551 lines
73 KiB
XML
551 lines
73 KiB
XML
<document id="DF0A8FF5CEB9D5A7F529719FF5AF8A0F" ID-CLB-Dataset="92619" ID-DOI="10.5852/ejt.2022.815.1749" ID-GBIF-Dataset="9a798cae-f10e-44c2-a65c-cdadbff28cb4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484153" ID-ZooBank="6AD0082E-7349-48DE-AFCA-1EE0BFBB3887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650913068508" checkinUser="felipe" docAuthor="Vranken, Nathan, Steenberge, Maarten Van, Heylen, Annelies, Decru, Eva & Snoeks, Jos" docDate="2022" docId="038F87A66126DA7EFDB2568F27FAFE3E" docLanguage="en" docName="ejt-815_1-94.pdf" docOrigin="European Journal of Taxonomy 815" docSource="http://dx.doi.org/10.5852/ejt.2022.815.1749" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplochromis glaucus Vranken & Steenberge & Heylen & Decru & Snoeks 2022, sp. nov." docType="treatment" docUuid="E5948349-E3F9-49A1-81CD-1C7D332E2F00" docUuidSource="ZooBank" docVersion="10" lastPageNumber="39" masterDocId="FFB6FFDE6107DA58FFD257322629FC0A" masterDocTitle="From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system" masterLastPageNumber="94" masterPageNumber="1" pageNumber="34" updateTime="1699329164302" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8A849C34D472382AB2CAA84540F87055">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="06113A38D673A40C4724BFD5668EC49F">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="69388C3DCC93874EAD39FC54AC4EF082">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. & Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="637F4427034CAAB83EFE1810FAA9E255" type="email">nathan.vranken@kuleuven.be</mods:nameIdentifier>
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<mods:namePart id="91B58A2FCBE26CBBAC07CC58AFAFD5F2">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="20156C588B55D9F0E5D1777C0741CEF3">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. & Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:nameIdentifier id="D0F5F0348D19E3BEB36F44E1F2E844CC" type="email">mvansteenberge@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="2D197A6BE995FB692DF95568CA28AFBE">Heylen, Annelies</mods:namePart>
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<mods:affiliation id="27159D8ECFA2E69C724181CCCCEEEA6F">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="1EA53730B8CCF9A853BA18E6D2176166" type="email">annelies.heylen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="8E293E6397E6D4BA618FBB69C4038E89">Decru, Eva</mods:namePart>
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<mods:affiliation id="0837703E7E2772322E602582661597EB">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="1814ADF5849E515834D49DFDBF448CAE" type="email">eva.decru.icht@gmail.com</mods:nameIdentifier>
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<mods:affiliation id="9082A8AD171431C3AF84E11EB43DC7C5">Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="51026EA20E6E960C6C8A4C8B83A8BCF6" type="email">jos.snoeks@africamuseum.be</mods:nameIdentifier>
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<mods:title id="B3A58707BEF0F08C533CEA62D00CBBFF">European Journal of Taxonomy</mods:title>
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<treatment id="038F87A66126DA7EFDB2568F27FAFE3E" ID-DOI="http://doi.org/10.5281/zenodo.6502611" ID-GBIF-Taxon="197197502" ID-Zenodo-Dep="6502611" LSID="urn:lsid:zoobank.org:act:E5948349-E3F9-49A1-81CD-1C7D332E2F00" httpUri="http://treatment.plazi.org/id/038F87A66126DA7EFDB2568F27FAFE3E" lastPageId="38" lastPageNumber="39" pageId="33" pageNumber="34">
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<taxonomicName id="4C264D336126DA79FDB2568F255AFDD2" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[608,883,445,472]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
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<emphasis id="B952EAA26126DA79FDB2568F255AFDD2" bold="true" box="[608,883,445,472]" italics="true" pageId="33" pageNumber="34">Haplochromis glaucus</emphasis>
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<taxonomicNameLabel id="A26157D96126DA79FCA8568C25FDFDD2" box="[890,980,446,472]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C33C653B6126DA79FEA656D622E9FDF5" box="[372,1216,484,511]" pageId="33" pageNumber="34" type="multiple">
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<subSubSection id="C33C653B6126DA79FD5A55382585FE2E" box="[648,940,522,548]" pageId="33" pageNumber="34" type="description">
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<paragraph id="8B9936B06126DA79FD5A55382585FE2E" blockId="33.[372,1216,445,548]" box="[648,940,522,548]" pageId="33" pageNumber="34">
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<figureCitation id="131D2A356126DA79FD5A553824D9FE2E" box="[648,752,522,548]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. ‘latifrons’ (), H. mentatus Regan, 1925 (), H. sp. ‘rex’ (), H. sp. ‘simba’ (▼), H. sp. ‘glaucus’ (), H. sp. ‘aquila’ (), H. sp. ‘kimondo’ (●), H. sp. ‘falcatus’ (), H. sp. ‘curvidens’ (), H. sp. ‘pardus’ (), H. sp. ‘quasimodo’ (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. ‘latifrons’ the female and for H. sp. ‘curvidens’ the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="33" pageNumber="34">Figs 1–2</figureCitation>
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,
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<figureCitation id="131D2A356126DA79FD2E55382560FE2E" box="[764,841,522,548]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="34.[189,232,1963,1989]" captionStartId-1="35.[189,232,1892,1918]" captionStartId-2="36.[189,232,1963,1989]" captionTargetBox-0="[189,1398,1206,1852]" captionTargetBox-1="[260,1324,283,1860]" captionTargetBox-2="[189,1389,1015,1868]" captionTargetId-0="figure-254@34.[189,1399,1206,1852]" captionTargetId-1="figure-12@35.[260,1324,265,1860]" captionTargetId-2="figure-239@36.[188,1391,1015,1871]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionTargetPageId-2="36" captionText-0="Fig. 17. Haplochromis glaucus sp. nov., holotype, ♂, 106.2 mm SL (RMCA 2019.002.P.0016). Drawn by N. Vranken." captionText-1="Fig. 18. Haplochromis glaucus sp. nov. a. Photograph of preserved holotype (RMCA 2019.002.P.0016; 106.2 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 19. Haplochromis glaucus sp. nov. (RMCA 2019.002.P.0020; 110.9 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484203" figureDoi-1="http://doi.org/10.5281/zenodo.6484205" figureDoi-2="http://doi.org/10.5281/zenodo.6484207" httpUri-0="https://zenodo.org/record/6484203/files/figure.png" httpUri-1="https://zenodo.org/record/6484205/files/figure.png" httpUri-2="https://zenodo.org/record/6484207/files/figure.png" pageId="33" pageNumber="34">17–19</figureCitation>
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;
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<tableCitation id="C6A4030B6126DA79FC8755382585FE2E" box="[853,940,522,548]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="33" pageNumber="34" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C653B6126DA7AFF6F557C2431FD63" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
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<paragraph id="8B9936B06126DA79FF6F557C27E0FE62" blockId="33.[189,457,590,616]" box="[189,457,590,616]" pageId="33" pageNumber="34">
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<heading id="D0D181DC6126DA79FF6F557C27E0FE62" bold="true" box="[189,457,590,616]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
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<emphasis id="B952EAA26126DA79FF6F557C27E0FE62" bold="true" box="[189,457,590,616]" pageId="33" pageNumber="34">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9936B06126DA79FF6F554D27A9FEEF" blockId="33.[188,1398,639,742]" pageId="33" pageNumber="34">Species with a piscivorous morphology; snout acute in lateral view; outer oral teeth few and large [UOT 25–47 (median 30)]; males grey with light blue flank and a dusky to black head; female colour pattern similar to males.</paragraph>
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<paragraph id="8B9936B06126DA79FF6F542224FFF807" blockId="33.[187,1398,783,1038]" pageId="33" pageNumber="34">
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Amongst piscivorous species from the Lake Edward system,
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<taxonomicName id="4C264D336126DA79FCAF542225D5FF23" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[893,1020,783,810]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
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<emphasis id="B952EAA26126DA79FCAF542225BBFF23" box="[893,914,784,809]" italics="true" pageId="33" pageNumber="34">H</emphasis>
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.
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<emphasis id="B952EAA26126DA79FC73543D25D5FF23" box="[929,1020,783,809]" italics="true" pageId="33" pageNumber="34">glaucus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157D96126DA79FBD354222273FF20" box="[1025,1114,784,810]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C264D336126DA79FB225422235FFF23" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[1264,1398,783,810]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
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<emphasis id="B952EAA26126DA79FB225422232CFF23" box="[1264,1285,784,809]" italics="true" pageId="33" pageNumber="34">H</emphasis>
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||
.
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<emphasis id="B952EAA26126DA79FAC6543D235FFF23" box="[1300,1398,783,809]" italics="true" pageId="33" pageNumber="34">latifrons</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A26157D96126DA79FF6E5404273FFF5A" box="[188,278,822,848]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C264D336126DA79FE9C540427F5FF45" authorityName="Regan" authorityYear="1925" box="[334,476,822,848]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="mentatus">
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<emphasis id="B952EAA26126DA79FE9C5404274AFF45" box="[334,355,822,847]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FEA0540427F5FF45" box="[370,476,822,847]" italics="true" pageId="33" pageNumber="34">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by a shorter caudal peduncle [CPL 13.4–16.1 (mean 14.8) vs 15.7–18.0 (16.6– 17.0) % SL]; further from
|
||
<taxonomicName id="4C264D336126DA79FE23546E245EFF7F" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[497,631,859,886]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FE23546E242FFF7F" box="[497,518,860,885]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FDC75469245EFF7F" box="[533,631,859,885]" italics="true" pageId="33" pageNumber="34">latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FDAF546E24FEFF7C" box="[637,727,860,886]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a longer anal fin base [AFB 17.3–20.3 (18.6) vs 14.7– 17.3 (15.7) % SL] and absence vs presence of a well-defined mid-lateral band; further from
|
||
<taxonomicName id="4C264D336126DA79FB3B54B0235FFF91" authorityName="Regan" authorityYear="1925" box="[1257,1398,898,924]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA26126DA79FB3B54B022D7FF91" box="[1257,1278,898,923]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FADE54B0235FFF91" box="[1292,1398,898,923]" italics="true" pageId="33" pageNumber="34">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by a broader lower pharyngeal bone [LPW 93.3–95.1 vs 83.6–85.7% LPL], a slightly longer pre-pectoral distance [PrP 36.4–39.4 (38.1) vs 33.1–38.2 (36.0) % SL], and dominant males uniformly light blue vs yellow-green with a red anterior part of flank.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06126DA79FF6F530A25F2F93C" blockId="33.[188,1400,1079,1334]" pageId="33" pageNumber="34">
|
||
It differs from
|
||
<taxonomicName id="4C264D336126DA79FEA1530A27E9F85B" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[371,448,1080,1106]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FEA1530A27A1F85B" box="[371,392,1080,1105]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE49530A27E9F85B" box="[411,448,1080,1105]" italics="true" pageId="33" pageNumber="34">rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE18530A2401F858" box="[458,552,1080,1106]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D336126DA79FDB5530A24FCF858" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[615,725,1080,1106]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FDB5530A2455F85B" box="[615,636,1080,1105]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FD5C530A24FCF858" box="[654,725,1080,1106]" italics="true" pageId="33" pageNumber="34">simba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FD0D530A2514F858" box="[735,829,1080,1106]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a broader interorbital area [IOW 50.9–57.1 (53.8) vs 44.9–52.7 (48.1–48.9) % HW]; further from
|
||
<taxonomicName id="4C264D336126DA79FCE7536C2557F87D" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[821,894,1118,1144]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="rex" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FCE7536C2563F87D" box="[821,842,1118,1143]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FC8B536C2557F87D" box="[857,894,1118,1143]" italics="true" pageId="33" pageNumber="34">rex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FC56536C25F7F872" box="[900,990,1118,1144]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a gentler sloping snout (30–40° vs 40–50°), acute vs rounded oral jaws in dorsal view, and dominant males light blue with a blackish operculum and a dusky snout vs cream-coloured with an orange operculum and a light blue snout; further from
|
||
<taxonomicName id="4C264D336126DA79FE8B53E227EEF8E0" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[345,455,1232,1258]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="simba" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FE8B53E22747F8E3" box="[345,366,1232,1257]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE5253E227EEF8E0" box="[384,455,1232,1258]" italics="true" pageId="33" pageNumber="34">simba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE0353E22407F8E0" box="[465,558,1232,1258]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by a broader lower pharyngeal bone [LPW 93.3–95.1 vs 83.8–87.9% LPL], absent or weakly developed vs strongly developed mental prominence, and dominant males uniformly light-blue vs yellow with an orange anterior part of flank.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06126DA79FF6F5252232CF9CC" blockId="33.[188,1397,1376,1478]" pageId="33" pageNumber="34">
|
||
It differs from
|
||
<taxonomicName id="4C264D336126DA79FEB4525327FFF970" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[358,470,1376,1402]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="aquila" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FEB452532752F970" box="[358,379,1377,1402]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE58525227FFF970" box="[394,470,1376,1402]" italics="true" pageId="33" pageNumber="34">aquila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE0E5252241CF970" box="[476,565,1376,1402]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a smaller eye [ED 23.2–28.7 (26.8) vs 30.0–31.5 (30.6) % HL], a narrower head [HW 38.9–40.9 (39.7) vs 40.1–43.7 (42.0) % HL], and dominant males light blue with crimson anal and caudal fins vs light grey with bright red anal and caudal fins.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06126DA79FF6F52C327EEFAA8" blockId="33.[189,1399,1520,1698]" pageId="33" pageNumber="34">
|
||
It differs from
|
||
<taxonomicName id="4C264D336126DA79FEB552C327C4FA00" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[359,493,1520,1547]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FEB552C32755FA00" box="[359,380,1521,1546]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE5852C227C4FA00" box="[394,493,1520,1546]" italics="true" pageId="33" pageNumber="34">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE2752C32478FA01" box="[501,593,1521,1547]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C264D336126DA79FD8E52C324D9FA00" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[604,752,1520,1547]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="curvidens" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FD8E52C32458FA00" box="[604,625,1521,1546]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FD5252C224D9FA00" box="[640,752,1520,1546]" italics="true" pageId="33" pageNumber="34">curvidens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FD2552C3257BFA01" box="[759,850,1521,1547]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C264D336126DA79FC5D52C32219FA00" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[911,1072,1520,1547]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FC5D52C3258DFA00" box="[911,932,1521,1546]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FC6052C22219FA00" box="[946,1072,1520,1546]" italics="true" pageId="33" pageNumber="34">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FBEA52C322B8FA01" box="[1080,1169,1521,1547]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a narrower head [HW 38.9–40.9 (39.7) vs 42.0–48.1 (43.4–45.3) % HL], large vs small outer oral teeth, and a smaller number of outer upper jaw teeth [UOT 25–47 (30) vs 43–71 (49–58)]; further from
|
||
<taxonomicName id="4C264D336126DA79FF6C5151276AFA76" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[190,323,1634,1661]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FF6C515126FBFA76" box="[190,210,1635,1660]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FF325150276AFA76" box="[224,323,1634,1660]" italics="true" pageId="33" pageNumber="34">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE985151278AFA77" box="[330,419,1635,1661]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D336126DA79FE0951512455FA76" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[475,636,1634,1661]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="quasimodo" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FE09515127D9FA76" box="[475,496,1635,1660]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE2C51502455FA76" box="[510,636,1634,1660]" italics="true" pageId="33" pageNumber="34">quasimodo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FD51515124F5FA77" box="[643,732,1635,1661]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by dominant males light blue vs grey dorsally and yellow or blue-black ventrally.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06126DA79FF6F51FF2414FB52" blockId="33.[189,1399,1740,1881]" pageId="33" pageNumber="34">
|
||
It differs from
|
||
<taxonomicName id="4C264D336126DA79FEBC51FF27D9FAEC" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[366,496,1740,1767]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="falcatus" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FEBC51FF27ABFAEC" box="[366,386,1741,1766]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE4051FE27D9FAEC" box="[402,496,1740,1766]" italics="true" pageId="33" pageNumber="34">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE2B51FF247CFAED" box="[505,597,1741,1767]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of a shorter pre-dorsal distance [PrD 35.4–37.0 (36.1) vs 36.9–41.1 (39.5) % SL], a steeper lower jaw side (35–45° vs 15–25°), weakly recurved vs strongly recurved outer oral teeth, and dominant males uniformly light blue vs olive-green with an orange-red anterior part of flank.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06126DA79FF6F50B125B1FBE2" blockId="33.[188,1399,1922,2025]" pageId="33" pageNumber="34">
|
||
It differs from
|
||
<taxonomicName id="4C264D336126DA79FEA750B127C7FB96" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[373,494,1922,1949]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="pardus" status="sp. nov.">
|
||
<emphasis id="B952EAA26126DA79FEA750B127A3FB96" box="[373,394,1923,1948]" italics="true" pageId="33" pageNumber="34">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26126DA79FE4F50B027C7FB96" box="[413,494,1922,1948]" italics="true" pageId="33" pageNumber="34">pardus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96126DA79FE2850B12470FB97" box="[506,601,1923,1949]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of deeper lacrimal [LaD 18.0–22.7 (19.8) vs 16.0–18.3 (17.3) % HL], a broader interorbital area [IOW 50.9–57.1 (53.8) vs 39.3–48.4 (44.6) % HW], and dominant males light blue vs speckled to uniformly black.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06125DA7AFF6F563B2431FD63" blockId="34.[189,1399,265,362]" pageId="34" pageNumber="35">
|
||
It further differs from
|
||
<taxonomicName id="4C264D336125DA7AFE1D563824BBFD29" authorityName="Regan" authorityYear="1921" box="[463,658,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="squamipinnis">
|
||
<emphasis id="B952EAA26125DA7AFE1D563827CDFD29" box="[463,484,266,291]" italics="true" pageId="34" pageNumber="35">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26125DA7AFE24563824BBFD29" box="[502,658,266,291]" italics="true" pageId="34" pageNumber="35">squamipinnis</emphasis>
|
||
</taxonomicName>
|
||
by the combination of a gentler gape inclination (20–30° vs 30–45°), absence vs presence of minute scales on proximal parts of dorsal and anal fins, and dominant males light blue vs slate blue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6125DA7AFF6F56A327ECFDF6" pageId="34" pageNumber="35" type="etymology">
|
||
<paragraph id="8B9936B06125DA7AFF6F56A3276DFDA1" blockId="34.[189,324,401,427]" box="[189,324,401,427]" pageId="34" pageNumber="35">
|
||
<heading id="D0D181DC6125DA7AFF6F56A3276DFDA1" bold="true" box="[189,324,401,427]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
|
||
<emphasis id="B952EAA26125DA7AFF6F56A3276DFDA1" bold="true" box="[189,324,401,427]" pageId="34" pageNumber="35">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06125DA7AFF6F568D27ECFDF6" blockId="34.[189,1398,447,508]" pageId="34" pageNumber="35">
|
||
Specific name from the Latin ‘
