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<document ID-DOI="10.11646/megataxa.7.1.1" ID-GBIF-Dataset="bd1f1932-a0fa-4daf-af9f-cc52c932ed40" ID-ISSN="2703-3090" ID-Zenodo-Dep="5964441" checkinTime="1643698274876" checkinUser="plazi" docAuthor="Iverson, John B." docDate="2022" docId="038F8788FF9DFFF5DAD1FABDFA0E248C" docLanguage="en" docName="44551-Article.Text-47635-51294-10-20220125.pdf.imf" docOrigin="Megataxa 7 (1)" docStyle="DocumentStyle:50B6B96D96A55C21793B52C23E95B310.3:Megataxa.2020-.journal_article" docStyleId="50B6B96D96A55C21793B52C23E95B310" docStyleName="Megataxa.2020-.journal_article" docStyleVersion="3" docTitle="Pelomedusa olivacea" docType="treatment" docVersion="9" lastPageNumber="12" masterDocId="FFB6FFF0FF96FFFEDA59FB57FF95276A" masterDocTitle="A review of Chelonian type specimens (order Testudines)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="12" updateTime="1680830866277" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Chelonian type specimens (order Testudines)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Iverson, John B.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Megataxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>7</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:end>85</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/megataxa.7.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">bd1f1932-a0fa-4daf-af9f-cc52c932ed40</mods:identifier>
<mods:identifier type="ISSN">2703-3090</mods:identifier>
<mods:identifier type="Zenodo-Dep">5964441</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6328813" ID-GBIF-Taxon="193856571" ID-Zenodo-Dep="6328813" LSID="urn:lsid:plazi:treatment:038F8788FF9DFFF5DAD1FABDFA0E248C" httpUri="http://treatment.plazi.org/id/038F8788FF9DFFF5DAD1FABDFA0E248C" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection box="[136,654,490,517]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph blockId="11.[136,778,490,677]" box="[136,654,490,517]" pageId="11" pageNumber="12">
<heading box="[136,654,490,517]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<treatmentCitationGroup box="[136,654,490,517]" pageId="11" pageNumber="12">
<treatmentCitation author="Schweigger, A. F." box="[136,654,490,517]" page="307" pageId="11" pageNumber="12" year="1812">
<taxonomicName authority="(Schweigger 1812: 307)" baseAuthorityName="Schweigger" baseAuthorityPageNumber="307" baseAuthorityYear="1812" box="[136,654,490,517]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="olivacea">
<emphasis bold="true" box="[136,654,490,517]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,378,490,517]" italics="true" pageId="11" pageNumber="12">Pelomedusa olivacea</emphasis>
(
<bibRefCitation author="Schweigger, A. F." box="[392,647,490,517]" pageId="11" pageNumber="12" pagination="406 - 462" refId="ref81144" refString="Schweigger, A. F. (1812) Prodromus monographiae Cheloniorum. Konigsberger Archiv fur Naturwissenschaft und Mathematik, 1, 271 - 368, 406 - 462." type="journal article" year="1812">Schweigger 1812:307</bibRefCitation>
)
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[136,778,490,677]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,245,523,549]" pageId="11" pageNumber="12">
<typeStatus box="[136,245,523,549]" pageId="11" pageNumber="12">Holotype</typeStatus>
</emphasis>
: MNHN 7971 (
<bibRefCitation author="Ceriaco, L. M. P. &amp; Bour, R." box="[426,712,523,549]" pageId="11" pageNumber="12" pagination="18 - 32" refId="ref68948" refString="Ceriaco, L. M. P. &amp; Bour, R. (2012. Schweigger's (1812) chelonian types from the extinct eighteenth century Portuguese &quot; Royal cabinet of natural history of Ajuda &quot;: some contributions for their identification and nomenclatural implications. Zootaxa, 3395, 18 - 32. https: // doi. org / 10.11646 / zootaxa. 3395.1.2" type="journal article" year="2012">Ceríaco &amp; Bour 2012:19</bibRefCitation>
;
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[232,289,555,581]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:504
</bibRefCitation>
); illustrated in
<bibRefCitation author="Oppel, M." box="[591,736,555,581]" pageId="11" pageNumber="12" refId="ref78561" refString="Oppel, M. (1811) Die Ordnungen, Familien und Gattungen der Reptilien als Prodrom einer Naturgeshichte derselben. J. Lindauer, Munchen, 86 pp. https: // doi. org / 10.5962 / bhl. title. 4911" type="book" year="1811">Oppel (1811</bibRefCitation>
: Pl. 25; reproduced in
<bibRefCitation author="Bour, R. &amp; Schmidtler, J." box="[442,765,587,613]" pageId="11" pageNumber="12" refId="ref68058" refString="Bour, R. &amp; Schmidtler, J. (2014) Les Dessins et Aquarelles de Nikolaus Michael Oppel. 2. Les Tortues. Privately printed, Munchen and Paris, 48 pp." type="book" year="2014">Bour &amp; Schmidtler 2014:31</bibRefCitation>
); photographed in
<bibRefCitation author="Bour, R." box="[432,623,619,645]" pageId="11" pageNumber="12" pagination="26 - 30" refId="ref67370" refString="Bour, R. (2009 a) Type specimens of Testudo subrufa Lacepede, 1788 and Emys olivacea Schweigger, 1812. Emys, 16 (1), 26 - 30." type="journal article" year="2009">Bour (2009a:28)</bibRefCitation>
and
<bibRefCitation author="Bour, R. &amp; Schmidtler, J." pageId="11" pageNumber="12" refId="ref68058" refString="Bour, R. &amp; Schmidtler, J. (2014) Les Dessins et Aquarelles de Nikolaus Michael Oppel. 2. Les Tortues. Privately printed, Munchen and Paris, 48 pp." type="book" year="2014">Bour &amp; Schmidtler (2014:23)</bibRefCitation>
.
</paragraph>
<paragraph blockId="11.[171,778,715,934]" box="[171,643,715,742]" pageId="11" pageNumber="12">
<taxonomicName authority="Rochebrune 1884: 25" authorityName="Rochebrune" authorityPageNumber="25" authorityYear="1884" box="[171,643,715,742]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="gasconi">
<emphasis box="[171,402,715,741]" italics="true" pageId="11" pageNumber="12">Pelomedusa gasconi</emphasis>
Rochebrune 1884:25
</taxonomicName>
</paragraph>
<paragraph blockId="11.[171,778,715,934]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[171,280,747,773]" pageId="11" pageNumber="12">
<typeStatus box="[171,280,747,773]" pageId="11" pageNumber="12">Holotype</typeStatus>
</emphasis>
: Not designated (see
<bibRefCitation author="Loveridge, A." box="[523,754,747,773]" pageId="11" pageNumber="12" pagination="465 - 524" refId="ref76845" refString="Loveridge, A. (1941) Revision of the African terrapins of the family Pelomedusidae. Bulletin of the Museum of Comparative Zoology, 88, 465 - 524." type="journal article" year="1941">Loveridge 1941:480</bibRefCitation>
), although illustrated in the original description, Pl. I, Figs. 12;
<collectionCode box="[373,451,811,837]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="12" type="Museum">ZFMK</collectionCode>
17076 designated as
<typeStatus box="[689,778,811,837]" pageId="11" pageNumber="12">neotype</typeStatus>
by
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[267,523,843,870]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[327,384,843,869]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014b:504
</bibRefCitation>
; photographed in
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[232,288,875,901]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:516
</bibRefCitation>
; sampled genetically by
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
<emphasis box="[232,288,907,933]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014
</bibRefCitation>
and
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[405,603,907,934]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[465,522,907,933]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="reference_group">
<paragraph blockId="11.[136,778,970,1253]" pageId="11" pageNumber="12">
<treatmentCitationGroup pageId="11" pageNumber="12">
<treatmentCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." page="539" pageId="11" pageNumber="12" year="2014">
<taxonomicName authority="Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Siroky, &amp; Fritz 2014 b: 539" authorityName="Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Siroky, &amp; Fritz" authorityPageNumber="539" authorityYear="2014" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schweinfurthi">
<emphasis bold="true" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,451,970,997]" italics="true" pageId="11" pageNumber="12">Pelomedusa schweinfurthi</emphasis>
Petzold, Vargas-Ramírez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Široký, &amp;
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">Fritz 2014b:539</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[136,778,970,1253]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,245,1099,1125]" pageId="11" pageNumber="12">
<typeStatus box="[136,245,1099,1125]" pageId="11" pageNumber="12">Holotype</typeStatus>
</emphasis>
:
<accessionNumber box="[264,402,1099,1126]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF56161" pageId="11" pageNumber="12">
<collectionCode box="[264,320,1099,1125]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
56161
</accessionNumber>
(photographed in
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
<emphasis box="[717,778,1099,1125]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014:538
</bibRefCitation>
, and
<bibRefCitation author="Vetter, H." box="[427,625,1131,1157]" pageId="11" pageNumber="12" refId="ref83967" refString="Vetter, H. (2018) Turtles of the World, Vol. 5: Australia and Oceania. Edition Chimaira, Frankfurt, 144 pp." type="book" year="2018">Vetter 2018:144</bibRefCitation>
);
<emphasis bold="true" box="[659,777,1131,1157]" pageId="11" pageNumber="12">
<typeStatus box="[659,777,1131,1157]" pageId="11" pageNumber="12">paratypes</typeStatus>
</emphasis>
(3):
<accessionNumber box="[284,422,1163,1190]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF56160" pageId="11" pageNumber="12">
<collectionCode box="[284,340,1163,1189]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
56160
</accessionNumber>
and 56162; and
<accessionNumber box="[628,775,1163,1190]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMB15697" pageId="11" pageNumber="12">
<collectionCode box="[628,689,1163,1189]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="11" pageNumber="12" type="Museum">ZMB</collectionCode>
15697
</accessionNumber>
. All types sampled genetically by
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
<emphasis box="[718,778,1195,1221]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="11.[136,778,1290,1573]" pageId="11" pageNumber="12">
<taxonomicName authority="Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Siroky, &amp; Fritz 2014: 540" authorityName="Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Siroky, &amp; Fritz" authorityPageNumber="540" authorityYear="2014" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="somalica">
<emphasis bold="true" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,413,1290,1317]" italics="true" pageId="11" pageNumber="12">Pelomedusa somalica</emphasis>
Petzold, Vargas-Ramírez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Široký, &amp; Fritz 2014:540
</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="11.[136,778,1290,1573]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,245,1419,1445]" pageId="11" pageNumber="12">
<typeStatus box="[136,245,1419,1445]" pageId="11" pageNumber="12">Holotype</typeStatus>
</emphasis>
:
<collectionCode box="[263,347,1419,1445]" pageId="11" pageNumber="12">BMNH</collectionCode>
1970.1481 (photographed in
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
<emphasis box="[232,308,1451,1477]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014:541
</bibRefCitation>
);
<emphasis bold="true" box="[483,601,1451,1477]" pageId="11" pageNumber="12">
<typeStatus box="[483,601,1451,1477]" pageId="11" pageNumber="12">paratypes</typeStatus>
</emphasis>
(5),
<collectionCode box="[694,778,1451,1477]" pageId="11" pageNumber="12">BMNH</collectionCode>
1931.7.20.412-414 and 1931.8.1.177; and
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/NMW24449" pageId="11" pageNumber="12">
<collectionCode box="[707,778,1483,1509]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
24449
</accessionNumber>
(sampled genetically by
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
<emphasis box="[714,778,1515,1541]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection box="[136,636,1610,1637]" pageId="11" pageNumber="12" type="reference_group">
<paragraph blockId="11.[136,778,1610,1766]" box="[136,636,1610,1637]" pageId="11" pageNumber="12">
<treatmentCitationGroup box="[136,636,1610,1637]" pageId="11" pageNumber="12">
<treatmentCitation author="Bonnaterre, P. - J." box="[136,636,1610,1637]" page="28" pageId="11" pageNumber="12" year="1789">
<taxonomicName authority="(Bonnaterre 1789: 28)" baseAuthorityName="Bonnaterre" baseAuthorityPageNumber="28" baseAuthorityYear="1789" box="[136,636,1610,1637]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="subrufa">
<emphasis bold="true" box="[136,636,1610,1637]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,373,1610,1637]" italics="true" pageId="11" pageNumber="12">Pelomedusa subrufa</emphasis>
(
<bibRefCitation author="Bonnaterre, P. - J." box="[388,627,1610,1637]" pageId="11" pageNumber="12" refId="ref65753" refString="Bonnaterre, P. - J. (1789) Tableau Encyclopedique et Methodique des Trois Regnes de la Nature. Erpetologie. Panckoucke, Hotel de Thou, Paris, 70 pp. https: // doi. org / 10.5962 / bhl. title. 11660" type="book" year="1789">Bonnaterre 1789:28</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[136,778,1610,1766]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[136,254,1643,1669]" pageId="11" pageNumber="12">
<typeStatus box="[136,249,1643,1669]" pageId="11" pageNumber="12">Holotype</typeStatus>
:
</emphasis>
MNHN7970(
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[413,650,1643,1670]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[471,523,1643,1669]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:504
</bibRefCitation>
); illustrated in
<bibRefCitation author="Lacepede, B. G. E. de" box="[266,454,1675,1701]" pageId="11" pageNumber="12" refId="ref76061" refString="Lacepede, B. G. E. de. (1788) Histoire Naturelle des Quadrupedes Ovipares et des Serpens. Tome Premier. Hotel de Thou, Paris, 651 pp. [octavo edition, 8 o]. https: // doi. org / 10.5962 / bhl. title. 5036" type="book" year="1788">Lacepède (1788</bibRefCitation>
:opp. p. 173); photographed in
<bibRefCitation author="Bour, R." box="[266,438,1707,1733]" pageId="11" pageNumber="12" pagination="26 - 30" refId="ref67370" refString="Bour, R. (2009 a) Type specimens of Testudo subrufa Lacepede, 1788 and Emys olivacea Schweigger, 1812. Emys, 16 (1), 26 - 30." type="journal article" year="2009">Bour 2009a:27</bibRefCitation>
and
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[503,765,1707,1734]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[568,629,1707,1733]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:507
</bibRefCitation>
); sampled genetically by
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[497,712,1739,1766]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[557,613,1739,1765]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014b)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="11.[171,571,1803,1861]" box="[171,571,1803,1829]" pageId="11" pageNumber="12">
<taxonomicName authority="Meuschen 1778: 11" authorityName="Meuschen" authorityPageNumber="11" authorityYear="1778" box="[171,571,1803,1829]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="planitia">
<emphasis box="[171,351,1803,1829]" italics="true" pageId="11" pageNumber="12">Testudo planitia</emphasis>
Meuschen 1778:11
</taxonomicName>
</paragraph>
<paragraph blockId="11.[171,571,1803,1861]" box="[171,434,1835,1861]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[171,289,1835,1861]" pageId="11" pageNumber="12">
<typeStatus box="[171,284,1835,1861]" pageId="11" pageNumber="12">Holotype</typeStatus>
:
</emphasis>
Not located.
</paragraph>
</subSubSection>
<subSubSection box="[171,577,1899,1925]" pageId="11" pageNumber="12" type="reference_group">
<paragraph blockId="11.[171,778,1899,2086]" box="[171,577,1899,1925]" pageId="11" pageNumber="12">
<treatmentCitationGroup box="[171,577,1899,1925]" pageId="11" pageNumber="12">
<treatmentCitation author="Lacepede, B. G. E. de" box="[171,577,1899,1925]" page="173" pageId="11" pageNumber="12" year="1788">
<taxonomicName authority="Lacepede 1788: 173" authorityName="Lacepede" authorityPageNumber="173" authorityYear="1788" box="[171,577,1899,1925]" class="Reptilia" family="Testudinidae" genus="Testudo" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="subrufa">
<emphasis box="[171,350,1899,1925]" italics="true" pageId="11" pageNumber="12">Testudo subrufa</emphasis>
<bibRefCitation author="Lacepede, B. G. E. de" box="[357,577,1899,1925]" pageId="11" pageNumber="12" refId="ref76061" refString="Lacepede, B. G. E. de. (1788) Histoire Naturelle des Quadrupedes Ovipares et des Serpens. Tome Premier. Hotel de Thou, Paris, 651 pp. [octavo edition, 8 o]. https: // doi. org / 10.5962 / bhl. title. 5036" type="book" year="1788">Lacepède 1788:173</bibRefCitation>
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</paragraph>
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<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[171,778,1899,2086]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[171,289,1931,1957]" pageId="11" pageNumber="12">
<typeStatus box="[171,284,1931,1957]" pageId="11" pageNumber="12">Holotype</typeStatus>
:
</emphasis>
MNHN 7970 (
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[459,708,1931,1958]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[521,577,1931,1957]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:504
</bibRefCitation>
); illustrated in the original description (opp. p. 173); photographed in
<bibRefCitation author="Bour, R." box="[496,665,1995,2021]" pageId="11" pageNumber="12" pagination="26 - 30" refId="ref67370" refString="Bour, R. (2009 a) Type specimens of Testudo subrufa Lacepede, 1788 and Emys olivacea Schweigger, 1812. Emys, 16 (1), 26 - 30." type="journal article" year="2009">Bour 2009a:27</bibRefCitation>
and
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[232,287,2027,2053]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:507
</bibRefCitation>
); sampled genetically by
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014b)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="11.[845,1440,171,262]" box="[845,1273,171,197]" pageId="11" pageNumber="12">
<taxonomicName authority="Cornalia 1849: 13" authorityName="Cornalia" authorityPageNumber="13" authorityYear="1849" box="[845,1273,171,197]" class="Reptilia" family="Pelomedusidae" genus="Pentonix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="americana">
<emphasis box="[845,1071,172,197]" italics="true" pageId="11" pageNumber="12">Pentonix americana</emphasis>
Cornalia 1849:13
</taxonomicName>
</paragraph>
<paragraph blockId="11.[845,1440,171,262]" pageId="11" pageNumber="12">
<materialsCitation ID-GBIF-Occurrence="3695119610" collectionCode="WWII" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[845,954,203,229]" pageId="11" pageNumber="12">
<typeStatus box="[845,954,203,229]" pageId="11" pageNumber="12">Holotype</typeStatus>
</emphasis>
: MSNM unnumbered (destroyed in
<collectionCode box="[1361,1436,203,229]" pageId="11" pageNumber="12">WWII</collectionCode>
;
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[906,1153,235,262]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[966,1022,235,261]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:513
</bibRefCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="reference_group">
<paragraph blockId="11.[845,1451,299,517]" box="[845,1403,299,326]" pageId="11" pageNumber="12">
<treatmentCitationGroup box="[845,1403,299,326]" pageId="11" pageNumber="12">
<treatmentCitation author="Hewitt, J." box="[845,1403,299,326]" page="338" pageId="11" pageNumber="12" year="1935">
<taxonomicName authority="Hewitt 1935: 338" authorityName="Hewitt" authorityPageNumber="338" authorityYear="1935" box="[845,1403,299,326]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="galeata" subSpecies="damarensis">
<emphasis box="[845,1208,299,325]" italics="true" pageId="11" pageNumber="12">Pelomedusa galeata damarensis</emphasis>
<bibRefCitation author="Hewitt, J." box="[1215,1403,299,326]" pageId="11" pageNumber="12" pagination="283 - 357" refId="ref74257" refString="Hewitt, J. (1935) Some new forms of batrachians and reptiles from South Africa. Records of the Albany Museum, 4, 283 - 357." type="journal article" year="1935">
Hewitt 1935:
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</paragraph>
<paragraph blockId="11.[845,1451,299,517]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[845,952,330,356]" pageId="11" pageNumber="12">Syntypes</emphasis>
(2; both sampled genetically by
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[1319,1435,331,357]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[1379,1435,331,357]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
</bibRefCitation>
2014b): PEM
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and
<accessionNumber box="[1196,1292,363,390]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R14954" pageId="11" pageNumber="12">R 14954</accessionNumber>
; PEM
<accessionNumber box="[1363,1451,363,390]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/R14953" pageId="11" pageNumber="12">R14953</accessionNumber>
designated
<typeStatus box="[1035,1138,395,421]" pageId="11" pageNumber="12">lectotype</typeStatus>
by
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[1184,1447,395,422]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[1247,1305,395,421]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014b:504
</bibRefCitation>
; photographed in
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[1097,1345,427,454]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[1158,1215,427,453]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b:517
</bibRefCitation>
; sampled genetically by
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[1076,1278,459,485]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[1139,1198,459,485]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014b
</bibRefCitation>
and
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
<emphasis italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
<paragraph blockId="11.[845,1451,555,710]" box="[845,1400,555,581]" pageId="11" pageNumber="12">
<treatmentCitationGroup box="[845,1400,555,581]" pageId="11" pageNumber="12">
<treatmentCitation author="Mertens, R." box="[845,1400,555,581]" page="141" pageId="11" pageNumber="12" year="1937">
<taxonomicName authority="Mertens 1937: 141" authorityName="Mertens" authorityPageNumber="141" authorityYear="1937" box="[845,1400,555,581]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="subrufa" subSpecies="wettsteini">
<emphasis box="[845,1188,555,581]" italics="true" pageId="11" pageNumber="12">Pelomedusa subrufa wettsteini</emphasis>
<bibRefCitation author="Mertens, R." box="[1195,1400,555,581]" pageId="11" pageNumber="12" pagination="139 - 142" refId="ref77648" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens 1937:141</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[845,1451,555,710]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[845,954,587,613]" pageId="11" pageNumber="12">
<typeStatus box="[845,954,587,613]" pageId="11" pageNumber="12">Holotype</typeStatus>
</emphasis>
:
<collectionCode box="[968,1024,587,613]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="11" pageNumber="12">SMF</collectionCode>
7958 (
<bibRefCitation author="Mertens, R." box="[1102,1298,587,614]" pageId="11" pageNumber="12" pagination="1 - 106" refId="ref77676" refString="Mertens, R. (1967) Die herpetologische Sektion des Natur- Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen. Senckenbergiana Biologica, 48, 1 - 106." type="journal article" year="1967">Mertens 1967:52</bibRefCitation>
); photographed in
<bibRefCitation author="Fritz, U. &amp; Petzold, A. &amp; Kehlmaier, C. &amp; Kindler, C. &amp; Campbell, P. &amp; Hofmeyr, M. D. &amp; Branch, W. R." box="[1130,1447,619,646]" pageId="11" pageNumber="12" pagination="501 - 522" refId="ref72240" refString="Fritz, U., Petzold, A., Kehlmaier, C., Kindler, C., Campbell, P., Hofmeyr, M. D. &amp; Branch, W. R. (2014 b) Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae). Zootaxa, 3795 (5), 501 - 522. https: // doi. org / 10.11646 / zootaxa. 3795.5.1" type="journal article" year="2014">
Fritz
<emphasis box="[1207,1281,619,645]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
(2014b:518)
</bibRefCitation>
;
<emphasis bold="true" box="[906,1024,651,677]" pageId="11" pageNumber="12">
<typeStatus box="[906,1024,651,677]" pageId="11" pageNumber="12">paratypes</typeStatus>
</emphasis>
(4 according to the original description), presumably in the
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<emphasis bold="true" box="[810,1089,746,773]" italics="true" pageId="11" pageNumber="12">Pelomedusa variabilis</emphasis>
Petzold, Vargas-Ramírez, Kehlmaier, Vamberger, Branch, Du Preez, Hofmeyr, Meyers, Schleicher, Široký, &amp; Fritz 2014:543
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<typeStatus box="[810,919,875,901]" pageId="11" pageNumber="12">Holotype</typeStatus>
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:
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(photographed in
<bibRefCitation author="Petzold, A. &amp; Vargas-Ramirez, M. &amp; Kehlmaier, C. &amp; Vamberger, M. &amp; Branch, W. R. &amp; du Preez, L. &amp; Hofmeyr, M. D. &amp; Meyer, L. &amp; Schleicher, A. &amp; Siroky, P. &amp; Fritz, U." pageId="11" pageNumber="12" pagination="523 - 548" refId="ref79181" refString="Petzold, A., Vargas-Ramirez, M., Kehlmaier, C., Vamberger, M., Branch, W. R., du Preez, L., Hofmeyr, M. D., Meyer, L., Schleicher, A., Siroky, P. &amp; Fritz, U. (2014) A revision of African Helmeted Terrapins (Testudines: Pelomedusidae: Pelomedusa), with descriptions of six new species. Zootaxa, 3795, 523 - 548. https: // doi. org / 10.11646 / zootaxa. 3795.5.2" type="journal article" year="2014">
Petzold
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2014:541
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</emphasis>
(10): MTD
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- 33819 and 35041,
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.27.1 and 1927.8.27.240-242, and
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