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<document id="D0AB09D73ECEFBF227385D493BC07DC7" ID-CLB-Dataset="92619" ID-DOI="10.5852/ejt.2022.815.1749" ID-GBIF-Dataset="9a798cae-f10e-44c2-a65c-cdadbff28cb4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484153" ID-ZooBank="6AD0082E-7349-48DE-AFCA-1EE0BFBB3887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650913068508" checkinUser="felipe" docAuthor="Vranken, Nathan, Steenberge, Maarten Van, Heylen, Annelies, Decru, Eva &amp; Snoeks, Jos" docDate="2022" docId="038F87A6613ADA1BFD9951FF256AFE20" docLanguage="en" docName="ejt-815_1-94.pdf" docOrigin="European Journal of Taxonomy 815" docSource="http://dx.doi.org/10.5852/ejt.2022.815.1749" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplochromis quasimodo Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks 2022, sp. nov." docType="treatment" docUuid="6AE722D9-3BF6-4DD5-8637-55B8400EDD11" docUuidSource="ZooBank" docVersion="11" lastPageNumber="68" masterDocId="FFB6FFDE6107DA58FFD257322629FC0A" masterDocTitle="From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system" masterLastPageNumber="94" masterPageNumber="1" pageNumber="62" updateTime="1699329164302" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="52A9B1C85F444F33BD05B5A6DAC50125">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="102479E9ACAA6D214709C47C83A70DDC">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="10C565C73B1FF94FA830057206750B52">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. &amp; Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:affiliation id="978989BAC072D6983C048ACE74701388">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. &amp; Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:affiliation id="4ABE6F9D7CAFA1A1CBABD4FC6D4711AD">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:namePart id="8B0932793F6518F9F1AC1239AF4145E3">Decru, Eva</mods:namePart>
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<mods:affiliation id="1D264AA0A38CECB3C88B68A8E52A6432">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:namePart id="33C47249DEB3A4C63F0508B50AE2C50A">Snoeks, Jos</mods:namePart>
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<heading id="D0D181DC613ADA65FD9951FF25C0FAE2" box="[587,1001,1741,1768]" centered="true" fontSize="11" level="2" pageId="61" pageNumber="62" reason="2">
<taxonomicName id="4C264D33613ADA65FD9951FF25AFFAE2" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[587,902,1741,1768]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA2613ADA65FD9951FF25AFFAE2" bold="true" box="[587,902,1741,1768]" italics="true" pageId="61" pageNumber="62">Haplochromis quasimodo</emphasis>
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<taxonomicNameLabel id="A26157D9613ADA65FC5C51FC25C0FAE2" box="[910,1001,1742,1768]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C33C653B613ADA65FD5A50262585FB24" box="[648,940,1812,1839]" pageId="61" pageNumber="62" type="description">
<paragraph id="8B9936B0613ADA65FD5A50262585FB24" blockId="61.[365,1222,1741,1839]" box="[648,940,1812,1839]" pageId="61" pageNumber="62">
<figureCitation id="131D2A35613ADA65FD5A502624D9FB24" box="[648,752,1812,1838]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. latifrons (), H. mentatus Regan, 1925 (), H. sp. rex (), H. sp. simba (▼), H. sp. glaucus (), H. sp. aquila (), H. sp. kimondo (●), H. sp. falcatus (), H. sp. curvidens (), H. sp. pardus (), H. sp. quasimodo (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. latifrons the female and for H. sp. curvidens the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="61" pageNumber="62">Figs 12</figureCitation>
,
<figureCitation id="131D2A35613ADA65FD2E50262560FB25" box="[764,841,1812,1839]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="62.[189,232,1963,1989]" captionStartId-1="64.[189,232,1892,1918]" captionStartId-2="66.[190,233,1963,1989]" captionTargetBox-0="[189,1398,1159,1876]" captionTargetBox-1="[288,1293,287,1843]" captionTargetBox-2="[189,1397,970,1866]" captionTargetId-0="figure-358@62.[189,1399,1159,1876]" captionTargetId-1="figure-18@64.[288,1293,265,1843]" captionTargetId-2="figure-223@66.[189,1399,970,1868]" captionTargetPageId-0="62" captionTargetPageId-1="64" captionTargetPageId-2="66" captionText-0="Fig. 35. Haplochromis quasimodo sp. nov., holotype, ♂, 120.4 mm SL (RMCA 2018.008.P.0336). Drawn by N. Vranken." captionText-1="Fig. 36.Haplochromis quasimodosp. nov. a. Photograph of preservedholotype (RMCA 2018.008.P.0336; 120.4 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL). d. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 37. Haplochromis quasimodo sp. nov. (RMCA 2018.008.P.0337; 126.2 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484240" figureDoi-1="http://doi.org/10.5281/zenodo.6484242" figureDoi-2="http://doi.org/10.5281/zenodo.6484244" httpUri-0="https://zenodo.org/record/6484240/files/figure.png" httpUri-1="https://zenodo.org/record/6484242/files/figure.png" httpUri-2="https://zenodo.org/record/6484244/files/figure.png" pageId="61" pageNumber="62">3537</figureCitation>
;
<tableCitation id="C6A4030B613ADA65FC8750262585FB24" box="[853,940,1812,1838]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="61" pageNumber="62" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
</paragraph>
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<subSubSection id="C33C653B613ADA67FF6F506725BAFF23" lastPageId="63" lastPageNumber="64" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="8B9936B0613ADA65FF6F506727E0FB65" blockId="61.[189,457,1877,1903]" box="[189,457,1877,1903]" pageId="61" pageNumber="62">
<heading id="D0D181DC613ADA65FF6F506727E0FB65" bold="true" box="[189,457,1877,1903]" fontSize="11" level="3" pageId="61" pageNumber="62" reason="3">
<emphasis id="B952EAA2613ADA65FF6F506727E0FB65" bold="true" box="[189,457,1877,1903]" pageId="61" pageNumber="62">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B0613ADA65FF6F50B625A9FBEE" blockId="61.[187,1398,1923,2021]" pageId="61" pageNumber="62">Species with a piscivorous morphology; body rather deep [BD 33.541.7 (mean 37.4) % SL]; interorbital area narrow [IOW 40.548.7 (43.9) % HL]; outer oral teeth many and small [UOT 4671 (median 58)]; dominant males light grey dorsally and blue-black ventrally.</paragraph>
<paragraph id="8B9936B06139DA66FF6F563B234EFDBA" blockId="62.[189,1399,265,432]" pageId="62" pageNumber="63">
Amongst piscivorous species from the Lake Edward system,
<taxonomicName id="4C264D336139DA66FC6E56382241FD29" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[956,1128,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA26139DA66FC6E563825F8FD29" box="[956,977,266,291]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FC34563B2241FD29" box="[998,1128,265,291]" italics="true" pageId="62" pageNumber="63">quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FBA4563B22FEFD29" box="[1142,1239,265,291]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C264D336139DA66FF6F561F276CFD4C" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[189,325,300,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
<emphasis id="B952EAA26139DA66FF6F561F26FBFD4C" box="[189,210,301,326]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FF33561E276CFD4C" box="[225,325,300,326]" italics="true" pageId="62" pageNumber="63">latifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FE9E561F2780FD4D" box="[332,425,301,327]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D336139DA66FE66561F246AFD4C" authorityName="Regan" authorityYear="1925" box="[436,579,301,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="mentatus">
<emphasis id="B952EAA26139DA66FE66561F27E0FD4C" box="[436,457,301,326]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FE0A561F246AFD4C" box="[472,579,301,326]" italics="true" pageId="62" pageNumber="63">mentatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C264D336139DA66FD80561F24B2FD4C" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[594,667,301,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
<emphasis id="B952EAA26139DA66FD80561F244EFD4C" box="[594,615,301,326]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FDA4561F24B2FD4C" box="[630,667,301,326]" italics="true" pageId="62" pageNumber="63">rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FD70561F24D6FD4D" box="[674,767,301,327]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D336139DA66FCD9561F255FFD4C" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[779,886,300,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA26139DA66FCD9561F2509FD4C" box="[779,800,301,326]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FCFD561E255FFD4C" box="[815,886,300,326]" italics="true" pageId="62" pageNumber="63">simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FCAF561F25F0FD4D" box="[893,985,301,327]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D336139DA66FC37561F224CFD4C" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[997,1125,300,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
<emphasis id="B952EAA26139DA66FC37561F25D3FD4C" box="[997,1018,301,326]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FBDB561E224CFD4C" box="[1033,1125,300,326]" italics="true" pageId="62" pageNumber="63">glaucus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FBBF561F22E3FD4D" box="[1133,1226,301,327]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C264D336139DA66FAD5561F235EFD4C" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1287,1399,300,327]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
<emphasis id="B952EAA26139DA66FAD5561F2335FD4C" box="[1287,1308,301,326]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FAF9561E235EFD4C" box="[1323,1399,300,326]" italics="true" pageId="62" pageNumber="63">aquila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FF6F56622730FD60" box="[189,281,336,362]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
by the combination of small vs large outer oral teeth and a larger number of outer upper jaw teeth [UOT 4671 (58) vs 2247 (2736)]; further from
<taxonomicName id="4C264D336139DA66FC9D564125F7FD86" authorityName="Regan" authorityYear="1925" box="[847,990,371,397]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="mentatus">
<emphasis id="B952EAA26139DA66FC9D5641254DFD86" box="[847,868,371,396]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FCA1564125F7FD86" box="[883,990,371,396]" italics="true" pageId="62" pageNumber="63">mentatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C264D336139DA66FC3E5641221CFD86" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1004,1077,371,397]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
<emphasis id="B952EAA26139DA66FC3E56412228FD86" box="[1004,1025,371,396]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FBC25641221CFD86" box="[1040,1077,371,396]" italics="true" pageId="62" pageNumber="63">rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FBEE564122B1FD87" box="[1084,1176,371,397]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D336139DA66FB7656412326FD86" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1188,1295,370,397]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA26139DA66FB7656412290FD86" box="[1188,1209,371,396]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FB1A56402326FD86" box="[1224,1295,370,396]" italics="true" pageId="62" pageNumber="63">simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FAC75641235BFD87" box="[1301,1394,371,397]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D336139DA66FF6F56A52714FDBA" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[189,317,406,432]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
<emphasis id="B952EAA26139DA66FF6F56A526FBFDBA" box="[189,210,407,432]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FF3356A42714FDBA" box="[225,317,406,432]" italics="true" pageId="62" pageNumber="63">glaucus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FE9756A4278BFDBA" box="[325,418,406,432]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C264D336139DA66FE0D56A52479FDBA" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[479,592,406,432]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
<emphasis id="B952EAA26139DA66FE0D56A527DDFDBA" box="[479,500,407,432]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FDD656A42479FDBA" box="[516,592,406,432]" italics="true" pageId="62" pageNumber="63">aquila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FD8556A42498FDBA" box="[599,689,406,432]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
by presence vs absence of a well-defined mid-lateral band.
</paragraph>
<paragraph id="8B9936B06139DA66FF6F56EA2200FE32" blockId="62.[189,1399,471,568]" pageId="62" pageNumber="63">
It further differs from
<taxonomicName id="4C264D336139DA66FE1756EA2464FDFB" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[453,589,471,498]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
<emphasis id="B952EAA26139DA66FE1756EA27F3FDFB" box="[453,474,472,497]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FE3B56E52464FDFB" box="[489,589,471,497]" italics="true" pageId="62" pageNumber="63">latifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FD8456EA2498FDF8" box="[598,689,472,498]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C264D336139DA66FD3956EA2553FDFB" authorityName="Regan" authorityYear="1925" box="[747,890,472,498]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="mentatus">
<emphasis id="B952EAA26139DA66FD3956EA2529FDFB" box="[747,768,472,497]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FCDD56EA2553FDFB" box="[783,890,472,497]" italics="true" pageId="62" pageNumber="63">mentatus</emphasis>
</taxonomicName>
by a deeper body [BD 33.541.7 (37.4) vs 27.232.3 (28.631.2) % SL]; further from
<taxonomicName id="4C264D336139DA66FD1156C92524FE1E" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[707,781,507,533]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
<emphasis id="B952EAA26139DA66FD1156C924FEFE1E" box="[707,727,507,532]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FD3A56C92524FE1E" box="[744,781,507,532]" italics="true" pageId="62" pageNumber="63">rex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FCC756C9255AFE1F" box="[789,883,507,533]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C264D336139DA66FC5256C925C5FE1E" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[896,1004,506,533]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
<emphasis id="B952EAA26139DA66FC5256C925BCFE1E" box="[896,917,507,532]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FC7756C825C5FE1E" box="[933,1004,506,532]" italics="true" pageId="62" pageNumber="63">simba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FC2656C9227BFE1F" box="[1012,1106,507,533]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C264D336139DA66FB4056C9233AFE1E" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[1170,1299,506,533]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
<emphasis id="B952EAA26139DA66FB4056C9228EFE1E" box="[1170,1191,507,532]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FB6556C8233AFE1E" box="[1207,1299,506,532]" italics="true" pageId="62" pageNumber="63">glaucus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FAC956C9235EFE1F" box="[1307,1399,507,533]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
by a broader head [HW 42.048.1 (45.3) vs 36.841.6 (39.240.8) % HL].
</paragraph>
<paragraph id="8B9936B06139DA66FF6F555226D1FF99" blockId="62.[189,1401,607,915]" pageId="62" pageNumber="63">
It differs from
<taxonomicName id="4C264D336139DA66FEA1555227D4FE73" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[371,509,607,634]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
<emphasis id="B952EAA26139DA66FEA1555227AFFE73" box="[371,390,608,633]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FE45556D27D4FE73" box="[407,509,607,633]" italics="true" pageId="62" pageNumber="63">kimondo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FDDB5552244FFE70" box="[521,614,608,634]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C264D336139DA66FD715552254FFE73" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[675,870,608,634]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="squamipinnis">
<emphasis id="B952EAA26139DA66FD715552249FFE73" box="[675,694,608,633]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FD1A5552254FFE73" box="[712,870,608,633]" italics="true" pageId="62" pageNumber="63">squamipinnis</emphasis>
</taxonomicName>
by a narrower interorbital area [IOW 40.5 48.7 (43.9) vs 48.658.5 (51.952.8) % HW]; further from
<taxonomicName id="4C264D336139DA66FC5D55B12235FE96" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[911,1052,642,669]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
<emphasis id="B952EAA26139DA66FC5D55B1258DFE96" box="[911,932,643,668]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FC6455B02235FE96" box="[950,1052,642,668]" italics="true" pageId="62" pageNumber="63">kimondo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FBF555B122ADFE97" box="[1063,1156,643,669]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
by the combination of a rhomboid vs pyriform body, a concave to weakly convex vs convex dorsal outline of head, a gentler sloping snout (3040° vs 4050°), and dominant males light grey dorsally and blue-black vs grey dorsally and yellow ventrally; further from
<taxonomicName id="4C264D336139DA66FD2A55DF2593FF0C" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[760,954,748,774]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="squamipinnis">
<emphasis id="B952EAA26139DA66FD2A55DF2524FF0C" box="[760,781,749,774]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FCCE55DF2593FF0C" box="[796,954,749,774]" italics="true" pageId="62" pageNumber="63">squamipinnis</emphasis>
</taxonomicName>
by a gentler gape inclination (2035° vs 3045°), a shorter lower jaw [LJL 44.249.6 (47.1) vs 47.858.6 (52.5) % HL], mostly absence vs presence of minute scales on proximal part of dorsal fin (rarely few rows of 14 scales present on dorsal fin in
<taxonomicName id="4C264D336139DA66FED054642780FF65" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[258,425,853,880]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA26139DA66FED05464273EFF65" box="[258,279,854,879]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FEF554672780FF65" box="[295,425,853,879]" italics="true" pageId="62" pageNumber="63">quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FE6054642439FF7A" box="[434,528,854,880]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
), and dominant males light grey dorsally and blue-black ventrally vs slate blue.
</paragraph>
<paragraph id="8B9936B06139DA67FF6F54892501FD4C" blockId="62.[189,1399,954,1016]" lastBlockId="63.[188,1398,265,327]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
It differs from
<taxonomicName id="4C264D336139DA66FEB9548927C7FFDE" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[363,494,954,981]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
<emphasis id="B952EAA26139DA66FEB9548927A9FFDE" box="[363,384,955,980]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B952EAA26139DA66FE42548827C7FFDE" box="[400,494,954,980]" italics="true" pageId="62" pageNumber="63">falcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FE2754892479FFDF" box="[501,592,955,981]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
by the combination of a shorter head [HL 33.937.2 (35.5) vs 36.6 39.6 (38.2) % SL], a longer pelvic fin [VL 26.233.7 (29.4) vs 21.625.7 (23.5) % SL], weakly recurved vs strongly recurved outer oral teeth, and dominant males light grey dorsally and blue-black ventrally vs olive-green with an orange-red anterior part of flank.
</paragraph>
<caption id="DF5966386139DA66FF6F509927EEFBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484240" ID-Zenodo-Dep="6484240" httpUri="https://zenodo.org/record/6484240/files/figure.png" pageId="62" pageNumber="63" startId="62.[189,232,1963,1989]" targetBox="[189,1398,1159,1876]" targetPageId="62">
<paragraph id="8B9936B06139DA66FF6F509927EEFBE2" blockId="62.[189,1399,1963,2025]" pageId="62" pageNumber="63">
<emphasis id="B952EAA26139DA66FF6F50992736FBCF" bold="true" box="[189,287,1963,1989]" pageId="62" pageNumber="63">Fig. 35.</emphasis>
<taxonomicName id="4C264D336139DA66FEF950992448FBCF" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[299,609,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA26139DA66FEF950992448FBCF" box="[299,609,1963,1989]" italics="true" pageId="62" pageNumber="63">Haplochromis quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96139DA66FDBF509E24E6FBCC" box="[621,719,1964,1990]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, 120.4 mm SL (RMCA 2018.008.P.0336). Drawn by N. Vranken.
</paragraph>
</caption>
<paragraph id="8B9936B06138DA67FF6F565C2374FE32" blockId="63.[188,1398,366,568]" pageId="63" pageNumber="64">
It differs from
<taxonomicName id="4C264D336138DA67FEAE565C2430FD82" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[380,537,366,392]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
<emphasis id="B952EAA26138DA67FEAE565C27B8FD8D" box="[380,401,366,391]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B952EAA26138DA67FE74565C2430FD82" box="[422,537,366,392]" italics="true" pageId="63" pageNumber="64">curvidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96138DA67FDF4565C24AEFD82" box="[550,647,366,392]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C264D336138DA67FD19565C256FFD82" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[715,838,366,392]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
<emphasis id="B952EAA26138DA67FD19565C24C9FD8D" box="[715,736,366,391]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B952EAA26138DA67FD27565C256FFD82" box="[757,838,366,392]" italics="true" pageId="63" pageNumber="64">pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96138DA67FC86565C259DFD82" box="[852,948,366,392]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
by a deeper cheek [ChD 24.832.9 (exceptionally
<quantity id="4CDE9B556138DA67FEA056A327E0FDA1" box="[370,457,401,427]" metricMagnitude="-1" metricUnit="m" metricValue="6.0198" pageId="63" pageNumber="64" unit="in" value="23.7">23.7 in</quantity>
<specimenCount id="9D20FD396138DA67FE0556A32456FDA6" box="[471,639,401,428]" pageId="63" pageNumber="64" type="generic">one specimen</specimenCount>
) (mean 27.5) vs 20.824.9 (22.523.2) % HL]; further from
<taxonomicName id="4C264D336138DA67FF6F5687277AFDC4" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[189,339,436,463]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
<emphasis id="B952EAA26138DA67FF6F568726FBFDC4" box="[189,210,437,462]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B952EAA26138DA67FF325686277AFDC4" box="[224,339,436,462]" italics="true" pageId="63" pageNumber="64">curvidens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96138DA67FE8B5687279BFDC5" box="[345,434,437,463]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
by a deeper body [BD 33.541.7 (37.4) vs 29.032.0 (30.8) % SL] and presence vs absence of a well-defined mid-lateral band; further from
<taxonomicName id="4C264D336138DA67FC9D56EA25EDFDFB" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[847,964,471,498]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
<emphasis id="B952EAA26138DA67FC9D56EA254DFDFB" box="[847,868,472,497]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B952EAA26138DA67FCA156E525EDFDFB" box="[883,964,471,497]" italics="true" pageId="63" pageNumber="64">pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96138DA67FC1856EA220AFDF8" box="[970,1059,472,498]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
by a larger number of caudal peduncle scales (CPS 1720, rarely 16 vs 16, rarely 17), a larger adult size (max. 165 vs
<quantity id="4CDE9B556138DA67FB3456C82313FE1F" box="[1254,1338,506,533]" metricMagnitude="-2" metricUnit="m" metricValue="9.6" pageId="63" pageNumber="64" unit="mm" value="96.0">96 mm</quantity>
SL), and colour pattern of small specimens (&lt;
<quantity id="4CDE9B556138DA67FD77552C252FFE32" box="[677,774,542,568]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="63" pageNumber="64" unit="mm" value="100.0">100 mm</quantity>
SL) light coloured vs speckled to uniformly black.
</paragraph>
<paragraph id="8B9936B06138DA67FF6F556D25BAFF23" blockId="63.[189,1400,607,810]" pageId="63" pageNumber="64">
Small specimens (&lt;
<quantity id="4CDE9B556138DA67FE61556D2421FE70" box="[435,520,607,634]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="63" pageNumber="64" unit="mm" value="90.0">90 mm</quantity>
SL) resemble
<taxonomicName id="4C264D336138DA67FD6F55522241FE73" authority="Greenwood, 1973" authorityName="Greenwood" authorityYear="1973" box="[701,1128,607,634]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="schubotziellus">
<emphasis id="B952EAA26138DA67FD6F555224F9FE73" box="[701,720,608,633]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B952EAA26138DA67FD33556D25A0FE73" box="[737,905,607,633]" italics="true" pageId="63" pageNumber="64">schubotziellus</emphasis>
<bibRefCitation id="EFB74B416138DA67FC41556D2241FE73" author="Greenwood P. H." box="[915,1128,607,633]" pageId="63" pageNumber="64" pagination="139 - 242" refId="ref48850" refString="Greenwood P. H. 1973. A revision of the Haplochromis and related species (Pisces: Cichlidae) from Lake George, Uganda. Bulletin of the British Museum (Natural History) Zoology 25: 139 - 242." type="journal article" year="1973">
Greenwood,
<quantity id="4CDE9B556138DA67FBFF556D2241FE73" box="[1069,1128,607,633]" metricMagnitude="1" metricUnit="m" metricValue="5.011419999999999" pageId="63" pageNumber="64" unit="in" value="1973.0">1973</quantity>
</bibRefCitation>
</taxonomicName>
in overall habitus and colour pattern. It differs from the
<typeStatus id="549D88126138DA67FD8655B02492FE96" box="[596,699,642,668]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C264D336138DA67FD3655B12599FE96" authorityName="Greenwood" authorityYear="1973" box="[740,944,642,669]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="schubotziellus">
<emphasis id="B952EAA26138DA67FD3655B124D1FE96" box="[740,760,643,668]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B952EAA26138DA67FCDA55B02599FE96" box="[776,944,642,668]" italics="true" pageId="63" pageNumber="64">schubotziellus</emphasis>
</taxonomicName>
(NHMUK 1972.6.2.351;
<specimenCount id="9D20FD396138DA67FB3455B122D5FE97" box="[1254,1276,643,669]" pageId="63" pageNumber="64" type="female"></specimenCount>
, 75.0 mm SL; Lake George, Kankurunga Island) by a smaller eye [ED 26.231.8 (29.2) vs 33.7% HL], a longer lower jaw [LJL 44.249.6 (47.1) vs 42.6% HL], a deeper cheek [ChD 23.732.9 (27.5) vs 24.0% HL], outer oral teeth with no to a small minor cusp vs a well-defined minor cusp, and inner oral teeth set in 12 weakly defined rows vs 2 well-defined rows in both jaws.
</paragraph>
</subSubSection>
<subSubSection id="C33C653B6138DA67FF6F546322CFFFB6" pageId="63" pageNumber="64" type="etymology">
<paragraph id="8B9936B06138DA67FF6F5463276DFF61" blockId="63.[189,324,849,875]" box="[189,324,849,875]" pageId="63" pageNumber="64">
<heading id="D0D181DC6138DA67FF6F5463276DFF61" bold="true" box="[189,324,849,875]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="B952EAA26138DA67FF6F5463276DFF61" bold="true" box="[189,324,849,875]" pageId="63" pageNumber="64">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06138DA67FF6F544D22CFFFB6" blockId="63.[189,1398,895,957]" pageId="63" pageNumber="64">Specific name from Quasimodo, hunchbacked character in Victor Hugos novel Notre-Dame de Paris (1831); referring to rather shallow head and deep and rhomboid bodies of large specimens.</paragraph>
</subSubSection>
<subSubSection id="C33C653B6138DA67FF6F54D122C8FBD3" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="8B9936B06138DA67FF6F54D12783FFF7" blockId="63.[189,426,995,1021]" box="[189,426,995,1021]" pageId="63" pageNumber="64">
<heading id="D0D181DC6138DA67FF6F54D12783FFF7" bold="true" box="[189,426,995,1021]" fontSize="11" level="3" pageId="63" pageNumber="64" reason="3">
<emphasis id="B952EAA26138DA67FF6F54D12783FFF7" bold="true" box="[189,426,995,1021]" pageId="63" pageNumber="64">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06138DA67FF38532025E5F878" blockId="63.[189,1398,1042,1139]" pageId="63" pageNumber="64">
<materialsCitation id="3B4E3CED6138DA67FF38532025E5F878" ID-GBIF-Occurrence="3861203309" collectingDate="2018-02-01" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="-30" latitude="-0.35880554" location="Lake Edward" longLatPrecision="1" longitude="29.721584" pageId="63" pageNumber="64" specimenCode="RMCA 2018.008.P.0336" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="holotype">
<emphasis id="B952EAA26138DA67FF3853202775F826" bold="true" box="[234,348,1042,1068]" pageId="63" pageNumber="64">
<typeStatus id="549D88126138DA67FF3853202775F826" box="[234,348,1042,1068]" pageId="63" pageNumber="64" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F33176206138DA67FF6F53072713F845" box="[189,314,1077,1103]" name="Uganda" pageId="63" pageNumber="64">UGANDA</collectingCountry>
<specimenCount id="9D20FD396138DA67FE8153042742F85A" box="[339,363,1078,1104]" pageId="63" pageNumber="64" type="male"></specimenCount>
,
<quantity id="4CDE9B556138DA67FEA5530727C7F85A" box="[375,494,1077,1104]" metricMagnitude="-1" metricUnit="m" metricValue="1.204" pageId="63" pageNumber="64" unit="mm" value="120.4">120.4 mm</quantity>
SL;
<collectingRegion id="49E2F8526138DA67FDF4530724EFF845" box="[550,710,1077,1103]" pageId="63" pageNumber="64">Lake Edward</collectingRegion>
;
<geoCoordinate id="EE1250776138DA67FD0353062542F845" box="[721,875,1076,1104]" degrees="0" direction="south" minutes="21" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="31.7" value="-0.35880554">0°2131.7″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FCAA5306220DF85A" box="[888,1060,1076,1104]" degrees="29" direction="east" minutes="43" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="17.7" value="29.721584">29°4317.7″ E</geoCoordinate>
; deep catch, open water ±
<quantity id="4CDE9B556138DA67FA8B530726FDF879" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="30.0">
<elevation id="000BD1836138DA67FA8B530726FDF879" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="30.0">30 m</elevation>
</quantity>
deep;
<date id="FF9810706138DA67FEF0536B2798F879" box="[290,433,1112,1139]" pageId="63" pageNumber="64" value="2018-02-01">
<collectingDate id="EFDCE9986138DA67FEF0536B2798F879" box="[290,433,1112,1139]" pageId="63" pageNumber="64" value="2018-02-01">1 Feb. 2018</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FE6F536B244BF878" box="[445,610,1112,1139]" pageId="63" pageNumber="64">HIPE3 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FD71536B25EEF878" box="[675,967,1112,1139]" pageId="63" pageNumber="64">RMCA 2018.008.P.0336</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9936B06138DA67FF3853A82727F9FB" blockId="63.[189,1399,1178,1521]" pageId="63" pageNumber="64">
<materialsCitation id="3B4E3CED6138DA67FF3853A8225EF914" ID-GBIF-Occurrence="3861203345" collectingDate="1953-06-01" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" latitude="-0.19416668" location="Kiavinionge" longLatPrecision="21" longitude="29.541945" pageId="63" pageNumber="64" specimenCode="IRSNB 13480" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
<emphasis id="B952EAA26138DA67FF3853A8274EF8BE" bold="true" box="[234,359,1178,1204]" pageId="63" pageNumber="64">
<typeStatus id="549D88126138DA67FF3853A8274EF8BE" box="[234,359,1178,1204]" pageId="63" pageNumber="64" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F33176206138DA67FF6F538F24D8F8DD" box="[189,753,1213,1240]" name="Democratic Republic of the Congo" pageId="63" pageNumber="64">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
<specimenCount id="9D20FD396138DA67FCDE538F2510F8D2" box="[780,825,1213,1240]" pageId="63" pageNumber="64" type="female">1 ♀</specimenCount>
,
<quantity id="4CDE9B556138DA67FC9B538F25E8F8D2" box="[841,961,1213,1240]" metricMagnitude="-1" metricUnit="m" metricValue="1.478" pageId="63" pageNumber="64" unit="mm" value="147.8">147.8 mm</quantity>
SL; “Lac Edouard:
<quantity id="4CDE9B556138DA67FB63538F2323F8DD" box="[1201,1290,1213,1239]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="63" pageNumber="64" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">23 km</quantity>
±
<quantity id="4CDE9B556138DA67FAF9538F235EF8D2" box="[1323,1399,1213,1240]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="64" unit="m" value="500.0">500 m</quantity>
au large à lOuest de Kiavinionge” [
<collectingRegion id="49E2F8526138DA67FDA753D22537F8F0" box="[629,798,1248,1274]" pageId="63" pageNumber="64">Lake Edward</collectingRegion>
:
<locationDeviation id="9A2B2F926138DA67FCFE53D3228FF8F1" box="[812,1190,1248,1275]" pageId="63" pageNumber="64">
<quantity id="4CDE9B556138DA67FCFE53D325AEF8F0" box="[812,903,1248,1275]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="63" pageNumber="64" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">23 km</quantity>
±
<quantity id="4CDE9B556138DA67FC7F53D225D3F8F1" box="[941,1018,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="63" pageNumber="64" unit="m" value="500.0">500 m</quantity>
offshore west
</locationDeviation>
of
<location id="8EF9606B6138DA67FB0153D22342F8F1" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FB0153D22342F8F1" box="[1235,1387,1248,1275]" country="Democratic Republic of the Congo" latitude="-0.19416668" longLatPrecision="21" longitude="29.541945" name="Kiavinionge" pageId="63" pageNumber="64" stateProvince="Lake Edward">Kiavinionge</location>
];
<geoCoordinate id="EE1250776138DA67FF6F52302714F914" box="[189,317,1282,1310]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="63" pageNumber="64" precision="15" seconds="39" value="-0.19416668">0°1139″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FE99523027F2F914" box="[331,475,1282,1310]" degrees="29" direction="east" minutes="32" orientation="longitude" pageId="63" pageNumber="64" precision="15" seconds="31" value="29.541945">29°3231″ E</geoCoordinate>
(inferred);
<date id="FF9810706138DA67FDB1523624D9F914" box="[611,752,1283,1310]" pageId="63" pageNumber="64" value="1953-06-01">
<collectingDate id="EFDCE9986138DA67FDB1523624D9F914" box="[611,752,1283,1310]" pageId="63" pageNumber="64" value="1953-06-01">1 Jun. 1953</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FD2E523625A4F914" box="[764,909,1284,1310]" pageId="63" pageNumber="64">KEA exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FC1D5236225EF914" box="[975,1143,1284,1310]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="63" pageNumber="64">IRSNB 13480</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FB435236241AF96E" ID-GBIF-Occurrence="3861203414" collectingDate="1953-06-01" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" location="Kaniatzi" pageId="63" pageNumber="64" specimenCode="IRSNB 13481" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
<specimenCount id="9D20FD396138DA67FB4352362297F914" box="[1169,1214,1284,1310]" pageId="63" pageNumber="64" type="female">1 ♀</specimenCount>
,
<quantity id="4CDE9B556138DA67FB1F5231236CF914" box="[1229,1349,1283,1310]" metricMagnitude="-1" metricUnit="m" metricValue="1.649" pageId="63" pageNumber="64" unit="mm" value="164.9">164.9 mm</quantity>
SL; “Lac Edouard: Kaniatzi (partie N. du lac)” [
<collectingRegion id="49E2F8526138DA67FD615215257EF94B" box="[691,855,1319,1345]" pageId="63" pageNumber="64">Lake Edward</collectingRegion>
:
<location id="8EF9606B6138DA67FCB2521525EAF94B" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FCB2521525EAF94B" box="[864,963,1319,1345]" country="Democratic Republic of the Congo" name="Kaniatzi" pageId="63" pageNumber="64" stateProvince="Lake Edward">Kaniatzi</location>
(northern part of Lake)];
<date id="FF9810706138DA67FB3B5215235BF948" box="[1257,1394,1319,1346]" pageId="63" pageNumber="64" value="1953-06-01">
<collectingDate id="EFDCE9986138DA67FB3B5215235BF948" box="[1257,1394,1319,1346]" pageId="63" pageNumber="64" value="1953-06-01">1 Jun. 1953</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FF6F52782765F96E" box="[189,332,1354,1380]" pageId="63" pageNumber="64">KEA exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FE5F5278241AF96E" box="[397,563,1354,1380]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="63" pageNumber="64">IRSNB 13481</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FD99527827DAF9A1" ID-GBIF-Occurrence="3861203316" collectingDate="1953-06-01" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" latitude="-0.19416668" location="Kiavinionge" longLatPrecision="21" longitude="29.541945" pageId="63" pageNumber="64" specimenCode="IRSNB 13485" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
<specimenCount id="9D20FD396138DA67FD9952782450F96F" box="[587,633,1354,1381]" pageId="63" pageNumber="64" type="male">1 ♂</specimenCount>
,
<quantity id="4CDE9B556138DA67FD57527824D5F96F" box="[645,764,1354,1381]" metricMagnitude="-1" metricUnit="m" metricValue="1.476" pageId="63" pageNumber="64" unit="mm" value="147.6">147.6 mm</quantity>
SL; “Lac Edouard:
<quantity id="4CDE9B556138DA67FC3452782217F96E" box="[998,1086,1354,1380]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="63" pageNumber="64" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">23 km</quantity>
à lOuest de Kiavinionge” [
<collectingRegion id="49E2F8526138DA67FF16525F2742F98D" box="[196,363,1389,1415]" pageId="63" pageNumber="64">Lake Edward</collectingRegion>
:
<locationDeviation id="9A2B2F926138DA67FEAA525F243DF982" box="[376,532,1389,1416]" pageId="63" pageNumber="64">
<quantity id="4CDE9B556138DA67FEAA525F27FAF98D" box="[376,467,1389,1415]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="63" pageNumber="64" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">23 km</quantity>
west
</locationDeviation>
of
<location id="8EF9606B6138DA67FD92525F24F1F982" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FD92525F24F1F982" box="[576,728,1389,1416]" country="Democratic Republic of the Congo" latitude="-0.19416668" longLatPrecision="21" longitude="29.541945" name="Kiavinionge" pageId="63" pageNumber="64" stateProvince="Lake Edward">Kiavinionge</location>
];
<geoCoordinate id="EE1250776138DA67FD3E525E2546F98D" box="[748,879,1388,1416]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="63" pageNumber="64" precision="15" seconds="39" value="-0.19416668">0°1139″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FCAC525E2238F982" box="[894,1041,1388,1416]" degrees="29" direction="east" minutes="32" orientation="longitude" pageId="63" pageNumber="64" precision="15" seconds="31" value="29.541945">29°3231″ E</geoCoordinate>
(inferred);
<date id="FF9810706138DA67FB4F525F2304F982" box="[1181,1325,1389,1416]" pageId="63" pageNumber="64" value="1953-06-01">
<collectingDate id="EFDCE9986138DA67FB4F525F2304F982" box="[1181,1325,1389,1416]" pageId="63" pageNumber="64" value="1953-06-01">1 Jun. 1953</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FAEE525F2722F9A0" pageId="63" pageNumber="64">KEA exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FE9E52A327DAF9A1" box="[332,499,1424,1451]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="63" pageNumber="64">IRSNB 13485</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FDDE52A32723F9FB" ID-GBIF-Occurrence="3861203398" collectingDate="1953-07-02" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" latitude="-0.68072224" location="Vitshumbi" longLatPrecision="1" longitude="29.38961" pageId="63" pageNumber="64" specimenCode="IRSNB 13488" specimenCount="2" specimenCount-female="2" stateProvince="Lake Edward" typeStatus="paratype">
<specimenCount id="9D20FD396138DA67FDDE52A32466F9A1" box="[524,591,1425,1451]" pageId="63" pageNumber="64" type="female">2 ♀♀</specimenCount>
,
<quantity id="4CDE9B556138DA67FD8F52A224FDF9A1" box="[605,724,1424,1451]" metricMagnitude="-1" metricUnit="m" metricValue="1.415" pageId="63" pageNumber="64" unit="mm" value="141.5">141.5 mm</quantity>
SL; “Lac Edouard: Vitshumbi (au Nord)” [
<collectingRegion id="49E2F8526138DA67FB1C52A2235AF9A0" box="[1230,1395,1424,1450]" pageId="63" pageNumber="64">Lake Edward</collectingRegion>
: north of
<location id="8EF9606B6138DA67FEF552812781F9C4" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FEF552812781F9C4" box="[295,424,1459,1486]" country="Democratic Republic of the Congo" latitude="-0.68072224" longLatPrecision="1" longitude="29.38961" name="Vitshumbi" pageId="63" pageNumber="64" stateProvince="Lake Edward">Vitshumbi</location>
