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<document ID-DOI="http://doi.org/10.5281/zenodo.3792550" ID-GBIF-Dataset="7112ded3-fa31-4c7b-a093-4d3dedafd571" ID-GBIF-Taxon="163755064" ID-ISSN="13132970" ID-Zenodo-Dep="3792550" approvalRequired="4" approvalRequired_for_bibRefs="2" approvalRequired_for_treatments="2" checkinTime="1587999565776" checkinUser="plazi" docAuthor="Solodovnikov, Alexey" docDate="2009" docId="038F87A1FFC7AA7C01CCFC3FFEF16E01" docLanguage="en" docName="ZK_article_1965.pdf" docOrigin="ZooKeys 5 (5)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@60a754c6" docStyleName="zookeys.2008.journal_article" docTitle="Philonthus neoniger Solodovnikov 2009, nom. nov." docType="treatment" docVersion="6" lastPageId="5" lastPageNumber="80" masterDocId="FFB6FFD9FFC5AA790140FFF0FFA66F4C" masterDocTitle="Xanthopygoides niger Cameron, 1951 (Xanthopygina) belongs to the genus Philonthus Stephens, 1829 (Philonthina): systematic and nomenclatural changes for the African Staphylinini (Coleoptera, Staphylinidae, Staphylininae, Staphylinini)" masterLastPageNumber="80" masterPageNumber="75" pageId="2" pageNumber="77" updateTime="1643568035591" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Xanthopygoides niger Cameron, 1951 (Xanthopygina) belongs to the genus Philonthus Stephens, 1829 (Philonthina): systematic and nomenclatural changes for the African Staphylinini (Coleoptera, Staphylinidae, Staphylininae, Staphylinini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solodovnikov, Alexey</mods:namePart>
<mods:affiliation>Natural History Museum of Denmark, Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-02-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>80</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.5.59</mods:identifier>
<mods:identifier type="GBIF-Dataset">7112ded3-fa31-4c7b-a093-4d3dedafd571</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576432</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792550" ID-GBIF-Taxon="163755064" ID-Zenodo-Dep="3792550" LSID="urn:lsid:plazi:treatment:038F87A1FFC7AA7C01CCFC3FFEF16E01" httpUri="http://treatment.plazi.org/id/038F87A1FFC7AA7C01CCFC3FFEF16E01" lastPageId="5" lastPageNumber="80" pageId="2" pageNumber="77">
<subSubSection box="[140,682,975,1002]" pageId="2" pageNumber="77" type="nomenclature">
<paragraph blockId="2.[140,1107,975,1108]" box="[140,682,975,1002]" pageId="2" pageNumber="77">
<heading box="[140,682,975,1002]" fontSize="11" level="2" pageId="2" pageNumber="77" reason="3">
<emphasis bold="true" box="[140,682,975,1002]" pageId="2" pageNumber="77">
<taxonomicName authority="Solodovnikov" authorityName="Solodovnikov" authorityYear="2009" box="[140,550,975,1002]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="neoniger" status="nom. nov.">
<emphasis bold="true" box="[140,378,975,1002]" italics="true" pageId="2" pageNumber="77">
<emphasis box="[140,269,975,1002]" inLineHeading="true" pageId="2" pageNumber="77" reason="1">Philonthus</emphasis>
neoniger
</emphasis>
Solodovnikov
</taxonomicName>
,
<taxonomicNameLabel box="[563,682,976,1002]" pageId="2" pageNumber="77">nom. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="77" type="reference_group">
<paragraph blockId="2.[140,1107,975,1108]" pageId="2" pageNumber="77">
=
<taxonomicName authority="(Cameron, 1948)" authorityName="Solodovnikov" authorityYear="2009" baseAuthorityName="Cameron" baseAuthorityYear="1948" box="[163,553,1011,1037]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="niger" status="comb. nov.">
<emphasis box="[163,342,1011,1037]" italics="true" pageId="2" pageNumber="77">Philonthus niger</emphasis>
(Cameron, 1948)
</taxonomicName>
,
<emphasis bold="true" box="[566,698,1011,1037]" pageId="2" pageNumber="77">
<taxonomicNameLabel box="[566,698,1011,1037]" pageId="2" pageNumber="77" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
for
<taxonomicName authority="(Cameron, 1951)" baseAuthorityName="Cameron" baseAuthorityYear="1951" class="Insecta" family="Staphylinidae" genus="Xanthopygoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[748,976,1011,1037]" italics="true" pageId="2" pageNumber="77">Xanthopygoides niger</emphasis>
(
<bibRefCitation author="Cameron M" pageId="2" pageNumber="77" pagination="398 - 407" refId="ref2285" refString="Cameron M (1951) New species of African Staphylinidae. Part II. Journal of the East Africa Natural History Society 19 (1950): 398 - 407." type="journal article" year="1951">Cameron, 1951</bibRefCitation>
)
</taxonomicName>
, secondary homonym of
<taxonomicName authority="(O. Muller, 1764)" baseAuthorityName="O. Muller" baseAuthorityYear="1764" box="[497,890,1046,1073]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[497,675,1046,1072]" italics="true" pageId="2" pageNumber="77">Philonthus niger</emphasis>
(O. Müller, 1764)
</taxonomicName>
; the latter is a nonvalid senior synonym of
<taxonomicName authority="(Fabricius, 1793)" authorityName="Herman" authorityYear="2001" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[417,845,1082,1108]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="splendens">
<emphasis box="[417,640,1082,1108]" italics="true" pageId="2" pageNumber="77">Philonthus splendens</emphasis>
(Fabricius, 1793)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="77" type="materials_examined">
<paragraph blockId="2.[140,1108,1151,1672]" pageId="2" pageNumber="77">
<emphasis bold="true" box="[140,436,1152,1178]" pageId="2" pageNumber="77">
<emphasis box="[140,198,1152,1178]" inLineHeading="true" pageId="2" pageNumber="77" reason="1">Type</emphasis>
material examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608994673" collectionCode="BM" elevation="1646" location="Natural History Museum in London" pageId="2" pageNumber="77" specimenCode="BM 1952-575" specimenCount="1" typeStatus="holotype">
Male:
<typeStatus box="[517,628,1152,1178]" pageId="2" pageNumber="77">Holotype</typeStatus>
: “[red margined circle]/ Emali Range Sultan Hamud
<quantity box="[285,448,1187,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.64592" metricValueMax="1.79832" metricValueMin="1.49352" pageId="2" pageNumber="77" unit="ft" value="5400.0" valueMax="5900.0" valueMin="4900.0">
<elevation box="[285,448,1187,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.64592" metricValueMax="1.79832" metricValueMin="1.49352" pageId="2" pageNumber="77" unit="ft" value="5400.0" valueMax="5900.0" valueMin="4900.0">4900-5900 ft.</elevation>
</quantity>
3-40/
<taxonomicName authorityName="Cameron" authorityYear="1951" box="[532,728,1188,1213]" class="Insecta" family="Heterogastridae" genus="Heterogaster" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[532,728,1188,1213]" italics="true" pageId="2" pageNumber="77">Heterogaster niger</emphasis>
</taxonomicName>
Type Cam. [handwritten label]/ Pres. by Com. Inst. Ent.
<specimenCode box="[421,586,1222,1249]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="2" pageNumber="77">BM 1952-575</specimenCode>
” (
<location LSID="urn:lsid:plazi:treatment:038F87A1FFC7AA7C01CCFC3FFEF16E01:8EF9606CFFC7AA7B0324FB36FBAD6BAD" box="[612,1035,1222,1249]" name="Natural History Museum in London" pageId="2" pageNumber="77">Natural History Museum in London</location>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="79" pageId="2" pageNumber="77" type="description">
<paragraph blockId="2.[140,1108,1151,1672]" pageId="2" pageNumber="77">
<emphasis bold="true" box="[185,360,1257,1283]" pageId="2" pageNumber="77">
<emphasis box="[185,354,1257,1283]" inLineHeading="true" pageId="2" pageNumber="77" reason="1">Redescription</emphasis>
.
</emphasis>
Body dark blackish brown (piceous). Mouthparts, tarsi, two basal segments of antennae paler, variously brownish. Bases of elytra from shoulder to base of scutellum (part mostly hidden under base of pronotum) yellowish. Head and pronotum smooth and glossy, elytra and abdomen dull, with rugose punctation. Body length
<quantity box="[219,305,1399,1425]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="77" unit="mm" value="12.0">12 mm</quantity>
. Habitus as in
<figureCitation box="[473,539,1398,1425]" captionStart="Fig" captionStartId="3.[140,174,1073,1095]" captionTargetBox="[395,850,171,1040]" captionTargetId="figure@3.[371,876,162,1049]" captionTargetPageId="3" captionText="Fig. 1. Philonthus neoniger Solodovnikov, nom. nov. Holotype. Habitus. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787017" httpUri="https://zenodo.org/record/3787017/files/figure.png" pageId="2" pageNumber="77">Fig. 1</figureCitation>
.
</paragraph>
<paragraph blockId="2.[140,1108,1151,1672]" lastBlockId="3.[140,1108,1152,1671]" lastPageId="3" lastPageNumber="78" pageId="2" pageNumber="77">
Head slightly transverse (width to length ratio 1.14), parallel-sided behind eyes, with broadly rounded but still distinct posterior angles. Eyes slightly shorter than temporae which, measured from posterior margin of eye to neck constriction, 1.32 times as long as eye. Surface of head glossy; transverse wavy microsculpture very distinct on frons only, hardly visible elsewhere. Temporae covered by numerous smaller and few large setiferous punctures. Two setiferous punctures on frons between eyes. Antennae (not entirely preserved in the
<typeStatus box="[478,583,1645,1672]" pageId="2" pageNumber="77">holotype</typeStatus>
): antennomere I slightly longer than antenno-mere II, antennomeres II and III about the same length; antennomeres IV-VII about as long as wide; antennomeres I-III with setae only, IV-VII (and presumably IV-XI) with setae and dense pubescence.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3787017" ID-Zenodo-Dep="3787017" box="[140,966,1073,1095]" httpUri="https://zenodo.org/record/3787017/files/figure.png" pageId="3" pageNumber="78" startId="3.[140,174,1073,1095]" targetBox="[395,850,171,1040]" targetPageId="3">
<paragraph blockId="3.[140,966,1073,1095]" box="[140,966,1073,1095]" pageId="3" pageNumber="78">
<emphasis bold="true" box="[140,204,1073,1095]" pageId="3" pageNumber="78">Fig. 1.</emphasis>
<taxonomicName authority="Solodovnikov" authorityName="Solodovnikov" authorityYear="2009" box="[210,519,1073,1095]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="neoniger" status="nom. nov.">
<emphasis box="[210,386,1073,1095]" italics="true" pageId="3" pageNumber="78">Philonthus neoniger</emphasis>
Solodovnikov
</taxonomicName>
,
<taxonomicNameLabel box="[529,624,1074,1095]" pageId="3" pageNumber="78">nom. nov.</taxonomicNameLabel>
<typeStatus box="[628,719,1073,1095]" pageId="3" pageNumber="78">Holotype</typeStatus>
. Habitus. Scale bar
<quantity box="[903,962,1073,1095]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="78" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph blockId="3.[140,1108,1152,1671]" pageId="3" pageNumber="78">Pronotum slightly longer than wide (length to width ratio 1.08) and slightly wider than head (pronotum width to head width ratio 1.08), with very distinct anterior angles and distinct but more rounded posterior angles, widest in the anterior third of its length; pronotal sides slightly converging and feebly sinuate in front and behind the maximal width of pronotum. Surface of pronotum smooth and glossy with hardly visible irregular miscrosculpture and micropunctation. Disk of pronotum with only one puncture in each dorsal “row”.</paragraph>
<paragraph blockId="3.[140,1108,1152,1671]" box="[185,728,1504,1531]" pageId="3" pageNumber="78">Scutellum closely, moderately roughly punctate.</paragraph>
<paragraph blockId="3.[140,1108,1152,1671]" pageId="3" pageNumber="78">Elytra longer and distinctly wider than pronotum, diverging posteriad (elytral width to length ratio 1.05). Their surface closely, strongly and coarsely punctate; interstices very narrow with visible irregular microsculpture. Pubescence of elytra short, close, brownish.</paragraph>
<paragraph blockId="4.[140,1109,166,897]" pageId="4" pageNumber="79">Abdomen very closely, evenly and rather finely punctate, its pubescence as on elytra. Abdominal tergites III-V with two (basal and subbasal) straight carinae; tergites VI-VII only with one (basal) straight carina. Apical margin of tergite VII with whitish seam.</paragraph>
<paragraph blockId="4.[140,1109,166,897]" pageId="4" pageNumber="79">Legs with tibiae pubescent and spinose. Anterior tarsi of male dilated (female unknown), dorsally setose, ventrally with dense cover of whitish adhesive setae. Middle and posterior tarsi simple, as long as middle and posterior tibiae, respectively.</paragraph>
<paragraph blockId="4.[140,1109,166,897]" pageId="4" pageNumber="79">
Aedeagus robust, short and wide, with broad blunt apex, with pair of slcerotized lobes of characteristic shape dorso-laterally; paramere deeply divided into two pointed branches which converge apico-medially, each of them apically with irregular group of sensory peg setae on the underside. Internal sac weakly sclerotized, bulb-like when everted (
<figureCitation box="[236,336,518,545]" captionStart="Figs 2-4" captionStartId="4.[140,184,1590,1612]" captionTargetBox="[226,1010,970,1566]" captionTargetId="figure@4.[225,1023,965,1570]" captionTargetPageId="4" captionText="Figs 2-4. Philonthus neoniger Solodovnikov, nom. nov. Details of structure: 2, prothorax (in vertral view, only left side illustrated, anterior leg removed); 3, aedeagus in dorsal (parameral) view (internal sac evert- ed); 4, aedeagus in lateral view (internal sac everted). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787019" httpUri="https://zenodo.org/record/3787019/files/figure.png" pageId="4" pageNumber="79">Figs 3, 4</figureCitation>
).
</paragraph>
<paragraph blockId="4.[140,1109,166,897]" box="[185,389,554,580]" pageId="4" pageNumber="79">Female unknown.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="79" type="discussion">
<paragraph blockId="4.[140,1109,166,897]" pageId="4" pageNumber="79">
<emphasis bold="true" box="[185,342,589,615]" pageId="4" pageNumber="79">
<emphasis box="[185,337,589,615]" inLineHeading="true" pageId="4" pageNumber="79" reason="1">Comparison</emphasis>
.
</emphasis>
Among the seven other species belonging to the subgenus
<taxonomicName authority="(Hromadka 2008)" baseAuthorityName="Hromadka" baseAuthorityYear="2008" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="subGenus" subGenus="Raucalius">
<emphasis box="[1002,1107,589,615]" italics="true" pageId="4" pageNumber="79">Raucalius</emphasis>
(
<bibRefCitation author="Hromadka L" box="[147,339,624,650]" pageId="4" pageNumber="79" pagination="51 - 65" refId="ref2248" refString="Hromadka L (2008) Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina). Acta Entomologica Musei Nationalis Pragae 48 (1): 51 - 65." type="journal article" year="2008">Hromádka 2008</bibRefCitation>
)
</taxonomicName>
<taxonomicName authority="Solodovnikov" authorityName="Solodovnikov" authorityYear="2009" box="[354,731,624,650]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="neoniger" status="nom. nov.">
<emphasis box="[354,568,624,650]" italics="true" pageId="4" pageNumber="79">Philonthus neoniger</emphasis>
Solodovnikov
</taxonomicName>
,
<taxonomicNameLabel box="[741,856,625,651]" pageId="4" pageNumber="79">nom. nov.</taxonomicNameLabel>
is most similar with
<taxonomicName authority="Tottenham, 1949" authorityName="Tottenham" authorityYear="1949" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="peripateticus">
<emphasis italics="true" pageId="4" pageNumber="79">P. peripateticus</emphasis>
<bibRefCitation author="Tottenham CE" box="[283,486,659,685]" pageId="4" pageNumber="79" pagination="291 - 362" refId="ref2460" refString="Tottenham CE (1949) Studies in the genus Philonthus Stephens (Coleoptera). The Transactions of the Royal Entomological Society of London 100 (12): 291 - 362." type="journal article" year="1949">Tottenham, 1949</bibRefCitation>
</taxonomicName>
. Both species however can be very easily distinguished by shape of the aedeagi (cf. with Figs 14, 15 in
<bibRefCitation author="Tottenham CE" box="[716,933,694,721]" pageId="4" pageNumber="79" pagination="291 - 362" refId="ref2460" refString="Tottenham CE (1949) Studies in the genus Philonthus Stephens (Coleoptera). The Transactions of the Royal Entomological Society of London 100 (12): 291 - 362." type="journal article" year="1949">Tottenham (1949)</bibRefCitation>
, Figs
<quantity box="[1005,1107,694,721]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" metricValueMax="8.128" metricValueMin="7.366" pageId="4" pageNumber="79" unit="in" value="30.5" valueMax="32.0" valueMin="29.0">29-32 in</quantity>
<bibRefCitation author="Hromadka L" box="[140,349,730,756]" pageId="4" pageNumber="79" pagination="51 - 65" refId="ref2248" refString="Hromadka L (2008) Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina). Acta Entomologica Musei Nationalis Pragae 48 (1): 51 - 65." type="journal article" year="2008">Hromádka (2008)</bibRefCitation>
, and with
<figureCitation box="[467,562,730,756]" captionStart="Figs 2-4" captionStartId="4.[140,184,1590,1612]" captionTargetBox="[226,1010,970,1566]" captionTargetId="figure@4.[225,1023,965,1570]" captionTargetPageId="4" captionText="Figs 2-4. Philonthus neoniger Solodovnikov, nom. nov. Details of structure: 2, prothorax (in vertral view, only left side illustrated, anterior leg removed); 3, aedeagus in dorsal (parameral) view (internal sac evert- ed); 4, aedeagus in lateral view (internal sac everted). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787019" httpUri="https://zenodo.org/record/3787019/files/figure.png" pageId="4" pageNumber="79">Figs 3, 4</figureCitation>
here). The aedeagus of
<taxonomicName box="[825,940,730,755]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="neoniger">
<emphasis box="[825,940,730,755]" italics="true" pageId="4" pageNumber="79">P. neoniger</emphasis>
</taxonomicName>
is more robust (relatively shorter and wider) than in
<emphasis box="[573,743,766,791]" italics="true" pageId="4" pageNumber="79">
<taxonomicName authorityName="Tottenham" authorityYear="1949" box="[573,739,766,791]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="peripateticus">P. peripateticus</taxonomicName>
.
</emphasis>
Also the peg setae on the paramere are arranged differently in both species: they are irregularly grouped near the apex of each parameral branch in
<taxonomicName box="[467,585,836,861]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="neoniger">
<emphasis box="[467,585,836,861]" italics="true" pageId="4" pageNumber="79">P. neoniger</emphasis>
</taxonomicName>
, but arranged in two longitudinal rows at the apex of each parameral branch in
<taxonomicName authorityName="Tottenham" authorityYear="1949" box="[521,682,871,896]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="peripateticus">
<emphasis box="[521,682,871,896]" italics="true" pageId="4" pageNumber="79">P. peripateticus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3787019" ID-Zenodo-Dep="3787019" httpUri="https://zenodo.org/record/3787019/files/figure.png" pageId="4" pageNumber="79" startId="4.[140,184,1590,1612]" targetBox="[226,1010,970,1566]" targetPageId="4">
<paragraph blockId="4.[140,1108,1590,1676]" pageId="4" pageNumber="79">
<emphasis bold="true" box="[140,231,1590,1612]" pageId="4" pageNumber="79">Figs 2-4.</emphasis>
<taxonomicName authority="Solodovnikov" authorityName="Solodovnikov" authorityYear="2009" box="[237,546,1590,1612]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="neoniger" status="nom. nov.">
<emphasis box="[237,412,1590,1612]" italics="true" pageId="4" pageNumber="79">Philonthus neoniger</emphasis>
Solodovnikov
</taxonomicName>
,
<taxonomicNameLabel box="[555,650,1591,1612]" pageId="4" pageNumber="79">nom. nov.</taxonomicNameLabel>
Details of structure: 2, prothorax (in vertral view, only left side illustrated, anterior leg removed); 3, aedeagus in dorsal (parameral) view (internal sac evert-ed); 4, aedeagus in lateral view (internal sac everted). Scale bar
<quantity box="[720,779,1654,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="79" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="80" type="distribution">
<paragraph blockId="5.[140,1108,166,333]" pageId="5" pageNumber="80">
<emphasis bold="true" box="[185,515,166,193]" pageId="5" pageNumber="80">
<emphasis box="[185,331,166,192]" inLineHeading="true" pageId="5" pageNumber="80" reason="1">Distribution</emphasis>
and bionomics.
</emphasis>
The species is known only from the
<typeStatus box="[905,1004,166,192]" pageId="5" pageNumber="80">holotype</typeStatus>
, from the type locality in
<collectingCountry box="[307,379,202,228]" name="Kenya" pageId="5" pageNumber="80">Kenya</collectingCountry>
, Eastern Africa. It was collected at an elevation of
<quantity box="[923,1078,201,228]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="5" pageNumber="80" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1500-1800 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="80" type="etymology">
<paragraph blockId="5.[140,1108,166,333]" pageId="5" pageNumber="80">
<emphasis bold="true" box="[185,320,237,263]" pageId="5" pageNumber="80">
<emphasis box="[185,315,237,263]" inLineHeading="true" pageId="5" pageNumber="80" reason="1">Etymology</emphasis>
.
</emphasis>
The new name is derived from the original name “
<taxonomicName baseAuthorityName="O. Muller" baseAuthorityYear="1764" box="[916,1095,237,263]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[916,1095,237,263]" italics="true" pageId="5" pageNumber="80">Philonthus niger</emphasis>
</taxonomicName>
” by adding the prefix “neo” [new] which refers to its new name. It is an adjective of masculine gender.
</paragraph>
</subSubSection>
</treatment>
</document>