423,442]" gridcol="5" gridrow="4" pageId="12" pageNumber="13">Moderate size, marginal</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FE3A64B1FE12" box="[189,1454,458,503]" gridrow="5" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FE3A61F0FE12" box="[189,239,458,503]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">5</th>
|
||
<td id="76C75D95FFBE006C2657FE3A60CEFE12" box="[265,465,458,503]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">Hepatic spine</td>
|
||
<td id="76C75D95FFBE006C26B9FE3A63D6FE12" box="[487,713,458,503]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">Situated behind the level of antennal spine</td>
|
||
<td id="76C75D95FFBE006C2581FE3A62DFFE12" box="[735,960,458,503]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">Robust, slightly lower level of antennal spine</td>
|
||
<td id="76C75D95FFBE006C2486FE3A65A9FE12" box="[984,1206,458,503]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">Robust, distinctly lower level of antennal spine</td>
|
||
<td id="76C75D95FFBE006C2391FE3A64B1FE12" box="[1231,1454,458,503]" gridcol="5" gridrow="5" pageId="12" pageNumber="13">Prominent, slightly lower level of antennal spine</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FDF864B1FDD1" box="[189,1454,520,564]" gridrow="6" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FDF861F0FDD1" box="[189,239,520,564]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">6</th>
|
||
<td id="76C75D95FFBE006C2657FDF860CEFDD1" box="[265,465,520,564]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">Scaphocerite (Length: width)</td>
|
||
<td id="76C75D95FFBE006C26B9FDF863D6FDD1" box="[487,713,520,564]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">2.25 times</td>
|
||
<td id="76C75D95FFBE006C2581FDF862DFFDD1" box="[735,960,520,564]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">2.3 times</td>
|
||
<td id="76C75D95FFBE006C2486FDF865A9FDD1" box="[984,1206,520,564]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">2.3 times</td>
|
||
<td id="76C75D95FFBE006C2391FDF864B1FDD1" box="[1231,1454,520,564]" gridcol="5" gridrow="6" pageId="12" pageNumber="13">2.1─2.3 times</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FDB564B1FD5A" box="[189,1454,581,703]" gridrow="7" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FDB561F0FD5A" box="[189,239,581,703]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">7</th>
|
||
<td id="76C75D95FFBE006C2657FDB560CEFD5A" box="[265,465,581,703]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">Antennular flagellum (outer)</td>
|
||
<td id="76C75D95FFBE006C26B9FDB563D6FD5A" box="[487,713,581,703]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">Biramous, proximal 3 segments fused, shorter free ramus with 3 segments and 8 groups of aesthetascs</td>
|
||
<td id="76C75D95FFBE006C2581FDB562DFFD5A" box="[735,960,581,703]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">Biramous,proximal 4 segments fused, shorter free ramus with 4 segments and 8─9 groups of aesthetascs</td>
|
||
<td id="76C75D95FFBE006C2486FDB565A9FD5A" box="[984,1206,581,703]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">Biramous, proximal 4 segments fused, shorter free ramus with 4 segments 7 groups of aesthetascs</td>
|
||
<td id="76C75D95FFBE006C2391FDB564B1FD5A" box="[1231,1454,581,703]" gridcol="5" gridrow="7" pageId="12" pageNumber="13">Biramous, proximal 3 segments fused, shorter free ramus with 7─8 segments and 12─14 groups of aesthetascs</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FD3E64B1FD19" box="[189,1454,718,764]" gridrow="8" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FD3E61F0FD19" box="[189,239,718,764]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">8</th>
|
||
<td id="76C75D95FFBE006C2657FD3E60CEFD19" box="[265,465,718,764]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">TeL: 6th somite a. TeL: Anterior width</td>
|
||
<td id="76C75D95FFBE006C26B9FD3E63D6FD19" box="[487,713,718,764]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">1.33 times 2.8 times</td>
|
||
<td id="76C75D95FFBE006C2581FD3E62DFFD19" box="[735,960,718,764]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">1.4 times 2.4 times</td>
|
||
<td id="76C75D95FFBE006C2486FD3E65A9FD19" box="[984,1206,718,764]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">1.5 times 2.9 times</td>
|
||
<td id="76C75D95FFBE006C2391FD3E64B1FD19" box="[1231,1454,718,764]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">1.5 times 3.5 times</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FCFD64B1FCB6" box="[189,1454,781,851]" gridrow="9" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FCFD61F0FCB6" box="[189,239,781,851]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">9</th>
|
||
<td id="76C75D95FFBE006C2657FCFD60CEFCB6" box="[265,465,781,851]" gridcol="1" gridrow="9" pageId="12" pageNumber="13">4th sternal plate</td>
|
||
<td id="76C75D95FFBE006C26B9FCFD63D6FCB6" box="[487,713,781,851]" gridcol="2" gridrow="9" pageId="12" pageNumber="13">Broad, low transverse ridge with an open median notch</td>
|
||
<td id="76C75D95FFBE006C2581FCFD62DFFCB6" box="[735,960,781,851]" gridcol="3" gridrow="9" pageId="12" pageNumber="13">Strongly elevated with a faint median notch</td>
|
||
<td id="76C75D95FFBE006C2486FCFD65A9FCB6" box="[984,1206,781,851]" gridcol="4" gridrow="9" pageId="12" pageNumber="13">Centrally elongated transverse ridge with a deep close median notch</td>
|
||
<td id="76C75D95FFBE006C2391FCFD64B1FCB6" box="[1231,1454,781,851]" gridcol="5" gridrow="9" pageId="12" pageNumber="13">Angular, broad lateral ridges with ‘V’ shaped median notch</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FC9264B1FC21" box="[189,1454,866,964]" gridrow="10" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FC9261F0FC21" box="[189,239,866,964]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">10</th>
|
||
<td id="76C75D95FFBE006C2657FC9260CEFC21" box="[265,465,866,964]" gridcol="1" gridrow="10" pageId="12" pageNumber="13">1st pereopod a. Fingers: palm b. Carpus: palm c. Merus: carpus</td>
|
||
<td id="76C75D95FFBE006C26B9FC9263D6FC21" box="[487,713,866,964]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">1.1 times 2.8 times Equal</td>
|
||
<td id="76C75D95FFBE006C2581FC9262DFFC21" box="[735,960,866,964]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">1.2 times 2.5 times Sub-equal</td>
|
||
<td id="76C75D95FFBE006C2486FC9265A9FC21" box="[984,1206,866,964]" gridcol="4" gridrow="10" pageId="12" pageNumber="13">Sub-equal 2.7 times Sub-equal</td>
|
||
<td id="76C75D95FFBE006C2391FC9264B1FC21" box="[1231,1454,866,964]" gridcol="5" gridrow="10" pageId="12" pageNumber="13">1.3─1.4 times 2.9─3.0 times 0.9─0.95 times</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FC2364B1FBD0" box="[189,1454,979,1077]" gridrow="11" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FC2361F0FBD0" box="[189,239,979,1077]" gridcol="0" gridrow="11" pageId="12" pageNumber="13">11</th>
|
||
<td id="76C75D95FFBE006C2657FC2360CEFBD0" box="[265,465,979,1077]" gridcol="1" gridrow="11" pageId="12" pageNumber="13">2nd pereopod a. Fingers: palm b. Carpus: palm c. Merus: palm</td>
|
||
<td id="76C75D95FFBE006C26B9FC2363D6FBD0" box="[487,713,979,1077]" gridcol="2" gridrow="11" pageId="12" pageNumber="13">≤ 0.5 times 0.35 times 0.71 times</td>
|
||
<td id="76C75D95FFBE006C2581FC2362DFFBD0" box="[735,960,979,1077]" gridcol="3" gridrow="11" pageId="12" pageNumber="13">≥0.5 times 0.22 times 0.75 times</td>
|
||
<td id="76C75D95FFBE006C2486FC2365A9FBD0" box="[984,1206,979,1077]" gridcol="4" gridrow="11" pageId="12" pageNumber="13">>0.5 times 0.25 times 0.72 times</td>
|
||
<td id="76C75D95FFBE006C2391FC2364B1FBD0" box="[1231,1454,979,1077]" gridcol="5" gridrow="11" pageId="12" pageNumber="13">0.45─0.47 times 0.33 times 0.7 times</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FBB364B1FB6E" box="[189,1454,1091,1163]" gridrow="12" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FBB361F0FB6E" box="[189,239,1091,1163]" gridcol="0" gridrow="12" pageId="12" pageNumber="13">12</th>
|
||
<td id="76C75D95FFBE006C2657FBB360CEFB6E" box="[265,465,1091,1163]" gridcol="1" gridrow="12" pageId="12" pageNumber="13">3rd – 5th pereopods a.Dactylus length: width b. Dactylus:propodus</td>
|
||
<td id="76C75D95FFBE006C26B9FBB363D6FB6E" box="[487,713,1091,1163]" gridcol="2" gridrow="12" pageId="12" pageNumber="13">2.6 times 0.20 times</td>
|
||
<td id="76C75D95FFBE006C2581FBB362DFFB6E" box="[735,960,1091,1163]" gridcol="3" gridrow="12" pageId="12" pageNumber="13">3.0 times 0.25 times</td>
|
||
<td id="76C75D95FFBE006C2486FBB365A9FB6E" box="[984,1206,1091,1163]" gridcol="4" gridrow="12" pageId="12" pageNumber="13">3.2 times 0.24 times</td>
|
||
<td id="76C75D95FFBE006C2391FBB364B1FB6E" box="[1231,1454,1091,1163]" gridcol="5" gridrow="12" pageId="12" pageNumber="13">2.1─2.5 times 0.25─0.28 times</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FB6C64B1FB2C" box="[189,1454,1180,1225]" gridrow="13" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FB6C61F0FB2C" box="[189,239,1180,1225]" gridcol="0" gridrow="13" pageId="12" pageNumber="13">13</th>
|
||
<td id="76C75D95FFBE006C2657FB6C60CEFB2C" box="[265,465,1180,1225]" gridcol="1" gridrow="13" pageId="12" pageNumber="13">Type locality</td>
|
||
<td id="76C75D95FFBE006C26B9FB6C63D6FB2C" box="[487,713,1180,1225]" gridcol="2" gridrow="13" pageId="12" pageNumber="13">Port Blair, Andaman</td>
|
||
<td id="76C75D95FFBE006C2581FB6C62DFFB2C" box="[735,960,1180,1225]" gridcol="3" gridrow="13" pageId="12" pageNumber="13">Eniwetok Atoll,Marshall Islands</td>
|
||
<td id="76C75D95FFBE006C2486FB6C65A9FB2C" box="[984,1206,1180,1225]" gridcol="4" gridrow="13" pageId="12" pageNumber="13">Lung Ha Wan, Hong Kong</td>
|
||
<td id="76C75D95FFBE006C2391FB6C64B1FB2C" box="[1231,1454,1180,1225]" gridcol="5" gridrow="13" pageId="12" pageNumber="13">Agatti Island, Lakshadweep</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FB2964B1FAE3" box="[189,1454,1241,1286]" gridrow="14" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FB2961F0FAE3" box="[189,239,1241,1286]" gridcol="0" gridrow="14" pageId="12" pageNumber="13">14</th>
|
||
<td id="76C75D95FFBE006C2657FB2960CEFAE3" box="[265,465,1241,1286]" gridcol="1" gridrow="14" pageId="12" pageNumber="13">Associated with</td>
|
||
<td id="76C75D95FFBE006C26B9FB2963D6FAE3" box="[487,713,1241,1286]" gridcol="2" gridrow="14" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F2571FB17639EFB1C" authorityName="Ruppell and Leuckart" authorityYear="1828" box="[559,641,1255,1273]" class="Anthozoa" family="Discosomidae" genus="Discosoma" kingdom="Animalia" order="Corallimorpharia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B952EAB9FFBEFF9F2571FB17639EFB1C" box="[559,641,1255,1273]" italics="true" pageId="12" pageNumber="13">Discosoma</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="76C75D95FFBE006C2581FB2962DFFAE3" box="[735,960,1241,1286]" gridcol="3" gridrow="14" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F25B2FB2A627CFB09" baseAuthorityName="Haddon and Shackleton" baseAuthorityYear="1893" box="[748,867,1241,1260]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="malu">
|
||
<emphasis id="B952EAB9FFBEFF9F25B2FB2A627CFB09" box="[748,867,1241,1260]" italics="true" pageId="12" pageNumber="13">Heteractis malu</emphasis>
|
||
</taxonomicName>
|
||
(formerly
|
||
<taxonomicName id="4C264D28FFBEFF9F2454FB036292FAE3" baseAuthorityName="Haddon & Shackleton" baseAuthorityYear="1893" box="[778,909,1267,1286]" class="Anthozoa" family="Stichodactylidae" genus="Radianthus" kingdom="Animalia" order="Actiniaria" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="malu">
|
||
<emphasis id="B952EAB9FFBEFF9F2454FB036292FAE3" box="[778,909,1267,1286]" italics="true" pageId="12" pageNumber="13">Radianthus malu</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</td>
|
||
<td id="76C75D95FFBE006C2486FB2965A9FAE3" box="[984,1206,1241,1286]" gridcol="4" gridrow="14" pageId="12" pageNumber="13">Actiniarian</td>
|
||
<td id="76C75D95FFBE006C2391FB2964B1FAE3" box="[1231,1454,1241,1286]" gridcol="5" gridrow="14" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C264D28FFBEFF9F23ACFB176493FB1C" box="[1266,1420,1254,1273]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFBEFF9F23ACFB176493FB1C" box="[1266,1420,1254,1273]" italics="true" pageId="12" pageNumber="13">Heteractis magnifica</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="351634E9FFBE006C27E3FAE664B1FACE" box="[189,1454,1302,1323]" gridrow="15" pageId="12" pageNumber="13">
|
||
<th id="76C75D95FFBE006C27E3FAE661F0FACE" box="[189,239,1302,1323]" gridcol="0" gridrow="15" pageId="12" pageNumber="13">15</th>
|
||
<td id="76C75D95FFBE006C2657FAE660CEFACE" box="[265,465,1302,1323]" gridcol="1" gridrow="15" pageId="12" pageNumber="13">Depth</td>
|
||
<td id="76C75D95FFBE006C26B9FAE663D6FACE" box="[487,713,1302,1323]" gridcol="2" gridrow="15" pageId="12" pageNumber="13">Shallow</td>
|
||
<td id="76C75D95FFBE006C2581FAE662DFFACE" box="[735,960,1302,1323]" gridcol="3" gridrow="15" pageId="12" pageNumber="13">1.0─2.0 m</td>
|
||
<td id="76C75D95FFBE006C2486FAE665A9FACE" box="[984,1206,1302,1323]" gridcol="4" gridrow="15" pageId="12" pageNumber="13">3.6 m</td>
|
||
<td id="76C75D95FFBE006C2391FAE664B1FACE" box="[1231,1454,1302,1323]" gridcol="5" gridrow="15" pageId="12" pageNumber="13">1.0─2.0 m</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBEFF9F2785FA8660F8F850" blockId="12.[180,805,1398,1973]" pageId="12" pageNumber="13">
|
||
Further,
|
||
<taxonomicName id="4C264D28FFBEFF9F2617FA8760DAFA76" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[329,453,1399,1427]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F2617FA8760DAFA76" box="[329,453,1399,1427]" italics="true" pageId="12" pageNumber="13">A. ornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C264D28FFBEFF9F255BFA876387FA76" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[517,664,1399,1427]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F255BFA876387FA76" box="[517,664,1399,1427]" italics="true" pageId="12" pageNumber="13">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
are similar to
|
||
<taxonomicName id="4C264D28FFBEFF9F2789FA6E6023FA5F" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[215,316,1437,1466]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBEFF9F2789FA6E6023FA5F" box="[215,316,1437,1466]" italics="true" pageId="12" pageNumber="13">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBEFF9F261AFA6E60BFFA5F" box="[324,416,1438,1466]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with characters like a marginal antennal slender spine; hepatic spine appears nearly behind the level of antennal spine; and the dactylus of ambulatory pereopods is slender and smooth. However,
|
||
<taxonomicName id="4C264D28FFBEFF9F2674F9C96081F9B3" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[298,414,1593,1622]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F2674F9C96081F9B3" box="[298,414,1593,1622]" italics="true" pageId="12" pageNumber="13">A. ornatus</emphasis>
|
||
</taxonomicName>
|
||
differs from the new species in the presence of a fourth thoracic sternite with a centrally elongated transverse ridge with a deep median notch (
|
||
<emphasis id="B952EAB9FFBEFF9F279EF95F61C2F92E" box="[192,221,1711,1739]" italics="true" pageId="12" pageNumber="13">vs.</emphasis>
|
||
angular broad lateral ridges in
|
||
<taxonomicName id="4C264D28FFBEFF9F250DF95E63AAF92E" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[595,693,1710,1739]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBEFF9F250DF95E63AAF92E" box="[595,693,1710,1739]" italics="true" pageId="12" pageNumber="13">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBEFF9F25E5F95E6207F92E" box="[699,792,1710,1739]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
<taxonomicName id="4C264D28FFBEFF9F27E8F9256083F917" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[182,412,1749,1778]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F27E8F9256083F917" box="[182,412,1749,1778]" italics="true" pageId="12" pageNumber="13">Actinimenes ornatus</emphasis>
|
||
</taxonomicName>
|
||
is known from the Pacific Ocean:
|
||
<collectingCountry id="F331763BFFBEFF9F27EAF90C605DF8FD" box="[180,322,1788,1816]" name="Hong Kong" pageId="12" pageNumber="13">Hong Kong</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F260EF90C60F6F8FC" author="Bruce, A. J." box="[336,489,1788,1817]" pageId="12" pageNumber="13" pagination="253 - 278" refId="ref8735" refString="Bruce, A. J. 1969. Preliminary descriptions of sixteen new species of the genus Periclimenes Costa, 1844 (Crustacea, Decapoda Natantia, Pontoniinae). Zoologische Mededelingen, 43: 253 - 278." type="journal article" year="1969">Bruce, 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F26AAF90C632CF8FC" author="Bruce, A. J." box="[500,563,1788,1817]" pageId="12" pageNumber="13" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">1979</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F331763BFFBEFF9F2518F90C6395F8FD" box="[582,650,1788,1816]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F25C7F90C607CF8A5" author="Suzuki, K. & Hayashi, K. I." pageId="12" pageNumber="13" pagination="193 - 208" refId="ref11233" refString="Suzuki, K. and Hayashi, K. I. 1977. Five caridean shrimps associated with sea anemones in central Japan. Publications of the Seto Marine Biological Laboratory, 24: 193 - 208." type="journal article" year="1977">Suzuki and Hayashi, 1977</bibRefCitation>
|
||
), Eniwetok Atoll, the
