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<document ID-DOI="10.11646/megataxa.7.1.1" ID-GBIF-Dataset="bd1f1932-a0fa-4daf-af9f-cc52c932ed40" ID-ISSN="2703-3090" ID-Zenodo-Dep="5964441" checkinTime="1643698274876" checkinUser="plazi" docAuthor="Iverson, John B." docDate="2022" docId="038F8788FFB1FFD6D973FFFCFE7226CC" docLanguage="en" docName="44551-Article.Text-47635-51294-10-20220125.pdf.imf" docOrigin="Megataxa 7 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.3:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="3" docTitle="Terrapene nelsoni subsp. klauberi Bogert 1943" docType="treatment" docVersion="10" lastPageNumber="41" masterDocId="FFB6FFF0FF96FFFEDA59FB57FF95276A" masterDocTitle="A review of Chelonian type specimens (order Testudines)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="40" updateTime="1680830866277" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Chelonian type specimens (order Testudines)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Iverson, John B.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Megataxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>85</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/megataxa.7.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">bd1f1932-a0fa-4daf-af9f-cc52c932ed40</mods:identifier>
<mods:identifier type="ISSN">2703-3090</mods:identifier>
<mods:identifier type="Zenodo-Dep">5964441</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6328658" ID-GBIF-Taxon="193856512" ID-Zenodo-Dep="6328658" LSID="urn:lsid:plazi:treatment:038F8788FFB1FFD6D973FFFCFE7226CC" httpUri="http://treatment.plazi.org/id/038F8788FFB1FFD6D973FFFCFE7226CC" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<subSubSection box="[810,1296,1194,1221]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph blockId="39.[810,1451,1194,1350]" box="[810,1296,1194,1221]" pageId="39" pageNumber="40">
<heading box="[810,1296,1194,1221]" fontSize="11" level="2" pageId="39" pageNumber="40" reason="2">
<taxonomicName authority="Bogert 1943: 2" authorityName="Bogert" authorityPageNumber="2" authorityYear="1943" box="[810,1296,1194,1221]" class="Reptilia" family="Emydidae" genus="Terrapene" kingdom="Animalia" order="Testudines" pageId="39" pageNumber="40" phylum="Chordata" rank="subSpecies" species="nelsoni" subSpecies="klauberi">
<emphasis bold="true" box="[810,1296,1194,1221]" pageId="39" pageNumber="40">
<emphasis bold="true" box="[810,1121,1194,1221]" italics="true" pageId="39" pageNumber="40">Terrapene nelsoni klauberi</emphasis>
Bogert 1943:2
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="materials_examined">
<paragraph blockId="39.[810,1451,1194,1350]" pageId="39" pageNumber="40">
<materialsCitation ID-GBIF-Occurrence="3695119350" collectionCode="AMNH, FMNH" pageId="39" pageNumber="40" specimenCode="AMNH 63752- 54, FMNH 41269" specimenCount="3" typeStatus="holotype">
<emphasis bold="true" box="[810,919,1227,1253]" pageId="39" pageNumber="40">
<typeStatus box="[810,919,1227,1253]" pageId="39" pageNumber="40">Holotype</typeStatus>
</emphasis>
: AMNH 63751;
<emphasis bold="true" box="[1155,1273,1227,1253]" pageId="39" pageNumber="40">
<typeStatus box="[1155,1273,1227,1253]" pageId="39" pageNumber="40">paratypes</typeStatus>
</emphasis>
(3 according to the original description),
<specimenCode collectionCode="AMNH" pageId="39" pageNumber="40">AMNH 63752- 54</specimenCode>
. However,
<specimenCode box="[1080,1241,1291,1317]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="39" pageNumber="40" type="Museum">FMNH 41269</specimenCode>
is also listed as a
<typeStatus box="[906,1002,1323,1349]" pageId="39" pageNumber="40">paratype</typeStatus>
by
<bibRefCitation author="Marx, H." box="[1044,1239,1323,1350]" pageId="39" pageNumber="40" pagination="409 - 496" refId="ref77120" refString="Marx, H. (1958) Catalogue of type specimens of reptiles and amphibians in Chicago Natural History Museum. Fieldiana Zoology, 36 (4), 409 - 496. https: // doi. org / 10.5962 / bhl. title. 3034" type="journal article" year="1958">Marx (1958:450)</bibRefCitation>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="reference_group">
<paragraph blockId="39.[810,1452,1386,1797]" box="[810,1240,1386,1413]" pageId="39" pageNumber="40">
<treatmentCitationGroup box="[810,1240,1386,1413]" pageId="39" pageNumber="40">
<treatmentCitation author="Agassiz, L." box="[810,1240,1386,1413]" page="445" pageId="39" pageNumber="40" year="1857">
<taxonomicName authority="(Agassiz 1857: 445)" baseAuthorityName="Agassiz" baseAuthorityPageNumber="445" baseAuthorityYear="1857" box="[810,1240,1386,1413]" class="Reptilia" family="Emydidae" genus="Terrapene" kingdom="Animalia" order="Testudines" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="ornata">
<emphasis bold="true" box="[810,1240,1386,1413]" italics="true" pageId="39" pageNumber="40">
Terrapene ornata (
<bibRefCitation author="Agassiz, L." box="[1025,1233,1386,1412]" pageId="39" pageNumber="40" pagination="1 - 452" refId="ref64046" refString="Agassiz, L. (1857) Contributions to the Natural History of the United States of America. First Monograph. Vol. I. Part I. Essay on Classification. Part II. North American Testudinata. Little, Brown &amp; Co., Boston, pp. 1 - 452." type="book chapter" year="1857">Agassiz 1857:445</bibRefCitation>
)
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="39.[810,1452,1386,1797]" pageId="39" pageNumber="40">
<emphasis bold="true" box="[810,917,1418,1444]" pageId="39" pageNumber="40">
<typeStatus box="[810,917,1418,1444]" pageId="39" pageNumber="40">Syntypes</typeStatus>
</emphasis>
(at least 7; see
<bibRefCitation author="Reynolds, R. P. &amp; Gotte, S. W." box="[1097,1410,1419,1445]" pageId="39" pageNumber="40" pagination="1 - 49" refId="ref79778" refString="Reynolds, R. P., Gotte, S. W. &amp; Ernst, C. H. (2007) Catalog of type specimens of Recent Crocodilia and Testudines in the National Museum of Natural History, Smithsonian Institution. Smithsonian Contributions to Zoology, 626, 1 - 49. https: // doi. org / 10.5479 / si. 00810282.626" type="journal article" year="2007">
Reynolds
<emphasis box="[1211,1269,1419,1445]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2007:12–13
</bibRefCitation>
for a discussion of the confusion surrounding the type series): USNM 7862 (formerly 57), 7541, 131837 (formerly one of three numbered 7541), 7547, 7692, and possibly USNM 7547 and 7740 (
<bibRefCitation author="Reynolds, R. P. &amp; Gotte, S. W." box="[913,1199,1579,1605]" pageId="39" pageNumber="40" pagination="1 - 49" refId="ref79778" refString="Reynolds, R. P., Gotte, S. W. &amp; Ernst, C. H. (2007) Catalog of type specimens of Recent Crocodilia and Testudines in the National Museum of Natural History, Smithsonian Institution. Smithsonian Contributions to Zoology, 626, 1 - 49. https: // doi. org / 10.5479 / si. 00810282.626" type="journal article" year="2007">
Reynolds
<emphasis box="[1032,1092,1579,1605]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2007:12
</bibRefCitation>
); BMNH 1873.8.13.3 (formerly one of three numbered USNM 7541); and MCZ 1536 (
<bibRefCitation author="Barbour, T. &amp; Loveridge, A." box="[1084,1438,1643,1669]" pageId="39" pageNumber="40" pagination="205 - 360" refId="ref64740" refString="Barbour, T. &amp; Loveridge, A. (1929) Typical reptiles and amphibians in the Museum of Comparative Zoology. Bulletin of the Museum of Comparative Zoology at Harvard College, 69 (10), 205 - 360." type="journal article" year="1929">Barbour &amp; Loveridge 1929:241</bibRefCitation>
); MCZ 1536 designated
<typeStatus box="[1187,1290,1675,1701]" pageId="39" pageNumber="40">lectotype</typeStatus>
by
<bibRefCitation author="Smith, H. M. &amp; Smith, R. B." pageId="39" pageNumber="40" refId="ref81902" refString="Smith, H. M. &amp; Smith, R. B. (1980 [&quot; 1979 &quot;]) Synopsis of the herptofauna of Mexico. Volume VI. Guide to Mexican turtles. Bibliographic addendum III. John Johnson, North Bennington, Vermont, 1044 pp. https: // doi. org / 10.2307 / 1444548" type="book" year="1980">Smith &amp; Smith (1980:587)</bibRefCitation>
.
<bibRefCitation author="Ward, J. P." box="[1146,1316,1707,1733]" pageId="39" pageNumber="40" pagination="1 - 4" refId="ref84557" refString="Ward, J. P. (1978) Terrapene ornata (Agassiz). Ornate Box Turtle. Catalogue of American Amphibians and Reptiles, 217, 1 - 4." type="journal article" year="1978">Ward (1978:1)</bibRefCitation>
and
<bibRefCitation author="Smith, H. M. &amp; Smith, R. B." pageId="39" pageNumber="40" refId="ref81902" refString="Smith, H. M. &amp; Smith, R. B. (1980 [&quot; 1979 &quot;]) Synopsis of the herptofauna of Mexico. Volume VI. Guide to Mexican turtles. Bibliographic addendum III. John Johnson, North Bennington, Vermont, 1044 pp. https: // doi. org / 10.2307 / 1444548" type="book" year="1980">Smith &amp; Smith (1980:586)</bibRefCitation>
also included USNM 7542 among the
<typeStatus box="[1030,1132,1771,1797]" pageId="39" pageNumber="40">syntypes</typeStatus>
.
</paragraph>
<paragraph blockId="39.[845,1397,1835,1893]" box="[845,1389,1835,1861]" pageId="39" pageNumber="40">
<taxonomicName authority="Cragin 1894: 37" authorityName="Cragin" authorityPageNumber="37" authorityYear="1894" box="[845,1389,1835,1861]" class="Reptilia" family="Emydidae" genus="Terrapene" kingdom="Animalia" order="Testudines" pageId="39" pageNumber="40" phylum="Chordata" rank="subSpecies" species="ornata" subSpecies="cimarronensis">
<emphasis box="[845,1206,1836,1861]" italics="true" pageId="39" pageNumber="40">Terrapene ornata cimarronensis</emphasis>
Cragin 1894:
<specimenCount box="[1360,1389,1835,1861]" pageId="39" pageNumber="40" type="generic" typeStatus="Holotype">37</specimenCount>
</taxonomicName>
</paragraph>
<paragraph blockId="39.[845,1397,1835,1893]" box="[845,1397,1867,1893]" pageId="39" pageNumber="40">
<emphasis bold="true" box="[845,954,1867,1893]" pageId="39" pageNumber="40">Holotype</emphasis>
: Not located; no
<typeStatus box="[1155,1262,1867,1893]" pageId="39" pageNumber="40">paratypes</typeStatus>
designated.
</paragraph>
<paragraph blockId="39.[845,1452,1931,2053]" box="[845,1267,1931,1957]" pageId="39" pageNumber="40">
<taxonomicName authorityName="Hay" authorityYear="1908" box="[845,1097,1931,1957]" class="Reptilia" family="Emydidae" genus="Terrapene" kingdom="Animalia" order="Testudines" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="longinsulae">
<emphasis box="[845,1097,1931,1957]" italics="true" pageId="39" pageNumber="40">Terrapene longinsulae</emphasis>
</taxonomicName>
<superScript attach="left" box="[1096,1101,1931,1945]" fontSize="6" pageId="39" pageNumber="40">t</superScript>
Hay 1908:166
</paragraph>
</subSubSection>
<subSubSection pageId="39" pageNumber="40" type="materials_examined">
<paragraph blockId="39.[845,1452,1931,2053]" pageId="39" pageNumber="40">
<emphasis bold="true" box="[845,954,1963,1989]" pageId="39" pageNumber="40">
<typeStatus box="[845,954,1963,1989]" pageId="39" pageNumber="40">Holotype</typeStatus>
</emphasis>
:
<collectionCode box="[968,1049,1963,1989]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40" type="Museum">USNM</collectionCode>
5983 (
<bibRefCitation author="Vlachos, E." box="[1127,1340,1963,1990]" pageId="39" pageNumber="40" pagination="3 - 94" refId="ref84153" refString="Vlachos, E. (2018) A review of the fossil record of North American turtles of the clade Pan-Testudinoidea. Bulletin of the Peabody Museum of Natural History, 59, 3 - 94. https: // doi. org / 10.3374 / 014.059.0101" type="journal article" year="2018">Vlachos (2018:63)</bibRefCitation>
; illustrated in the original description (Pl.
<collectionCode box="[1372,1444,1995,2021]" pageId="39" pageNumber="40">XXVI</collectionCode>
) and in
<bibRefCitation author="Joyce, W. G. &amp; Lyson, T. R. &amp; Czaplewski, N. J." box="[982,1184,2027,2053]" pageId="39" pageNumber="40" pagination="177 - 190" refId="ref75321" refString="Joyce, W. G., Petric ˇ evici, A., Lyson, T. R. &amp; Czaplewski, N. J. (2012) A new box turtle from the Miocene / Pliocene boundary (Latest Hemphillian) of Oklahoma and a refined chronology of box turtle diversification. Journal of Paleontology, 86 (1), 177 - 190. https: // doi. org / 10.1666 / 11 - 073.1" type="journal article" year="2012">
Joyce
<emphasis box="[1053,1109,2027,2053]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
(2012
</bibRefCitation>
:Figs 3–4).
</paragraph>
</subSubSection>
<subSubSection box="[171,530,171,197]" pageId="40" pageNumber="41" type="reference_group">
<paragraph blockId="40.[171,778,171,294]" box="[171,530,171,197]" pageId="40" pageNumber="41">
<treatmentCitationGroup box="[171,530,171,197]" pageId="40" pageNumber="41">
<taxonomicName authorityName="Hay" authorityYear="1916" box="[171,387,171,197]" class="Reptilia" family="Emydidae" genus="Terrapene" kingdom="Animalia" order="Testudines" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="whitneyi">
<emphasis box="[171,387,171,197]" italics="true" pageId="40" pageNumber="41">Terrapene whitneyi</emphasis>
</taxonomicName>
<superScript attach="left" box="[387,392,171,185]" fontSize="6" pageId="40" pageNumber="41">t</superScript>
<treatmentCitation author="Hay, O. P." box="[399,530,171,197]" page="8" pageId="40" pageNumber="41" year="1916">
<bibRefCitation author="Hay, O. P." box="[399,530,171,197]" pageId="40" pageNumber="41" pagination="39 - 76" refId="ref73982" refString="Hay, O. P. (1916) Descriptions of some Floridian fossil vertebrates, belonging mostly to the Pleistocene. Annual Report of the Florida State Geological Survey, 8, 39 - 76." type="journal article" year="1916">Hay 1916:8</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="40" pageNumber="41" type="materials_examined">
<paragraph blockId="40.[171,778,171,294]" pageId="40" pageNumber="41">
<materialsCitation ID-GBIF-Occurrence="3695119705" collectionCode="USNM" pageId="40" pageNumber="41" specimenCode="USNM 8617" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[171,280,203,229]" pageId="40" pageNumber="41">
<typeStatus box="[171,280,203,229]" pageId="40" pageNumber="41">Holotype</typeStatus>
</emphasis>
:
<specimenCode box="[295,439,203,229]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="40" pageNumber="41" type="Museum">USNM 8617</specimenCode>
(
<bibRefCitation author="Vlachos, E." box="[453,667,203,230]" pageId="40" pageNumber="41" pagination="3 - 94" refId="ref84153" refString="Vlachos, E. (2018) A review of the fossil record of North American turtles of the clade Pan-Testudinoidea. Bulletin of the Peabody Museum of Natural History, 59, 3 - 94. https: // doi. org / 10.3374 / 014.059.0101" type="journal article" year="2018">Vlachos (2018:64)</bibRefCitation>
; illustrated in the original description (
<collectorName box="[659,709,235,261]" pageId="40" pageNumber="41">Pl. I</collectorName>
,
<collectorName box="[722,778,235,261]" pageId="40" pageNumber="41">Figs.</collectorName>
4–5,
<collectorName box="[288,346,267,293]" pageId="40" pageNumber="41">Pl. II</collectorName>
,
<collectorName box="[358,403,267,293]" pageId="40" pageNumber="41">Fig.</collectorName>
1).
</materialsCitation>
</paragraph>
<paragraph blockId="40.[171,761,331,422]" box="[171,750,331,358]" pageId="40" pageNumber="41">
<taxonomicName authority="Smith &amp; Ramsay 1952: 45" authorityName="Smith &amp; Ramsay" authorityPageNumber="45" authorityYear="1952" box="[171,750,331,358]" class="Reptilia" family="Emydidae" genus="Terrapene" kingdom="Animalia" order="Testudines" pageId="40" pageNumber="41" phylum="Chordata" rank="subSpecies" species="ornata" subSpecies="luteola">
<emphasis box="[171,452,331,357]" italics="true" pageId="40" pageNumber="41">Terrapene ornata luteola</emphasis>
Smith &amp; Ramsay 1952:
<specimenCount box="[721,750,331,358]" pageId="40" pageNumber="41" type="generic" typeStatus="Holotype">45</specimenCount>
</taxonomicName>
</paragraph>
<paragraph blockId="40.[171,761,331,422]" pageId="40" pageNumber="41">
<emphasis bold="true" box="[171,280,363,389]" pageId="40" pageNumber="41">Holotype</emphasis>
:
<specimenCode box="[294,416,363,389]" collectionCode="TCU" pageId="40" pageNumber="41">TCU 1280</specimenCode>
;
<emphasis bold="true" box="[428,535,363,389]" pageId="40" pageNumber="41">
<typeStatus box="[428,535,363,389]" pageId="40" pageNumber="41">paratype</typeStatus>
</emphasis>
(1),
<materialsCitation ID-GBIF-Occurrence="3695119763" box="[589,761,363,389]" collectionCode="UIMNH" pageId="40" pageNumber="41" specimenCode="UIMNH 18039" specimenCount="1" typeStatus="paratype">
<specimenCode box="[589,761,363,389]" collectionCode="UIMNH" country="USA" httpUri="http://grbio.org/cool/zrdp-fspx" name="University of Illinois, Museum of Natural History" pageId="40" pageNumber="41">UIMNH 18039</specimenCode>
</materialsCitation>
(
<bibRefCitation author="Smith, H. M. &amp; Langebartel, D. A. &amp; Williams, K. L." box="[240,475,395,422]" pageId="40" pageNumber="41" pagination="1 - 80" refId="ref82018" refString="Smith, H. M., Langebartel, D. A. &amp; Williams, K. L. (1964) Herpetological type-specimens in the University of Illinois Museum of Natural History. Illinois Biological Monographs, 32, 1 - 80. https: // doi. org / 10.5962 / bhl. title. 50204" type="journal article" year="1964">
Smith
<emphasis box="[315,365,395,421]" italics="true" pageId="40" pageNumber="41">et al</emphasis>
. 1964:36
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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