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<document ID-DOI="http://doi.org/10.5281/zenodo.3792544" ID-GBIF-Dataset="7112ded3-fa31-4c7b-a093-4d3dedafd571" ID-GBIF-Taxon="163755067" ID-ISSN="13132970" ID-Zenodo-Dep="3792544" approvalRequired="4" approvalRequired_for_bibRefs="2" approvalRequired_for_treatments="2" checkinTime="1587999565776" checkinUser="plazi" docAuthor="Solodovnikov, Alexey" docDate="2009" docId="038F87A1FFC4AA7B01CCFA3CFC356DC6" docLanguage="en" docName="ZK_article_1965.pdf" docOrigin="ZooKeys 5 (5)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@60a754c6" docStyleName="zookeys.2008.journal_article" docTitle="Philonthus neoniger Solodovnikov 2009, nom. nov." docType="treatment" docVersion="6" lastPageId="2" lastPageNumber="77" masterDocId="FFB6FFD9FFC5AA790140FFF0FFA66F4C" masterDocTitle="Xanthopygoides niger Cameron, 1951 (Xanthopygina) belongs to the genus Philonthus Stephens, 1829 (Philonthina): systematic and nomenclatural changes for the African Staphylinini (Coleoptera, Staphylinidae, Staphylininae, Staphylinini)" masterLastPageNumber="80" masterPageNumber="75" pageId="1" pageNumber="76" updateTime="1643568035591" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Xanthopygoides niger Cameron, 1951 (Xanthopygina) belongs to the genus Philonthus Stephens, 1829 (Philonthina): systematic and nomenclatural changes for the African Staphylinini (Coleoptera, Staphylinidae, Staphylininae, Staphylinini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solodovnikov, Alexey</mods:namePart>
<mods:affiliation>Natural History Museum of Denmark, Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-02-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>80</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.5.59</mods:identifier>
<mods:identifier type="GBIF-Dataset">7112ded3-fa31-4c7b-a093-4d3dedafd571</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576432</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792544" ID-GBIF-Taxon="163755067" ID-Zenodo-Dep="3792544" LSID="urn:lsid:plazi:treatment:038F87A1FFC4AA7B01CCFA3CFC356DC6" httpUri="http://treatment.plazi.org/id/038F87A1FFC4AA7B01CCFA3CFC356DC6" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">
<subSubSection box="[140,869,1484,1511]" pageId="1" pageNumber="76" type="nomenclature">
<paragraph blockId="1.[140,869,1484,1511]" box="[140,869,1484,1511]" pageId="1" pageNumber="76">
<heading box="[140,869,1484,1511]" fontSize="11" level="2" pageId="1" pageNumber="76" reason="3">
<emphasis bold="true" box="[140,869,1484,1511]" pageId="1" pageNumber="76">
<taxonomicName authority="Solodovnikov" authorityName="Solodovnikov" authorityYear="2009" box="[140,583,1484,1511]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="neoniger" status="nom. nov.">
<emphasis bold="true" box="[140,395,1484,1511]" italics="true" pageId="1" pageNumber="76">Philonthus neoniger</emphasis>
Solodovnikov
</taxonomicName>
,
<taxonomicNameLabel box="[595,722,1484,1511]" pageId="1" pageNumber="76">nom. nov.</taxonomicNameLabel>
Taxonomy
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76" type="discussion">
<paragraph blockId="1.[140,1108,1539,1671]" lastBlockId="2.[140,1108,166,651]" lastPageId="2" lastPageNumber="77" pageId="1" pageNumber="76">
<taxonomicName box="[140,353,1539,1565]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="neoniger">
<emphasis box="[140,353,1539,1565]" italics="true" pageId="1" pageNumber="76">Philonthus neoniger</emphasis>
</taxonomicName>
(the original combination
<taxonomicName baseAuthorityName="Cameron" baseAuthorityYear="1951" box="[653,878,1539,1565]" class="Insecta" family="Staphylinidae" genus="Xanthopygoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[653,878,1539,1565]" italics="true" pageId="1" pageNumber="76">Xanthopygoides niger</emphasis>
</taxonomicName>
) is here moved from the subtribe
<taxonomicName authorityName="Sharp" authorityYear="1884" box="[284,444,1574,1600]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
to
<taxonomicName authorityName="Stephens" authorityYear="1829" box="[484,600,1574,1600]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis box="[484,600,1574,1600]" italics="true" pageId="1" pageNumber="76">Philonthus</emphasis>
</taxonomicName>
, the
<typeStatus box="[661,709,1575,1601]" pageId="1" pageNumber="76">type</typeStatus>
genus of the subtribe
<taxonomicName authorityName="Kirby" authorityYear="1837" box="[966,1104,1574,1600]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
. This transfer is based on the following complex of character states shared by
<taxonomicName box="[992,1107,1610,1635]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="neoniger">
<emphasis box="[992,1107,1610,1635]" italics="true" pageId="1" pageNumber="76">P. neoniger</emphasis>
</taxonomicName>
with other members of
<emphasis box="[411,535,1645,1671]" italics="true" pageId="1" pageNumber="76">
<taxonomicName authorityName="Stephens" authorityYear="1829" box="[411,529,1645,1671]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
:
</emphasis>
lack of empodial setae; pronotal hypomeron with superior marginal line deflexed under anterior angle of pronotum, and inferior marginal line becoming obsolete near lateral margin of procoxal cavity, both lines not joining each other (
<figureCitation box="[318,386,237,263]" captionStart="Figs 2-4" captionStartId="4.[140,184,1590,1612]" captionTargetBox="[226,1010,970,1566]" captionTargetId="figure@4.[225,1023,965,1570]" captionTargetPageId="4" captionText="Figs 2-4. Philonthus neoniger Solodovnikov, nom. nov. Details of structure: 2, prothorax (in vertral view, only left side illustrated, anterior leg removed); 3, aedeagus in dorsal (parameral) view (internal sac evert- ed); 4, aedeagus in lateral view (internal sac everted). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787019" httpUri="https://zenodo.org/record/3787019/files/figure.png" pageId="2" pageNumber="77">Fig. 2</figureCitation>
); prosternum fused to pronotum, notosternal sutures indistinct (
<figureCitation box="[148,215,272,298]" captionStart="Figs 2-4" captionStartId="4.[140,184,1590,1612]" captionTargetBox="[226,1010,970,1566]" captionTargetId="figure@4.[225,1023,965,1570]" captionTargetPageId="4" captionText="Figs 2-4. Philonthus neoniger Solodovnikov, nom. nov. Details of structure: 2, prothorax (in vertral view, only left side illustrated, anterior leg removed); 3, aedeagus in dorsal (parameral) view (internal sac evert- ed); 4, aedeagus in lateral view (internal sac everted). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787019" httpUri="https://zenodo.org/record/3787019/files/figure.png" pageId="2" pageNumber="77">Fig. 2</figureCitation>
); no translucent postcoxal process of pronotal hypomeron (
<figureCitation box="[879,947,272,298]" captionStart="Figs 2-4" captionStartId="4.[140,184,1590,1612]" captionTargetBox="[226,1010,970,1566]" captionTargetId="figure@4.[225,1023,965,1570]" captionTargetPageId="4" captionText="Figs 2-4. Philonthus neoniger Solodovnikov, nom. nov. Details of structure: 2, prothorax (in vertral view, only left side illustrated, anterior leg removed); 3, aedeagus in dorsal (parameral) view (internal sac evert- ed); 4, aedeagus in lateral view (internal sac everted). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3787019" httpUri="https://zenodo.org/record/3787019/files/figure.png" pageId="2" pageNumber="77">Fig. 2</figureCitation>
); entire ligula; no infraorbital and mandibular ridges on head capsule. Such features of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="neoniger">
<emphasis italics="true" pageId="2" pageNumber="77">Philonthus neoniger</emphasis>
</taxonomicName>
as smooth pronotal disk with only one puncture in each of two dorsal “rows”, dense and rugose punctation of elytra and abdomen, and, especially, a rather peculiar shape of the aedeagus (Figs 3,4) allow it to be unambiguously placed in the subgenus
<taxonomicName authority="Tottenham, 1949" authorityName="Tottenham" authorityYear="1949" box="[140,453,448,474]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="subGenus" subGenus="Raucalius">
<emphasis box="[140,245,448,474]" italics="true" pageId="2" pageNumber="77">Raucalius</emphasis>
<bibRefCitation author="Tottenham CE" box="[251,453,448,474]" pageId="2" pageNumber="77" pagination="291 - 362" refId="ref2460" refString="Tottenham CE (1949) Studies in the genus Philonthus Stephens (Coleoptera). The Transactions of the Royal Entomological Society of London 100 (12): 291 - 362." type="journal article" year="1949">Tottenham, 1949</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[140,1108,166,651]" pageId="2" pageNumber="77">
Members of the subtribe
<taxonomicName authorityName="Sharp" authorityYear="1884" box="[487,650,483,509]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
, instead, are usually characterized by a complete postmandibular ridge; the inferior marginal line of pronotal hypomeron well separated from the superior marginal line, both lines continuing uninterrupted and separated along the prosternal margin. Also, unlike
<taxonomicName authorityName="Kirby" authorityYear="1837" box="[726,864,589,615]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
, all members of
<taxonomicName authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
have a pair of empodial setae (
<bibRefCitation author="Smetana A &amp; Davies A" box="[601,903,624,650]" pageId="2" pageNumber="77" pagination="1 - 88" refId="ref2376" refString="Smetana A, Davies A (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates 3287: 1 - 88." type="journal article" year="2000">Smetana and Davies 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>