|
||
<collectingCountry id="F331763BFFBEFF9F2501F8D3623AF8A5" box="[607,805,1827,1856]" name="Marshall Islands" pageId="12" pageNumber="13">Marshall Islands</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F27E2F8BA604DF882" author="Bruce, A. J." box="[188,338,1866,1895]" pageId="12" pageNumber="13" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce, 1979</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F331763BFFBEFF9F263BF8B960C0F883" box="[357,479,1865,1894]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F331763BFFBEFF9F26B6F8BA632BF883" box="[488,564,1866,1894]" name="South Korea" pageId="12" pageNumber="13">Korea</collectingCountry>
|
||
, the Red Sea (
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F2586F8BA60BBF86B" author="Bruce, A. J. & Svoboda, A." pageId="12" pageNumber="13" refId="ref9319" refString="Bruce, A. J. and Svoboda, A. 1983. Observations upon some pontoniine shrimps from Aqaba, Jordan. Zoologische Verhandelingen, 205: l - 44." type="journal volume" year="1983">Bruce and Svoboda, 1983</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F331763BFFBEFF9F26E5F8816313F868" box="[443,524,1905,1933]" name="Kenya" pageId="12" pageNumber="13">Kenya</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F331763BFFBEFF9F250EF88063A5F868" box="[592,698,1904,1933]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F2592F88061E7F851" author="Marin, I. N." pageId="12" pageNumber="13" pagination="321 - 340" refId="ref10560" refString="Marin, I. N. 2006. Description of Crinotonia anastasiae, new genus, new species, a new crinoid associated pontoniine shrimp (Crustacea: Caridea) from Nha Trang Bay, Vietnam, with inclusion of Periclimenes attenuatus Bruce, 1971, in the new genus. The Raffles Bulletin of Zoology, 54: 321 - 340." type="journal article" year="2006">Marin, 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F265AF86860C5F850" author="Lee, K. H. & Ko, H. S." box="[260,474,1943,1973]" pageId="12" pageNumber="13" pagination="239 - 245" refId="ref10470" refString="Lee, K. H. and Ko, H. S. 2011. First records of two pontoniid shrimps (Crustacea: Decapoda: Caridea: Palaemonidae) from Korea. Animal Systematics, Evolution and Diversity, 27: 239 - 245." type="journal article" year="2011">Lee and Ko, 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBEFF9F242FFA8665C4F86B" blockId="12.[841,1468,1398,1934]" pageId="12" pageNumber="13">
|
||
Similarly,
|
||
<taxonomicName id="4C264D28FFBEFF9F24B8FA87656EFA76" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[998,1137,1399,1427]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F24B8FA87656EFA76" box="[998,1137,1399,1427]" italics="true" pageId="12" pageNumber="13">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
varied from
|
||
<taxonomicName id="4C264D28FFBEFF9F225CFA87647FFA76" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1282,1376,1398,1427]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koyas" status="sp.nov.">
|
||
<emphasis id="B952EAB9FFBEFF9F225CFA87647FFA76" box="[1282,1376,1398,1427]" italics="true" pageId="12" pageNumber="13">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBEFF9F223BFA8764A3FA76" box="[1381,1468,1399,1427]" pageId="12" pageNumber="13" rank="species">sp.nov.</taxonomicNameLabel>
|
||
by having a broad low transverse ridge, with an open median notch in the fourth thoracic sternite (angular broad lateral ridges with deep median incision in
|
||
<taxonomicName id="4C264D28FFBEFF9F22C0FA1C6293F9CA" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBEFF9F22C0FA1C6293F9CA" italics="true" pageId="12" pageNumber="13">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBEFF9F24CBF9E362EBF9CA" box="[917,1012,1554,1583]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) (
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F2353F9E265DCF9CA" author="Fransen, C. H. J. M." box="[1037,1219,1554,1583]" pageId="12" pageNumber="13" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F238EF9E36478F9CA" author="Bruce, A. J." box="[1232,1383,1554,1583]" pageId="12" pageNumber="13" pagination="20 - 36" refId="ref9236" refString="Bruce, A. J. 2010. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the Coral reefs 2009 Heron Island expedition. Zootaxa, 2604: 20 - 36." type="journal article" year="2010">Bruce, 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F222AF9E4653CF9B3" author="Duris, Z." pageId="12" pageNumber="13" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D28FFBEFF9F2366F9C9645DF9B3" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[1080,1346,1593,1622]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBEFF9F2366F9C9645DF9B3" box="[1080,1346,1593,1622]" italics="true" pageId="12" pageNumber="13">Actinimenes inornatus</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed in
|
||
<collectingCountry id="F331763BFFBEFF9F24A9F991655AF998" box="[1015,1093,1633,1661]" name="Kenya" pageId="12" pageNumber="13">Kenya</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F230DF99065F4F998" author="Bruce, A. J." box="[1107,1259,1632,1661]" pageId="12" pageNumber="13" pagination="89 - 153" refId="ref8864" refString="Bruce, A. J. 1976. A report on some pontoniinid shrimps collected from the Seychelle Islands by the FRV Manihine, 1972, with a review of the Seychelles pontoniinid shrimp fauna. Zoological Journal of the Linnean Society, 59: 89 - 153." type="journal article" year="1976">Bruce, 1976</bibRefCitation>
|
||
), Grand Comoro Island near
|
||
<collectingCountry id="F331763BFFBEFF9F24BFF9776569F941" box="[993,1142,1671,1700]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F23D6F977643BF941" author="Bruce, A. J." box="[1160,1316,1671,1700]" pageId="12" pageNumber="13" pagination="1 - 13" refId="ref8774" refString="Bruce, A. J. 1971. Pontoniinid shrimps from the ninth cruise of RV Anton Bruun, IIOE, 1964: I. Palaemonella Dana and Perclimenes Costa. Smithsonian Contributions to Zoology, 82: 1 - 13." type="journal article" year="1971">Bruce, 1971</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F331763BFFBEFF9F2265F97764A4F941" box="[1339,1467,1671,1700]" name="Seychelles" pageId="12" pageNumber="13">Seychelles</collectingCountry>
|
||
Islands(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F24F5F95E6525F92E" author="Bruce, A. J." box="[939,1082,1710,1739]" pageId="12" pageNumber="13" pagination="1 - 13" refId="ref8774" refString="Bruce, A. J. 1971. Pontoniinid shrimps from the ninth cruise of RV Anton Bruun, IIOE, 1964: I. Palaemonella Dana and Perclimenes Costa. Smithsonian Contributions to Zoology, 82: 1 - 13." type="journal article" year="1971">Bruce, 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F231CF95E6560F92E" author="Bruce, A. J." box="[1090,1151,1710,1739]" pageId="12" pageNumber="13" pagination="89 - 153" refId="ref8864" refString="Bruce, A. J. 1976. A report on some pontoniinid shrimps collected from the Seychelle Islands by the FRV Manihine, 1972, with a review of the Seychelles pontoniinid shrimp fauna. Zoological Journal of the Linnean Society, 59: 89 - 153." type="journal article" year="1976">1976</bibRefCitation>
|
||
),Andaman Islands(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F2234F95E6298F917" author="Kemp, S. W." pageId="12" pageNumber="13" pagination="113 - 288" refId="ref10211" refString="Kemp, S. W. 1922. Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24: 113 - 288." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F331763BFFBEFF9F24C6F925651AF917" box="[920,1029,1749,1778]" name="Maldives" pageId="12" pageNumber="13">Maldives</collectingCountry>
|
||
and
|
||
<collectingCountry id="F331763BFFBEFF9F2364F9256413F917" box="[1082,1292,1749,1778]" name="India" pageId="12" pageNumber="13">Laccadive Islands</collectingCountry>
|
||
, Bay of Bengal, Mariana Island (
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F2349F90C65AEF8FC" author="Bruce, A. J." box="[1047,1201,1788,1817]" pageId="12" pageNumber="13" pagination="89 - 153" refId="ref8864" refString="Bruce, A. J. 1976. A report on some pontoniinid shrimps collected from the Seychelle Islands by the FRV Manihine, 1972, with a review of the Seychelles pontoniinid shrimp fauna. Zoological Journal of the Linnean Society, 59: 89 - 153." type="journal article" year="1976">Bruce, 1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F23E1F90C65E0F8FC" author="Bruce, A. J." box="[1215,1279,1788,1817]" pageId="12" pageNumber="13" pagination="141 - 169" refId="ref9106" refString="Bruce, A. J. 1984. Marine caridean shrimps of the Seychelles. p. 141 - 169. In: D. R. Stoddart (ed), Biogeography and Ecology of the Seychelles Islands. The Hague, Boston, Lancaster, Junk Publishers." type="book chapter" year="1984">1984</bibRefCitation>
|
||
), Heron Island (
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F240AF8D362ECF8A5" author="Bruce, A. J." box="[852,1011,1827,1856]" pageId="12" pageNumber="13" pagination="20 - 36" refId="ref9236" refString="Bruce, A. J. 2010. More pontoniine shrimps (Crustacea: Decapoda: Palaemonidae) from the Coral reefs 2009 Heron Island expedition. Zootaxa, 2604: 20 - 36." type="journal article" year="2010">Bruce, 2010</bibRefCitation>
|
||
), Ryukyu Islands,
|
||
<collectingCountry id="F331763BFFBEFF9F23BDF8D3647EF8A5" box="[1251,1377,1827,1856]" name="Indonesia" pageId="12" pageNumber="13">Indonesia</collectingCountry>
|
||
, South
|
||
<collectingCountry id="F331763BFFBEFF9F2415F8B96286F883" box="[843,921,1865,1894]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
|
||
Sea, Great Barrier Reef,
|
||
<collectingCountry id="F331763BFFBEFF9F2385F8B96419F883" box="[1243,1286,1865,1894]" name="Fiji" pageId="12" pageNumber="13">Fiji</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F331763BFFBEFF9F2212F8B962BDF868" name="Micronesia" pageId="12" pageNumber="13">Caroline Islands</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBEFF9F24EEF88065D1F86B" author="Chace, F. A. Jr. & Bruce, A. J." box="[944,1230,1904,1934]" pageId="12" pageNumber="13" pagination="1 - 152" refId="ref9351" refString="Chace, F. A. Jr. and Bruce, A. J. 1993. The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine expedition 1907 - 1910, Part 6: Superfamily Palaemonoidea. Smithsonian Contributions to Zoology, 543: 1 - 152." type="journal article" year="1993">Chace and Bruce, 1993</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBFFF9E2785FF676382FB67" blockId="13.[180,807,151,1154]" pageId="13" pageNumber="14">
|
||
Color patterns are one of the important taxonomic characters for palaemonid shrimps and are mostly species-specific for the
|
||
<taxonomicName id="4C264D28FFBFFF9E2690FF156344FEE4" authorityName="Duris and Horka" authorityYear="2017" box="[462,603,229,257]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAB9FFBFFF9E2690FF156344FEE4" box="[462,603,229,257]" italics="true" pageId="13" pageNumber="14">Actinimenes</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E25E7FF1561E7FECC" author="Fransen, C. H. J. M." pageId="13" pageNumber="14" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2658FEFE6080FECC" author="Duris, Z." box="[262,415,268,302]" pageId="13" pageNumber="14" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš, 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E26F3FEFE63C4FECC" author="Duris, Z." box="[429,731,268,302]" pageId="13" pageNumber="14" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
). The coloration of
|
||
<taxonomicName id="4C264D28FFBFFF9E2600FEC360DBFEB5" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[350,452,307,336]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2600FEC360DBFEB5" box="[350,452,307,336]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E2693FEC36334FEAA" box="[461,555,307,335]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to that of
|
||
<taxonomicName id="4C264D28FFBFFF9E27EBFEAA6056FE92" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[181,329,346,375]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E27EBFEAA6056FE92" box="[181,329,346,375]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C264D28FFBFFF9E26D7FEAA633EFE92" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[393,545,346,375]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E26D7FEAA633EFE92" box="[393,545,346,375]" italics="true" pageId="13" pageNumber="14">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
, where the carapace, rostrum, and abdominal region have a transparent and faint brownish color. However,
|
||
<taxonomicName id="4C264D28FFBFFF9E253CFE5863EDFE20" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[610,754,424,453]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E253CFE5863EDFE20" box="[610,754,424,453]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C264D28FFBFFF9E27EBFE3F6057FE0E" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[181,328,462,491]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E27EBFE3F6057FE0E" box="[181,328,462,491]" italics="true" pageId="13" pageNumber="14">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
are distinguished from
|
||
<taxonomicName id="4C264D28FFBFFF9E2538FE3F63D8FE0E" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[614,711,462,491]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2538FE3F63D8FE0E" box="[614,711,462,491]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E2592FE3F623AFE0E" box="[716,805,463,491]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the presence of a broad dorsal white band on the eyestalks, cardiac region, and presence of a broad medial ventral white band extended longitudinally from the posterior end of the stomach to the anterior end of the fifth abdominal segment (absent on
|
||
<taxonomicName id="4C264D28FFBFFF9E2780FD49605BFD30" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[222,324,696,725]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2780FD49605BFD30" box="[222,324,696,725]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E2613FD4960B4FD30" box="[333,427,696,725]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) (
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E269BFD48637FFD30" author="Bruce, A. J." box="[453,608,696,725]" pageId="13" pageNumber="14" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce, 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2530FD49623EFD30" author="Fransen, C. H. J. M." box="[622,801,696,725]" pageId="13" pageNumber="14" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E27EBFD2F60F4FD19" author="Salvat, B. & Bacchet, P." box="[181,491,735,764]" pageId="13" pageNumber="14" refId="ref11170" refString="Salvat, B. and Bacchet, P. 2011. Guide des recifs coralliens de Tahiti et ses iles. Papeete, Au Vent des Iles, 365 p." type="book" year="2011">Salvat and Bacchet, 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E26A6FD116206FD19" author="Duris, Z." box="[504,793,735,769]" pageId="13" pageNumber="14" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D28FFBFFF9E27E6FCF660BAFCC6" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[184,421,774,803]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatu">
|
||
<emphasis id="B952EAB9FFBFFF9E27E6FCF660BAFCC6" box="[184,421,774,803]" italics="true" pageId="13" pageNumber="14">Actinimenes ornatu</emphasis>
|
||
</taxonomicName>
|
||
s differs from
|
||
<taxonomicName id="4C264D28FFBFFF9E2500FCF663E6FCC6" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[606,761,774,803]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E2500FCF663E6FCC6" box="[606,761,774,803]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E2454FCF66032FCAF" baseAuthorityName="Bruce" baseAuthorityYear="1979" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E2454FCF66032FCAF" italics="true" pageId="13" pageNumber="14">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C264D28FFBFFF9E2626FCDD60FEFCAF" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[376,481,813,842]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2626FCDD60FEFCAF" box="[376,481,813,842]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E26B5FCDE6353FCAF" box="[491,588,814,842]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having finely striated, longitudinally and alternating rows of minute red and white chromatophores extended over the pleura, the antennal peduncles and scaphocerite with purple and white spots (especially along the dorsum of the eyestalk and across the ophthalmic somite) transparent pereopods and uropods with numerous paired purple and whitish chromatophores (
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E258DFBCE61EAFB67" author="Bruce, A. J." pageId="13" pageNumber="14" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce, 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E265EFB9560B0FB67" author="Fransen, C. H. J. M." box="[256,431,1125,1154]" pageId="13" pageNumber="14" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E26E4FB95638EFB67" author="Lee, K. H. & Ko, H. S." box="[442,657,1124,1154]" pageId="13" pageNumber="14" pagination="239 - 245" refId="ref10470" refString="Lee, K. H. and Ko, H. S. 2011. First records of two pontoniid shrimps (Crustacea: Decapoda: Caridea: Palaemonidae) from Korea. Animal Systematics, Evolution and Diversity, 27: 239 - 245." type="journal article" year="2011">Lee and Ko, 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936ABFFBFFF9E2785FB5A6527F8D3" blockId="13.[180,806,1194,1846]" lastBlockId="13.[841,1469,151,1846]" pageId="13" pageNumber="14">
|
||
<emphasis id="B952EAB9FFBFFF9E2785FB5A6046FB23" box="[219,345,1194,1222]" italics="true" pageId="13" pageNumber="14">Host range</emphasis>
|
||
.
|
||
<taxonomicName id="4C264D28FFBFFF9E2636FB5A6321FB22" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[360,574,1194,1223]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2636FB5A6321FB22" box="[360,574,1194,1223]" italics="true" pageId="13" pageNumber="14">Actinimenes koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E2518FB5A63BEFB23" box="[582,673,1194,1222]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found with the magnificent sea anemone,
|
||
<taxonomicName id="4C264D28FFBFFF9E25D8FB216239FB08" box="[646,806,1233,1261]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFBFFF9E25D8FB216239FB08" box="[646,806,1233,1261]" italics="true" pageId="13" pageNumber="14">H. magnifica</emphasis>
|
||
</taxonomicName>
|
||
along with another anemone-associated shrimp
|
||
<taxonomicName id="4C264D28FFBFFF9E2455FB086056FADE" class="Malacostraca" family="Palaemonidae" genus="Ancylocaris" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevicarpalis">
|
||
<emphasis id="B952EAB9FFBFFF9E2455FB086056FADE" italics="true" pageId="13" pageNumber="14">A. brevicarpalis</emphasis>
|
||
</taxonomicName>
|
||
in the Agatti Island of
|
||
<collectingCountry id="F331763BFFBFFF9E250CFAEE63E6FADE" box="[594,761,1310,1339]" name="India" pageId="13" pageNumber="14">Lakshadweep</collectingCountry>
|
||
at a depth of
|
||
<quantity id="4CDE9B4EFFBFFF9E2640FAB56077FA87" box="[286,360,1349,1378]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="14" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">
|
||
1–
|
||
<specimenCount id="9D20FD22FFBFFF9E2664FAB5606FFA87" box="[314,368,1349,1378]" count="2" pageId="13" pageNumber="14" type="generic">2 m.</specimenCount>
|
||
</quantity>
|
||
Similarly,
|
||
<taxonomicName id="4C264D28FFBFFF9E26B1FAB66360FA87" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[495,639,1350,1378]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E26B1FAB66360FA87" box="[495,639,1350,1378]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
was observed on
|
||
<taxonomicName id="4C264D28FFBFFF9E2782FA9D609EFA6C" box="[220,385,1389,1417]" class="Anthozoa" family="Discosomidae" genus="Discosoma" kingdom="Animalia" order="Corallimorpharia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B952EAB9FFBFFF9E2782FA9D6046FA6C" box="[220,345,1389,1417]" italics="true" pageId="13" pageNumber="14">Discosoma</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C264D28FFBFFF9E2698FA9D6363FA6C" box="[454,636,1388,1417]" class="Malacostraca" family="Palaemonidae" genus="Ancylocaris" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevicarpalis">
|
||
<emphasis id="B952EAB9FFBFFF9E2698FA9D6363FA6C" box="[454,636,1388,1417]" italics="true" pageId="13" pageNumber="14">A. brevicarpalis</emphasis>
|
||
</taxonomicName>
|
||
and occurred with
|
||
<taxonomicName id="4C264D28FFBFFF9E27ACFA646098FA4A" box="[242,391,1427,1456]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFBFFF9E27ACFA646098FA4A" box="[242,391,1427,1456]" italics="true" pageId="13" pageNumber="14">H. magnifica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E26CAFA6361E3FA32" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="damicornis">
|
||
<emphasis id="B952EAB9FFBFFF9E26CAFA636383FA55" box="[404,668,1427,1456]" italics="true" pageId="13" pageNumber="14">Pocillopora damicornis</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E265CFA4A623FFA32" authority="(Ellis and Solander, 1786)" baseAuthorityName="Ellis and Solander" baseAuthorityYear="1786" box="[258,800,1466,1495]" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="verrucosa">
|
||
<emphasis id="B952EAB9FFBFFF9E265CFA4A60F0FA32" box="[258,495,1466,1495]" italics="true" pageId="13" pageNumber="14">Pocillopora verrucosa</emphasis>
|
||
(Ellis and Solander, 1786)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E27EAFA126334FA1B" authority="Dana, 1846" authorityName="Dana" authorityYear="1846" box="[180,555,1505,1534]" class="Anthozoa" family="Pocilloporidae" genus="Seriatopora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="hystrix">
|
||
<emphasis id="B952EAB9FFBFFF9E27EAFA12608BFA1B" box="[180,404,1505,1534]" italics="true" pageId="13" pageNumber="14">Seriatopora hystrix</emphasis>
|
||
Dana, 1846
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E2569FA116044F9C0" authority="(Esper, 1792)" baseAuthorityName="Esper" baseAuthorityYear="1792" class="Anthozoa" family="Pocilloporidae" genus="Stylophora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="pistillata">
|
||
<emphasis id="B952EAB9FFBFFF9E2569FA11623CFA1B" box="[567,803,1505,1534]" italics="true" pageId="13" pageNumber="14">Stylophora pistillata</emphasis>
|
||
(Esper, 1792)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C264D28FFBFFF9E26C2F9F8635FF9C0" box="[412,576,1544,1573]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B952EAB9FFBFFF9E26C2F9F86317F9C1" box="[412,520,1544,1572]" italics="true" pageId="13" pageNumber="14">Acropora</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E25DEF9F9623FF9C0" author="Patton, W. K." box="[640,800,1544,1573]" pageId="13" pageNumber="14" pagination="271 - 295" refId="ref11034" refString="Patton, W. K. 1966. Decapod Crustacea commensal with Queensland branching corals. Crustaceana, 10: 271 - 295." type="journal article" year="1966">Patton, 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E27EBF9C0607CF9A9" author="Fransen, C. H. J. M." box="[181,355,1583,1612]" pageId="13" pageNumber="14" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D28FFBFFF9E262AF9DF637CF9A9" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[372,611,1583,1612]" class="Malacostraca" family="Palaemonidae" genus="Actinemenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E262AF9DF637CF9A9" box="[372,611,1583,1612]" italics="true" pageId="13" pageNumber="14">Actinemenes ornatus</emphasis>
|
||
</taxonomicName>
|
||
specimens were noticed from the actinian sea anemones:
|
||
<taxonomicName id="4C264D28FFBFFF9E25F1F9A761E7F97F" baseAuthorityName="Hemprich & Ehrenberg in Ehrenberg" baseAuthorityYear="1834" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="crispa">
|
||
<emphasis id="B952EAB9FFBFFF9E25F1F9A761E7F97F" italics="true" pageId="13" pageNumber="14">Heteractis crispa</emphasis>
|
||
</taxonomicName>
|
||
(Hemprich and Ehrenberg in Ehrenberg, 1834),
|
||
<taxonomicName id="4C264D28FFBFFF9E27EAF955604AF925" box="[180,341,1700,1729]" class="Anthozoa" family="Stichodactylidae" genus="Heteractis" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="magnifica">
|
||
<emphasis id="B952EAB9FFBFFF9E27EAF955604AF925" box="[180,341,1700,1729]" italics="true" pageId="13" pageNumber="14">H. magnifica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B952EAB9FFBFFF9E2636F955623AF924" box="[360,805,1700,1729]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C264D28FFBFFF9E2636F95560CEF924" baseAuthorityName="Haddon & Shackleton" baseAuthorityYear="1893" box="[360,465,1700,1729]" class="Anthozoa" family="Stichodactylidae" genus="Radianthus" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="malu">H. malu</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E26BFF955623AF924" box="[481,805,1700,1729]" class="Anthozoa" family="Actiniidae" genus="Entacmaea" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="actinostoloides">Entacmaea actinostoloides</taxonomicName>
|
||
</emphasis>
|
||
(Leuckart in Rüppell and Leuckart, 1828),
|
||
<taxonomicName id="4C264D28FFBFFF9E25F8F93C61F4F8EB" class="Anthozoa" family="Actiniidae" genus="Entacmaea" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B952EAB9FFBFFF9E25F8F93C623AF90D" box="[678,805,1740,1768]" italics="true" pageId="13" pageNumber="14">Entacmaea</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E27A0F90260AAF8EB" box="[254,437,1778,1806]" class="Anthozoa" family="Actiniidae" genus="Gyrostoma" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="undetermined">
|
||
<emphasis id="B952EAB9FFBFFF9E27A0F9026098F8EB" box="[254,391,1778,1806]" italics="true" pageId="13" pageNumber="14">Gyrostoma</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E2694F902608FF8D3" authority="(Wassilieff, 1908)" baseAuthorityName="Wassilieff" baseAuthorityYear="1908" class="Anthozoa" family="Actinostolidae" genus="Parasicyonis" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="actinostoloides">
|
||
<emphasis id="B952EAB9FFBFFF9E2694F902623BF8EA" box="[458,804,1778,1807]" italics="true" pageId="13" pageNumber="14">Parasicyonis actinostoloides</emphasis>
|
||
(Wassilieff, 1908)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E26C2F8E9628BFF51" authority="(Wassilieff, 1908)" baseAuthorityName="Wassilieff" baseAuthorityYear="1908" box="[412,916,151,1845]" class="Anthozoa" family="Actinostolidae" genus="Parasicyonis" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="maxima">
|
||
<emphasis id="B952EAB9FFBFFF9E26C2F8E9638CF8D0" box="[412,659,1817,1845]" italics="true" pageId="13" pageNumber="14">Parasicyonis maxima</emphasis>
|
||
(Wassilieff, 1908)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C264D28FFBFFF9E2491FF67628CFF39" authority="(Klunzinger, 1877)" baseAuthorityName="Klunzinger" baseAuthorityYear="1877" class="Anthozoa" family="Thalassianthidae" genus="Cryptodendrum" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="adhaesivum">
|
||
<emphasis id="B952EAB9FFBFFF9E2491FF676404FF51" box="[975,1307,151,180]" italics="true" pageId="13" pageNumber="14">Cryptodendrum adhaesivum</emphasis>
|
||
(Klunzinger, 1877)
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E248CFF4F6577FF39" author="Bruce, A. J." box="[978,1128,191,220]" pageId="13" pageNumber="14" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce, 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E232AFF4F64A9FF39" author="Suzuki, K. & Hayashi, K. I." box="[1140,1462,191,220]" pageId="13" pageNumber="14" pagination="193 - 208" refId="ref11233" refString="Suzuki, K. and Hayashi, K. I. 1977. Five caridean shrimps associated with sea anemones in central Japan. Publications of the Seto Marine Biological Laboratory, 24: 193 - 208." type="journal article" year="1977">Suzuki and Hayashi, 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2415FF166533FEE1" author="Marin, I. N. & Britayev, T. A. & Anker, A." box="[843,1068,230,260]" pageId="13" pageNumber="14" pagination="199 - 218" refId="ref10683" refString="Marin, I. N.; Britayev, T. A. and Anker, A. 2004. Pontoniine shrimps associated with cnidarians: new records and list of species from coastal waters of Viet Nam. Arthropoda Selecta. Russian Journal of Arthropoda Research, 13: 199 - 218." type="journal article" year="2004">
|
||
Marin
|
||
<emphasis id="B952EAB9FFBFFF9E24FEFF1762C6FEE6" box="[928,985,230,259]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D28FFBFFF9E2361FF176458FEE6" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1087,1351,230,259]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E2361FF176458FEE6" box="[1087,1351,230,259]" italics="true" pageId="13" pageNumber="14">Actinimenes ornatellus</emphasis>
|
||
</taxonomicName>
|
||
was only found on
|
||
<taxonomicName id="4C264D28FFBFFF9E2490FEFF652AFECE" baseAuthorityName="Haddon & Shackleton" baseAuthorityYear="1893" box="[974,1077,270,299]" class="Anthozoa" family="Stichodactylidae" genus="Radianthus" kingdom="Animalia" order="Actiniaria" pageId="13" pageNumber="14" phylum="Cnidaria" rank="species" species="malu">
|
||
<emphasis id="B952EAB9FFBFFF9E2490FEFF652AFECE" box="[974,1077,270,299]" italics="true" pageId="13" pageNumber="14">H. malu</emphasis>
|
||
</taxonomicName>
|
||
. According to earlier reports, the species of the genus
|
||
<taxonomicName id="4C264D28FFBFFF9E2323FEC66414FEB7" authorityName="Duris and Horka" authorityYear="2017" box="[1149,1291,310,338]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAB9FFBFFF9E2323FEC66414FEB7" box="[1149,1291,310,338]" italics="true" pageId="13" pageNumber="14">Actinimenes</emphasis>
|
||
</taxonomicName>
|
||
are generalist symbionts, inhabiting a significant number of hosts. In the molecular analysis, the
|
||
<collectionCode id="ED37AE6EFFBFFF9E2250FE766421FE47" box="[1294,1342,390,418]" country="France" name="Musee de Lectoure" pageId="13" pageNumber="14">ML</collectionCode>
|
||
tree was reconstructed using the mitochondrial COI gene (
|
||
<figureCitation id="131D2A2EFFBFFF9E2409FE2462B6FE17" box="[855,937,468,498]" captionStart="Figure 7" captionStartId="14.[181,250,1151,1174]" captionTargetBox="[181,1460,193,1089]" captionTargetId="figure-40@14.[173,1466,151,1095]" captionTargetPageId="14" captionText="Figure 7. Maximum Likelihood phylogenetic tree based on partial COI (A) and 16S (B) sequences of Actinimenes koyas sp. nov. and two other Actinimenes species obtained from GenBank (Scale bar = 10 % divergence; significant bootstrap value> 60)." figureDoi="http://doi.org/10.5281/zenodo.10904886" httpUri="https://zenodo.org/record/10904886/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
|
||
), which revealed that
|
||
<taxonomicName id="4C264D28FFBFFF9E23EEFE25640DFE14" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1200,1298,468,497]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E23EEFE25640DFE14" box="[1200,1298,468,497]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E2248FE256470FE14" box="[1302,1391,469,497]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
forms a monophyletic clade and has a close relationship to
|
||
<taxonomicName id="4C264D28FFBFFF9E2415FDD462C0FDA4" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[843,991,548,577]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E2415FDD462C0FDA4" box="[843,991,548,577]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B952EAB9FFBFFF9E2340FDD465BEFDA4" box="[1054,1185,548,577]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C264D28FFBFFF9E2340FDD46582FDA4" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[1054,1181,548,577]" class="Malacostraca" family="Palaemonidae" genus="Actinemenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatus">A. ornatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with significant nodal support (bootstrap values 72 %and 99 %respectively). The sequences of
|
||
<taxonomicName id="4C264D28FFBFFF9E2344FD846496FD75" authority="(Kemp, 1922)" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[1050,1417,627,656]" class="Malacostraca" family="Palaemonidae" genus="Zenopontonia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rex">
|
||
<emphasis id="B952EAB9FFBFFF9E2344FD8465FEFD75" box="[1050,1249,628,656]" italics="true" pageId="13" pageNumber="14">Zenopontonia rex</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E23B3FD83649EFD75" author="Kemp, S. W." box="[1261,1409,627,656]" pageId="13" pageNumber="14" pagination="113 - 288" refId="ref10211" refString="Kemp, S. W. 1922. Notes on Crustacea Decapoda in the Indian Museum. XV. Pontoniinae. Records of the Indian Museum, 24: 113 - 288." type="journal article" year="1922">Kemp,1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C264D28FFBFFF9E2415FD6B6519FD5D" box="[843,1030,667,696]" class="Malacostraca" family="Palaemonidae" genus="Ancylocaris" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brevicarpalis">
|
||
<emphasis id="B952EAB9FFBFFF9E2415FD6B6519FD5D" box="[843,1030,667,696]" italics="true" pageId="13" pageNumber="14">A. brevicarpalis</emphasis>
|
||
</taxonomicName>
|
||
were used as the outgroup species (
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E240AFD336510FD3A" author="Gan, Z. B. & Li, X. Z. & Chan, T. Y. & Chu, K. H. & Kou, Q." box="[852,1039,706,736]" pageId="13" pageNumber="14" pagination="282 - 290" refId="ref9818" refString="Gan, Z. B.; Li, X. Z.; Chan, T. Y.; Chu, K. H. and Kou, Q. 2016. Phylogeny of Indo-West Pacific pontoniine shrimps (Crustacea: Decapoda: Caridea) based on multilocus analysis. Journal of Zoological Systematics and Evolutionary Research, 53: 282 - 290." type="journal article" year="2016">
|
||
Gan
|
||
<emphasis id="B952EAB9FFBFFF9E24D1FD3362D4FD3A" box="[911,971,706,735]" italics="true" pageId="13" pageNumber="14">et al.,</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2347FD3265F2FD3A" author="Horka, I. & De Grave, S. & Fransen, C. H. & Petrusek, A. & Duris, Z." box="[1049,1261,706,735]" pageId="13" pageNumber="14" pagination="26486" refId="ref10018" refString="Horka, I.; De Grave, S.; Fransen, C. H.; Petrusek, A. and Duris, Z. 2016. Multiple host switching events shape the evolution of symbiotic palaemonid shrimps (Crustacea: Decapoda). Scientific Reports, 6: 26486." type="journal article" year="2016">
|
||
Horká
|
||
<emphasis id="B952EAB9FFBFFF9E2332FD3365B7FD3A" box="[1132,1192,706,735]" italics="true" pageId="13" pageNumber="14">et al.,</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E23A8FD326428FD3A" author="Horka, I. & De Grave, S. & Fransen, C. H. & Petrusek, A. & Duris, Z." box="[1270,1335,706,735]" pageId="13" pageNumber="14" pagination="71 - 81" refId="ref10071" refString="Horka, I.; De Grave, S.; Fransen, C. H.; Petrusek, A. and Duris, Z. 2018. Multiple origins and strong phenotypic convergence in fish-cleaning palaemonid shrimp lineages. Molecular Phylogenetics and Evolution, 124: 71 - 81." type="journal article" year="2018">2018</bibRefCitation>
|
||
). Similarly, the tree topology of 16S gene sequences showed similar clade patterns for
|
||
<taxonomicName id="4C264D28FFBFFF9E2320FCE265FEFCCA" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1150,1249,786,815]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2320FCE265FEFCCA" box="[1150,1249,786,815]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E23B8FCE2645FFCCB" box="[1254,1344,786,814]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and other
|
||
<taxonomicName id="4C264D28FFBFFF9E2415FCCA6533FCB3" box="[843,1068,825,854]" pageId="13" pageNumber="14">
|
||
<emphasis id="B952EAB9FFBFFF9E2415FCCA62CBFCB3" box="[843,980,826,854]" italics="true" pageId="13" pageNumber="14">Actinimenes</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131D2A2EFFBFFF9E2362FCC96594FCB3" box="[1084,1163,825,854]" captionStart="Figure 7" captionStartId="14.[181,250,1151,1174]" captionTargetBox="[181,1460,193,1089]" captionTargetId="figure-40@14.[173,1466,151,1095]" captionTargetPageId="14" captionText="Figure 7. Maximum Likelihood phylogenetic tree based on partial COI (A) and 16S (B) sequences of Actinimenes koyas sp. nov. and two other Actinimenes species obtained from GenBank (Scale bar = 10 % divergence; significant bootstrap value> 60)." figureDoi="http://doi.org/10.5281/zenodo.10904886" httpUri="https://zenodo.org/record/10904886/files/figure.png" pageId="13" pageNumber="14">Fig. 7B</figureCitation>
|
||
). Unfortunately, COI and 16S sequences are not available for
|
||
<taxonomicName id="4C264D28FFBFFF9E225FFC916488FC9B" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1281,1431,865,894]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E225FFC916488FC9B" box="[1281,1431,865,894]" italics="true" pageId="13" pageNumber="14">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
in GenBank.The genetic divergence between
|
||
<taxonomicName id="4C264D28FFBFFF9E2265FC796488FC43" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1339,1431,905,934]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2265FC796488FC43" box="[1339,1431,905,934]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E22C5FC796262FC28" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and the other species of
|
||
<taxonomicName id="4C264D28FFBFFF9E23C6FC41643BFC28" authorityName="Duris and Horka" authorityYear="2017" box="[1176,1316,945,973]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B952EAB9FFBFFF9E23C6FC41643BFC28" box="[1176,1316,945,973]" italics="true" pageId="13" pageNumber="14">Actinimenes</emphasis>
|
||
</taxonomicName>
|
||
ranged from 15.7 to 18.7% for COI (
|
||
<tableCitation id="C6A40310FFBFFF9E2307FC286581FC10" box="[1113,1182,984,1013]" captionStart="Table 3" captionStartId="14.[181,241,1317,1340]" captionTargetPageId="14" captionText="Table 3.Pairwise geneticdistances calculated using Kimura2-parametermodel with COI sequencesof genus Actinimenes,Ancylocaris and Zenopontonia." pageId="13" pageNumber="14">Tab.3</tableCitation>
|
||
) and 1.2 to 6.5% for 16S (
|
||
<tableCitation id="C6A40310FFBFFF9E2408FBF06282FBF8" box="[854,925,1024,1053]" captionStart="Table 4" captionStartId="15.[181,241,152,175]" captionTargetPageId="15" captionText="Table 4.Pairwise genetic distances calculated usingKimura 2-parameter model with 16S sequencesof genus Actinimenes, Ancylocaris and Zenopontonia." pageId="13" pageNumber="14">Tab. 4</tableCitation>
|
||
). These high genetic differences are generally considered species-specific for decapod crustaceans (
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E240CFBBF6572FB89" author="Malay, M. C. M. D. & Paulay, G." box="[850,1133,1103,1132]" pageId="13" pageNumber="14" pagination="634 - 662" refId="ref10516" refString="Malay, M. C. M. D. and Paulay, G. 2010. Peripatric speciation drives diversification and distributional pattern of reef hermit crabs (Decapoda: Diogenidae: Calcinus). Evolution, 64: 634 - 662." type="journal article" year="2010">Malay and Paulay, 2010</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E232BFBBF64AAFB89" author="Komai, T. & Tsuchida, S." box="[1141,1461,1103,1132]" pageId="13" pageNumber="14" pagination="1789 - 1824" refId="ref10244" refString="Komai, T. and Tsuchida, S. 2015. New records of Alvinocarididae (Crustacea: Decapoda: Caridea) from the southwestern Pacific hydrothermal vents, with descriptions of one new genus and three new species. Journal of Natural History, 49: 1789 - 1824." type="journal article" year="2015">Komai and Tsuchida, 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2417FB87657AFB76" author="Chakraborty, R. D. & Chan, T. Y. & Maheswarudu, G. & Kuberan, G. & Purushothaman, P. & Chang, S. C. & Jomon, S." box="[841,1125,1142,1172]" pageId="13" pageNumber="14" pagination="923 - 930" refId="ref9402" refString="Chakraborty, R. D.; Chan, T. Y.; Maheswarudu, G.; Kuberan, G.; Purushothaman, P.; Chang, S. C. and Jomon, S. 2015. On Plesionika quasigrandis Chace, 1985 (Decapoda, Caridea, Pandalidae) from southwestern India. Crustaceana, 88: 923 - 930." type="journal article" year="2015">
|
||
Chakraborty
|
||
<emphasis id="B952EAB9FFBFFF9E24B7FB87653CFB76" box="[1001,1059,1143,1172]" italics="true" pageId="13" pageNumber="14">et al.,</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2330FB8764AFFB71" author="Purushothaman, P. & Chakraborty, R. D. & Kuberan, G. & Maheswarudu, G." box="[1134,1456,1143,1172]" pageId="13" pageNumber="14" pagination="12" refId="ref11058" refString="Purushothaman, P.; Chakraborty, R. D.; Kuberan, G. and Maheswarudu, G. 2019. Integrative taxonomy of commercially important deep-water penaeoid shrimps from India. Journal of Genetics, 98: 12." type="journal article" year="2019">
|
||
Purushothaman
|
||
<emphasis id="B952EAB9FFBFFF9E226DFB876472FB76" box="[1331,1389,1143,1172]" italics="true" pageId="13" pageNumber="14">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
). The present molecular analyses are strongly supported by the morphological identification of
|
||
<taxonomicName id="4C264D28FFBFFF9E2277FB366491FB06" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1321,1422,1222,1251]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2277FB366491FB06" box="[1321,1422,1222,1251]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E22C9FB376263FAEF" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from
|
||
<collectingCountry id="F331763BFFBFFF9E24E0FB1E657CFAEE" box="[958,1123,1262,1291]" name="India" pageId="13" pageNumber="14">Lakshadweep</collectingCountry>
|
||
,
|
||
<collectingCountry id="F331763BFFBFFF9E2332FB1E65B1FAEE" box="[1132,1198,1262,1291]" name="India" pageId="13" pageNumber="14">India</collectingCountry>
|
||
.Furthermore,
|
||
<taxonomicName id="4C264D28FFBFFF9E2202FB1E64A5FAEE" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1372,1466,1262,1291]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E2202FB1E64A5FAEE" box="[1372,1466,1262,1291]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E2415FAE662B8FAD7" box="[843,935,1302,1330]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs considerably both in morphological traits and coloration from its sister species. These sea anemone-associated shrimps exhibit remarkable color patterns;
|
||
<taxonomicName id="4C264D28FFBFFF9E24E4FA7D6558FA4C" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[954,1095,1421,1449]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E24E4FA7D6558FA4C" box="[954,1095,1421,1449]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C264D28FFBFFF9E2323FA7D6412FA4C" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1149,1293,1420,1449]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E2323FA7D6412FA4C" box="[1149,1293,1420,1449]" italics="true" pageId="13" pageNumber="14">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
have eyestalks with a broad dorsal white band and
|
||
<taxonomicName id="4C264D28FFBFFF9E2254FA446463FA34" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[1290,1404,1460,1489]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatu">
|
||
<emphasis id="B952EAB9FFBFFF9E2254FA446463FA34" box="[1290,1404,1460,1489]" italics="true" pageId="13" pageNumber="14">A. ornatu</emphasis>
|
||
</taxonomicName>
|
||
s has minute red and white chromatophores on the body (
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E240AF9F362F9F9C5" author="Bruce, A. J." box="[852,998,1539,1568]" pageId="13" pageNumber="14" pagination="209 - 230" refId="ref8978" refString="Bruce, A. J. 1979. A report on a small collection of pontoniine shrimps from Eniwetok Atoll. Crustaceana, Supplement 5: 209 - 230." type="journal article" year="1979">Bruce,1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E24AEF9F46587F9C5" author="Fransen, C. H. J. M." box="[1008,1176,1539,1568]" pageId="13" pageNumber="14" pagination="131 - 147" refId="ref9778" refString="Fransen, C. H. J. M. 1989. Notes on caridean shrimps collected during the Snellius-II Expedition. I. Associates of Anthozoa. Netherlands Journal of Sea Research, 23: 131 - 147." type="journal article" year="1989">Fransen, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E23FFF9F46473F9C5" author="Lee, K. H. & Ko, H. S." box="[1185,1388,1539,1568]" pageId="13" pageNumber="14" pagination="239 - 245" refId="ref10470" refString="Lee, K. H. and Ko, H. S. 2011. First records of two pontoniid shrimps (Crustacea: Decapoda: Caridea: Palaemonidae) from Korea. Animal Systematics, Evolution and Diversity, 27: 239 - 245." type="journal article" year="2011">Lee and Ko, 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E222AF9F36500F9AD" author="Salvat, B. & Bacchet, P." pageId="13" pageNumber="14" refId="ref11170" refString="Salvat, B. and Bacchet, P. 2011. Guide des recifs coralliens de Tahiti et ses iles. Papeete, Au Vent des Iles, 365 p." type="book" year="2011">Salvat and Bacchet, 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB74B5AFFBFFF9E2376F9DD6428F9AD" author="Duris, Z." box="[1064,1335,1579,1613]" pageId="13" pageNumber="14" pagination="1147 - 1161" refId="ref9580" refString="Duris, Z. 2017. Palaemonid shrimps (Crustacea: Decapoda) of Saudi Arabia from the ' Red Sea biodiversity survey' 2011 - 2013, with 11 new records for the Red Sea. Marine Biodiversity, 47: 1147 - 1161." type="journal article" year="2017">Ďuriš and Horká, 2017</bibRefCitation>
|
||
). The color patterns of the new species are rather dull and without band patterns. Furthermore,
|
||
<taxonomicName id="4C264D28FFBFFF9E23C1F98B65E0F972" authority="Paramasivam & Dhinakaran & Ajith Kumar & Lal, 2022" authorityName="Paramasivam & Dhinakaran & Ajith Kumar & Lal" authorityYear="2022" box="[1183,1279,1658,1687]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="koyas" status="sp. nov.">
|
||
<emphasis id="B952EAB9FFBFFF9E23C1F98B65E0F972" box="[1183,1279,1658,1687]" italics="true" pageId="13" pageNumber="14">A. koyas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157C2FFBFFF9E225AF98B6444F972" box="[1284,1371,1659,1687]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
showed significant differences with
|
||
<taxonomicName id="4C264D28FFBFFF9E23D3F9526407F95A" baseAuthorityName="Kemp" baseAuthorityYear="1922" box="[1165,1304,1698,1727]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="inornatus">
|
||
<emphasis id="B952EAB9FFBFFF9E23D3F9526407F95A" box="[1165,1304,1698,1727]" italics="true" pageId="13" pageNumber="14">A. inornatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D28FFBFFF9E227BF95264ABF95A" baseAuthorityName="Bruce" baseAuthorityYear="1979" box="[1317,1460,1698,1727]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatellus">
|
||
<emphasis id="B952EAB9FFBFFF9E227BF95264ABF95A" box="[1317,1460,1698,1727]" italics="true" pageId="13" pageNumber="14">A. ornatellus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C264D28FFBFFF9E2422F93A62F6F903" baseAuthorityName="Bruce" baseAuthorityYear="1969" box="[892,1001,1738,1766]" class="Malacostraca" family="Palaemonidae" genus="Actinimenes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatu">
|
||
<emphasis id="B952EAB9FFBFFF9E2422F93A62F6F903" box="[892,1001,1738,1766]" italics="true" pageId="13" pageNumber="14">A. ornatu</emphasis>
|
||
</taxonomicName>
|
||
s in morphological characters related to the antennular f lagellum, sternal plate, and length of pereopods (
|
||
<tableCitation id="C6A40310FFBFFF9E2482F8E9653AF8D3" box="[988,1061,1817,1846]" captionStart="Table 2" captionStartId="12.[181,242,152,175]" captionTargetPageId="12" captionText="Table 2.Comparison of morphologicalfeatures of Actinimenes koyas sp.nov.and other Actinimenes spp.:Anp–Antennular peduncle and TeL–Telson length." pageId="13" pageNumber="14">Tab. 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |