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<document id="F3F16922DE091B5C9064E2EF12E805A4" ID-CLB-Dataset="45825" ID-DOI="10.5281/zenodo.206943" ID-GBIF-Dataset="1f8d9fa0-f9a1-4029-bd2b-efa8b99537dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="206943" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460417791106" checkinUser="plazi" docAuthor="Tri, Ngo Van &amp; Onn, Chan Kin" docDate="2011" docId="038F87B1FFCBFC03FF77F98D0F5C9770" docLanguage="en" docName="zt03125p063.pdf" docOrigin="Zootaxa 3125" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyrtodactylus cucphuongensis Tri &amp; Onn, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="57" masterDocId="FFB6FFC9FFCAFC05FFE0FFD40B18956B" masterDocTitle="A new karstic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from Northern Vietnam" masterLastPageNumber="63" masterPageNumber="51" pageNumber="52" updateTime="1698329184312" updateUser="ExternalLinkService">
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<mods:title id="0B1F39502F77F7A360A728DD056E7F0C">A new karstic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from Northern Vietnam</mods:title>
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<mods:namePart id="0953609332907CB7886F263C33F0E6B7">Tri, Ngo Van</mods:namePart>
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<mods:namePart id="4C8521EB2FEC9A61785D45A32549EC58">Onn, Chan Kin</mods:namePart>
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<mods:date id="B367C84D2947839243036DC88D0B4CD0">2011</mods:date>
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<treatment id="038F87B1FFCBFC03FF77F98D0F5C9770" ID-DOI="http://doi.org/10.5281/zenodo.5688290" ID-GBIF-Taxon="119425610" ID-Zenodo-Dep="5688290" LSID="urn:lsid:plazi:treatment:038F87B1FFCBFC03FF77F98D0F5C9770" httpUri="http://treatment.plazi.org/id/038F87B1FFCBFC03FF77F98D0F5C9770" lastPageId="6" lastPageNumber="57" pageId="1" pageNumber="52">
<subSubSection id="C33C652CFFCBFC04FF77F98D0A3B93FF" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="8B9936A7FFCBFC04FF77F98D09749319" blockId="1.[151,620,1624,1684]" box="[151,620,1624,1651]" pageId="1" pageNumber="52">
<heading id="D0D181CBFFCBFC04FF77F98D09749319" bold="true" box="[151,620,1624,1651]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="B952EAB5FFCBFC04FF77F98D09749319" bold="true" box="[151,620,1624,1651]" pageId="1" pageNumber="52">
<taxonomicName id="4C264D24FFCBFC04FF77F98D091F9318" box="[151,519,1625,1651]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="52" phylum="Chordata" rank="species" species="cucphuongensis" status="sp. nov.">
<emphasis id="B952EAB5FFCBFC04FF77F98D091F9318" bold="true" box="[151,519,1625,1651]" italics="true" pageId="1" pageNumber="52">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157CEFFCBFC04FDEFF98C09749319" box="[527,620,1624,1650]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936A7FFCBFC04FF77F9A80A3B93FF" blockId="1.[151,620,1624,1684]" box="[151,291,1660,1684]" pageId="1" pageNumber="52">Figures: 13</paragraph>
</subSubSection>
<subSubSection id="C33C652CFFCBFC04FF77F91009AF924C" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="8B9936A7FFCBFC04FF77F91009AF924C" blockId="1.[151,1437,1731,2011]" pageId="1" pageNumber="52">
<emphasis id="B952EAB5FFCBFC04FF77F9100AB193B7" bold="true" box="[151,425,1731,1756]" pageId="1" pageNumber="52">
<typeStatus id="549D8805FFCBFC04FF77F9100A1E93B7" box="[151,262,1732,1756]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
.
<collectionCode id="ED37AE62FFCBFC04FEF2F9170A7193B7" box="[274,361,1731,1756]" pageId="1" pageNumber="52">ITBCZ</collectionCode>
2206
</emphasis>
, adult male collected by Ngo Van Tri on
<date id="FF981067FFCBFC04FC9BF9100F1493B7" box="[891,1036,1732,1756]" pageId="1" pageNumber="52" value="2008-07-20">20 July 2008</date>
from the karst outcrops adjacent to the main road in Cuc Phuong National Park, Nho Quan district, Ninh Binh province, Northern
<collectingCountry id="F3317637FFCBFC04FADDF9330E829394" box="[1341,1434,1767,1791]" name="Vietnam" pageId="1" pageNumber="52">Vietnam</collectingCountry>
(
<geoCoordinate id="EE125060FFCBFC04FF7FF8DB0A36924C" box="[159,302,1804,1831]" direction="north" orientation="latitude" pageId="1" pageNumber="52" precision="1" value="20.2554">20º15.324N</geoCoordinate>
,
<geoCoordinate id="EE125060FFCBFC04FED9F8DB0AC9924C" box="[313,465,1804,1831]" direction="east" orientation="longitude" pageId="1" pageNumber="52" precision="1" value="105.63433">105º38.060E</geoCoordinate>
) at
<quantity id="4CDE9B42FFCBFC04FE1BF8DB095A924C" box="[507,578,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="52" unit="m" value="100.0">100 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="C33C652CFFCBFC07FF27F8E60E1995BF" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="8B9936A7FFCBFC07FF27F8E60E1995BF" blockId="1.[151,1437,1731,2011]" lastBlockId="2.[151,1437,151,717]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<emphasis id="B952EAB5FFCBFC04FF27F8E60A599220" bold="true" box="[199,321,1842,1867]" pageId="1" pageNumber="52">Diagnosis.</emphasis>
<taxonomicName id="4C264D24FFCBFC04FEA8F8E709B89220" box="[328,672,1843,1867]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="52" phylum="Chordata" rank="species" species="cucphuongensis" status="sp. nov.">
<emphasis id="B952EAB5FFCBFC04FEA8F8E709B89220" box="[328,672,1843,1867]" italics="true" pageId="1" pageNumber="52">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157CEFFCBFC04FD46F8E708189220" box="[678,768,1843,1867]" pageId="1" pageNumber="52" rank="species">
<emphasis id="B952EAB5FFCBFC04FD46F8E708189220" bold="true" box="[678,768,1843,1867]" pageId="1" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all other congeners by the following combination of characters: SVL 96.0 mm; large, dark spots on top of head; wide nuchal band which tapers at both ends; five or six irregular, dark brown bands between limb insertions; precloacal pores absent; 14 relatively enlarged scales beneath thighs in discontinuous arrangement, separated medially by eight smaller scales; 18 interorbital scales on frontal bone; 18 scales in a straight line between eye and nostril; 42 rows of ventral scales between ventrolateral folds; 10 irregular, longitudinal rows of conical tubercles at midbody between lateral folds; 27 paravertebral tubercles between limb insertions; 19 subdigital lamellae on first toe; 24 subdigital lamellae on fourth toe.
</paragraph>
</subSubSection>
<subSubSection id="C33C652CFFC8FC00FF27FF0B0E9D9637" lastPageId="5" lastPageNumber="56" pageId="2" pageNumber="53" type="description">
<paragraph id="8B9936A7FFC8FC07FF27FF0B090C97A7" blockId="2.[151,1437,151,717]" pageId="2" pageNumber="53">
<emphasis id="B952EAB5FFC8FC07FF27FF0B0AF29593" bold="true" box="[199,490,223,248]" pageId="2" pageNumber="53">
Description of
<typeStatus id="549D8805FFC8FC07FE9CFF340AFD9593" box="[380,485,224,248]" pageId="2" pageNumber="53" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, SVL 96.0 mm (
<figureCitation id="131D2A22FFC8FC07FCDBFF0B088C9593" box="[827,916,223,248]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1842,1865]" captionTargetBox="[158,1427,762,1830]" captionTargetId="figure@2.[151,1436,756,1839]" captionTargetPageId="2" captionText="FIGURE 1. A: Live male holotype of Cyrtodactylus cucphuongensis (ITBZC 2206); B: preserved holotype of Cyrtodactylus cucphuongensis (ITBZC 2206)." httpUri="https://zenodo.org/record/206944/files/figure.png" pageId="2" pageNumber="53">
Fig. 1
<emphasis id="B952EAB5FFC8FC07FC60FF0B088C9593" bold="true" box="[896,916,223,248]" pageId="2" pageNumber="53">A</emphasis>
</figureCitation>
). Head moderately long (HeadL/SVL: 0.27), relatively narrow (HeadW/HeadL: 0.65), depressed (HeadH/HeadL: 0.37), distinct from neck; lores and interorbital region inflated, canthus rostralis present, frontonasal region concave; snout elongate (SnEye/HeadL: 0.40), pointed, longer than eye diameter (OrbD/SnEye: 0.66); scales on snout small, rounded, granular, homogeneous, larger than those on occipital region. Eye large (OrbD/HeadL: 0.27), pupils reddish; supraciliaries short, bearing tiny conical spines posteriorly. Ear opening oval, oblique, small (EarL/HeadL: 0.05); eye to ear distance greater than diameter of eye (EyeEar/OrbD: 1.06). Rostral scales smooth, incompletely divided posteriorly by a shallow dorsal groove; two enlarged supranasals in broad contact posteriorly with one smaller intersupranasal (
<figureCitation id="131D2A22FFC8FC07FF7FFE2B0BFC977C" box="[159,228,511,535]" captionStart="FIGURE 2" captionStartId="3.[151,249,1962,1985]" captionTargetBox="[188,1405,360,1926]" captionTargetId="figure@3.[179,1415,351,1936]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cyrtodactylus cucphuongensis showing: A: Rostral scales; B: Mental scales; C: Tubercle rows on dorsum; D: Ventral scales; E: Cloacal and femoral region; F: Enlarged subcaudal scales." httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="2" pageNumber="53">Fig. 2</figureCitation>
A); rostral in contact with first supralabial and nostril; nostril oval, surrounded by supranasal, rostral, first supralabial, and three enlarged postnasals; 13 rows of small scales separates orbit from supralabials. Mental triangular, wider (3.0 mm) than deep (2.2 mm); one pair of enlarged postmentals in broad contact medially, bordered anteromedially by mental, bordered anterolaterally by first infralabial, and posterolaterally by six enlarged lateral chinshields (
<figureCitation id="131D2A22FFC8FC07FE37FD5B090397CC" box="[471,539,655,679]" captionStart="FIGURE 2" captionStartId="3.[151,249,1962,1985]" captionTargetBox="[188,1405,360,1926]" captionTargetId="figure@3.[179,1415,351,1936]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cyrtodactylus cucphuongensis showing: A: Rostral scales; B: Mental scales; C: Tubercle rows on dorsum; D: Ventral scales; E: Cloacal and femoral region; F: Enlarged subcaudal scales." httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="2" pageNumber="53">Fig. 2</figureCitation>
B). Twelve supralabials on both sides; 16 interorbital scale rows on frontal bone; 20 scales between eye and nostril.
</paragraph>
<caption id="DF59662FFFC8FC07FF77F8E60AC1920C" httpUri="https://zenodo.org/record/206944/files/figure.png" pageId="2" pageNumber="53" targetBox="[158,1427,762,1830]" targetPageId="2">
<paragraph id="8B9936A7FFC8FC07FF77F8E60AC1920C" blockId="2.[151,1436,1842,1895]" pageId="2" pageNumber="53">
<emphasis id="B952EAB5FFC8FC07FF77F8E60A2C9223" bold="true" box="[151,308,1842,1865]" pageId="2" pageNumber="53">FIGURE 1. A:</emphasis>
Live male holotype of
<taxonomicName id="4C264D24FFC8FC07FDC4F8E708439222" box="[548,859,1843,1865]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="cucphuongensis">
<emphasis id="B952EAB5FFC8FC07FDC4F8E708439222" box="[548,859,1843,1865]" italics="true" pageId="2" pageNumber="53">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
(ITBZC 2206);
<emphasis id="B952EAB5FFC8FC07FBE2F8E70F039223" bold="true" box="[1026,1051,1843,1864]" pageId="2" pageNumber="53">B:</emphasis>
preserved holotype of
<taxonomicName id="4C264D24FFC8FC07FAEEF8E70A21920C" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="cucphuongensis">
<emphasis id="B952EAB5FFC8FC07FAEEF8E70A21920C" italics="true" pageId="2" pageNumber="53">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
(ITBZC 2206).
</paragraph>
</caption>
<paragraph id="8B9936A7FFC8FC06FF25F84009099454" blockId="2.[151,1436,1940,2036]" lastBlockId="3.[151,1436,151,319]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
Body slender, elongate (TrunkL/SVL: 0.43). Dorsal scales granular; large, regularly distributed conical tubercles (57 times size of adjacent scales) extend from occipital region to base of tail; larger tubercles arranged in 10 rows at midbody between ventrolateral folds (
<figureCitation id="131D2A22FFC8FC07FD48F80809F6929F" box="[680,750,2012,2036]" captionStart="FIGURE 2" captionStartId="3.[151,249,1962,1985]" captionTargetBox="[188,1405,360,1926]" captionTargetId="figure@3.[179,1415,351,1936]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cyrtodactylus cucphuongensis showing: A: Rostral scales; B: Mental scales; C: Tubercle rows on dorsum; D: Ventral scales; E: Cloacal and femoral region; F: Enlarged subcaudal scales." httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="2" pageNumber="53">Fig. 2</figureCitation>
C); tubercles are smallest on flanks and occipital region; 27 paravertebral tubercles between limb insertions. Ventral scales larger than those on dorsum, smooth, relatively round, subimbricate, largest posteriorly; 42 scale rows across belly between ventrolateral folds (
<figureCitation id="131D2A22FFC9FC06FB0EFF680E2C95BF" box="[1262,1332,188,212]" captionStart="FIGURE 2" captionStartId="3.[151,249,1962,1985]" captionTargetBox="[188,1405,360,1926]" captionTargetId="figure@3.[179,1415,351,1936]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cyrtodactylus cucphuongensis showing: A: Rostral scales; B: Mental scales; C: Tubercle rows on dorsum; D: Ventral scales; E: Cloacal and femoral region; F: Enlarged subcaudal scales." httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="3" pageNumber="54">Fig. 2</figureCitation>
D); gular region with relatively homogeneous, smooth scales. Precloacal groove absent; precloacal scales enlarged; precloacal pores absent (
<figureCitation id="131D2A22FFC9FC06FEB9FED00A859477" box="[345,413,260,284]" captionStart="FIGURE 2" captionStartId="3.[151,249,1962,1985]" captionTargetBox="[188,1405,360,1926]" captionTargetId="figure@3.[179,1415,351,1936]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cyrtodactylus cucphuongensis showing: A: Rostral scales; B: Mental scales; C: Tubercle rows on dorsum; D: Ventral scales; E: Cloacal and femoral region; F: Enlarged subcaudal scales." httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="3" pageNumber="54">Fig. 2</figureCitation>
E). Scales on palm and hind limbs smooth, granular with scattered, conical tubercles which are smaller than those on dorsum.
</paragraph>
<caption id="DF59662FFFC9FC06FF77F87E0F02928B" httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="3" pageNumber="54" targetBox="[188,1405,360,1926]" targetPageId="3">
<paragraph id="8B9936A7FFC9FC06FF77F87E0F02928B" blockId="3.[151,1436,1962,2016]" pageId="3" pageNumber="54">
<emphasis id="B952EAB5FFC9FC06FF77F87E0A0D92AB" bold="true" box="[151,277,1962,1985]" pageId="3" pageNumber="54">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="4C264D24FFC9FC06FE41F87F09C292AA" box="[417,730,1963,1985]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="cucphuongensis">
<emphasis id="B952EAB5FFC9FC06FE41F87F09C292AA" box="[417,730,1963,1985]" italics="true" pageId="3" pageNumber="54">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
showing:
<emphasis id="B952EAB5FFC9FC06FCA7F87F087A92AB" bold="true" box="[839,866,1963,1984]" pageId="3" pageNumber="54">A:</emphasis>
Rostral scales;
<emphasis id="B952EAB5FFC9FC06FBE8F87F0F3A92AB" bold="true" box="[1032,1058,1963,1984]" pageId="3" pageNumber="54">B:</emphasis>
Mental scales;
<emphasis id="B952EAB5FFC9FC06FB26F87E0FC092AA" bold="true" box="[1222,1240,1962,1985]" pageId="3" pageNumber="54">C</emphasis>
: Tubercle rows on dorsum;
<emphasis id="B952EAB5FFC9FC06FF10F81E0A1292B4" bold="true" box="[240,266,1994,2015]" pageId="3" pageNumber="54">D:</emphasis>
Ventral scales;
<emphasis id="B952EAB5FFC9FC06FE4AF81E0ADB92B4" bold="true" box="[426,451,1994,2015]" pageId="3" pageNumber="54">E:</emphasis>
Cloacal and femoral region;
<emphasis id="B952EAB5FFC9FC06FD0CF81E081C92B4" bold="true" box="[748,772,1994,2015]" pageId="3" pageNumber="54">F:</emphasis>
Enlarged subcaudal scales.
</paragraph>
</caption>
<caption id="DF59662FFFCEFC01FF77F8160FAC92B3" box="[151,1204,1986,2009]" httpUri="https://zenodo.org/record/206946/files/figure.png" pageId="4" pageNumber="55" targetBox="[167,1423,206,1951]" targetPageId="4">
<paragraph id="8B9936A7FFCEFC01FF77F8160FAC92B3" blockId="4.[151,1204,1986,2009]" box="[151,1204,1986,2009]" pageId="4" pageNumber="55">
<emphasis id="B952EAB5FFCEFC01FF77F8160A2B92B3" bold="true" box="[151,307,1986,2009]" pageId="4" pageNumber="55">FIGURE 3. A:</emphasis>
Manus of
<taxonomicName id="4C264D24FFCEFC01FE40F81709CF92B2" box="[416,727,1987,2009]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="55" phylum="Chordata" rank="species" species="cucphuongensis" status="sp. nov.">
<emphasis id="B952EAB5FFCEFC01FE40F81709CF92B2" box="[416,727,1987,2009]" italics="true" pageId="4" pageNumber="55">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157CEFFCEFC01FD3DF817083592B3" box="[733,813,1987,2008]" pageId="4" pageNumber="55" rank="species">
<emphasis id="B952EAB5FFCEFC01FD3DF817083592B3" bold="true" box="[733,813,1987,2008]" pageId="4" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
;
<emphasis id="B952EAB5FFCEFC01FCDBF817085492B3" bold="true" box="[827,844,1987,2008]" pageId="4" pageNumber="55">B</emphasis>
: Pes of
<taxonomicName id="4C264D24FFCEFC01FC7EF8170F4692B2" box="[926,1118,1987,2009]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="55" phylum="Chordata" rank="species" species="cucphuongensis" status="sp. nov.">
<emphasis id="B952EAB5FFCEFC01FC7EF8170F4692B2" box="[926,1118,1987,2009]" italics="true" pageId="4" pageNumber="55">C. cucphuongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157CEFFCEFC01FB84F8170FAC92B3" box="[1124,1204,1987,2008]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B9936A7FFCFFC00FF25FF4C0BFC94C7" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="56">
Fore and hindlimbs slender (ForeL/SVL: 0.17; CrusL/SVL: 0.19); digits slender, strongly inflected at basal interphalangeal joints, all bearing slightly curved claws; basal subdigital lamellae nearly as broad as digit, without scansorial surface: 78878 manus;
<date id="FF981067FFCFFC00FDB0FF0B09D4959C" box="[592,716,223,247]" pageId="5" pageNumber="56" value="1911-07-07" valueMax="1911-10-07" valueMin="1911-07-07">771011</date>
10 pes; narrow lamellae distal to digital inflection, not including ventral claw sheath:
<date id="FF981067FFCFFC00FE51FED009539477" box="[433,587,260,284]" pageId="5" pageNumber="56" value="1914-12-12" valueMax="1914-12-14" valueMin="1914-12-12">12121414</date>
12 manus;
<date id="FF981067FFCFFC00FD38FED008699477" box="[728,881,260,284]" pageId="5" pageNumber="56" value="1913-12-13" valueMax="1913-12-14" valueMin="1913-12-13">12131413</date>
16 pes; one or two rows of small, non lamellar granules between basal and distal lamellar series; two or eight small, non lamellar granules between broad and narrow lamellae of manus and pes (
<figureCitation id="131D2A22FFCFFC00FDE1FE980941940F" box="[513,601,332,356]" captionStart="FIGURE 3. A" captionStartId="4.[151,250,1986,2009]" captionTargetBox="[167,1423,206,1951]" captionTargetId="figure@4.[151,1436,193,1965]" captionTargetPageId="4" captionText="FIGURE 3. A: Manus of Cyrtodactylus cucphuongensis sp. nov.; B: Pes of C. cucphuongensis sp. nov." httpUri="https://zenodo.org/record/206946/files/figure.png" pageId="5" pageNumber="56">Fig. 3A</figureCitation>
, B); interdigital webbing present but weakly developed. Relative length of digits in mm: (manus) III (8.9)&gt; IV (8.5)&gt; V (8.0)&gt; II (7.8)&gt; I (5.8); (pes): V (10.5)&gt; IV (10.2)&gt; III (9.4)&gt; II (7.1)&gt; I (5.8).
</paragraph>
<paragraph id="8B9936A7FFCFFC00FF25FE6309959757" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="56">
Tail regenerated at base, tail length 79.3 mm, slender, tapering to a point; two smooth, yellowish-white postcloacal tubercles at base; base of tail with four longitudinal rows of weakly keeled tubercles dorsally; dorsal surface of tail with heterogeneous scales; subcaudal scales smooth, juxtaposed; median row of narrow, enlarged subcaudal scales throughout the length of tail (
<figureCitation id="131D2A22FFCFFC00FDCAFDF009759757" box="[554,621,548,572]" captionStart="FIGURE 2" captionStartId="3.[151,249,1962,1985]" captionTargetBox="[188,1405,360,1926]" captionTargetId="figure@3.[179,1415,351,1936]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cyrtodactylus cucphuongensis showing: A: Rostral scales; B: Mental scales; C: Tubercle rows on dorsum; D: Ventral scales; E: Cloacal and femoral region; F: Enlarged subcaudal scales." httpUri="https://zenodo.org/record/206945/files/figure.png" pageId="5" pageNumber="56">Fig. 2</figureCitation>
F).
</paragraph>
<paragraph id="8B9936A7FFCFFC00FF25FD930E9D9637" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="56">
<emphasis id="B952EAB5FFCFFC00FF25FD930A8C970B" bold="true" box="[197,404,583,608]" pageId="5" pageNumber="56">Coloration in life</emphasis>
(
<figureCitation id="131D2A22FFCFFC00FE45FD9309189734" box="[421,512,583,607]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1842,1865]" captionTargetBox="[158,1427,762,1830]" captionTargetId="figure@2.[151,1436,756,1839]" captionTargetPageId="2" captionText="FIGURE 1. A: Live male holotype of Cyrtodactylus cucphuongensis (ITBZC 2206); B: preserved holotype of Cyrtodactylus cucphuongensis (ITBZC 2206)." httpUri="https://zenodo.org/record/206944/files/figure.png" pageId="5" pageNumber="56">Fig. 1A</figureCitation>
). Base color yellowish-brown; large, dark brown spots on top of head; eye rings yellowish; wide, dark brown nuchal band with tapering ends, outlined by yellow both anterior and posteriorly; postorbital stripes in contact with tapered ends of the nuchal band; two yellow-edged stripes on lateral surface of neck from posterior end of jaw, terminating at the shoulders; nape with a series of four oval blotches, the middle two connected, forming a butterfly shape. Five irregular, dark bands on dorsum, one immediately posterior to hind limbs in the form of two connected blotches; small dark blotches scattered on flanks and space between dorsal bands; dorsal surface of limbs, manus and pes yellowish-brown, scattered with dark bands and blotches; regenerated tail is dark brown. Ventral surface of body pinkish, scattered with small yellowish and dark-brown blotches.
</paragraph>
</subSubSection>
<subSubSection id="C33C652CFFCFFC00FF27FCBC0F0C96CE" pageId="5" pageNumber="56" type="etymology">
<paragraph id="8B9936A7FFCFFC00FF27FCBC0F0C96CE" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="56">
<emphasis id="B952EAB5FFCFFC00FF27FCBC0A5696EB" bold="true" box="[199,334,872,896]" pageId="5" pageNumber="56">Etymology.</emphasis>
The specific epithet is named after the Cúc Phơ ng National Park. Suggested common name: Cúc Phơ ng Benttoed Gecko; Vietnamese name: Thằn lằn chân ngón Cúc Phơ ng.
</paragraph>
</subSubSection>
<caption id="DF59662FFFCFFC00FF77F87E0A8992B4" httpUri="https://zenodo.org/record/206947/files/figure.png" pageId="5" pageNumber="56" targetBox="[151,1435,1062,1941]" targetPageId="5">
<paragraph id="8B9936A7FFCFFC00FF77F87E0A8992B4" blockId="5.[151,1436,1962,2015]" pageId="5" pageNumber="56">
<emphasis id="B952EAB5FFCFFC00FF77F87E0A0A92AB" bold="true" box="[151,274,1962,1985]" pageId="5" pageNumber="56">FIGURE 4.</emphasis>
Macrohabitat from the type locality of
<taxonomicName id="4C264D24FFCFFC00FD41F87E08C092AB" box="[673,984,1962,1984]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="56" phylum="Chordata" rank="species" species="cucphuongensis" status="sp. nov.">
<emphasis id="B952EAB5FFCFFC00FD41F87E08C092AB" box="[673,984,1962,1984]" italics="true" pageId="5" pageNumber="56">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157CEFFCFFC00FC3DF87F0F3692AB" box="[989,1070,1963,1984]" pageId="5" pageNumber="56" rank="species">
<emphasis id="B952EAB5FFCFFC00FC3DF87F0F3692AB" bold="true" box="[989,1070,1963,1984]" pageId="5" pageNumber="56">sp. nov.</emphasis>
</taxonomicNameLabel>
in Cuc Phuong National Park, Ninh Binh province, Vietnam.
</paragraph>
</caption>
<subSubSection id="C33C652CFFCCFC03FF27FF430F5C9770" pageId="6" pageNumber="57" type="distribution">
<paragraph id="8B9936A7FFCCFC03FF27FF430880940F" blockId="6.[151,1436,151,356]" pageId="6" pageNumber="57">
<emphasis id="B952EAB5FFCCFC03FF27FF43094B95DB" bold="true" box="[199,595,151,176]" pageId="6" pageNumber="57">Distribution and natural history.</emphasis>
<taxonomicName id="4C264D24FFCCFC03FDBCFF4C08AD95DB" box="[604,949,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="cucphuongensis" status="sp. nov.">
<emphasis id="B952EAB5FFCCFC03FDBCFF4C08AD95DB" box="[604,949,152,176]" italics="true" pageId="6" pageNumber="57">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157CEFFCCFC03FC5DFF4C0F0195DB" box="[957,1049,152,176]" pageId="6" pageNumber="57" rank="species">
<emphasis id="B952EAB5FFCCFC03FC5DFF4C0F0195DB" bold="true" box="[957,1049,152,176]" pageId="6" pageNumber="57">sp. nov.</emphasis>
</taxonomicNameLabel>
is currently known only from the cave in the karst forests of Cuc Phuong National Park in the Nho Quan district, Ninh Binh province and from the ceiling of a water-filled cave at Yen Thanh commune, Thach Thanh district, Thanh Hoa province, Northern
<collectingCountry id="F3317637FFCCFC03FA84FF0B0BDF9477" name="Vietnam" pageId="6" pageNumber="57">Vietnam</collectingCountry>
(
<bibRefCitation id="EFB74B56FFCCFC03FF36FED00AAD9477" author="Nguyen" box="[214,437,260,284]" pageId="6" pageNumber="57" refString="Nguyen, V. S., Ho, T. C., Nguyen, Q. T. &amp; Le, T. D. (2003) Herpetological diversity in Cuc Phuong National Park. Agriculture Publisher. Hanoi." type="book" year="2003">
Nguyen
<emphasis id="B952EAB5FFCCFC03FED8FED10A699477" box="[312,369,260,284]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
2003
</bibRefCitation>
). The specimen from the latter locality was caught during the dry season when the water was low. Other individuals were seen at the entrance of the cave of the
<typeStatus id="549D8805FFCCFC03FC5AFEFC08F2942B" box="[954,1002,296,320]" pageId="6" pageNumber="57">type</typeStatus>
locality but eluded capture. The original tail of the observed lizards was yellowish-white with dark bands.
</paragraph>
<caption id="DF59662FFFCCFC03FF77FE420E8494C0" ID-Table-UUID="DF59662FFFCCFC03FF77FE420E8494C0" box="[151,1436,405,428]" httpUri="http://table.plazi.org/id/DF59662FFFCCFC03FF77FE420E8494C0" pageId="6" pageNumber="57" targetBox="[151,1436,559,1367]" targetIsTable="true" targetPageId="6">
<paragraph id="8B9936A7FFCCFC03FF77FE420E8494C0" blockId="6.[151,1436,405,539]" box="[151,1436,405,428]" pageId="6" pageNumber="57">
<emphasis id="B952EAB5FFCCFC03FF77FE420A1D94C0" bold="true" box="[151,261,406,427]" pageId="6" pageNumber="57">TABLE 1.</emphasis>
Mensural and meristic data for the type series of
<taxonomicName id="4C264D24FFCCFC03FD1EFE420F2D94C7" box="[766,1077,406,428]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="cucphuongensis">
<emphasis id="B952EAB5FFCCFC03FD1EFE420F2D94C7" box="[766,1077,406,428]" italics="true" pageId="6" pageNumber="57">Cyrtodactylus cucphuongensis</emphasis>
</taxonomicName>
Abbreviations are stated in materi-
</paragraph>
</caption>
<paragraph id="8B9936A7FFCCFC03FF77FE610F1394A0" blockId="6.[151,1436,405,539]" box="[151,1035,437,459]" pageId="6" pageNumber="57">als and methods. Measurements in millimeters (mm); M= male; Reg = regenerated tail.</paragraph>
<paragraph id="8B9936A7FFCCFC03FE6FFE320F5C9770" blockId="6.[151,1436,405,539]" pageId="6" pageNumber="57">
<bibRef id="5286A251FFCCFC03FE6FFE320F5C9770" pageId="6" pageNumber="57" type="journal article">
<author id="539BDE42FFCCFC03FE6FFE320AC09771" fns="I" npo="LF" pageId="6" pageNumber="57">
<typeStatus id="549D8805FFCCFC03FE6FFE320F079497" box="[399,1055,486,508]" pageId="6" pageNumber="57" type="holotype">Holotype Holotype</typeStatus>
ITBCZ
</author>
<part id="FF829C3AFFCCFC03FE3FFDD109099770" box="[479,529,517,539]" pageId="6" pageNumber="57" type="volume">2206</part>
<title id="F922DD91FFCCFC03FC21FDD10F129771" box="[961,1034,517,538]" pageId="6" pageNumber="57">ITBCZ</title>
<pagination id="B2EFC853FFCCFC03FBF1FDD10F5C9770" box="[1041,1092,517,539]" pageId="6" pageNumber="57" type="pagination">2206</pagination>
</bibRef>
</paragraph>
</subSubSection>
</treatment>
</document>