];
<geoCoordinate id="EE1250776138DA67FE6C52802471F9C4" box="[446,600,1458,1486]" degrees="0" direction="south" minutes="40" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="50.6" value="-0.68072224">0°4050.6″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FDBA52802539F9C4" box="[616,784,1458,1486]" degrees="29" direction="east" minutes="23" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="22.6" value="29.38961">29°2322.6″ E</geoCoordinate>
(inferred);
<date id="FF9810706138DA67FC4F5286221EF9C4" box="[925,1079,1459,1486]" pageId="63" pageNumber="64" value="1953-07-02">
<collectingDate id="EFDCE9986138DA67FC4F5286221EF9C4" box="[925,1079,1459,1486]" pageId="63" pageNumber="64" value="1953-07-02">02 Jul. 1953</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FB94528622F0F9C4" box="[1094,1241,1460,1486]" pageId="63" pageNumber="64">KEA exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FAF252862723F9FB" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="63" pageNumber="64">IRSNB 13488</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9936B06138DA67FF6F512A22C8FBD3" blockId="63.[189,1399,1560,2009]" pageId="63" pageNumber="64">
<materialsCitation id="3B4E3CED6138DA67FF6F512A27C3FA73" ID-GBIF-Occurrence="3861203381" collectingDate="2016-10-20" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" latitude="-0.162" location="Katwe" longLatPrecision="1" longitude="29.888" pageId="63" pageNumber="64" specimenCode="RMCA 2016.035.P.0208, RMCA 2016.035.P.0209, RMCA 2016.035.P.0210, RMCA 2016.035.P.0211" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Lake Edward">
<collectingCountry id="F33176206138DA67FF6F512A2713FA38" box="[189,314,1560,1586]" name="Uganda" pageId="63" pageNumber="64">UGANDA</collectingCountry>
<emphasis id="B952EAA26138DA67FEB4512A2432FA38" bold="true" box="[358,539,1560,1586]" pageId="63" pageNumber="64">
<collectingRegion id="49E2F8526138DA67FEB4512A2432FA38" box="[358,539,1560,1586]" pageId="63" pageNumber="64">Lake Edward</collectingRegion>
</emphasis>
<specimenCount id="9D20FD396138DA67FD90512A24A4FA39" box="[578,653,1560,1587]" pageId="63" pageNumber="64" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9D20FD396138DA67FD70512A24C4FA39" box="[674,749,1560,1587]" pageId="63" pageNumber="64" type="male">2 ♂♂</specimenCount>
,
<quantity id="4CDE9B556138DA67FCD3512A25E9FA39" box="[769,960,1560,1587]" metricMagnitude="-1" metricUnit="m" metricValue="1.035" metricValueMax="1.143" metricValueMin="0.9269999999999999" pageId="63" pageNumber="64" unit="mm" value="103.5" valueMax="114.3" valueMin="92.7">92.7114.3 mm</quantity>
SL; Katoko breeding ground, soft substrate offshore of
<location id="8EF9606B6138DA67FE6C510E2427FA5C" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FE6C510E2427FA5C" box="[446,526,1596,1622]" country="Uganda" latitude="-0.162" longLatPrecision="1" longitude="29.888" municipality="Katoko" name="Katwe" pageId="63" pageNumber="64" stateProvince="Lake Edward">Katwe</location>
;
<geoCoordinate id="EE1250776138DA67FDCF5108249EFA5C" box="[541,695,1594,1622]" degrees="0" direction="south" minutes="09" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="43.2" value="-0.162">0°0943.2″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FD155108255BFA5C" box="[711,882,1594,1622]" degrees="29" direction="east" minutes="53" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="16.8" value="29.888">29°5316.8″ E</geoCoordinate>
;
<date id="FF9810706138DA67FC50510E220AFA5F" box="[898,1059,1595,1622]" pageId="63" pageNumber="64" value="2016-10-20">
<collectingDate id="EFDCE9986138DA67FC50510E220AFA5F" box="[898,1059,1595,1622]" pageId="63" pageNumber="64" value="2016-10-20">20 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FBE0510E22F3FA5F" box="[1074,1242,1595,1622]" pageId="63" pageNumber="64">HIPE1 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FAF0510E27C3FA73" pageId="63" pageNumber="64">RMCA 2016.035.P.0208 to 0211</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FDDE516D2360FA96" ID-GBIF-Occurrence="3861203415" collectingDate="2016-10-23" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" latitude="-0.192" location="Mweya" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" pageId="63" pageNumber="64" specimenCode="RMCA 2016.035.P.0213, RMCA 2016.035.P.0214" specimenCount="2" specimenCount-female="2" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FDDE516D247AFA73" box="[524,595,1631,1657]" pageId="63" pageNumber="64" type="female">2 ♀♀</specimenCount>
, 83.4,
<quantity id="4CDE9B556138DA67FD7E516D2533FA73" box="[684,794,1631,1657]" metricMagnitude="-2" metricUnit="m" metricValue="9.309999999999999" pageId="63" pageNumber="64" unit="mm" value="93.1">93.1 mm</quantity>
SL;
<collectingMunicipality id="6BFDACCA6138DA67FC89516D220FFA73" box="[859,1062,1631,1657]" pageId="63" pageNumber="64">Mukutu Kihinga</collectingMunicipality>
, rocky offshore of
<location id="8EF9606B6138DA67FAC4516D235BFA73" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FAC4516D235BFA73" box="[1302,1394,1631,1657]" country="Uganda" latitude="-0.192" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" name="Mweya" pageId="63" pageNumber="64" stateProvince="Lake Edward">Mweya</location>
;
<geoCoordinate id="EE1250776138DA67FF6F51B2277FFA96" box="[189,342,1664,1692]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="31.2" value="-0.192">0°1131.2″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FEB451B22438FA96" box="[358,529,1664,1692]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="26.4" value="29.874">29°5226.4″ E</geoCoordinate>
;
<date id="FF9810706138DA67FDF251B024EBFA96" box="[544,706,1666,1692]" pageId="63" pageNumber="64" value="2016-10-23">
<collectingDate id="EFDCE9986138DA67FDF251B024EBFA96" box="[544,706,1666,1692]" pageId="63" pageNumber="64" value="2016-10-23">23 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FD0351B02550FA96" box="[721,889,1666,1692]" pageId="63" pageNumber="64">HIPE1 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FC1251B02360FA96" box="[960,1353,1666,1693]" pageId="63" pageNumber="64">RMCA 2016.035.P.0213 to 0214</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FABA51B02536FAE9" ID-GBIF-Occurrence="3861203382" collectingDate="2016-10-23" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" latitude="-0.192" location="Mweya" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" pageId="63" pageNumber="64" specimenCode="RMCA 2016.035.P.0212" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FABA51B026FCFACA" pageId="63" pageNumber="64" type="male">1 ♂</specimenCount>
,
<quantity id="4CDE9B556138DA67FF375197277BFACA" box="[229,338,1701,1728]" metricMagnitude="-2" metricUnit="m" metricValue="7.89" pageId="63" pageNumber="64" unit="mm" value="78.9">78.9 mm</quantity>
SL;
<collectingMunicipality id="6BFDACCA6138DA67FE4051972475FAB5" box="[402,604,1701,1727]" pageId="63" pageNumber="64">Mukutu Kihinga</collectingMunicipality>
, rocky offshore of
<location id="8EF9606B6138DA67FC9B5194258CFACA" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FC9B5194258CFACA" box="[841,933,1702,1728]" country="Uganda" latitude="-0.192" longLatPrecision="1" longitude="29.874" municipality="Mukutu Kihinga" name="Mweya" pageId="63" pageNumber="64" stateProvince="Lake Edward">Mweya</location>
;
<geoCoordinate id="EE1250776138DA67FC6751912266FAB5" box="[949,1103,1699,1727]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="31.2" value="-0.192">0°1131.2″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FB8D51912325FACA" box="[1119,1292,1699,1728]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="26.4" value="29.874">29°5226.4″ E</geoCoordinate>
;
<date id="FF9810706138DA67FACE519726D2FAE8" pageId="63" pageNumber="64" value="2016-10-23">
<collectingDate id="EFDCE9986138DA67FACE519726D2FAE8" pageId="63" pageNumber="64" value="2016-10-23">23 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FED951FA279AFAE8" box="[267,435,1736,1762]" pageId="63" pageNumber="64">HIPE1 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FE2951FA2536FAE9" box="[507,799,1736,1763]" pageId="63" pageNumber="64">RMCA 2016.035.P.0212</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FCED51FA2730FB23" ID-GBIF-Occurrence="3861203327" collectingDate="2017-03-26" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" latitude="-0.40158334" location="Rwenshama" longLatPrecision="1" longitude="29.776417" pageId="63" pageNumber="64" specimenCode="RMCA 2017.006.P.0349, RMCA 2017.006.P.0350, RMCA 2017.006.P.0351" specimenCount="3" specimenCount-female="3" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FCED51FA25AFFAE9" box="[831,902,1736,1763]" pageId="63" pageNumber="64" type="female">3 ♀♀</specimenCount>
,
<quantity id="4CDE9B556138DA67FC4551FA227AFAE9" box="[919,1107,1736,1763]" metricMagnitude="-2" metricUnit="m" metricValue="9.475" metricValueMax="11.01" metricValueMin="7.94" pageId="63" pageNumber="64" unit="mm" value="94.75" valueMax="110.1" valueMin="79.4">79.4110.1 mm</quantity>
SL;
<location id="8EF9606B6138DA67FB4051FA230DFAE8" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FB4051FA230DFAE8" box="[1170,1316,1736,1762]" country="Uganda" latitude="-0.40158334" longLatPrecision="1" longitude="29.776417" name="Rwenshama" pageId="63" pageNumber="64" stateProvince="Lake Edward">Rwenshama</location>
, rocky shore;
<geoCoordinate id="EE1250776138DA67FEC651D8279AFB0C" box="[276,435,1770,1798]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="05.7" value="-0.40158334">0°2405.7″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FE1551D82451FB0C" box="[455,632,1770,1798]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="35.1" value="29.776417">29°4635.1″ E</geoCoordinate>
;
<date id="FF9810706138DA67FD5E51D92515FB0C" box="[652,828,1771,1798]" pageId="63" pageNumber="64" value="2017-03-26">
<collectingDate id="EFDCE9986138DA67FD5E51D92515FB0C" box="[652,828,1771,1798]" pageId="63" pageNumber="64" value="2017-03-26">26 Mar. 2017</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FC8251DE25D4FB0F" box="[848,1021,1771,1798]" pageId="63" pageNumber="64">HIPE2 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FB9C51DE2730FB23" pageId="63" pageNumber="64">RMCA 2017.006.P.0349 to 0351</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FEE5503D2459FB46" ID-GBIF-Occurrence="3861203348" collectingDate="2018-01-18" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.16802779" location="Katwe" longLatPrecision="1" longitude="29.874277" pageId="63" pageNumber="64" specimenCode="RMCA 2018.008.P.0334" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FEE5503D274EFB23" box="[311,359,1807,1833]" pageId="63" pageNumber="64" type="male">1 ♂</specimenCount>
,
<quantity id="4CDE9B556138DA67FEA5503D27CBFB23" box="[375,482,1807,1833]" metricMagnitude="-2" metricUnit="m" metricValue="8.91" pageId="63" pageNumber="64" unit="mm" value="89.1">89.1 mm</quantity>
SL; islands near
<location id="8EF9606B6138DA67FD6A503D2521FB23" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FD6A503D2521FB23" box="[696,776,1807,1833]" country="Uganda" latitude="-0.16802779" longLatPrecision="1" longitude="29.874277" name="Katwe" pageId="63" pageNumber="64" stateProvince="Lake Edward">Katwe</location>
;
<geoCoordinate id="EE1250776138DA67FCC4503F2599FB23" box="[790,944,1805,1833]" degrees="0" direction="south" minutes="10" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="04.9" value="-0.16802779">0°1004.9″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FC12503F2242FB23" box="[960,1131,1805,1833]" degrees="29" direction="east" minutes="52" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="27.4" value="29.874277">29°5227.4″ E</geoCoordinate>
;
<date id="FF9810706138DA67FBA8503D2331FB23" box="[1146,1304,1807,1833]" pageId="63" pageNumber="64" value="2018-01-18">
<collectingDate id="EFDCE9986138DA67FBA8503D2331FB23" box="[1146,1304,1807,1833]" pageId="63" pageNumber="64" value="2018-01-18">18 Jan. 2018</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FAF5503D2722FB46" pageId="63" pageNumber="64">HIPE3 exped.</collectorName>
leg.;
<specimenCode id="DB809ECB6138DA67FE9C50002459FB46" box="[334,624,1842,1868]" pageId="63" pageNumber="64">RMCA 2018.008.P.0334</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FD595000225BFB7A" ID-GBIF-Occurrence="3861203394" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" latitude="-0.40444446" location="Rwenshama" longLatPrecision="1" longitude="29.773556" pageId="63" pageNumber="64" specimenCode="RMCA 2018.008.P.0335" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FD5950002491FB46" box="[651,696,1842,1868]" pageId="63" pageNumber="64" type="female">1 ♀</specimenCount>
,
<quantity id="4CDE9B556138DA67FD1550002569FB46" box="[711,832,1842,1868]" metricMagnitude="-1" metricUnit="m" metricValue="1.307" pageId="63" pageNumber="64" unit="mm" value="130.7">130.7 mm</quantity>
SL;
<geoCoordinate id="EE1250776138DA67FCA85002223BFB46" box="[890,1042,1840,1868]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="16.0" value="-0.40444446">0°2416.0″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FBF2500222E3FB46" box="[1056,1226,1840,1868]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="24.8" value="29.773556">29°4624.8″ E</geoCoordinate>
;
<date id="FF9810706138DA67FB055000235BFB46" box="[1239,1394,1842,1868]" pageId="63" pageNumber="64" value="2018-01-24">
<collectingDate id="EFDCE9986138DA67FB055000235BFB46" box="[1239,1394,1842,1868]" pageId="63" pageNumber="64" value="2018-01-24">24 Jan. 2018</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FF6F5067274AFB65" box="[189,355,1877,1903]" pageId="63" pageNumber="64">HIPE3 exped.</collectorName>
leg.; bought at
<location id="8EF9606B6138DA67FDCD50672487FB65" LSID="urn:lsid:plazi:treatment:038F87A6613ADA1BFD9951FF256AFE20:8EF9606B6138DA67FDCD50672487FB65" box="[543,686,1877,1903]" country="Uganda" latitude="-0.40444446" longLatPrecision="1" longitude="29.773556" name="Rwenshama" pageId="63" pageNumber="64" stateProvince="Lake Edward">Rwenshama</location>
landing site;
<specimenCode id="DB809ECB6138DA67FC835067225BFB7A" box="[849,1138,1877,1904]" pageId="63" pageNumber="64">RMCA 2018.008.P.0335</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FB5C506727F6FBBC" ID-GBIF-Occurrence="3861203337" collectingDate="2018-02-01" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="30" latitude="-0.35880554" longLatPrecision="1" longitude="29.721584" pageId="63" pageNumber="64" specimenCode="RMCA 2018.008.P.0339" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FB5C50672292FB7A" box="[1166,1211,1877,1904]" pageId="63" pageNumber="64" type="female">1 ♀</specimenCount>
,
<quantity id="4CDE9B556138DA67FB195067236DFB7A" box="[1227,1348,1877,1904]" metricMagnitude="-1" metricUnit="m" metricValue="1.488" pageId="63" pageNumber="64" unit="mm" value="148.8">148.8 mm</quantity>
SL;
<geoCoordinate id="EE1250776138DA67FF6F5045277EFB98" box="[189,343,1911,1939]" degrees="0" direction="south" minutes="21" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="31.7" value="-0.35880554">0°2131.7″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FEB55045243BFB99" box="[359,530,1911,1939]" degrees="29" direction="east" minutes="43" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="17.7" value="29.721584">29°4317.7″ E</geoCoordinate>
;
<date id="FF9810706138DA67FDF0504A249FFB98" box="[546,694,1912,1938]" pageId="63" pageNumber="64" value="2018-02-01">
<collectingDate id="EFDCE9986138DA67FDF0504A249FFB98" box="[546,694,1912,1938]" pageId="63" pageNumber="64" value="2018-02-01">1 Feb. 2018</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FD17504A2544FB98" box="[709,877,1912,1938]" pageId="63" pageNumber="64">HIPE3 exped.</collectorName>
leg.; deep catch, open water ±
<quantity id="4CDE9B556138DA67FB3C504A2304FB99" box="[1262,1325,1912,1939]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="30.0">
<elevation id="000BD1836138DA67FB3C504A2304FB99" box="[1262,1325,1912,1939]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="30.0">30 m</elevation>
</quantity>
deep;
<specimenCode id="DB809ECB6138DA67FF6F50AE27F6FBBC" box="[189,479,1948,1974]" pageId="63" pageNumber="64">RMCA 2018.008.P.0339</specimenCode>
</materialsCitation>
<materialsCitation id="3B4E3CED6138DA67FE2E50AE22F4FBD3" ID-GBIF-Occurrence="3861203391" collectingDate="2018-02-01" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="30" latitude="-0.35880554" longLatPrecision="1" longitude="29.721584" pageId="63" pageNumber="64" specimenCode="RMCA 2018.008.P.0337, RMCA 2018.008.P.0338" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward">
<specimenCount id="9D20FD396138DA67FE2E50AE2403FBBC" box="[508,554,1948,1974]" pageId="63" pageNumber="64" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D20FD396138DA67FDE850AE2443FBBC" box="[570,618,1948,1974]" pageId="63" pageNumber="64" type="male">1 ♂</specimenCount>
, 117.9,
<quantity id="4CDE9B556138DA67FD1850A9256AFBBC" box="[714,835,1947,1974]" metricMagnitude="-1" metricUnit="m" metricValue="1.262" pageId="63" pageNumber="64" unit="mm" value="126.2">126.2 mm</quantity>
SL;
<geoCoordinate id="EE1250776138DA67FCAD50A82231FBBC" box="[895,1048,1946,1974]" degrees="0" direction="south" minutes="21" orientation="latitude" pageId="63" pageNumber="64" precision="1" seconds="31.7" value="-0.35880554">0°2131.7″ S</geoCoordinate>
,
<geoCoordinate id="EE1250776138DA67FBF550A822F8FBBC" box="[1063,1233,1946,1974]" degrees="29" direction="east" minutes="43" orientation="longitude" pageId="63" pageNumber="64" precision="1" seconds="17.7" value="29.721584">29°4317.7″ E</geoCoordinate>
;
<date id="FF9810706138DA67FB0D50AE235BFBBC" box="[1247,1394,1947,1974]" pageId="63" pageNumber="64" value="2018-02-01">
<collectingDate id="EFDCE9986138DA67FB0D50AE235BFBBC" box="[1247,1394,1947,1974]" pageId="63" pageNumber="64" value="2018-02-01">1 Feb. 2018</collectingDate>
</date>
;
<collectorName id="26D353666138DA67FF6F508D274BFBD3" box="[189,354,1983,2009]" pageId="63" pageNumber="64">HIPE3 exped.</collectorName>
leg.; deep catch, open water ±
<quantity id="4CDE9B556138DA67FD1D508D2522FBD3" box="[719,779,1983,2009]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="30.0">
<elevation id="000BD1836138DA67FD1D508D2522FBD3" box="[719,779,1983,2009]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="63" pageNumber="64" unit="m" value="30.0">30 m</elevation>
</quantity>
deep;
<specimenCode id="DB809ECB6138DA67FC8B508D22F4FBD3" box="[857,1245,1983,2009]" pageId="63" pageNumber="64">RMCA 2018.008.P.0337 to 0338</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5966386147DA18FF6F50562212FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484242" ID-Zenodo-Dep="6484242" httpUri="https://zenodo.org/record/6484242/files/figure.png" pageId="64" pageNumber="65" startId="64.[189,232,1892,1918]" targetBox="[288,1293,287,1843]" targetPageId="64">
<paragraph id="8B9936B06147DA18FF6F50562212FBE2" blockId="64.[185,1399,1892,2025]" pageId="64" pageNumber="65">
<emphasis id="B952EAA26147DA18FF6F5056273EFB75" bold="true" box="[189,279,1892,1919]" pageId="64" pageNumber="65">Fig. 36.</emphasis>
<taxonomicName id="4C264D336147DA18FEC850572460FB75" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[282,585,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="64" pageNumber="65" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA26147DA18FEC850572460FB75" box="[282,585,1893,1919]" italics="true" pageId="64" pageNumber="65">Haplochromis quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96147DA18FD9B50572489FB75" box="[585,672,1893,1919]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B952EAA26147DA18FD755057249FFB75" bold="true" box="[679,694,1893,1919]" pageId="64" pageNumber="65">a</emphasis>
. Photograph of preserved holotype (RMCA 2018.008.P.0336; 120.4 mm SL).
<emphasis id="B952EAA26147DA18FEA150BA27AAFBA8" bold="true" box="[371,387,1928,1954]" pageId="64" pageNumber="65">b</emphasis>
. X-ray image of holotype.
<emphasis id="B952EAA26147DA18FD6C50BA24C2FBA8" bold="true" box="[702,747,1928,1954]" pageId="64" pageNumber="65">cd</emphasis>
. Photographs of freshly caught specimens.
<emphasis id="B952EAA26147DA18FB3750BA22DBFBA8" bold="true" box="[1253,1266,1928,1954]" pageId="64" pageNumber="65">c</emphasis>
. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL).
<emphasis id="B952EAA26147DA18FCC950992502FBCF" bold="true" box="[795,811,1963,1989]" pageId="64" pageNumber="65">d</emphasis>
. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
</paragraph>
</caption>
<subSubSection id="C33C653B6146DA1BFF6F563A24C3FD9A" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66" type="description">
<paragraph id="8B9936B06146DA19FF6F563A2767FD28" blockId="65.[189,334,264,290]" box="[189,334,264,290]" pageId="65" pageNumber="66">
<heading id="D0D181DC6146DA19FF6F563A2767FD28" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B952EAA26146DA19FF6F563A2767FD28" bold="true" box="[189,334,264,290]" pageId="65" pageNumber="66">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06146DA19FF6F5605227FFEDF" blockId="65.[188,1400,311,725]" pageId="65" pageNumber="66">
Based on
<specimenCount id="9D20FD396146DA19FEEB560527F5FD5B" box="[313,476,311,337]" pageId="65" pageNumber="66" type="generic">21 specimens</specimenCount>
(
<quantity id="4CDE9B556146DA19FE2256052499FD58" box="[496,688,311,338]" metricMagnitude="-1" metricUnit="m" metricValue="1.219" metricValueMax="1.649" metricValueMin="0.789" pageId="65" pageNumber="66" unit="mm" value="121.9" valueMax="164.9" valueMin="78.9">78.9164.9 mm</quantity>
SL); body average in depth in comparison to generalised
<taxonomicName id="4C264D336146DA19FF6F56692712FD7E" authorityName="Trewavas" authorityYear="1933" box="[189,315,346,373]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="elegans">
<emphasis id="B952EAA26146DA19FF6F566926FBFD7E" box="[189,210,347,372]" italics="true" pageId="65" pageNumber="66">H</emphasis>
.
<emphasis id="B952EAA26146DA19FF3356682712FD7E" box="[225,315,346,372]" italics="true" pageId="65" pageNumber="66">elegans</emphasis>
</taxonomicName>
(but deep for a piscivorous species;
<tableCitation id="C6A4030B6146DA19FD3556682515FD7E" box="[743,828,346,372]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="65" pageNumber="66" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
) and rhomboid (
<figureCitation id="131D2A356146DA19FBD056682273FD7F" box="[1026,1114,346,373]" captionStart="Fig" captionStartId="62.[189,232,1963,1989]" captionTargetBox="[189,1398,1159,1876]" captionTargetId="figure-358@62.[189,1399,1159,1876]" captionTargetPageId="62" captionText="Fig. 35. Haplochromis quasimodo sp. nov., holotype, ♂, 120.4 mm SL (RMCA 2018.008.P.0336). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484240" httpUri="https://zenodo.org/record/6484240/files/figure.png" pageId="65" pageNumber="66">Fig. 35</figureCitation>
). Head average in width and with a concave to weakly convex dorsal outline; eye small; interorbital area very narrow; cheek and lacrimal deep; lacrimal somewhat convex and hereby protrudes somewhat laterally. Snout average in length, acute, and slopes gently at 3040°; premaxillary pedicel long and prominent. Jaws isognathous, long, slim, narrow, and rounded in dorsal view; gape large and slopes gently at 2035°; maxilla extends to between verticals through anterior margins of orbit and pupil. Lower jaw shallow and with a straight ventral outline in lateral view, mental prominence absent or weakly developed, and lower jaw side nearly flat with an inclination of 1525° to horizontal in anterior view. Upper jaw not expanded. Lips and oral mucosa thin. Neurocranium average in depth, ethmo-vomerine block decurved to horizontally inclined, preorbital region shallow (2124% NL), orbital region average in depth (3032% NL), and supraoccipital crest deep and pyramidical or weakly wedge-shaped (
<figureCitation id="131D2A356146DA19FC3055882260FEDF" box="[994,1097,698,725]" captionStart="Fig" captionStartId="64.[189,232,1892,1918]" captionTargetBox="[288,1293,287,1843]" captionTargetId="figure-18@64.[288,1293,265,1843]" captionTargetPageId="64" captionText="Fig. 36.Haplochromis quasimodosp. nov. a. Photograph of preservedholotype (RMCA 2018.008.P.0336; 120.4 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL). d. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484242" httpUri="https://zenodo.org/record/6484242/files/figure.png" pageId="65" pageNumber="66">Fig. 36b</figureCitation>
).
</paragraph>
<paragraph id="8B9936B06146DA19FF6F55CE244FF825" blockId="65.[189,1400,764,1071]" pageId="65" pageNumber="66">
Outer oral teeth numerous and very small. Necks stout, conical, and straight; crowns weakly recurved, unicuspid in large specimens (&gt;
<quantity id="4CDE9B556146DA19FD94542D24B2FF33" box="[582,667,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="66" unit="mm" value="80.0">80 mm</quantity>
SL), anteriorly unicuspid and posteriorly mostly bi-, weakly bi-, and weakly tricuspid in small specimens (&lt;
<quantity id="4CDE9B556146DA19FCF054702552FF57" box="[802,891,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="66" unit="mm" value="80.0">80 mm</quantity>
SL), all acutely pointed. Dental arcades rounded. Outer teeth closely and regularly set with neck-distances of ½1 neck-width. In upper jaw, 13 posteriormost teeth slightly enlarged. Inner teeth small, straight, an admixture of acutely pointed unicuspids and weakly tricuspids. Tooth bands very slender crescent-shaped with 12 rows of inner teeth, and narrow posteriorly until only outer row remains past ⅔ length of tooth band. Inner teeth closely and regularly set on 13/2 outer neck-widths from outer row; implantation mostly recumbent; size uniform throughout tooth band.
</paragraph>
<paragraph id="8B9936B06146DA19FF6F5365242FF92B" blockId="65.[189,1401,1111,1313]" pageId="65" pageNumber="66">
Lower pharyngeal bone average in length, triangular, slim, and shallow over whole length (
<figureCitation id="131D2A356146DA19FADD53652340F87B" box="[1295,1385,1111,1137]" captionStart="Fig" captionStartId="66.[190,233,1963,1989]" captionTargetBox="[189,1397,970,1866]" captionTargetId="figure-223@66.[189,1399,970,1868]" captionTargetPageId="66" captionText="Fig. 37. Haplochromis quasimodo sp. nov. (RMCA 2018.008.P.0337; 126.2 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484244" httpUri="https://zenodo.org/record/6484244/files/figure.png" pageId="65" pageNumber="66">Fig. 37</figureCitation>
). Pharyngeal teeth relatively large and slender; major cusps acutely pointed; cusp gaps nearly straight; minor cusps and cusp protuberances mostly absent. Teeth in two median longitudinal rows equal in size and form to lateral teeth,
<quantity id="4CDE9B556146DA19FDCC53F224ACF8D0" box="[542,645,1216,1242]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="65" pageNumber="66" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">1113 in</quantity>
each row. Posterior transverse row with 2021 teeth, implanted erectly with a lateral inclination; major cusps nearly straight, bluntly pointed, and laterally compressed; minor cusps mostly present.
</paragraph>
<paragraph id="8B9936B06146DA19FF6F527A235EF968" blockId="65.[189,1399,1352,1379]" box="[189,1399,1352,1379]" pageId="65" pageNumber="66">Chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin.</paragraph>
<paragraph id="8B9936B06146DA19FF6F52BB2561FA90" blockId="65.[189,1399,1417,1691]" pageId="65" pageNumber="66">
Caudal fin emarginate; dorsal and anal fins reach to between verticals through caudal-fin base and two scales posterior to this vertical. In about half of all specimens (10 of
<specimenCount id="9D20FD396146DA19FC34529F2217F9CD" box="[998,1086,1453,1479]" pageId="65" pageNumber="66" type="generic" typeStatus="type">21 type</specimenCount>
specimens), some minute, ellipsoid scales present on basal part of membrane of anal fin; between some pairs of fin rays, up to two rows of 15 scales extend from body onto fin; scales very variable in distribution and invisible to naked eye (
<figureCitation id="131D2A356146DA19FF2651242762FA3B" box="[244,331,1558,1585]" captionStart="Fig" captionStartId="62.[189,232,1963,1989]" captionTargetBox="[189,1398,1159,1876]" captionTargetId="figure-358@62.[189,1399,1159,1876]" captionTargetPageId="62" captionText="Fig. 35. Haplochromis quasimodo sp. nov., holotype, ♂, 120.4 mm SL (RMCA 2018.008.P.0336). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484240" httpUri="https://zenodo.org/record/6484240/files/figure.png" pageId="65" pageNumber="66">Fig. 35</figureCitation>
). Dorsal fin rarely with a few isolated rows of 14 minute scales (in 3 of
<specimenCount id="9D20FD396146DA19FB5D512422CFFA3B" box="[1167,1254,1558,1585]" pageId="65" pageNumber="66" type="generic" typeStatus="type">21 type</specimenCount>
specimens). Pectoral fin long and reaches to between first anal-fin spine and second anal fin branched ray; pelvic fin reaches to between first and second anal-fin spine in females, to second anal fin branched ray in males; first branched pelvic-fin ray elongated in all specimens.
</paragraph>
<paragraph id="8B9936B06146DA19FF6F51F3243DFAF5" blockId="65.[189,1398,1729,1791]" pageId="65" pageNumber="66">All gill rakers in outer row of first gill arch short, relatively stout, and mostly simple, but sometimes anvil-shaped or weakly bifid.</paragraph>
<paragraph id="8B9936B06146DA19FF6F5014278AFB4A" blockId="65.[189,419,1830,1856]" box="[189,419,1830,1856]" pageId="65" pageNumber="66">
<heading id="D0D181DC6146DA19FF6F5014278AFB4A" bold="true" box="[189,419,1830,1856]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B952EAA26146DA19FF6F5014278AFB4A" bold="true" box="[189,419,1830,1856]" pageId="65" pageNumber="66">Colouration in life</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06146DA1AFF6F506627E8FD87" blockId="65.[189,1400,1876,2009]" lastBlockId="66.[189,1399,265,397]" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
Dominant males: body and dorsal part of head light grey with faint yellow sheen; ventral half of body blue-black; belly and chest black; cheek, lower jaw, and lips white; snout dusky; eye with (dark) grey outer ring and silver inner ring (
<figureCitation id="131D2A356146DA19FD9150A92482FBBC" box="[579,683,1947,1974]" captionStart="Fig" captionStartId="64.[189,232,1892,1918]" captionTargetBox="[288,1293,287,1843]" captionTargetId="figure-18@64.[288,1293,265,1843]" captionTargetPageId="64" captionText="Fig. 36.Haplochromis quasimodosp. nov. a. Photograph of preservedholotype (RMCA 2018.008.P.0336; 120.4 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL). d. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484242" httpUri="https://zenodo.org/record/6484242/files/figure.png" pageId="65" pageNumber="66">Fig. 36c</figureCitation>
). Flank with a well-defined mid-lateral band, an interrupted dorsal-lateral band, and 56 vertical stripes; lacrimal and vertical preopercular stripes well-defined; posterior margin of operculum black. Pectoral fin hyaline; pelvic fin black; dorsal fin dusky and with black lappets, base, and posterior part; anal fin black and with 27 very large egg spots (i.e., three times distance between rays) with dusky rings; caudal fin dusky and with black base, hyaline distal part, and maculated dorsal part.
</paragraph>
<paragraph id="8B9936B06145DA1AFF6F568622C8FE51" blockId="66.[188,1399,436,603]" pageId="66" pageNumber="67">
Females and juveniles: body and head uniformly white, except for light grey dorsum and dorsal part of head and a dusky snout; eye with (dark) grey outer ring and silver inner ring (
<figureCitation id="131D2A356145DA1AFB9056E52280FDF8" box="[1090,1193,471,498]" captionStart="Fig" captionStartId="64.[189,232,1892,1918]" captionTargetBox="[288,1293,287,1843]" captionTargetId="figure-18@64.[288,1293,265,1843]" captionTargetPageId="64" captionText="Fig. 36.Haplochromis quasimodosp. nov. a. Photograph of preservedholotype (RMCA 2018.008.P.0336; 120.4 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL). d. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484242" httpUri="https://zenodo.org/record/6484242/files/figure.png" pageId="66" pageNumber="67">Fig. 36d</figureCitation>
). Flank with faint to well-defined mid-lateral and interrupted dorsal-lateral bands. Pectoral and dorsal fins hyaline; dorsal fin with black lappets and dusky base and distal part; anal and caudal fins yellow; anal fin with 35 spots resembling egg spots; caudal fin with a dusky base and a dusky and maculated dorsal part.
</paragraph>
<paragraph id="8B9936B06145DA1AFF6F55B027F9FE96" blockId="66.[189,464,642,668]" box="[189,464,642,668]" pageId="66" pageNumber="67">
<heading id="D0D181DC6145DA1AFF6F55B027F9FE96" bold="true" box="[189,464,642,668]" fontSize="11" level="3" pageId="66" pageNumber="67" reason="3">
<emphasis id="B952EAA26145DA1AFF6F55B027F9FE96" bold="true" box="[189,464,642,668]" pageId="66" pageNumber="67">Preserved colouration</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06145DA1BFF6F558324C3FD9A" blockId="66.[189,1400,689,821]" lastBlockId="67.[189,1399,265,400]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
Body and operculum with yellowish dorsal part, white ventral part; transition gradual; in dominant males, ventral part of body overlain black (
<figureCitation id="131D2A356145DA1AFD1C55E6251EFEE5" box="[718,823,724,751]" captionStart="Fig" captionStartId="64.[189,232,1892,1918]" captionTargetBox="[288,1293,287,1843]" captionTargetId="figure-18@64.[288,1293,265,1843]" captionTargetPageId="64" captionText="Fig. 36.Haplochromis quasimodosp. nov. a. Photograph of preservedholotype (RMCA 2018.008.P.0336; 120.4 mm SL). b. X-ray image of holotype. cd. Photographs of freshly caught specimens. c. Dominant male (RMCA 2018.008.P(HP3072); 123.7 mm SL). d. Female (RMCA 2018.008.P(HP3064); 116.6 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484242" httpUri="https://zenodo.org/record/6484242/files/figure.png" pageId="66" pageNumber="67">Fig. 36a</figureCitation>
). Flank with mostly mid-lateral and interrupted dorsal-lateral bands; in dominant males, with 56 vertical stripes. Cheek yellowish, snout dusky, and lower jaw whitish. Nostril and interorbital stripes and nape band faint in all specimens; lacrimal stripe faint in females, broad and well-defined in males; vertical preopercular stripes well-defined in males. Pectoral fin hyaline; pelvic and anal fins yellowish in females and black in males; anal fin with 27 large egg-spots in males. Caudal and dorsal fins dusky and dorsal fin with black lappets; in males, base of caudal fin and posterior part of dorsal fin black.
</paragraph>
</subSubSection>
<caption id="DF5966386145DA1AFF6C509925D5FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484244" ID-Zenodo-Dep="6484244" httpUri="https://zenodo.org/record/6484244/files/figure.png" pageId="66" pageNumber="67" startId="66.[190,233,1963,1989]" targetBox="[189,1397,970,1866]" targetPageId="66">
<paragraph id="8B9936B06145DA1AFF6C509925D5FBE2" blockId="66.[189,1398,1963,2025]" pageId="66" pageNumber="67">
<emphasis id="B952EAA26145DA1AFF6C50992733FBCF" bold="true" box="[190,282,1963,1989]" pageId="66" pageNumber="67">Fig. 37.</emphasis>
<taxonomicName id="4C264D336145DA1AFEF35099247AFBCF" authority="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks, 2022" authorityName="Vranken &amp; Steenberge &amp; Heylen &amp; Decru &amp; Snoeks" authorityYear="2022" box="[289,595,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
<emphasis id="B952EAA26145DA1AFEF35099247AFBCF" box="[289,595,1963,1989]" italics="true" pageId="66" pageNumber="67">Haplochromis quasimodo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26157D96145DA1AFD88509E249DFBCC" box="[602,692,1964,1990]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
(RMCA 2018.008.P.0337; 126.2 mm SL).
<emphasis id="B952EAA26145DA1AFB7950992293FBCF" bold="true" box="[1195,1210,1963,1989]" pageId="66" pageNumber="67">a</emphasis>
. Dorsal view of the lower pharyngeal jaw.
<emphasis id="B952EAA26145DA1AFE2150FC242AFBE2" bold="true" box="[499,515,1998,2024]" pageId="66" pageNumber="67">b</emphasis>
. Lateral view of the lower pharyngeal jaw.
</paragraph>
</caption>
<subSubSection id="C33C653B6144DA1BFF6F568A256AFE20" pageId="67" pageNumber="68" type="distribution">
<paragraph id="8B9936B06144DA1BFF6F568A27D8FDD8" blockId="67.[189,497,440,466]" box="[189,497,440,466]" pageId="67" pageNumber="68">
<heading id="D0D181DC6144DA1BFF6F568A27D8FDD8" bold="true" box="[189,497,440,466]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="B952EAA26144DA1BFF6F568A27D8FDD8" bold="true" box="[189,497,440,466]" pageId="67" pageNumber="68">Distribution and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B06144DA1BFF6F56DA256AFE20" blockId="67.[188,1398,488,554]" pageId="67" pageNumber="68">Endemic to Lake Edward system; found in offshore, benthic areas in shallow and deep waters. Based on its morphology, most probably a piscivorous species.</paragraph>
</subSubSection>
</treatment>
</document>