|
||
<taxonomicName id="4C264D336125DA7AFDF2568D2455FDD3" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[544,636,447,473]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="glaucus">
|
||
<emphasis id="B952EAA26125DA7AFDF2568D2455FDD3" box="[544,636,447,473]" italics="true" pageId="34" pageNumber="35">glaucus</emphasis>
|
||
</taxonomicName>
|
||
’ for ‘greyish blue’; referring to grey and light-blue colour pattern of all adult specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6125DA7CFF6F5516246CFD4C" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="materials_examined">
|
||
<paragraph id="8B9936B06125DA7AFF6F55162783FE34" blockId="34.[189,426,548,574]" box="[189,426,548,574]" pageId="34" pageNumber="35">
|
||
<heading id="D0D181DC6125DA7AFF6F55162783FE34" bold="true" box="[189,426,548,574]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
|
||
<emphasis id="B952EAA26125DA7AFF6F55162783FE34" bold="true" box="[189,426,548,574]" pageId="34" pageNumber="35">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06125DA7AFF385561253FFEB9" blockId="34.[189,1399,595,691]" pageId="34" pageNumber="35">
|
||
<materialsCitation id="3B4E3CED6125DA7AFF385561253FFEB9" ID-GBIF-Occurrence="3861203370" collectingDate="2019-03-21" collectionCode="RMCA" collectorName="HIPE 4 exped." country="Uganda" latitude="-0.093" location="Kayanja" longLatPrecision="1" longitude="29.758" pageId="34" pageNumber="35" specimenCode="RMCA 2019.002.P.0016" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="holotype">
|
||
<emphasis id="B952EAA26125DA7AFF3855612775FE67" bold="true" box="[234,348,595,621]" pageId="34" pageNumber="35">
|
||
<typeStatus id="549D88126125DA7AFF3855612775FE67" box="[234,348,595,621]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33176206125DA7AFF6F55442713FE9A" box="[189,314,630,656]" name="Uganda" pageId="34" pageNumber="35">UGANDA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD396125DA7AFE8455442747FE9A" box="[342,366,630,656]" pageId="34" pageNumber="35" type="male">♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556125DA7AFEAE554427DCFE9A" box="[380,501,630,656]" metricMagnitude="-1" metricUnit="m" metricValue="1.062" pageId="34" pageNumber="35" unit="mm" value="106.2">106.2 mm</quantity>
|
||
SL;
|
||
<collectingRegion id="49E2F8526125DA7AFDE2554424FBFE9A" box="[560,722,630,656]" pageId="34" pageNumber="35">Lake Edward</collectingRegion>
|
||
,
|
||
<location id="8EF9606B6125DA7AFD0D55442568FE9A" LSID="urn:lsid:plazi:treatment:038F87A66126DA7EFDB2568F27FAFE3E:8EF9606B6125DA7AFD0D55442568FE9A" box="[735,833,630,656]" country="Uganda" latitude="-0.093" longLatPrecision="1" longitude="29.758" name="Kayanja" pageId="34" pageNumber="35" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE1250776125DA7AFC695546227DFE9A" box="[955,1108,628,656]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="34" pageNumber="35" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776125DA7AFBB055462325FE9A" box="[1122,1292,628,656]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="34" pageNumber="35" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706125DA7AFAC8554426D2FEB9" pageId="34" pageNumber="35" value="2019-03-21">
|
||
<collectingDate id="EFDCE9986125DA7AFAC8554426D2FEB9" pageId="34" pageNumber="35" value="2019-03-21">21 Mar. 2019</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666125DA7AFED555AB2785FEB9" box="[263,428,665,691]" pageId="34" pageNumber="35">HIPE4 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6125DA7AFE3C55AB253BFEB9" box="[494,786,665,691]" pageId="34" pageNumber="35">RMCA 2019.002.P.0016</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06125DA7AFF3855E8257CFFAE" blockId="34.[189,1399,730,933]" pageId="34" pageNumber="35">
|
||
<materialsCitation id="3B4E3CED6125DA7AFF3855E827DCFF54" ID-GBIF-Occurrence="3861203344" collectingDate="1953-05-25" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" latitude="-0.49499997" location="Kigera river" longLatPrecision="21" longitude="29.637222" pageId="34" pageNumber="35" specimenCode="IRSNB 13477" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<emphasis id="B952EAA26125DA7AFF3855E8274EFEFE" bold="true" box="[234,359,730,756]" pageId="34" pageNumber="35">
|
||
<typeStatus id="549D88126125DA7AFF3855E8274EFEFE" box="[234,359,730,756]" pageId="34" pageNumber="35" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33176206125DA7AFF6F55CC24C6FF12" box="[189,751,766,792]" name="Democratic Republic of the Congo" pageId="34" pageNumber="35">DEMOCRATIC REPUBLIC OF THE CONGO</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD396125DA7AFCDB55CC2511FF12" box="[777,824,766,792]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556125DA7AFC9755CF2594FF12" box="[837,957,765,792]" metricMagnitude="-1" metricUnit="m" metricValue="1.583" pageId="34" pageNumber="35" unit="mm" value="158.3">158.3 mm</quantity>
|
||
SL; “Lac Edouard: au large de la riv. Kigera” [
|
||
<collectingRegion id="49E2F8526125DA7AFEFA541327E4FF31" box="[296,461,801,827]" pageId="34" pageNumber="35">Lake Edward</collectingRegion>
|
||
: offshore
|
||
<location id="8EF9606B6125DA7AFD93541324FFFF31" LSID="urn:lsid:plazi:treatment:038F87A66126DA7EFDB2568F27FAFE3E:8EF9606B6125DA7AFD93541324FFFF31" box="[577,726,800,827]" country="Democratic Republic of the Congo" latitude="-0.49499997" longLatPrecision="21" longitude="29.637222" name="Kigera river" pageId="34" pageNumber="35" stateProvince="Lake Edward">Kigera river</location>
|
||
];
|
||
<geoCoordinate id="EE1250776125DA7AFD3B542D2543FF31" box="[745,874,799,827]" degrees="0" direction="south" minutes="29" orientation="latitude" pageId="34" pageNumber="35" precision="15" seconds="42" value="-0.49499997">0°29′42″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776125DA7AFCAA542D2221FF31" box="[888,1032,799,827]" degrees="29" direction="east" minutes="38" orientation="longitude" pageId="34" pageNumber="35" precision="15" seconds="14" value="29.637222">29°38′14″ E</geoCoordinate>
|
||
(inferred);
|
||
<date id="FF9810706125DA7AFB5D54132306FF36" box="[1167,1327,801,828]" pageId="34" pageNumber="35" value="1953-05-25">
|
||
<collectingDate id="EFDCE9986125DA7AFB5D54132306FF36" box="[1167,1327,801,828]" pageId="34" pageNumber="35" value="1953-05-25">25 May 1953</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666125DA7AFAEE54132722FF54" pageId="34" pageNumber="35">KEA exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6125DA7AFE9C547627DCFF54" box="[334,501,836,862]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="34" pageNumber="35">IRSNB 13477</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6125DA7AFDC254762579FFAE" ID-GBIF-Occurrence="3861203352" collectingDate="1953-06-01" collectionCode="IRSNB" collectorName="KEA exped." country="Democratic Republic of the Congo" latitude="-0.19416668" location="Kiavinionge" longLatPrecision="21" longitude="29.541945" pageId="34" pageNumber="35" specimenCode="IRSNB 13480" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396125DA7AFDC254762416FF54" box="[528,575,836,862]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556125DA7AFD9F547624ECFF54" box="[589,709,836,863]" metricMagnitude="-1" metricUnit="m" metricValue="1.509" pageId="34" pageNumber="35" unit="mm" value="150.9">150.9 mm</quantity>
|
||
SL; “Lac Edouard:
|
||
<quantity id="4CDE9B556125DA7AFC6654762224FF54" box="[948,1037,836,862]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="34" pageNumber="35" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2–3 km</quantity>
|
||
±
|
||
<quantity id="4CDE9B556125DA7AFBFC54762250FF54" box="[1070,1145,836,863]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="35" unit="m" value="500.0">500 m</quantity>
|
||
au large à l’Ouest de Kiavinionge” [
|
||
<collectingRegion id="49E2F8526125DA7AFEB854552426FF8B" box="[362,527,871,897]" pageId="34" pageNumber="35">Lake Edward</collectingRegion>
|
||
:
|
||
<locationDeviation id="9A2B2F926125DA7AFDC8545525ACFF88" box="[538,901,871,898]" pageId="34" pageNumber="35">
|
||
<quantity id="4CDE9B556125DA7AFDC85455245AFF8B" box="[538,627,871,897]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="34" pageNumber="35" unit="km" value="2.5" valueMax="3.0" valueMin="2.0">2–3 km</quantity>
|
||
±
|
||
<quantity id="4CDE9B556125DA7AFD46545524F6FF88" box="[660,735,871,898]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="35" unit="m" value="500.0">500 m</quantity>
|
||
offshore west
|
||
</locationDeviation>
|
||
of
|
||
<location id="8EF9606B6125DA7AFC7F5455226CFF88" LSID="urn:lsid:plazi:treatment:038F87A66126DA7EFDB2568F27FAFE3E:8EF9606B6125DA7AFC7F5455226CFF88" box="[941,1093,871,898]" country="Democratic Republic of the Congo" latitude="-0.19416668" longLatPrecision="21" longitude="29.541945" name="Kiavinionge" pageId="34" pageNumber="35" stateProvince="Lake Edward">Kiavinionge</location>
|
||
];
|
||
<geoCoordinate id="EE1250776125DA7AFB8A545422F0FF8B" box="[1112,1241,870,898]" degrees="0" direction="south" minutes="11" orientation="latitude" pageId="34" pageNumber="35" precision="15" seconds="39" value="-0.19416668">0°11′39″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776125DA7AFB345454235EFF88" box="[1254,1399,870,898]" degrees="29" direction="east" minutes="32" orientation="longitude" pageId="34" pageNumber="35" precision="15" seconds="31" value="29.541945">29°32′31″ E</geoCoordinate>
|
||
(inferred);
|
||
<date id="FF9810706125DA7AFEEF54B827E1FFAF" box="[317,456,906,933]" pageId="34" pageNumber="35" value="1953-06-01">
|
||
<collectingDate id="EFDCE9986125DA7AFEEF54B827E1FFAF" box="[317,456,906,933]" pageId="34" pageNumber="35" value="1953-06-01">1 Jun. 1953</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666125DA7AFE0654B9244DFFAE" box="[468,612,906,933]" pageId="34" pageNumber="35">KEA exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6125DA7AFD7454B82579FFAE" box="[678,848,906,932]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="34" pageNumber="35">IRSNB 13480</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06125DA7CFF6F54FE246CFD4C" blockId="34.[189,1399,970,1104]" lastBlockId="36.[189,1398,265,327]" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
|
||
<materialsCitation id="3B4E3CED6125DA7AFF6F54FE230BF803" ID-GBIF-Occurrence="3861203330" collectingDate="2018-02-04" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" lastPageId="34" lastPageNumber="37" latitude="-0.21391667" location="Kagoro" longLatPrecision="1" longitude="29.82214" pageId="34" pageNumber="35" specimenCode="RMCA 2018.008.P.0365, RMCA 2018.008.P.0366" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<collectingCountry id="F33176206125DA7AFF6F54FE2713FFEC" box="[189,314,972,998]" name="Uganda" pageId="34" pageNumber="35">UGANDA</collectingCountry>
|
||
–
|
||
<emphasis id="B952EAA26125DA7AFEB354FE243AFFEC" bold="true" box="[353,531,972,998]" pageId="34" pageNumber="35">
|
||
<collectingRegion id="49E2F8526125DA7AFEB354FE243AFFEC" box="[353,531,972,998]" pageId="34" pageNumber="35">Lake Edward</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D20FD396125DA7AFDE754FE244FFFEC" box="[565,614,972,998]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD396125DA7AFDAB54FE2485FFEC" box="[633,684,972,998]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556125DA7AFD6F54FE2545FFEC" box="[701,876,972,998]" metricMagnitude="-2" metricUnit="m" metricValue="9.215" metricValueMax="9.36" metricValueMin="9.07" pageId="34" pageNumber="35" unit="mm" value="92.15" valueMax="93.6" valueMin="90.7">90.7–93.6 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B6125DA7AFC7C54FE222FFFEC" LSID="urn:lsid:plazi:treatment:038F87A66126DA7EFDB2568F27FAFE3E:8EF9606B6125DA7AFC7C54FE222FFFEC" box="[942,1030,972,998]" country="Uganda" latitude="-0.21391667" longLatPrecision="1" longitude="29.82214" name="Kagoro" pageId="34" pageNumber="35" stateProvince="Lake Edward">Kagoro</location>
|
||
fishing ground;
|
||
<geoCoordinate id="EE1250776125DA7AFB0754F82358FFEC" box="[1237,1393,970,998]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="34" pageNumber="35" precision="1" seconds="50.1" value="-0.21391667">0°12′50.1″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776125DA7AFF6F54DF2776F803" box="[189,351,1005,1033]" degrees="29" direction="east" minutes="49" orientation="longitude" pageId="34" pageNumber="35" precision="1" seconds="19.7" value="29.82214">29°49′19.7″E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706125DA7AFEBC54DD242BF803" box="[366,514,1007,1033]" pageId="34" pageNumber="35" value="2018-02-04">
|
||
<collectingDate id="EFDCE9986125DA7AFEBC54DD242BF803" box="[366,514,1007,1033]" pageId="34" pageNumber="35" value="2018-02-04">4 Feb. 2018</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666125DA7AFDC354DD2490F803" box="[529,697,1007,1033]" pageId="34" pageNumber="35">HIPE3 exped.</collectorName>
|
||
leg.; open water;
|
||
<specimenCode id="DB809ECB6125DA7AFC4A54DD230BF803" box="[920,1314,1007,1034]" pageId="34" pageNumber="35">RMCA 2018.008.P.0365 to 0366</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6125DA7AFA9254DD2251F826" ID-GBIF-Occurrence="3861203342" collectingDate="2018-02-04" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" lastPageId="34" lastPageNumber="37" latitude="-0.21391667" location="Kagoro" longLatPrecision="1" longitude="29.82214" pageId="34" pageNumber="35" specimenCode="RMCA 2018.008.P.0367" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD396125DA7AFA9254DD2346F800" box="[1344,1391,1007,1034]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556125DA7AFF6F53202702F827" box="[189,299,1042,1069]" metricMagnitude="-2" metricUnit="m" metricValue="9.26" pageId="34" pageNumber="35" unit="mm" value="92.6">92.6 mm</quantity>
|
||
SL; same collection data as for preceding;
|
||
<specimenCode id="DB809ECB6125DA7AFC8053202251F826" box="[850,1144,1042,1068]" pageId="34" pageNumber="35">RMCA 2018.008.P.0367</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6125DA7CFB49532027C1FD29" ID-GBIF-Occurrence="3861203341" collectingDate="2019-03-21" collectionCode="RMCA" collectorName="HIPE 4 exped." country="Uganda" lastPageId="36" lastPageNumber="37" latitude="-0.093" location="Kayanja" longLatPrecision="1" longitude="29.758" pageId="34" pageNumber="35" specimenCode="RMCA 2019.002.P.0017, RMCA 2019.002.P.0018" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD396125DA7AFB49532022E5F827" box="[1179,1228,1042,1069]" pageId="34" pageNumber="35" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD396125DA7AFB325320233DF827" box="[1248,1300,1042,1069]" pageId="34" pageNumber="35" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556125DA7AFAF45320271AF85A" metricMagnitude="-1" metricUnit="m" metricValue="1.034" metricValueMax="1.047" metricValueMin="1.021" pageId="34" pageNumber="35" unit="mm" value="103.4" valueMax="104.7" valueMin="102.1">102.1– 104.7 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B6125DA7AFEBB530727E2F845" LSID="urn:lsid:plazi:treatment:038F87A66126DA7EFDB2568F27FAFE3E:8EF9606B6125DA7AFEBB530727E2F845" box="[361,459,1077,1103]" country="Uganda" latitude="-0.093" longLatPrecision="1" longitude="29.758" name="Kayanja" pageId="34" pageNumber="35" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE1250776125DA7AFD93530624FEF845" box="[577,727,1076,1104]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="34" pageNumber="35" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776125DA7AFD31530625A3F85A" box="[739,906,1076,1104]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="34" pageNumber="35" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706125DA7AFC475307221DF845" box="[917,1076,1077,1104]" pageId="34" pageNumber="35" value="2019-03-21">
|
||
<collectingDate id="EFDCE9986125DA7AFC475307221DF845" box="[917,1076,1077,1104]" pageId="34" pageNumber="35" value="2019-03-21">21 Mar. 2019</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D353666125DA7AFBED530722CBF845" box="[1087,1250,1077,1103]" pageId="34" pageNumber="35">HIPE4 exped.</collectorName>
|
||
leg.;
|
||
<specimenCode id="DB809ECB6125DA7CFAF0530727C1FD29" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">RMCA 2019.002.P.0017 to 0018</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6123DA7CFDD5563B2468FD4C" ID-GBIF-Occurrence="3861203401" collectingDate="2019-03-21" collectionCode="RMCA" collectorName="HIPE 4 exped." country="Uganda" lastPageId="36" lastPageNumber="37" latitude="-0.093" location="Kayanja" longLatPrecision="1" longitude="29.758" pageId="36" pageNumber="35" specimenCode="RMCA 2019.002.P.0019, RMCA 2019.002.P.0020, RMCA 2019.002.P.0021" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Lake Edward">
|
||
<specimenCount id="9D20FD396123DA7CFDD5563B2410FD29" box="[519,569,265,291]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D20FD396123DA7CFD9B563B24A6FD29" box="[585,655,265,291]" pageId="36" pageNumber="37" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556123DA7CFD72563B2542FD29" box="[672,875,265,291]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.223" metricValueMin="1.077" pageId="36" pageNumber="37" unit="mm" value="115.0" valueMax="122.3" valueMin="107.7">107.7–122.3 mm</quantity>
|
||
SL; same collection data as for preceding;
|
||
<specimenCode id="DB809ECB6123DA7CFF6F561E2468FD4C" box="[189,577,300,327]" pageId="36" pageNumber="37">RMCA 2019.002.P.0019 to 0021</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5966386125DA7AFF6F50992759FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484203" ID-Zenodo-Dep="6484203" httpUri="https://zenodo.org/record/6484203/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1963,1989]" targetBox="[189,1398,1206,1852]" targetPageId="34">
|
||
<paragraph id="8B9936B06125DA7AFF6F50992759FBE2" blockId="34.[189,1399,1963,2025]" pageId="34" pageNumber="35">
|
||
<emphasis id="B952EAA26125DA7AFF6F50992732FBCF" bold="true" box="[189,283,1963,1989]" pageId="34" pageNumber="35">Fig. 17.</emphasis>
|
||
<taxonomicName id="4C264D336125DA7AFEF050992406FBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[290,559,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA26125DA7AFEF050992406FBCF" box="[290,559,1963,1989]" italics="true" pageId="34" pageNumber="35">Haplochromis glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96125DA7AFDE5509E24BCFBCC" box="[567,661,1964,1990]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂, 106.2 mm SL (RMCA 2019.002.P.0016). Drawn by N. Vranken.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5966386124DA7BFF6F505624E5FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484205" ID-Zenodo-Dep="6484205" httpUri="https://zenodo.org/record/6484205/files/figure.png" pageId="35" pageNumber="36" startId="35.[189,232,1892,1918]" targetBox="[260,1324,283,1860]" targetPageId="35">
|
||
<paragraph id="8B9936B06124DA7BFF6F505624E5FBE2" blockId="35.[188,1398,1892,2025]" pageId="35" pageNumber="36">
|
||
<emphasis id="B952EAA26124DA7BFF6F50562733FB75" bold="true" box="[189,282,1892,1919]" pageId="35" pageNumber="36">Fig. 18.</emphasis>
|
||
<taxonomicName id="4C264D336124DA7BFEF350572405FB75" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[289,556,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA26124DA7BFEF350572405FB75" box="[289,556,1893,1919]" italics="true" pageId="35" pageNumber="36">Haplochromis glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96124DA7BFDE1505724A4FB75" box="[563,653,1893,1919]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B952EAA26124DA7BFD465057248AFB75" bold="true" box="[660,675,1893,1919]" pageId="35" pageNumber="36">a</emphasis>
|
||
. Photograph of preserved holotype (RMCA 2019.002.P.0016; 106.2 mm SL).
|
||
<emphasis id="B952EAA26124DA7BFEA750BA27ACFBA8" bold="true" box="[373,389,1928,1954]" pageId="35" pageNumber="36">b</emphasis>
|
||
. X-ray image of holotype
|
||
<emphasis id="B952EAA26124DA7BFD6950BA24CEFBA8" bold="true" box="[699,743,1928,1954]" pageId="35" pageNumber="36">c–d</emphasis>
|
||
. Photographs of freshly caught specimens.
|
||
<emphasis id="B952EAA26124DA7BFB3750BA22DBFBA8" bold="true" box="[1253,1266,1928,1954]" pageId="35" pageNumber="36">c</emphasis>
|
||
. Dominant male, the holotype.
|
||
<emphasis id="B952EAA26124DA7BFE62509927E9FBCF" bold="true" box="[432,448,1963,1989]" pageId="35" pageNumber="36">d</emphasis>
|
||
. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33C653B6123DA7EFF6F565F276FFD99" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8B9936B06123DA7CFF6F565F2767FD8D" blockId="36.[189,334,365,391]" box="[189,334,365,391]" pageId="36" pageNumber="37">
|
||
<heading id="D0D181DC6123DA7CFF6F565F2767FD8D" bold="true" box="[189,334,365,391]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
|
||
<emphasis id="B952EAA26123DA7CFF6F565F2767FD8D" bold="true" box="[189,334,365,391]" pageId="36" pageNumber="37">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06123DA7CFF6F56AE2710FF1D" blockId="36.[188,1400,412,791]" pageId="36" pageNumber="37">
|
||
Based on
|
||
<specimenCount id="9D20FD396123DA7CFEE756AE27FCFDBC" box="[309,469,412,438]" pageId="36" pageNumber="37" type="generic">11 specimens</specimenCount>
|
||
(
|
||
<quantity id="4CDE9B556123DA7CFE3556AE248DFDBD" box="[487,676,412,439]" metricMagnitude="-1" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.583" metricValueMin="0.907" pageId="36" pageNumber="37" unit="mm" value="124.5" valueMax="158.3" valueMin="90.7">90.7–158.3 mm</quantity>
|
||
SL); body shallow (
|
||
<tableCitation id="C6A4030B6123DA7CFC4956AE25DCFDBC" box="[923,1013,412,438]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="36" pageNumber="37" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
|
||
) and oval (
|
||
<figureCitation id="131D2A356123DA7CFB5056AE22F2FDBC" box="[1154,1243,412,438]" captionStart="Fig" captionStartId="34.[189,232,1963,1989]" captionTargetBox="[189,1398,1206,1852]" captionTargetId="figure-254@34.[189,1399,1206,1852]" captionTargetPageId="34" captionText="Fig. 17. Haplochromis glaucus sp. nov., holotype, ♂, 106.2 mm SL (RMCA 2019.002.P.0016). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484203" httpUri="https://zenodo.org/record/6484203/files/figure.png" pageId="36" pageNumber="37">Fig. 17</figureCitation>
|
||
). Head long, very narrow, and with a straight dorsal outline; eye small; interorbital area narrow; lacrimal and cheek deep. Snout long, rounded in dorsal view, very narrow, acute, and slopes gently at 30–40°; premaxillary pedicel long and weakly prominent. Jaws long, relatively stout, narrow, isognathous, and acute in dorsal view; gape large and slopes gently at 20–30°; maxilla extends (almost) to vertical through anterior margin of orbit. Lower jaw relatively stout and with a straight to slightly convex ventral outline in lateral view, mental prominence absent or weakly developed, and lower jaw side steep with an inclination of 35–45° to horizontal in anterior view. Upper jaw expanded anteriorly and weakly ventrally. Lips and oral mucosa large. Neurocranium shallow, ethmo-vomerine block decurved, preorbital region shallow (20–23% NL), orbital region shallow (28–31% NL), and supraoccipital crest shallow and wedge-shaped (
|
||
<figureCitation id="131D2A356123DA7CFF1755CE2702FF1D" box="[197,299,764,791]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[260,1324,283,1860]" captionTargetId="figure-12@35.[260,1324,265,1860]" captionTargetPageId="35" captionText="Fig. 18. Haplochromis glaucus sp. nov. a. Photograph of preserved holotype (RMCA 2019.002.P.0016; 106.2 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484205" httpUri="https://zenodo.org/record/6484205/files/figure.png" pageId="36" pageNumber="37">Fig. 18b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936B06123DA7DFF6F540C249FFD84" blockId="36.[189,1399,829,927]" lastBlockId="37.[189,1400,265,398]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">Outer oral teeth few, unicuspid, and very large. Necks conical, stout, and straight; crowns recurved and acutely pointed. Dental arcades rounded. Outer teeth widely and irregularly set with neck-distances of 1–4 neck-widths. In upper jaw, 1–3 posteriormost teeth slightly enlarged. Inner teeth small, strongly recurved, unicuspid, and acutely pointed. Tooth bands very slender crescent-shaped with 1–3 rows of inner teeth, and narrow posteriorly until only outer row remains past ¾ length of tooth band. Inner teeth closely and regularly set on 1–2 outer neck-widths from outer row; implantation recumbent; size decreases slightly buccally and posteriorly.</paragraph>
|
||
<caption id="DF5966386123DA7CFF6F509925F8FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484207" ID-Zenodo-Dep="6484207" httpUri="https://zenodo.org/record/6484207/files/figure.png" pageId="36" pageNumber="37" startId="36.[189,232,1963,1989]" targetBox="[189,1389,1015,1868]" targetPageId="36">
|
||
<paragraph id="8B9936B06123DA7CFF6F509925F8FBE2" blockId="36.[188,1399,1963,2025]" pageId="36" pageNumber="37">
|
||
<emphasis id="B952EAA26123DA7CFF6F50992733FBCF" bold="true" box="[189,282,1963,1989]" pageId="36" pageNumber="37">Fig. 19.</emphasis>
|
||
<taxonomicName id="4C264D336123DA7CFEF250992405FBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[288,556,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="glaucus" status="sp. nov.">
|
||
<emphasis id="B952EAA26123DA7CFEF250992405FBCF" box="[288,556,1963,1989]" italics="true" pageId="36" pageNumber="37">Haplochromis glaucus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96123DA7CFDE0509E24A4FBCC" box="[562,653,1964,1990]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(RMCA 2019.002.P.0020; 110.9 mm SL).
|
||
<emphasis id="B952EAA26123DA7CFB53509922B9FBCF" bold="true" box="[1153,1168,1963,1989]" pageId="36" pageNumber="37">a</emphasis>
|
||
. Dorsal view of the lower pharyngeal jaw.
|
||
<emphasis id="B952EAA26123DA7CFE1A50FC27F1FBE2" bold="true" box="[456,472,1998,2024]" pageId="36" pageNumber="37">b</emphasis>
|
||
. Lateral view of the lower pharyngeal jaw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936B06122DA7DFF6F568427B3FE88" blockId="37.[189,1399,438,643]" pageId="37" pageNumber="38">
|
||
Lower pharyngeal bone average in length, triangular, slim, and shallow over whole length (
|
||
<figureCitation id="131D2A356122DA7DFADD56842341FDDA" box="[1295,1384,438,464]" captionStart="Fig" captionStartId="36.[189,232,1963,1989]" captionTargetBox="[189,1389,1015,1868]" captionTargetId="figure-239@36.[188,1391,1015,1871]" captionTargetPageId="36" captionText="Fig. 19. Haplochromis glaucus sp. nov. (RMCA 2019.002.P.0020; 110.9 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484207" httpUri="https://zenodo.org/record/6484207/files/figure.png" pageId="37" pageNumber="38">Fig. 19</figureCitation>
|
||
). Pharyngeal teeth relatively large and slender; major cusps acutely pointed; cusp gaps straight; minor cusps and cusp protuberances mostly absent. Teeth in two median longitudinal rows equal in size and form to lateral teeth,
|
||
<quantity id="4CDE9B556122DA7DFE285513244FFE31" box="[506,614,545,571]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="37" pageNumber="38" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">10–12 in</quantity>
|
||
each row. Posterior transverse row with 15–16 teeth, implanted recumbently with a lateral inclination; major cusps recurved, bluntly pointed, and laterally compressed; minor cusps small.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F5598235EFECE" blockId="37.[189,1399,682,709]" box="[189,1399,682,709]" pageId="37" pageNumber="38">Chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin.</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F55DE2577FF44" blockId="37.[189,1399,748,846]" pageId="37" pageNumber="38">Caudal fin emarginate; dorsal and anal fins reach to between verticals through one scale anterior to and two scales posterior to caudal-fin base. Pectoral and pelvic fins reach to between anal opening and first anal-fin spine; first branched pelvic-fin ray not elongated.</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F5447224FFFB9" blockId="37.[189,1399,885,948]" pageId="37" pageNumber="38">Ceratobranchial gill rakers in outer row of first gill arch short, stout, and simple; posteriormost rakers anvil-shaped to rarely weakly trifid. Epibranchial gill rakers slender and simple.</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F54E9278AFFFF" blockId="37.[189,419,987,1013]" box="[189,419,987,1013]" pageId="37" pageNumber="38">
|
||
<heading id="D0D181DC6122DA7DFF6F54E9278AFFFF" bold="true" box="[189,419,987,1013]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B952EAA26122DA7DFF6F54E9278AFFFF" bold="true" box="[189,419,987,1013]" pageId="37" pageNumber="38">Colouration in life</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F533824E1F948" blockId="37.[189,1399,1034,1346]" pageId="37" pageNumber="38">
|
||
Dominant males: flank and caudal peduncle light-blue; dorsum grey with a yellow sheen; belly, chest, and operculum blackish with a yellow sheen; flank with 5–7 faint vertical stripes (
|
||
<figureCitation id="131D2A356122DA7DFB85531C2295F842" box="[1111,1212,1070,1096]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[260,1324,283,1860]" captionTargetId="figure-12@35.[260,1324,265,1860]" captionTargetPageId="35" captionText="Fig. 18. Haplochromis glaucus sp. nov. a. Photograph of preserved holotype (RMCA 2019.002.P.0016; 106.2 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484205" httpUri="https://zenodo.org/record/6484205/files/figure.png" pageId="37" pageNumber="38">Fig. 18c</figureCitation>
|
||
). Preoperculum black; cheek black with blue sheen; lacrimal and snout dusky; lower jaw blackish; branchiostegal membrane black; eye with dark grey outer ring and silver inner ring. Nostril, interorbital, supraorbital, and lacrimal stripes well-defined; nape band, vertical preopercular stripe, and mental blotch present; operculum with black posterior border. Pectoral and dorsal fins hyaline; dorsal fin with black lappets, dusky base, and dusky and maculated crimson posterior part. Anal and caudal fins crimson; anal fin with dusky posterior part and 1–2 large orange egg-spots with dusky rings; caudal fin with dusky base, hyaline posterior part, and crimson maculae.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F52582309FB3B" blockId="37.[189,1399,1386,1841]" pageId="37" pageNumber="38">
|
||
Females: strikingly similar to dominant males. Body, dorsum, and caudal peduncle grey with a yellow sheen; flanks light blue and with 5–7 faint vertical stripes; belly and chest blackish with a yellow sheen (
|
||
<figureCitation id="131D2A356122DA7DFF1452832719F9C6" box="[198,304,1457,1484]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[260,1324,283,1860]" captionTargetId="figure-12@35.[260,1324,265,1860]" captionTargetPageId="35" captionText="Fig. 18. Haplochromis glaucus sp. nov. a. Photograph of preserved holotype (RMCA 2019.002.P.0016; 106.2 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484205" httpUri="https://zenodo.org/record/6484205/files/figure.png" pageId="37" pageNumber="38">Fig. 18d</figureCitation>
|
||
). Operculum, preoperculum, cheek, lacrimal, and snout grey; operculum with blue sheen; cheek with black sheen; lower jaw blackish; branchiostegal membrane black; eye with dark grey outer ring and silver inner ring. Nostril, interorbital, supraorbital, and lacrimal stripes well-defined; nape band, vertical preopercular stripe, and mental blotch faint; operculum with faint black posterior border. Pectoral and dorsal fins hyaline; dorsal fin with black lappets, dusky base, and dusky posterior part. Anal and caudal fins yellow with a red sheen; anal fin with hyaline posterior part and 1–2 spots resembling egg-spots; caudal fin with dusky base, hyaline posterior part, and dusky maculae. Juveniles: dorsum greyish, belly, chest, operculum, and cheek white; transition gradual. Nostril, interorbital, supraorbital, and lacrimal stripes, nape band, and mental blotch faint. Pectoral, pelvic, dorsal, and anal fins hyaline; dorsal fin with black lappets and dusky base and posterior part; anal fin with a yellow sheen and 1–2 spots resembling egg-spots. Caudal fin dusky and with black maculae and a yellow ventral part.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06122DA7DFF6F506A27F9FB78" blockId="37.[189,464,1880,1906]" box="[189,464,1880,1906]" pageId="37" pageNumber="38">
|
||
<heading id="D0D181DC6122DA7DFF6F506A27F9FB78" bold="true" box="[189,464,1880,1906]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
|
||
<emphasis id="B952EAA26122DA7DFF6F506A27F9FB78" bold="true" box="[189,464,1880,1906]" pageId="37" pageNumber="38">Preserved colouration</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06122DA7EFF6F50B5276FFD99" blockId="37.[189,1400,1927,2024]" lastBlockId="38.[189,1399,265,403]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||
Dorsum brown; belly and chest speckled black; flank dark yellowish and with 6–8 faint vertical stripes. Cheek yellow and speckled black; snout dusky; lower jaw and pre- and subopercula black (
|
||
<figureCitation id="131D2A356122DA7DFAD150992340FBCF" box="[1283,1385,1963,1989]" captionStart="Fig" captionStartId="35.[189,232,1892,1918]" captionTargetBox="[260,1324,283,1860]" captionTargetId="figure-12@35.[260,1324,265,1860]" captionTargetPageId="35" captionText="Fig. 18. Haplochromis glaucus sp. nov. a. Photograph of preserved holotype (RMCA 2019.002.P.0016; 106.2 mm SL). b. X-ray image of holotype c–d. Photographs of freshly caught specimens. c. Dominant male, the holotype. d. Female (RMCA 2019.002.P.0017; 102.1 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484205" httpUri="https://zenodo.org/record/6484205/files/figure.png" pageId="37" pageNumber="38">Fig. 18a</figureCitation>
|
||
). Nostril, interorbital, and vertical preopercular stripes well-defined; lacrimal stripe broad and welldefined; supraorbital stripe and nape band faint. Pectoral and anal fins hyaline; anal fin with black lappets, a dusky posterior margin, and 1–2 large egg-spots; pelvic fin black; dorsal fin dusky and with black lappets and a maculated posterior part; caudal fin dorsally dusky, ventrally hyaline, and with a dusky base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6121DA7EFF6F568E27FAFE3E" pageId="38" pageNumber="39" type="distribution">
|
||
<paragraph id="8B9936B06121DA7EFF6F568E27D8FDDC" blockId="38.[189,497,444,470]" box="[189,497,444,470]" pageId="38" pageNumber="39">
|
||
<heading id="D0D181DC6121DA7EFF6F568E27D8FDDC" bold="true" box="[189,497,444,470]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B952EAA26121DA7EFF6F568E27D8FDDC" bold="true" box="[189,497,444,470]" pageId="38" pageNumber="39">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06121DA7EFF6F56C227FAFE3E" blockId="38.[189,1398,496,564]" pageId="38" pageNumber="39">Only known from Lake Edward, found over sandy substrates. Based on its morphology, most probably a piscivorous species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |