424 lines
54 KiB
XML
424 lines
54 KiB
XML
<document id="61F3FBDC7578AEBEB20ACE4A6EA7422A" ID-CLB-Dataset="92619" ID-DOI="10.5852/ejt.2022.815.1749" ID-GBIF-Dataset="9a798cae-f10e-44c2-a65c-cdadbff28cb4" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484153" ID-ZooBank="6AD0082E-7349-48DE-AFCA-1EE0BFBB3887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650913068508" checkinUser="felipe" docAuthor="Vranken, Nathan, Steenberge, Maarten Van, Heylen, Annelies, Decru, Eva & Snoeks, Jos" docDate="2022" docId="038F87A6610ADA49FD8E52CD2781FACE" docLanguage="en" docName="ejt-815_1-94.pdf" docOrigin="European Journal of Taxonomy 815" docSource="http://dx.doi.org/10.5852/ejt.2022.815.1749" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haplochromis latifrons Vranken & Steenberge & Heylen & Decru & Snoeks 2022, sp. nov." docType="treatment" docUuid="F6BB9E11-AF52-476E-87CE-5C2602859D1B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="18" masterDocId="FFB6FFDE6107DA58FFD257322629FC0A" masterDocTitle="From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system" masterLastPageNumber="94" masterPageNumber="1" pageNumber="14" updateTime="1699329164302" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="49EE19C7DAF631B13505EF52DF7A3CC1">From a pair to a dozen: the piscivorous species of Haplochromis (Cichlidae) from the Lake Edward system</mods:title>
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<mods:namePart id="F37FEAE6B25DE13D3F261F587258BE2A">Vranken, Nathan</mods:namePart>
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<mods:affiliation id="A2BC61FE7350286F572393A499AD494A">Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium. & Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="BAD998C3D47110502F15BABE53EC8851" type="email">nathan.vranken@kuleuven.be</mods:nameIdentifier>
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<mods:namePart id="E19C66B8EC6834EF655DB61EF2CD92F0">Steenberge, Maarten Van</mods:namePart>
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<mods:affiliation id="45527A7EBCE23E638429F6F86BBFCCF2">Royal Belgian Institute of Natural Sciences, Operational Directorate Taxonomy and Phylogeny, Vautierstraat 29, 1000 Brussels, Belgium. & Masaryk University, Department of Botany and Zoology, Kotlářská 2, 611 37 Brno, Czech Republic.</mods:affiliation>
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<mods:nameIdentifier id="B76C72A4EF58C1FC7EB4882ED7374030" type="email">mvansteenberge@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="699949A42CAA24EBEE011D950F9CF14E">Heylen, Annelies</mods:namePart>
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<mods:affiliation id="28AC96F3E7789A8478484DF0ED4F2A3E">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="8F8702802F673958651A2496E432D8B9" type="email">annelies.heylen@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="D1BC9A9F0AB07EDF15C759B98C8BE658">Decru, Eva</mods:namePart>
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<mods:affiliation id="7846755F03F4CA65202930C1A02C2AFD">KU Leuven, Laboratory of Biodiversity and Evolutionary Genomics, Department of Biology, Charles Deberiotstraat 32, 3000 Leuven, Belgium.</mods:affiliation>
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<mods:affiliation id="934A8A2177E5A7BCD92798ED6D3B8E74">Royal Museum for Central Africa, Biology department, Section Vertebrates, Leuvensesteenweg 13, 3080 Tervuren, Belgium.</mods:affiliation>
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<mods:nameIdentifier id="A563B8117F8B399716B1EE2D209AB2C7" type="email">jos.snoeks@africamuseum.be</mods:nameIdentifier>
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<mods:title id="8F28454E8318385731F2A65C59EF7FC6">European Journal of Taxonomy</mods:title>
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<treatment id="038F87A6610ADA49FD8E52CD2781FACE" ID-DOI="http://doi.org/10.5281/zenodo.6484167" ID-GBIF-Taxon="197197510" ID-Zenodo-Dep="6484167" LSID="urn:lsid:zoobank.org:act:F6BB9E11-AF52-476E-87CE-5C2602859D1B" httpUri="http://treatment.plazi.org/id/038F87A6610ADA49FD8E52CD2781FACE" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
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<paragraph id="8B9936B0610ADA55FD8E52CD25F1FA10" blockId="13.[368,1220,1535,1633]" box="[604,984,1535,1562]" pageId="13" pageNumber="14">
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<heading id="D0D181DC610ADA55FD8E52CD25F1FA10" box="[604,984,1535,1562]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
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<taxonomicName id="4C264D33610ADA55FD8E52CD255EFA10" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[604,887,1535,1562]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
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<emphasis id="B952EAA2610ADA55FD8E52CD255EFA10" bold="true" box="[604,887,1535,1562]" italics="true" pageId="13" pageNumber="14">Haplochromis latifrons</emphasis>
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<taxonomicNameLabel id="A26157D9610ADA55FCAC513225F1FA10" box="[894,984,1536,1562]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B9936B0610ADA55FEA2511122EDFA34" blockId="13.[368,1220,1535,1633]" box="[368,1220,1571,1598]" pageId="13" pageNumber="14">
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<uuid id="FF800C65610ADA55FD40511122EDFA34" box="[658,1220,1571,1598]" pageId="13" pageNumber="14">F6BB9E11-AF52-476E-87CE-5C2602859D1B</uuid>
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</paragraph>
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<subSubSection id="C33C653B610ADA55FD45517425B4FA6A" box="[663,925,1606,1633]" pageId="13" pageNumber="14" type="description">
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<paragraph id="8B9936B0610ADA55FD45517425B4FA6A" blockId="13.[368,1220,1535,1633]" box="[663,925,1606,1633]" pageId="13" pageNumber="14">
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<figureCitation id="131D2A35610ADA55FD45517424D6FA6A" box="[663,767,1606,1632]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[190,234,1822,1848]" captionStartId-1="6.[189,231,1892,1918]" captionTargetBox-0="[193,1398,734,1726]" captionTargetBox-1="[208,1335,272,1830]" captionTargetId-0="figure-217@5.[189,1399,730,1726]" captionTargetId-1="figure-18@6.[208,1335,272,1830]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 1. The Lake Edward system, consisting of Lakes Edward and George that are connected through the Kazinga Channel. The catch localities of all examined specimens are indicated: Haplochromis sp. ‘latifrons’ (), H. mentatus Regan, 1925 (), H. sp. ‘rex’ (), H. sp. ‘simba’ (▼), H. sp. ‘glaucus’ (), H. sp. ‘aquila’ (), H. sp. ‘kimondo’ (●), H. sp. ‘falcatus’ (), H. sp. ‘curvidens’ (), H. sp. ‘pardus’ (), H. sp. ‘quasimodo’ (▲), and H. squamipinnis Regan, 1921 (█). The type localities of H. mentatus and H. squamipinnis are not shown as the exact locations are unknown." captionText-1="Fig. 2. a. Differences in total number of outer teeth and premaxillary dentition between the macro- and microdontic groups of piscivores from the Lake Edward system.b. Schematic presentation of the dominant male colour patterns in life, in the various groups studied. For H. sp. ‘latifrons’ the female and for H. sp. ‘curvidens’ the non-dominant colour patterns are shown, as those of dominant males are unknown." figureDoi-0="http://doi.org/10.5281/zenodo.6484155" figureDoi-1="http://doi.org/10.5281/zenodo.6484157" httpUri-0="https://zenodo.org/record/6484155/files/figure.png" httpUri-1="https://zenodo.org/record/6484157/files/figure.png" pageId="13" pageNumber="14">Figs 1–2</figureCitation>
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,
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<figureCitation id="131D2A35610ADA55FCD951742513FA6B" box="[779,826,1606,1633]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="14.[189,232,1963,1989]" captionStartId-1="15.[189,232,1892,1918]" captionStartId-2="16.[190,233,1963,1989]" captionTargetBox-0="[189,1398,1298,1854]" captionTargetBox-1="[189,1398,287,1830]" captionTargetBox-2="[203,1382,975,1878]" captionTargetId-0="figure-19@14.[189,1399,1298,1854]" captionTargetId-1="figure-12@15.[186,1402,287,1830]" captionTargetId-2="figure-18@16.[203,1384,975,1880]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="Fig. 5. Haplochromis latifrons sp. nov., holotype, ♀, 158.2 mm SL (RMCA 2018.008.P.0330). Drawn by N. Vranken." captionText-1="Fig. 6. Haplochromis latifrons sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0330; 158.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Holotype. d. Another female (RMCA 2017.006.P.0340; 105.4 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." captionText-2="Fig. 7. Haplochromis latifrons sp. nov. (RMCA 2018.008.P.0329; 155.7 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi-0="http://doi.org/10.5281/zenodo.6484169" figureDoi-1="http://doi.org/10.5281/zenodo.6484173" figureDoi-2="http://doi.org/10.5281/zenodo.6484177" httpUri-0="https://zenodo.org/record/6484169/files/figure.png" httpUri-1="https://zenodo.org/record/6484173/files/figure.png" httpUri-2="https://zenodo.org/record/6484177/files/figure.png" pageId="13" pageNumber="14">5–7</figureCitation>
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;
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<tableCitation id="C6A4030B610ADA55FC94517425B4FA6A" box="[838,925,1606,1632]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="13" pageNumber="14" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C653B610ADA56FF6F51B524EEFE1F" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="8B9936B0610ADA55FF6F51B527E0FAAB" blockId="13.[189,457,1671,1697]" box="[189,457,1671,1697]" pageId="13" pageNumber="14">
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<heading id="D0D181DC610ADA55FF6F51B527E0FAAB" bold="true" box="[189,457,1671,1697]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
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<emphasis id="B952EAA2610ADA55FF6F51B527E0FAAB" bold="true" box="[189,457,1671,1697]" pageId="13" pageNumber="14">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9936B0610ADA55FF6F518424BDFB30" blockId="13.[189,1398,1718,1850]" pageId="13" pageNumber="14">Species with a piscivorous morphology; body very shallow [BD 27.2–30.1 (mean 28.6) % SL]; interorbital area flat and broad [IOW 57.4–63.3 (60.0) % HW]; outer oral teeth few and large [UOT 24–42 (median 31)]; females green dorsally, white ventrally, and with a well-defined mid-lateral band; dominant male colour pattern unknown.</paragraph>
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<paragraph id="8B9936B0610ADA55FF6F5053231DFBEF" blockId="13.[188,1398,1889,2022]" pageId="13" pageNumber="14">
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Amongst piscivorous species from the Lake Edward system,
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<taxonomicName id="4C264D33610ADA55FC5250502221FB71" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[896,1032,1889,1916]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
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<emphasis id="B952EAA2610ADA55FC52505025BCFB71" box="[896,917,1890,1915]" italics="true" pageId="13" pageNumber="14">H</emphasis>
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.
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<emphasis id="B952EAA2610ADA55FC7650532221FB71" box="[932,1032,1889,1915]" italics="true" pageId="13" pageNumber="14">latifrons</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157D9610ADA55FBDF5050224EFB76" box="[1037,1127,1890,1916]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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differs from all, except
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<taxonomicName id="4C264D33610ADA55FF6E50B72762FB94" authorityName="Regan" authorityYear="1925" box="[188,331,1925,1951]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="mentatus">
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<emphasis id="B952EAA2610ADA55FF6E50B726F8FB94" box="[188,209,1925,1950]" italics="true" pageId="13" pageNumber="14">H</emphasis>
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.
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<emphasis id="B952EAA2610ADA55FF3250B72762FB94" box="[224,331,1925,1950]" italics="true" pageId="13" pageNumber="14">mentatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C264D33610ADA55FE5150B72424FB94" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[387,525,1924,1951]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
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<emphasis id="B952EAA2610ADA55FE5150B727B1FB94" box="[387,408,1925,1950]" italics="true" pageId="13" pageNumber="14">H</emphasis>
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.
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<emphasis id="B952EAA2610ADA55FE7550B62424FB94" box="[423,525,1924,1950]" italics="true" pageId="13" pageNumber="14">kimondo</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D9610ADA55FDC150B72459FB95" box="[531,624,1925,1951]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, by the combination of a broader interorbital area [IOW 57.4–63.3 (60.0) vs 39.3–57.1 (43.9–53.8) % HW], a shorter anal fin base [AFB 14.7–17.3 (15.7) vs 17.1–22.2 (18.0–20.5) % SL], and a smaller number of branched anal-fin rays [AFRr 7–8 vs 9–11, rarely 8].
|
||
</paragraph>
|
||
<paragraph id="8B9936B06109DA56FF6F563B220AFD86" blockId="14.[189,1400,265,397]" pageId="14" pageNumber="15">
|
||
It differs from
|
||
<taxonomicName id="4C264D336109DA56FEA356382429FD29" authorityName="Regan" authorityYear="1925" box="[369,512,265,291]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="mentatus">
|
||
<emphasis id="B952EAA26109DA56FEA3563827ACFD29" box="[369,389,266,291]" italics="true" pageId="14" pageNumber="15">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26109DA56FE4756382429FD29" box="[405,512,266,291]" italics="true" pageId="14" pageNumber="15">mentatus</emphasis>
|
||
</taxonomicName>
|
||
by the combination of a shorter dorsal-fin base [DFB 47.2–50.1 (49.0) vs 50.3–54.2 (52.3) % SL], a strongly vs weakly prominent premaxillary pedicel, a gentler sloping lower jaw side (25–30° vs 30–45°), juveniles and females green dorsally and white ventrally vs uniformly yellow-green, and presence vs absence of a well-defined mid-lateral band.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06109DA56FF6F568724EEFE1F" blockId="14.[189,1398,436,533]" pageId="14" pageNumber="15">
|
||
It differs from
|
||
<taxonomicName id="4C264D336109DA56FEA656872428FDC4" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[372,513,436,463]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="kimondo" status="sp. nov.">
|
||
<emphasis id="B952EAA26109DA56FEA6568727A0FDC4" box="[372,393,437,462]" italics="true" pageId="14" pageNumber="15">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26109DA56FE4956862428FDC4" box="[411,513,436,462]" italics="true" pageId="14" pageNumber="15">kimondo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96109DA56FDDE56872443FDC5" box="[524,618,437,463]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the combination of large vs small outer oral teeth, a smaller number of outer upper jaw teeth [UOT 24–42 (31) vs 43–70 (56)], and a shorter anal fin base [AFB 14.7–17.3 (15.7) vs 17.0–19.2 (18.0) % SL].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6109DA56FF6F550E2437FEAD" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="8B9936B06109DA56FF6F550E276DFE5C" blockId="14.[189,324,572,598]" box="[189,324,572,598]" pageId="14" pageNumber="15">
|
||
<heading id="D0D181DC6109DA56FF6F550E276DFE5C" bold="true" box="[189,324,572,598]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B952EAA26109DA56FF6F550E276DFE5C" bold="true" box="[189,324,572,598]" pageId="14" pageNumber="15">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06109DA56FF6F55582437FEAD" blockId="14.[189,1398,618,680]" pageId="14" pageNumber="15">
|
||
Specific name from Latin ‘
|
||
<emphasis id="B952EAA26109DA56FE2655582407FE8E" box="[500,558,618,644]" italics="true" pageId="14" pageNumber="15">latus</emphasis>
|
||
’ for ‘wide’ and ‘
|
||
<emphasis id="B952EAA26109DA56FD3C55582503FE8E" box="[750,810,618,644]" italics="true" pageId="14" pageNumber="15">frons</emphasis>
|
||
’ for ‘forehead’; referring to very broad interorbital area for a piscivorous species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6109DA56FF6F55FD27CCF8B3" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="8B9936B06109DA56FF6F55FD2783FEE3" blockId="14.[189,426,719,745]" box="[189,426,719,745]" pageId="14" pageNumber="15">
|
||
<heading id="D0D181DC6109DA56FF6F55FD2783FEE3" bold="true" box="[189,426,719,745]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="B952EAA26109DA56FF6F55FD2783FEE3" bold="true" box="[189,426,719,745]" pageId="14" pageNumber="15">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06109DA56FF3855CC25EDFF54" blockId="14.[189,1399,766,862]" pageId="14" pageNumber="15">
|
||
<materialsCitation id="3B4E3CED6109DA56FF3855CC25EDFF54" ID-GBIF-Occurrence="3861203404" collectingDate="2018-01-24" collectionCode="RMCA" country="Uganda" latitude="-0.40444446" location="Lake Edward" longLatPrecision="1" longitude="29.773556" pageId="14" pageNumber="15" specimenCode="RMCA 2018.008.P.0330" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<emphasis id="B952EAA26109DA56FF3855CC2775FF12" bold="true" box="[234,348,766,792]" pageId="14" pageNumber="15">
|
||
<typeStatus id="549D88126109DA56FF3855CC2775FF12" box="[234,348,766,792]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33176206109DA56FF6F54132713FF31" box="[189,314,801,827]" name="Uganda" pageId="14" pageNumber="15">UGANDA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D20FD396109DA56FE8154132740FF31" box="[339,361,801,827]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556109DA56FEAA541327C6FF31" box="[376,495,801,828]" metricMagnitude="-1" metricUnit="m" metricValue="1.5819999999999999" pageId="14" pageNumber="15" unit="mm" value="158.2">158.2 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B6109DA56FDFA541324E0FF31" LSID="urn:lsid:plazi:treatment:038F87A6610ADA49FD8E52CD2781FACE:8EF9606B6109DA56FDFA541324E0FF31" box="[552,713,801,827]" country="Uganda" latitude="-0.40444446" longLatPrecision="1" longitude="29.773556" name="Lake Edward" pageId="14" pageNumber="15">Lake Edward</location>
|
||
;
|
||
<geoCoordinate id="EE1250776109DA56FD06542D2542FF31" box="[724,875,799,827]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="16.0" value="-0.40444446">0°24′16.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776109DA56FCAA542D2208FF31" box="[888,1057,799,827]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="24.8" value="29.773556">29°46′24.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706109DA56FBFF541322EFFF31" box="[1069,1222,801,827]" pageId="14" pageNumber="15" value="2018-01-24">
|
||
<collectingDate id="EFDCE9986109DA56FBFF541322EFFF31" box="[1069,1222,801,827]" pageId="14" pageNumber="15" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
; HIPE3 exped. leg.; bought at Rwenshama landing site;
|
||
<specimenCode id="DB809ECB6109DA56FD4954762596FF54" box="[667,959,836,862]" pageId="14" pageNumber="15">RMCA 2018.008.P.0330</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06109DA56FF3854B727CCF8B3" blockId="14.[189,1399,901,1209]" pageId="14" pageNumber="15">
|
||
<materialsCitation id="3B4E3CED6109DA56FF3854B7230BFFEC" ID-GBIF-Occurrence="3861203332" collectingDate="2016-10-23" collectionCode="RMCA" collectorName="HIPE 1 exped." country="Uganda" elevation="-20" latitude="-0.2" location="Lake Edward" longLatPrecision="1" longitude="29.793999" pageId="14" pageNumber="15" specimenCode="RMCA 2016.035.P.0199, RMCA 2016.035.P.0200, RMCA 2016.035.P.0201" specimenCount="3" specimenCount-female="3" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<emphasis id="B952EAA26109DA56FF3854B7274EFF95" bold="true" box="[234,359,901,927]" pageId="14" pageNumber="15">
|
||
<typeStatus id="549D88126109DA56FF3854B7274EFF95" box="[234,359,901,927]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F33176206109DA56FF6F549B2713FFC9" box="[189,314,937,963]" name="Uganda" pageId="14" pageNumber="15">UGANDA</collectingCountry>
|
||
–
|
||
<emphasis id="B952EAA26109DA56FE8F549B2427FFC9" bold="true" box="[349,526,937,963]" pageId="14" pageNumber="15">
|
||
<collectingRegion id="49E2F8526109DA56FE8F549B2427FFC9" box="[349,526,937,963]" pageId="14" pageNumber="15">Lake Edward</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D20FD396109DA56FDFE549B245BFFC9" box="[556,626,937,963]" pageId="14" pageNumber="15" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556109DA56FD50549B2564FFC9" box="[642,845,937,963]" metricMagnitude="-1" metricUnit="m" metricValue="1.2885" metricValueMax="1.349" metricValueMin="1.228" pageId="14" pageNumber="15" unit="mm" value="128.85" valueMax="134.9" valueMin="122.8">122.8–134.9 mm</quantity>
|
||
SL;
|
||
<geoCoordinate id="EE1250776109DA56FC595495220CFFC9" box="[907,1061,935,963]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="00.0" value="-0.2">0°12′00.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776109DA56FBE7549522C8FFC9" box="[1077,1249,935,963]" degrees="29" direction="east" minutes="47" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="38.4" value="29.793999">29°47′38.4″ E</geoCoordinate>
|
||
; deep catch, open water ±
|
||
<quantity id="4CDE9B556109DA56FEB454FE278DFFEC" box="[358,420,972,998]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="20.0">
|
||
<elevation id="000BD1836109DA56FEB454FE278DFFEC" box="[358,420,972,998]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="20.0">20 m</elevation>
|
||
</quantity>
|
||
deep;
|
||
<date id="FF9810706109DA56FE2A54FE24B3FFEC" box="[504,666,972,998]" pageId="14" pageNumber="15" value="2016-10-23">
|
||
<collectingDate id="EFDCE9986109DA56FE2A54FE24B3FFEC" box="[504,666,972,998]" pageId="14" pageNumber="15" value="2016-10-23">23 Oct. 2016</collectingDate>
|
||
</date>
|
||
; HIPE1 exped. leg.;
|
||
<specimenCode id="DB809ECB6109DA56FC4A54FE2295FFED" box="[920,1212,972,999]" pageId="14" pageNumber="15">RMCA 2016.035.P.0199</specimenCode>
|
||
to
|
||
<specimenCode id="DB809ECB6109DA56FB3554FE230BFFEC" box="[1255,1314,972,998]" pageId="14" pageNumber="15">0201</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6109DA56FA9254FE2408F826" ID-GBIF-Occurrence="3861203302" collectingDate="2016-10-23" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" elevation="20" latitude="-0.2" location="Rwenshama" longLatPrecision="1" longitude="29.793999" pageId="14" pageNumber="15" specimenCode="RMCA 2017.006.P.0340" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396109DA56FA9254FE2346FFEC" box="[1344,1391,972,998]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556109DA56FF6F54DD271EF800" box="[189,311,1007,1034]" metricMagnitude="-1" metricUnit="m" metricValue="1.054" pageId="14" pageNumber="15" unit="mm" value="105.4">105.4 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B6109DA56FEA454DD2421F803" LSID="urn:lsid:plazi:treatment:038F87A6610ADA49FD8E52CD2781FACE:8EF9606B6109DA56FEA454DD2421F803" box="[374,520,1007,1033]" country="Uganda" latitude="-0.40158334" longLatPrecision="1" longitude="29.776417" name="Rwenshama" pageId="14" pageNumber="15" stateProvince="Lake Edward">Rwenshama</location>
|
||
, rocky shore;
|
||
<geoCoordinate id="EE1250776109DA56FD6754DF2566F803" box="[693,847,1005,1033]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="05.7" value="-0.40158334">0°24′05.7″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776109DA56FC8D54DF2223F800" box="[863,1034,1005,1034]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="35.1" value="29.776417">29°46′35.1″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706109DA56FBC854DD22E9F803" box="[1050,1216,1007,1034]" pageId="14" pageNumber="15" value="2017-03-25">
|
||
<collectingDate id="EFDCE9986109DA56FBC854DD22E9F803" box="[1050,1216,1007,1034]" pageId="14" pageNumber="15" value="2017-03-25">25 Mar. 2017</collectingDate>
|
||
</date>
|
||
; HIPE2 exped. leg.;
|
||
<specimenCode id="DB809ECB6109DA56FF2E53202408F826" box="[252,545,1042,1068]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="14" pageNumber="15" type="Museum">RMCA 2017.006.P.0340</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6109DA56FD91532025A7F845" ID-GBIF-Occurrence="3861203397" collectingDate="2017-03-31" collectionCode="RMCA" collectorName="HIPE 2 exped." country="Uganda" elevation="20" latitude="-0.2" location="Kayanja" longLatPrecision="1" longitude="29.793999" pageId="14" pageNumber="15" specimenCode="RMCA 2017.006.P.0341" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396109DA56FD915320245CF827" box="[579,629,1042,1069]" pageId="14" pageNumber="15" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556109DA56FD54532024DAF827" box="[646,755,1042,1069]" metricMagnitude="-2" metricUnit="m" metricValue="8.22" pageId="14" pageNumber="15" unit="mm" value="82.2">82.2 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B6109DA56FCE7532025BEF826" LSID="urn:lsid:plazi:treatment:038F87A6610ADA49FD8E52CD2781FACE:8EF9606B6109DA56FCE7532025BEF826" box="[821,919,1042,1068]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Kayanja" pageId="14" pageNumber="15" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE1250776109DA56FBC55323229AF826" box="[1047,1203,1041,1069]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="34.8" value="-0.093">0°05′34.8″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776109DA56FB1653232358F827" box="[1220,1393,1041,1069]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="28.8" value="29.758">29°45′28.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706109DA56FF6F5307274FF845" box="[189,358,1077,1104]" pageId="14" pageNumber="15" value="2017-03-31">
|
||
<collectingDate id="EFDCE9986109DA56FF6F5307274FF845" box="[189,358,1077,1104]" pageId="14" pageNumber="15" value="2017-03-31">31 Mar. 2017</collectingDate>
|
||
</date>
|
||
; HIPE2 exped. leg.;
|
||
<specimenCode id="DB809ECB6109DA56FDBB530725A7F845" box="[617,910,1077,1103]" pageId="14" pageNumber="15">RMCA 2017.006.P.0341</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6109DA56FC7D5307229AF879" ID-GBIF-Occurrence="3861203386" collectingDate="2018-01-20" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="20" latitude="-0.2" location="Kayanja" longLatPrecision="1" longitude="29.793999" pageId="14" pageNumber="15" specimenCode="RMCA 2018.008.P.0328" specimenCount="1" specimenCount-male="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396109DA56FC7D530725C8F85A" box="[943,993,1077,1104]" pageId="14" pageNumber="15" type="male">1 ♂</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556109DA56FC2053072276F85A" box="[1010,1119,1077,1104]" metricMagnitude="-2" metricUnit="m" metricValue="7.529999999999999" pageId="14" pageNumber="15" unit="mm" value="75.3">75.3 mm</quantity>
|
||
SL;
|
||
<location id="8EF9606B6109DA56FB725307232BF845" LSID="urn:lsid:plazi:treatment:038F87A6610ADA49FD8E52CD2781FACE:8EF9606B6109DA56FB725307232BF845" box="[1184,1282,1077,1103]" country="Uganda" latitude="-0.2" longLatPrecision="1" longitude="29.793999" name="Kayanja" pageId="14" pageNumber="15" stateProvince="Lake Edward">Kayanja</location>
|
||
offshore;
|
||
<geoCoordinate id="EE1250776109DA56FF6F5365277AF879" box="[189,339,1111,1139]" degrees="0" direction="south" minutes="05" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="31.2" value="-0.092">0°05′31.2″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776109DA56FE8C5365242FF879" box="[350,518,1111,1139]" degrees="29" direction="east" minutes="45" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="30.3" value="29.758417">29°45′30.3″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706109DA56FDC3536B248FF879" box="[529,678,1113,1139]" pageId="14" pageNumber="15" value="2018-01-20">
|
||
<collectingDate id="EFDCE9986109DA56FDC3536B248FF879" box="[529,678,1113,1139]" pageId="14" pageNumber="15" value="2018-01-20">20 Jan. 2018</collectingDate>
|
||
</date>
|
||
; HIPE3 exped. leg.;
|
||
<specimenCode id="DB809ECB6109DA56FC46536B229AF879" box="[916,1203,1113,1139]" pageId="14" pageNumber="15">RMCA 2018.008.P.0328</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B4E3CED6109DA56FB1B536B27C8F8B3" ID-GBIF-Occurrence="3861203395" collectingDate="2018-01-24" collectionCode="RMCA" collectorName="HIPE 3 exped." country="Uganda" elevation="20" latitude="-0.2" location="Rwenshama" longLatPrecision="1" longitude="29.793999" pageId="14" pageNumber="15" specimenCode="RMCA 2018.008.P.0329" specimenCount="1" specimenCount-female="1" stateProvince="Lake Edward" typeStatus="paratype">
|
||
<specimenCount id="9D20FD396109DA56FB1B536B22DDF879" box="[1225,1268,1113,1139]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
|
||
,
|
||
<quantity id="4CDE9B556109DA56FAD3536A235EF879" box="[1281,1399,1112,1139]" metricMagnitude="-1" metricUnit="m" metricValue="1.557" pageId="14" pageNumber="15" unit="mm" value="155.7">155.7 mm</quantity>
|
||
SL;
|
||
<geoCoordinate id="EE1250776109DA56FF3D534827AEF89C" box="[239,391,1146,1174]" degrees="0" direction="south" minutes="24" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="16.0" value="-0.40444446">0°24′16.0″ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1250776109DA56FE4653482414F89C" box="[404,573,1146,1174]" degrees="29" direction="east" minutes="46" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="24.8" value="29.773556">29°46′24.8″ E</geoCoordinate>
|
||
;
|
||
<date id="FF9810706109DA56FD98534E24CDF89C" box="[586,740,1148,1174]" pageId="14" pageNumber="15" value="2018-01-24">
|
||
<collectingDate id="EFDCE9986109DA56FD98534E24CDF89C" box="[586,740,1148,1174]" pageId="14" pageNumber="15" value="2018-01-24">24 Jan. 2018</collectingDate>
|
||
</date>
|
||
; HIPE3 exped. leg.; bought at
|
||
<location id="8EF9606B6109DA56FB9D534E22F7F89C" LSID="urn:lsid:plazi:treatment:038F87A6610ADA49FD8E52CD2781FACE:8EF9606B6109DA56FB9D534E22F7F89C" box="[1103,1246,1148,1174]" pageId="14" pageNumber="15">Rwenshama</location>
|
||
landing site;
|
||
<specimenCode id="DB809ECB6109DA56FF6F53AD27C8F8B3" box="[189,481,1183,1209]" pageId="14" pageNumber="15">RMCA 2018.008.P.0329</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5966386109DA56FF6F50992759FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484169" ID-Zenodo-Dep="6484169" httpUri="https://zenodo.org/record/6484169/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1963,1989]" targetBox="[189,1398,1298,1854]" targetPageId="14">
|
||
<paragraph id="8B9936B06109DA56FF6F50992759FBE2" blockId="14.[189,1399,1963,2025]" pageId="14" pageNumber="15">
|
||
<emphasis id="B952EAA26109DA56FF6F50992725FBCF" bold="true" box="[189,268,1963,1989]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C264D336109DA56FEC750992403FBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[277,554,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26109DA56FEC750992403FBCF" box="[277,554,1963,1989]" italics="true" pageId="14" pageNumber="15">Haplochromis latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96109DA56FDE1509E24B8FBCC" box="[563,657,1964,1990]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♀, 158.2 mm SL (RMCA 2018.008.P.0330). Drawn by N. Vranken.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5966386108DA57FF6F50562407FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484173" ID-Zenodo-Dep="6484173" httpUri="https://zenodo.org/record/6484173/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1892,1918]" targetBox="[189,1398,287,1830]" targetPageId="15">
|
||
<paragraph id="8B9936B06108DA57FF6F50562407FBE2" blockId="15.[188,1399,1892,2025]" pageId="15" pageNumber="16">
|
||
<emphasis id="B952EAA26108DA57FF6F50562722FB75" bold="true" box="[189,267,1892,1919]" pageId="15" pageNumber="16">Fig. 6.</emphasis>
|
||
<taxonomicName id="4C264D336108DA57FEC15057240EFB75" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[275,551,1893,1919]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26108DA57FEC15057240EFB75" box="[275,551,1893,1919]" italics="true" pageId="15" pageNumber="16">Haplochromis latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96108DA57FDFC505724A0FB75" box="[558,649,1893,1919]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B952EAA26108DA57FD42505724B6FB75" bold="true" box="[656,671,1893,1919]" pageId="15" pageNumber="16">a</emphasis>
|
||
. Photograph of preserved holotype (RMCA 2018.008.P.0330; 158.2 mm SL).
|
||
<emphasis id="B952EAA26108DA57FEA150BA27AAFBA8" bold="true" box="[371,387,1928,1954]" pageId="15" pageNumber="16">b</emphasis>
|
||
. X-ray image of holotype.
|
||
<emphasis id="B952EAA26108DA57FD6C50BA24C2FBA8" bold="true" box="[702,747,1928,1954]" pageId="15" pageNumber="16">c–d</emphasis>
|
||
. Photographs of freshly caught specimens.
|
||
<emphasis id="B952EAA26108DA57FB3450BA22DAFBA8" bold="true" box="[1254,1267,1928,1954]" pageId="15" pageNumber="16">c</emphasis>
|
||
. Holotype.
|
||
<emphasis id="B952EAA26108DA57FF6E509926E5FBCF" bold="true" box="[188,204,1963,1989]" pageId="15" pageNumber="16">d</emphasis>
|
||
. Another female (RMCA 2017.006.P.0340; 105.4 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33C653B6117DA49FF6F563A27FDFA27" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8B9936B06117DA48FF6F563A2767FD28" blockId="16.[189,334,264,290]" box="[189,334,264,290]" pageId="16" pageNumber="17">
|
||
<heading id="D0D181DC6117DA48FF6F563A2767FD28" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="B952EAA26117DA48FF6F563A2767FD28" bold="true" box="[189,334,264,290]" pageId="16" pageNumber="17">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06117DA48FF6F560524A7FEDF" blockId="16.[188,1400,311,725]" pageId="16" pageNumber="17">
|
||
Based on
|
||
<specimenCount id="9D20FD396117DA48FEE0560527EBFD5B" box="[306,450,311,337]" pageId="16" pageNumber="17" type="generic">8 specimens</specimenCount>
|
||
(
|
||
<quantity id="4CDE9B556117DA48FE00560524A7FD58" box="[466,654,311,338]" metricMagnitude="-1" metricUnit="m" metricValue="1.1675" metricValueMax="1.5819999999999999" metricValueMin="0.7529999999999999" pageId="16" pageNumber="17" unit="mm" value="116.75" valueMax="158.2" valueMin="75.3">75.3–158.2 mm</quantity>
|
||
SL); body very shallow (
|
||
<tableCitation id="C6A4030B6117DA48FC695605223AFD5B" box="[955,1043,311,337]" captionStart="Table 1" captionStartId="9.[182,252,548,576]" captionTargetPageId="9" captionText="Table 1. (continued on the next three pages). Measurements and counts of the piscivorous species of Haplochromis from the Lake Edward system." httpUri="http://table.plazi.org/id/DF596638610EDA51FF6455162158FE4A" pageId="16" pageNumber="17" tableUuid="DF596638610EDA51FF6455162158FE4A">Table 1</tableCitation>
|
||
) and oval to pyriform; caudal peduncle long (
|
||
<figureCitation id="131D2A356117DA48FEA8566827ECFD7F" box="[378,453,346,373]" captionStart="Fig" captionStartId="14.[189,232,1963,1989]" captionTargetBox="[189,1398,1298,1854]" captionTargetId="figure-19@14.[189,1399,1298,1854]" captionTargetPageId="14" captionText="Fig. 5. Haplochromis latifrons sp. nov., holotype, ♀, 158.2 mm SL (RMCA 2018.008.P.0330). Drawn by N. Vranken." figureDoi="http://doi.org/10.5281/zenodo.6484169" httpUri="https://zenodo.org/record/6484169/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
|
||
). Head narrow, dorsally flat, and with straight dorsal outline; interorbital area flat and average in width in comparison to generalised
|
||
<taxonomicName id="4C264D336117DA48FC88564C25F2FD9D" authorityName="Trewavas" authorityYear="1933" box="[858,987,381,408]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B952EAA26117DA48FC88564C2546FD9D" box="[858,879,382,407]" italics="true" pageId="16" pageNumber="17">H</emphasis>
|
||
.
|
||
<emphasis id="B952EAA26117DA48FC53564F25F2FD9D" box="[897,987,381,407]" italics="true" pageId="16" pageNumber="17">elegans</emphasis>
|
||
</taxonomicName>
|
||
(but very broad for a piscivorous species); eye very small and high on head; cheek and lacrimal deep. Snout very long, acute, and slopes gently at 30–40°; premaxillary pedicel long and strongly prominent. Jaws isognathous, long, stout, very narrow, and rounded in dorsal view; gape large and slopes gently at 15–25°; maxilla (almost) extends to vertical through anterior margin of orbit. Lower jaw relatively deep and with a weakly convex ventral outline in lateral view, mental prominence absent or weakly developed, and lower jaw side nearly flat with an inclination of 25–30° to horizontal in anterior view. Upper jaw expanded anteriorly. Lips and oral mucosa large. Neurocranium shallow, ethmo-vomerine block decurved, preorbital region very shallow (19–21% NL), orbital region shallow (27–32% NL), and supraoccipital crest very shallow and pyramidical or wedge-shaped (
|
||
<figureCitation id="131D2A356117DA48FDFA558824A9FEDF" box="[552,640,698,725]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,287,1830]" captionTargetId="figure-12@15.[186,1402,287,1830]" captionTargetPageId="15" captionText="Fig. 6. Haplochromis latifrons sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0330; 158.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Holotype. d. Another female (RMCA 2017.006.P.0340; 105.4 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484173" httpUri="https://zenodo.org/record/6484173/files/figure.png" pageId="16" pageNumber="17">Fig. 6b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936B06117DA49FF6F55CE2277FD86" blockId="16.[187,1399,764,896]" lastBlockId="17.[189,1400,265,397]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
Outer oral teeth few, unicuspid, and large. Necks stout, conical, and straight; crowns recurved and acutely pointed. Dental arcades rounded. Outer teeth widely and very irregularly set with neck-distances of 1–5 neck-widths. In upper jaw, 2–3 posteriormost teeth enlarged. Inner teeth relatively large, recurved, unicuspid in large specimens (>
|
||
<quantity id="4CDE9B556117DA48FDEE545724A6FF8A" box="[572,655,869,896]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="17" unit="mm" value="75.0">75 mm</quantity>
|
||
SL), tricuspid in small specimens (<
|
||
<quantity id="4CDE9B556117DA48FB9F54572288FF8A" box="[1101,1185,869,896]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="17" unit="mm" value="75.0">75 mm</quantity>
|
||
SL). Tooth bands very slender crescent-shaped with 1–3 rows of inner teeth and narrow posteriorly until only outer row remains past ⅔ length of tooth band in upper jaw, past ½ length of tooth band in lower jaw. Inner teeth widely and irregularly set on ½–1 outer neck-width from outer row; implantation erect in first row and recumbent in subsequent rows; size decreases slightly buccally and posteriorly.
|
||
</paragraph>
|
||
<caption id="DF5966386117DA48FF6C509925F8FBE2" ID-DOI="http://doi.org/10.5281/zenodo.6484177" ID-Zenodo-Dep="6484177" httpUri="https://zenodo.org/record/6484177/files/figure.png" pageId="16" pageNumber="17" startId="16.[190,233,1963,1989]" targetBox="[203,1382,975,1878]" targetPageId="16">
|
||
<paragraph id="8B9936B06117DA48FF6C509925F8FBE2" blockId="16.[189,1399,1963,2025]" pageId="16" pageNumber="17">
|
||
<emphasis id="B952EAA26117DA48FF6C50992722FBCF" bold="true" box="[190,267,1963,1989]" pageId="16" pageNumber="17">Fig. 7.</emphasis>
|
||
<taxonomicName id="4C264D336117DA48FEC05099240FFBCF" authority="Vranken & Steenberge & Heylen & Decru & Snoeks, 2022" authorityName="Vranken & Steenberge & Heylen & Decru & Snoeks" authorityYear="2022" box="[274,550,1963,1989]" class="Actinopterygii" family="Cichlidae" genus="Haplochromis" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="latifrons" status="sp. nov.">
|
||
<emphasis id="B952EAA26117DA48FEC05099240FFBCF" box="[274,550,1963,1989]" italics="true" pageId="16" pageNumber="17">Haplochromis latifrons</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26157D96117DA48FDFF509E24A1FBCC" box="[557,648,1964,1990]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(RMCA 2018.008.P.0329; 155.7 mm SL).
|
||
<emphasis id="B952EAA26117DA48FB52509922A6FBCF" bold="true" box="[1152,1167,1963,1989]" pageId="16" pageNumber="17">a</emphasis>
|
||
. Dorsal view of the lower pharyngeal jaw.
|
||
<emphasis id="B952EAA26117DA48FE1A50FC27F1FBE2" bold="true" box="[456,472,1998,2024]" pageId="16" pageNumber="17">b</emphasis>
|
||
. Lateral view of the lower pharyngeal jaw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9936B06116DA49FF6F5687230CFE51" blockId="17.[189,1401,436,603]" pageId="17" pageNumber="18">
|
||
Lower pharyngeal bone long, narrow, slim, and shallow over entire length (
|
||
<figureCitation id="131D2A356116DA49FB9F568622BEFDC5" box="[1101,1175,436,463]" captionStart="Fig" captionStartId="16.[190,233,1963,1989]" captionTargetBox="[203,1382,975,1878]" captionTargetId="figure-18@16.[203,1384,975,1880]" captionTargetPageId="16" captionText="Fig. 7. Haplochromis latifrons sp. nov. (RMCA 2018.008.P.0329; 155.7 mm SL). a. Dorsal view of the lower pharyngeal jaw. b. Lateral view of the lower pharyngeal jaw." figureDoi="http://doi.org/10.5281/zenodo.6484177" httpUri="https://zenodo.org/record/6484177/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
|
||
). Pharyngeal teeth relatively large and slender; major cusps acutely pointed; cusp gaps concave; minor cusps and cusp protuberances very small. Teeth in two median longitudinal rows equal in size and form to lateral teeth, about
|
||
<quantity id="4CDE9B556116DA49FEDA552C276FFE32" box="[264,326,542,568]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="18" unit="in" value="10.0">10 in</quantity>
|
||
each row. Posterior transverse row with 15–16 teeth, implanted recumbently with lateral inclination; major cusps recurved, bluntly pointed, and laterally compressed; minor cusps small.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F55B0235EFE96" blockId="17.[189,1399,642,669]" box="[189,1399,642,669]" pageId="17" pageNumber="18">Chest scales small; transition to larger flank scales gradual. Minute scales on proximal half of caudal fin.</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F55F6255BFF2E" blockId="17.[189,1399,708,805]" pageId="17" pageNumber="18">Caudal fin emarginate; dorsal and anal fins reach to vertical through 2–4 scales anterior to caudal-fin base. Pectoral and pelvic fins reach to just anterior to genital opening in females, unknown in males; first branched pelvic-fin ray slightly elongated in all specimens.</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F5479221DFF83" blockId="17.[189,1399,843,905]" pageId="17" pageNumber="18">Ceratobranchial gill rakers in outer row of first gill arch short, stout, and simple; posteriormost rakers simple to trifid or anvil-shaped. Epibranchial gill rakers slender and simple.</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F5482278AFFC0" blockId="17.[189,419,944,970]" box="[189,419,944,970]" pageId="17" pageNumber="18">
|
||
<heading id="D0D181DC6116DA49FF6F5482278AFFC0" bold="true" box="[189,419,944,970]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B952EAA26116DA49FF6F5482278AFFC0" bold="true" box="[189,419,944,970]" pageId="17" pageNumber="18">Colouration in life</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F54ED2487FFF3" blockId="17.[189,686,991,1017]" box="[189,686,991,1017]" pageId="17" pageNumber="18">Dominant males: colour pattern unknown.</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F531222F1F93B" blockId="17.[189,1400,1056,1329]" pageId="17" pageNumber="18">
|
||
Females and juveniles: dorsal parts of body and operculum olive-green; ventral parts of body and operculum white; abrupt transition at height of lower lateral line (
|
||
<figureCitation id="131D2A356116DA49FC135371221CF854" box="[961,1077,1091,1118]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,287,1830]" captionTargetId="figure-12@15.[186,1402,287,1830]" captionTargetPageId="15" captionText="Fig. 6. Haplochromis latifrons sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0330; 158.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Holotype. d. Another female (RMCA 2017.006.P.0340; 105.4 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484173" httpUri="https://zenodo.org/record/6484173/files/figure.png" pageId="17" pageNumber="18">Fig. 6c–d</figureCitation>
|
||
). Flank with a well-defined black mid-lateral band from posterior margin of eye to caudal-fin base. Dorsum with a blue sheen; cheek and lower jaw white; lacrimal, snout, and lips dusky olive-green; eye with dark grey outer ring and golden inner ring. Nostril and interorbital stripes faint; lacrimal blotch large anteroventrally of eye. Pectoral fin yellowish; pelvic fin hyaline with a greenish sheen; dorsal and caudal fins uniformly dusky, dorsal fin with black lappets (i.e., distal extensions of membrane between spines); anal fin with a hyaline-white base, a dusky-yellow distal part, and 1–2 small spots resembling egg-spots.
|
||
</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F526A27F9F978" blockId="17.[189,464,1368,1394]" box="[189,464,1368,1394]" pageId="17" pageNumber="18">
|
||
<heading id="D0D181DC6116DA49FF6F526A27F9F978" bold="true" box="[189,464,1368,1394]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B952EAA26116DA49FF6F526A27F9F978" bold="true" box="[189,464,1368,1394]" pageId="17" pageNumber="18">Preserved colouration</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F52B427FDFA27" blockId="17.[189,1401,1414,1581]" pageId="17" pageNumber="18">
|
||
Females and juveniles: dorsal part of body brown; ventral part of body, cheek, and lower jaw white; lacrimal, snout and lips dusky (
|
||
<figureCitation id="131D2A356116DA49FDE0529824A0F9CF" box="[562,649,1450,1477]" captionStart="Fig" captionStartId="15.[189,232,1892,1918]" captionTargetBox="[189,1398,287,1830]" captionTargetId="figure-12@15.[186,1402,287,1830]" captionTargetPageId="15" captionText="Fig. 6. Haplochromis latifrons sp. nov. a. Photograph of preserved holotype (RMCA 2018.008.P.0330; 158.2 mm SL). b. X-ray image of holotype. c–d. Photographs of freshly caught specimens. c. Holotype. d. Another female (RMCA 2017.006.P.0340; 105.4 mm SL) to illustrate the live colour patterns. The contrast was slightly enhanced." figureDoi="http://doi.org/10.5281/zenodo.6484173" httpUri="https://zenodo.org/record/6484173/files/figure.png" pageId="17" pageNumber="18">Fig. 6a</figureCitation>
|
||
). Flank with a mid-lateral band from posterior margin of eye to caudal-fin base. Nostril and interorbital stripes faint, lacrimal blotch present. Pectoral fin hyaline; pelvic fin white; dorsal and caudal fins uniformly dusky, dorsal fin with black lappets; anal fin with a white base and dusky distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C653B6116DA49FF6F51662781FACE" pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph id="8B9936B06116DA49FF6F516627D8FA65" blockId="17.[189,497,1620,1647]" box="[189,497,1620,1647]" pageId="17" pageNumber="18">
|
||
<heading id="D0D181DC6116DA49FF6F516627D8FA65" bold="true" box="[189,497,1620,1647]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="B952EAA26116DA49FF6F516627D8FA65" bold="true" box="[189,497,1620,1647]" pageId="17" pageNumber="18">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9936B06116DA49FF6F51B12781FACE" blockId="17.[189,1398,1667,1732]" pageId="17" pageNumber="18">Only known from Lake Edward, found in offshore areas. Based on its morphology, most probably a piscivorous species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |