468 lines
68 KiB
XML
468 lines
68 KiB
XML
<document id="871EAA6753BC1D12976DAA829AAEB93D" ID-CLB-Dataset="5952" ID-DOI="10.11646/zootaxa.3760.2.1" ID-GBIF-Dataset="7ba5f9ac-4317-459f-9829-2975de0a42ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="4908324" ID-ZooBank="C0E4CA6E-6D93-477E-99F6-14FFB652A2A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1623095615282" checkinUser="felipe" docAuthor="Michonneau, François & Paulay, Gustav" docDate="2014" docId="038F8787C101435DFF6CEF04FAB5FAA8" docLanguage="en" docName="zt03760p140.pdf" docOrigin="Zootaxa 3760 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyrella mookiei Michonneau & Paulay 2014, sp. nov." docType="treatment" docUuid="D3E02B0E-308A-4C67-A143-05DB64692E75" docUuidSource="ZooBank" docVersion="7" lastPageNumber="124" masterDocId="FFB6FFFFC110434AFFFBEA5BFFEEFFFB" masterDocTitle="Revision of the genus Phyrella (Holothuroidea: Dendrochirotida) with the description of a new species from Guam" masterLastPageNumber="140" masterPageNumber="101" pageNumber="118" updateTime="1699002177088" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F2E1579CD60FCA30DC9C1D0D6B1850C9">Revision of the genus Phyrella (Holothuroidea: Dendrochirotida) with the description of a new species from Guam</mods:title>
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<mods:namePart id="F5EA5B694B2DF8237C33EBB0731E1E27">Michonneau, François</mods:namePart>
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<mods:namePart id="6E315AB4DB5ACC211434AC868ED704FE">Paulay, Gustav</mods:namePart>
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<mods:title id="B2CDA4BF46169EEAD5D72B7C1FC44F40">Zootaxa</mods:title>
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<mods:part id="8D977DF71E4DB27BC466BB41AFD0376D">
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<mods:date id="911CE0CBFBACFF6DFA43DDFCD883D199">2014</mods:date>
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<mods:number id="EBEDAB956D9B56EE19DBB1D89FEDB108">2014-01-31</mods:number>
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<mods:number id="5DFF5751D8F97585450D2EB51165CA30">3760</mods:number>
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<treatment id="038F8787C101435DFF6CEF04FAB5FAA8" ID-DOI="http://doi.org/10.5281/zenodo.4908310" ID-GBIF-Taxon="183202678" ID-Zenodo-Dep="4908310" LSID="urn:lsid:zoobank.org:act:D3E02B0E-308A-4C67-A143-05DB64692E75" httpUri="http://treatment.plazi.org/id/038F8787C101435DFF6CEF04FAB5FAA8" lastPageId="23" lastPageNumber="124" pageId="17" pageNumber="118">
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<subSubSection id="C33C651AC101435BFF6CEF04FE27FA81" box="[151,457,1375,1402]" pageId="17" pageNumber="118" type="nomenclature">
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<paragraph id="8B993691C101435BFF6CEF04FE27FA81" blockId="17.[151,457,1375,1435]" box="[151,457,1375,1402]" pageId="17" pageNumber="118">
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<heading id="D0D181FDC101435BFF6CEF04FE27FA81" bold="true" box="[151,457,1375,1402]" fontSize="11" level="1" pageId="17" pageNumber="118" reason="1">
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<emphasis id="B952EA83C101435BFF6CEF04FE27FA81" bold="true" box="[151,457,1375,1402]" pageId="17" pageNumber="118">
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<taxonomicName id="4C264D12C101435BFF6CEF04FE8BFA82" ID-CoL="6VGG5" authority="Michonneau & Paulay, 2014" authorityName="Michonneau & Paulay" authorityYear="2014" box="[151,357,1375,1401]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="17" pageNumber="118" phylum="Echinodermata" rank="species" species="mookiei" status="sp. nov.">
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<emphasis id="B952EA83C101435BFF6CEF04FE8BFA82" bold="true" box="[151,357,1375,1401]" italics="true" pageId="17" pageNumber="118">Phyrella mookiei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26157F8C101435BFE97EF3BFE27FA81" box="[364,457,1376,1402]" pageId="17" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C651AC101435BFF6CEFD9FCD4FA1B" pageId="17" pageNumber="118" type="description">
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<paragraph id="8B993691C101435BFF6CEFD9FE96FA60" blockId="17.[151,457,1375,1435]" box="[151,376,1410,1435]" pageId="17" pageNumber="118">
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<figureCitation id="131D2A14C101435BFF6CEFD9FED5FA60" box="[151,315,1410,1435]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStart-6="FIGURE 16" captionStartId-0="18.[151,249,1358,1380]" captionStartId-1="19.[151,249,1542,1564]" captionStartId-2="19.[151,249,1978,2000]" captionStartId-3="20.[151,249,1777,1799]" captionStartId-4="21.[151,249,1909,1931]" captionStartId-5="22.[151,249,909,931]" captionStartId-6="22.[151,249,1941,1963]" captionTargetBox-0="[151,1436,373,1337]" captionTargetBox-1="[189,1400,177,1525]" captionTargetBox-2="[153,1398,1644,1956]" captionTargetBox-3="[195,1389,196,1753]" captionTargetBox-4="[196,1390,199,1881]" captionTargetBox-5="[168,1418,196,886]" captionTargetBox-6="[388,1201,1020,1915]" captionTargetId-0="figure-360@18.[151,1436,373,1337]" captionTargetId-1="figure-70@19.[189,1400,169,1525]" captionTargetId-2="figure-74@19.[150,1436,1644,1956]" captionTargetId-3="figure-54@20.[192,1394,193,1756]" captionTargetId-4="figure-59@21.[189,1397,193,1887]" captionTargetId-5="figure-60@22.[164,1423,193,888]" captionTargetId-6="figure-64@22.[381,1206,1012,1920]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="19" captionTargetPageId-3="20" captionTargetPageId-4="21" captionTargetPageId-5="22" captionTargetPageId-6="22" captionText-0="FIGURE 10. Phyrella mookiei in situ, Guam. The animal is buried in sediment; the oral disc with extended tentacles is visible. Oral disc diameter approximately 1 cm in diameter. Photograph by David Burdick." captionText-1="FIGURE 11. Live appearance of Phyrella mookiei UF 10336 (Holotype). A: Dorsal view, about 65 mm long; B: close-up view of oral region." captionText-2="FIGURE 12. Rosettes of Phyrella mookiei. A: UF 10336 (Holotype), mid-dorsal body wall; B: UF 4770 (Paratype), introvert; C: UF 4770 (Paratype), cloacal region. Scale bar, 20 µm." captionText-3="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." captionText-4="FIGURE 14. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert." captionText-5="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." captionText-6="FIGURE 16. Ossicle assemblage of Phyrella mookiei UF 11359 (Paratype). a–d: rods in podia that flank anal teeth." figureDoi-0="http://doi.org/10.5281/zenodo.4908360" figureDoi-1="http://doi.org/10.5281/zenodo.4908364" figureDoi-2="http://doi.org/10.5281/zenodo.4908368" figureDoi-3="http://doi.org/10.5281/zenodo.4908372" figureDoi-4="http://doi.org/10.5281/zenodo.4908376" figureDoi-5="http://doi.org/10.5281/zenodo.4908382" figureDoi-6="http://doi.org/10.5281/zenodo.4908384" httpUri-0="https://zenodo.org/record/4908360/files/figure.png" httpUri-1="https://zenodo.org/record/4908364/files/figure.png" httpUri-2="https://zenodo.org/record/4908368/files/figure.png" httpUri-3="https://zenodo.org/record/4908372/files/figure.png" httpUri-4="https://zenodo.org/record/4908376/files/figure.png" httpUri-5="https://zenodo.org/record/4908382/files/figure.png" httpUri-6="https://zenodo.org/record/4908384/files/figure.png" pageId="17" pageNumber="118">Figures 10–16</figureCitation>
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;
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<figureCitation id="131D2A14C101435BFEBDEFD9FE96FA60" box="[326,376,1410,1435]" captionStart="FIGURE 27" captionStartId="35.[151,249,1914,1936]" captionTargetBox="[177,1410,534,1892]" captionTargetId="figure-173@35.[177,1410,531,1892]" captionTargetPageId="35" captionText="FIGURE 27. Calcareous rings. a–c: Phyrella fragilis (a: UF 4096, b: WAM Z26012, c: WAM Z26013); d–e: Phyrella trapeza, syntypes (d: MCZ HOL1488, thick body wall; e: MCZ HOL1488, thin body wall); f: Phyrella thyonoides, holotype (MCZ HOL1654); g: Phyrella cf. thyonoides (UF 9601); h: Phyrella mookiei, paratype (UF 4770); i: Phyllophorus aculeatus, holotype (USNM 1871)." figureDoi="http://doi.org/10.5281/zenodo.4908418" httpUri="https://zenodo.org/record/4908418/files/figure.png" pageId="17" pageNumber="118">27 h</figureCitation>
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</paragraph>
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http://zoobank.org/
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<emphasis id="B952EA83C101435BFF6CEC56FE6FF9DD" bold="true" box="[151,385,1549,1574]" inLineHeading="true" pageId="17" pageNumber="118" reason="1">Material examined.</emphasis>
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<materialsCitation id="3B4E3CCCC101435BFE72EC56FBBFF9B1" ID-GBIF-Occurrence="3311708301" collectingDate="2010-06-12" collectionCode="UF" collectorName="Nathaniel Evans & Francois Michonneau & Gustav Paulay & Arthur Anker." country="Guam" county="Gun Beach" elevation="5" location="North" municipality="Tumon Bay" pageId="17" pageNumber="118" specimenCode="UF 10336" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549D8833C101435BFE72EC56FE18F9DD" box="[393,502,1549,1574]" pageId="17" pageNumber="118" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DB809EEAC101435BFDFAEC55FD9BF9DD" box="[513,629,1549,1574]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="17" pageNumber="118" type="Museum">UF 10336</specimenCode>
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end of
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, reef flat
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, coll.
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,
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<collectorName id="26D35347C101435BFE16EC6AFD06F9B1" box="[493,744,1585,1610]" pageId="17" pageNumber="118">François Michonneau</collectorName>
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,
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<collectorName id="26D35347C101435BFD0CEC6AFC71F9B1" box="[759,927,1585,1610]" pageId="17" pageNumber="118">Gustav Paulay</collectorName>
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,
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<collectorName id="26D35347C101435BFC55EC6AFBBFF9B1" box="[942,1105,1585,1610]" pageId="17" pageNumber="118">Arthur Anker.</collectorName>
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:
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,
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<collectingCountry id="F3317601C101435BFAABEC6AFA79F9B1" box="[1360,1431,1585,1610]" name="Guam" pageId="17" pageNumber="118">Guam</collectingCountry>
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outer reef flat,
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, coll.
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<collectorName id="26D35347C101435BFD3DEC0EFC83F995" box="[710,877,1621,1646]" pageId="17" pageNumber="118">Gustav Paulay</collectorName>
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;
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<materialsCitation id="3B4E3CCCC101435BFC82EC0DFCCCF969" ID-GBIF-Occurrence="3311708339" collectingDate="2011-09-17" collectionCode="UF" collectorName="David Burdick." country="Guam" elevation="2" location="Piti Power" pageId="17" pageNumber="118" specimenCode="UF 11539" specimenCount="1" typeStatus="paratype">
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,
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|
||
plant intake tunnel under road,
|
||
<quantity id="4CDE9B74C101435BFEE6EC22FEA4F96A" box="[285,330,1657,1682]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="118" unit="m" value="2.0">
|
||
<elevation id="000BD1A2C101435BFEE6EC22FEA4F96A" box="[285,330,1657,1682]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="118" unit="m" value="2.0">2 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF981051C101435BFEADEC22FDDCF969" box="[342,562,1657,1682]" pageId="17" pageNumber="118" value="2011-09-17">
|
||
<collectingDate id="EFDCE9B9C101435BFEADEC22FDDCF969" box="[342,562,1657,1682]" pageId="17" pageNumber="118" value="2011-09-17">17 September 2011</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26D35347C101435BFD8EEC22FCCCF969" box="[629,802,1657,1682]" pageId="17" pageNumber="118">David Burdick.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C651AC101435DFF3CECC6FDAAFC3C" lastPageId="23" lastPageNumber="124" pageId="17" pageNumber="118" type="description">
|
||
<paragraph id="8B993691C1014358FF3CECC6FEE5FEBB" blockId="17.[151,1436,1549,2006]" lastBlockId="18.[151,1436,151,320]" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118">
|
||
<emphasis id="B952EA83C101435BFF3CECC6FD9DF94D" bold="true" box="[199,627,1693,1718]" pageId="17" pageNumber="118">
|
||
<emphasis id="B952EA83C101435BFF3CECC6FEB7F94D" box="[199,345,1693,1718]" inLineHeading="true" pageId="17" pageNumber="118" reason="1">Description.</emphasis>
|
||
External morphology.
|
||
</emphasis>
|
||
Body wall soft, fairly thin, densely covered with tube feet. Beige (UF 4770, UF 11539) to off-white (UF 10336), with well-defined burgundy (UF 4770, UF 10336) to dark brown (UF 11539) spots either restricted to venter (UF 4770) or across entire body (UF 10336, UF 11539); area around introvert and cloaca with diffuse burgundy coloration in UF 10336 (
|
||
<figureCitation id="131D2A14C101435BFC76ED52FC1AF8D9" box="[909,1012,1801,1826]" captionStart="FIGURE 11" captionStartId="19.[151,249,1542,1564]" captionTargetBox="[189,1400,177,1525]" captionTargetId="figure-70@19.[189,1400,169,1525]" captionTargetPageId="19" captionText="FIGURE 11. Live appearance of Phyrella mookiei UF 10336 (Holotype). A: Dorsal view, about 65 mm long; B: close-up view of oral region." figureDoi="http://doi.org/10.5281/zenodo.4908364" httpUri="https://zenodo.org/record/4908364/files/figure.png" pageId="17" pageNumber="118">Fig. 11A</figureCitation>
|
||
). Coloration in preservative similar to live after 3 years. Oral disc marbled with burgundy and white (
|
||
<figureCitation id="131D2A14C101435BFC96ED76FC51F8BD" box="[877,959,1837,1862]" captionStart="FIGURE 10" captionStartId="18.[151,249,1358,1380]" captionTargetBox="[151,1436,373,1337]" captionTargetId="figure-360@18.[151,1436,373,1337]" captionTargetPageId="18" captionText="FIGURE 10. Phyrella mookiei in situ, Guam. The animal is buried in sediment; the oral disc with extended tentacles is visible. Oral disc diameter approximately 1 cm in diameter. Photograph by David Burdick." figureDoi="http://doi.org/10.5281/zenodo.4908360" httpUri="https://zenodo.org/record/4908360/files/figure.png" pageId="17" pageNumber="118">Fig. 10</figureCitation>
|
||
,
|
||
<figureCitation id="131D2A14C101435BFC36ED76FBDFF8BD" box="[973,1073,1837,1862]" captionStart="FIGURE 11" captionStartId="19.[151,249,1542,1564]" captionTargetBox="[189,1400,177,1525]" captionTargetId="figure-70@19.[189,1400,169,1525]" captionTargetPageId="19" captionText="FIGURE 11. Live appearance of Phyrella mookiei UF 10336 (Holotype). A: Dorsal view, about 65 mm long; B: close-up view of oral region." figureDoi="http://doi.org/10.5281/zenodo.4908364" httpUri="https://zenodo.org/record/4908364/files/figure.png" pageId="17" pageNumber="118">Fig. 11B</figureCitation>
|
||
). Tentacles dendritic with longstalk, with small dark spots at their extremities (
|
||
<figureCitation id="131D2A14C101435BFD33ED0AFCF5F891" box="[712,795,1873,1898]" captionStart="FIGURE 10" captionStartId="18.[151,249,1358,1380]" captionTargetBox="[151,1436,373,1337]" captionTargetId="figure-360@18.[151,1436,373,1337]" captionTargetPageId="18" captionText="FIGURE 10. Phyrella mookiei in situ, Guam. The animal is buried in sediment; the oral disc with extended tentacles is visible. Oral disc diameter approximately 1 cm in diameter. Photograph by David Burdick." figureDoi="http://doi.org/10.5281/zenodo.4908360" httpUri="https://zenodo.org/record/4908360/files/figure.png" pageId="17" pageNumber="118">Fig. 10</figureCitation>
|
||
,
|
||
<figureCitation id="131D2A14C101435BFCD1ED0AFC60F891" box="[810,910,1873,1898]" captionStart="FIGURE 11" captionStartId="19.[151,249,1542,1564]" captionTargetBox="[189,1400,177,1525]" captionTargetId="figure-70@19.[189,1400,169,1525]" captionTargetPageId="19" captionText="FIGURE 11. Live appearance of Phyrella mookiei UF 10336 (Holotype). A: Dorsal view, about 65 mm long; B: close-up view of oral region." figureDoi="http://doi.org/10.5281/zenodo.4908364" httpUri="https://zenodo.org/record/4908364/files/figure.png" pageId="17" pageNumber="118">Fig. 11B</figureCitation>
|
||
). Body relatively straight when fully relaxed, contracting to U-shaped.
|
||
<typeStatus id="549D8833C101435BFE40ED2EFDCDF875" box="[443,547,1909,1934]" pageId="17" pageNumber="118" type="holotype">Holotype</typeStatus>
|
||
<quantity id="4CDE9B74C101435BFDD6ED2EFD92F876" box="[557,636,1909,1934]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="118" unit="mm" value="60.0">60 mm</quantity>
|
||
long along dorsum,
|
||
<quantity id="4CDE9B74C101435BFC94ED2EFC53F876" box="[879,957,1909,1934]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="17" pageNumber="118" unit="mm" value="64.0">64 mm</quantity>
|
||
along venter and
|
||
<quantity id="4CDE9B74C101435BFB6AED2EFB0FF876" box="[1169,1249,1909,1934]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="118" unit="mm" value="18.0">18 mm</quantity>
|
||
wide; UF 4770, more contracted,
|
||
<quantity id="4CDE9B74C101435BFE90EDC2FE57F84A" box="[363,441,1945,1970]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="118" unit="mm" value="40.0">40 mm</quantity>
|
||
long along dorsum,
|
||
<quantity id="4CDE9B74C101435BFD3AEDC2FCFEF84A" box="[705,784,1945,1970]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="17" pageNumber="118" unit="mm" value="57.0">57 mm</quantity>
|
||
long along venter, and
|
||
<quantity id="4CDE9B74C101435BFBBBEDC2FB61F84A" box="[1088,1167,1945,1970]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="118" unit="mm" value="23.0">23 mm</quantity>
|
||
wide; UF 11539 very contracted,
|
||
<quantity id="4CDE9B74C101435BFEE4EDE6FE80F82E" box="[287,366,1981,2006]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="17" pageNumber="118" unit="mm" value="37.0">37 mm</quantity>
|
||
long along dorsum,
|
||
<quantity id="4CDE9B74C101435BFD90EDE6FD57F82E" box="[619,697,1981,2006]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="17" pageNumber="118" unit="mm" value="45.0">45 mm</quantity>
|
||
long along venter, and
|
||
<quantity id="4CDE9B74C101435BFC27EDE6FBC5F82E" box="[988,1067,1981,2006]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="118" unit="mm" value="16.0">16 mm</quantity>
|
||
wide. Introvert retracted in all specimens,
|
||
<quantity id="4CDE9B74C1024358FED9EACCFE8CFF54" box="[290,354,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="119" unit="mm" value="4.0">4 mm</quantity>
|
||
(UF 10336),
|
||
<quantity id="4CDE9B74C1024358FDF0EACCFDA5FF54" box="[523,587,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="119" unit="mm" value="6.0">6 mm</quantity>
|
||
(UF 4770), and
|
||
<quantity id="4CDE9B74C1024358FCE6EACCFC82FF54" box="[797,876,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="119" unit="mm" value="12.0">12 mm</quantity>
|
||
(UF 11539) long. Tube feet abundant, evenly dispersed across radial and inter-radial areas, slightly denser ventrally, generally same color as body wall near base, lightening distally. Eighteen tentacles (10 on outer circle, 8 on inner circle) in UF 4770, ~17 tentacles in
|
||
<typeStatus id="549D8833C1024358FAC1EA84FA72FF03" box="[1338,1436,223,248]" pageId="18" pageNumber="119" type="holotype">holotype</typeStatus>
|
||
based on live pictures (introvert retracted and not dissected). Cloacal membrane white; surrounded by 5 calcified anal teeth.
|
||
</paragraph>
|
||
<caption id="DF596619C1024358FF6CEF15FC30FA78" ID-DOI="http://doi.org/10.5281/zenodo.4908360" ID-Zenodo-Dep="4908360" httpUri="https://zenodo.org/record/4908360/files/figure.png" pageId="18" pageNumber="119" startId="18.[151,249,1358,1380]" targetBox="[151,1436,373,1337]" targetPageId="18">
|
||
<paragraph id="8B993691C1024358FF6CEF15FC30FA78" blockId="18.[151,1436,1358,1411]" pageId="18" pageNumber="119">
|
||
<emphasis id="B952EA83C1024358FF6CEF15FEF1FA98" bold="true" box="[151,287,1358,1380]" pageId="18" pageNumber="119">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C264D12C1024358FEDEEF15FE3CFA9F" authorityName="Michonneau & Paulay" authorityYear="2014" box="[293,466,1358,1380]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="18" pageNumber="119" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C1024358FEDEEF15FE3CFA9F" box="[293,466,1358,1380]" italics="true" pageId="18" pageNumber="119">Phyrella mookiei</emphasis>
|
||
</taxonomicName>
|
||
in situ, Guam. The animal is buried in sediment; the oral disc with extended tentacles is visible. Oral disc diameter approximately 1 cm in diameter. Photograph by David Burdick.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B993691C1024358FF3CEFF1FC4CF8FC" blockId="18.[151,1436,1450,1979]" pageId="18" pageNumber="119">
|
||
<emphasis id="B952EA83C1024358FF3CEFF1FD5FFA38" bold="true" box="[199,689,1450,1475]" inLineHeading="true" pageId="18" pageNumber="119" reason="1">Internal anatomy (UF 4770, UF 11539).</emphasis>
|
||
Polian vesicle single, white,
|
||
<quantity id="4CDE9B74C1024358FBE7EFF1FB97FA39" box="[1052,1145,1450,1475]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="18" pageNumber="119" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
|
||
long. Stone canal short (
|
||
<quantity id="4CDE9B74C1024358FF64EF95FF0BFA1C" box="[159,229,1486,1511]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="119" unit="mm" value="2.0">2 mm</quantity>
|
||
), partially embedded in membrane surrounding calcareous ring. Madreporite free, small (<
|
||
<quantity id="4CDE9B74C1024358FAB0EF95FA7EFA1C" box="[1355,1424,1486,1511]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="119" unit="mm" value="1.0">1 mm</quantity>
|
||
), spherical. Gonads well developed in both dissected specimens, both tufts extending almost entire length of animal; gonadal tubules unbranched to dichotomously branched 1–3 times; white with minute burgundy spots (similar color to body wall), with spots more common proximally and distally (UF 4770); in UF 10336, oocytes are clearly discernible, white, aligned in a row, encased in the thin, translucent gonadal tube wall, with abundant, minute, burgundy spots. Both dissected specimens have eviscerated, retaining gonads, anterior fragment of esophagus, cloaca and both respiratory trees. Intestine (retained separately for UF 4770) distended where filled with fine sand and other debris. Respiratory trees white with minute burgundy spots that are much less abundant in UF 4770 than in UF 11549, both branches extending almost entire length of animal.
|
||
</paragraph>
|
||
<paragraph id="8B993691C1024358FF3CED49FA66F840" blockId="18.[151,1436,1450,1979]" pageId="18" pageNumber="119">
|
||
<emphasis id="B952EA83C1024358FF3CED49FE61F8D0" bold="true" box="[199,399,1810,1835]" inLineHeading="true" pageId="18" pageNumber="119" reason="1">Calcareous ring.</emphasis>
|
||
Calcareous ring embedded in a fairly thin membrane, with highly fragmented radial and inter-radial elements (
|
||
<figureCitation id="131D2A14C1024358FE63ED6DFE14F8B4" box="[408,506,1846,1871]" captionStart="FIGURE 27" captionStartId="35.[151,249,1914,1936]" captionTargetBox="[177,1410,534,1892]" captionTargetId="figure-173@35.[177,1410,531,1892]" captionTargetPageId="35" captionText="FIGURE 27. Calcareous rings. a–c: Phyrella fragilis (a: UF 4096, b: WAM Z26012, c: WAM Z26013); d–e: Phyrella trapeza, syntypes (d: MCZ HOL1488, thick body wall; e: MCZ HOL1488, thin body wall); f: Phyrella thyonoides, holotype (MCZ HOL1654); g: Phyrella cf. thyonoides (UF 9601); h: Phyrella mookiei, paratype (UF 4770); i: Phyllophorus aculeatus, holotype (USNM 1871)." figureDoi="http://doi.org/10.5281/zenodo.4908418" httpUri="https://zenodo.org/record/4908418/files/figure.png" pageId="18" pageNumber="119">Fig. 27h</figureCitation>
|
||
). Anterior margins of radial plates unequally divided by well-formed notches. Anterior margins of inter-radial plates arrow-head shaped, more pointed and elongated in UF 4770 than UF 10336. Radial and inter-radial elements closely associated along most of their length. Inter-radials each composed of 5–12 elements. Radials each composed of 14–16 elements. Radials form posteriorly tails that curve to point anteriorly.
|
||
</paragraph>
|
||
<caption id="DF596619C1034359FF6CEC5DFEC6F9C0" ID-DOI="http://doi.org/10.5281/zenodo.4908364" ID-Zenodo-Dep="4908364" httpUri="https://zenodo.org/record/4908364/files/figure.png" pageId="19" pageNumber="120" startId="19.[151,249,1542,1564]" targetBox="[189,1400,177,1525]" targetPageId="19">
|
||
<paragraph id="8B993691C1034359FF6CEC5DFEC6F9C0" blockId="19.[151,1435,1542,1595]" pageId="19" pageNumber="120">
|
||
<emphasis id="B952EA83C1034359FF6CEC5DFEF0F9E0" bold="true" box="[151,286,1542,1564]" pageId="19" pageNumber="120">FIGURE 11.</emphasis>
|
||
Live appearance of
|
||
<taxonomicName id="4C264D12C1034359FE17EC5DFD50F9E7" authority="UF" authorityName="UF" box="[492,702,1542,1564]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="19" pageNumber="120" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C1034359FE17EC5DFD76F9E7" box="[492,664,1542,1564]" italics="true" pageId="19" pageNumber="120">Phyrella mookiei</emphasis>
|
||
UF
|
||
</taxonomicName>
|
||
10336 (Holotype). A: Dorsal view, about 65 mm long; B: close-up view of oral region.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596619C1034359FF6CEDE1FD36F814" ID-DOI="http://doi.org/10.5281/zenodo.4908368" ID-Zenodo-Dep="4908368" httpUri="https://zenodo.org/record/4908368/files/figure.png" pageId="19" pageNumber="120" startId="19.[151,249,1978,2000]" targetBox="[153,1398,1644,1956]" targetPageId="19">
|
||
<paragraph id="8B993691C1034359FF6CEDE1FD36F814" blockId="19.[151,1436,1978,2031]" pageId="19" pageNumber="120">
|
||
<emphasis id="B952EA83C1034359FF6CEDE1FEF1F834" bold="true" box="[151,287,1978,2000]" pageId="19" pageNumber="120">FIGURE 12.</emphasis>
|
||
Rosettes of
|
||
<taxonomicName id="4C264D12C1034359FE65EDE1FDA5F82B" box="[414,587,1978,2000]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="19" pageNumber="120" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C1034359FE65EDE1FDA5F82B" box="[414,587,1978,2000]" italics="true" pageId="19" pageNumber="120">Phyrella mookiei</emphasis>
|
||
</taxonomicName>
|
||
. A: UF 10336 (Holotype), mid-dorsal body wall; B: UF 4770 (Paratype), introvert; C: UF 4770 (Paratype), cloacal region. Scale bar, 20
|
||
<emphasis id="B952EA83C1034359FD54ED81FD50F814" box="[687,702,2010,2031]" italics="true" pageId="19" pageNumber="120">µ</emphasis>
|
||
m.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596619C104435EFF6CECAAFB7DF8DD" ID-DOI="http://doi.org/10.5281/zenodo.4908372" ID-Zenodo-Dep="4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="20" pageNumber="121" startId="20.[151,249,1777,1799]" targetBox="[195,1389,196,1753]" targetPageId="20">
|
||
<paragraph id="8B993691C104435EFF6CECAAFB7DF8DD" blockId="20.[151,1436,1777,1830]" pageId="20" pageNumber="121">
|
||
<emphasis id="B952EA83C104435EFF6CECAAFEF1F8FD" bold="true" box="[151,287,1777,1799]" pageId="20" pageNumber="121">FIGURE 13.</emphasis>
|
||
Ossicle assemblage of
|
||
<taxonomicName id="4C264D12C104435EFDF6ECAAFD0FF8FC" authority="UF" authorityName="UF" box="[525,737,1777,1799]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="20" pageNumber="121" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C104435EFDF6ECAAFD54F8FC" box="[525,698,1777,1799]" italics="true" pageId="20" pageNumber="121">Phyrella mookiei</emphasis>
|
||
UF
|
||
</taxonomicName>
|
||
10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596619C105435FFF6CED2EFE44F833" ID-DOI="http://doi.org/10.5281/zenodo.4908376" ID-Zenodo-Dep="4908376" httpUri="https://zenodo.org/record/4908376/files/figure.png" pageId="21" pageNumber="122" startId="21.[151,249,1909,1931]" targetBox="[196,1390,199,1881]" targetPageId="21">
|
||
<paragraph id="8B993691C105435FFF6CED2EFE44F833" blockId="21.[151,1436,1909,1992]" pageId="21" pageNumber="122">
|
||
<emphasis id="B952EA83C105435FFF6CED2EFECFF871" bold="true" box="[151,289,1909,1931]" pageId="21" pageNumber="122">FIGURE 14.</emphasis>
|
||
Ossicle assemblage of
|
||
<taxonomicName id="4C264D12C105435FFDEDED2EFD03F870" authority="UF" authorityName="UF" box="[534,749,1909,1931]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="21" pageNumber="122" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C105435FFDEDED2EFD2BF870" box="[534,709,1909,1931]" italics="true" pageId="21" pageNumber="122">Phyrella mookiei</emphasis>
|
||
UF
|
||
</taxonomicName>
|
||
4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596619C106435CFF6CE9D6FDF8FC39" ID-DOI="http://doi.org/10.5281/zenodo.4908382" ID-Zenodo-Dep="4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="22" pageNumber="123" startId="22.[151,249,909,931]" targetBox="[168,1418,196,886]" targetPageId="22">
|
||
<paragraph id="8B993691C106435CFF6CE9D6FDF8FC39" blockId="22.[151,1435,909,962]" pageId="22" pageNumber="123">
|
||
<emphasis id="B952EA83C106435CFF6CE9D6FEF1FC58" bold="true" box="[151,287,909,931]" pageId="22" pageNumber="123">FIGURE 15.</emphasis>
|
||
Ossicle assemblage of
|
||
<taxonomicName id="4C264D12C106435CFDF4E9D6FD0CFC58" authority="UF" authorityName="UF" box="[527,738,909,931]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="123" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C106435CFDF4E9D6FD55FC58" box="[527,699,909,931]" italics="true" pageId="22" pageNumber="123">Phyrella mookiei</emphasis>
|
||
UF
|
||
</taxonomicName>
|
||
4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF596619C106435CFF6CEDCEFAC3F850" ID-DOI="http://doi.org/10.5281/zenodo.4908384" ID-Zenodo-Dep="4908384" box="[151,1325,1941,1963]" httpUri="https://zenodo.org/record/4908384/files/figure.png" pageId="22" pageNumber="123" startId="22.[151,249,1941,1963]" targetBox="[388,1201,1020,1915]" targetPageId="22">
|
||
<paragraph id="8B993691C106435CFF6CEDCEFAC3F850" blockId="22.[151,1325,1941,1963]" box="[151,1325,1941,1963]" pageId="22" pageNumber="123">
|
||
<emphasis id="B952EA83C106435CFF6CEDCEFEF1F851" bold="true" box="[151,287,1941,1963]" pageId="22" pageNumber="123">FIGURE 16.</emphasis>
|
||
Ossicle assemblage of
|
||
<taxonomicName id="4C264D12C106435CFDF6EDCEFD0EF850" authority="UF" authorityName="UF" box="[525,736,1941,1963]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="22" pageNumber="123" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C106435CFDF6EDCEFD57F850" box="[525,697,1941,1963]" italics="true" pageId="22" pageNumber="123">Phyrella mookiei</emphasis>
|
||
UF
|
||
</taxonomicName>
|
||
11359 (Paratype). a–d: rods in podia that flank anal teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B993691C107435DFF3CEACCFDAAFC3C" blockId="23.[151,1436,151,1365]" pageId="23" pageNumber="124">
|
||
<emphasis id="B952EA83C107435DFF3CEACCFCF0FF4B" bold="true" box="[199,798,151,176]" pageId="23" pageNumber="124">
|
||
<emphasis id="B952EA83C107435DFF3CEACCFE5CFF4B" box="[199,434,151,176]" inLineHeading="true" pageId="23" pageNumber="124" reason="1">Ossicle assemblage.</emphasis>
|
||
Dorsal and ventral body wall
|
||
</emphasis>
|
||
with similar ossicle complements of tables and rosettes (
|
||
<figureCitation id="131D2A14C107435DFF64EAE7FEE4FF2E" box="[159,266,188,213]" captionStart="FIGURE 12" captionStartId="19.[151,249,1978,2000]" captionTargetBox="[153,1398,1644,1956]" captionTargetId="figure-74@19.[150,1436,1644,1956]" captionTargetPageId="19" captionText="FIGURE 12. Rosettes of Phyrella mookiei. A: UF 10336 (Holotype), mid-dorsal body wall; B: UF 4770 (Paratype), introvert; C: UF 4770 (Paratype), cloacal region. Scale bar, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4908368" httpUri="https://zenodo.org/record/4908368/files/figure.png" pageId="23" pageNumber="124">Fig. 12A</figureCitation>
|
||
), latter more abundant dorsally than ventrally. Tables of variable forms; disc
|
||
<quantity id="4CDE9B74C107435DFB7CEAE7FB1BFF2F" box="[1159,1269,188,213]" metricMagnitude="1" metricUnit="m" metricValue="9.75" metricValueMax="12.5" metricValueMin="7.0" pageId="23" pageNumber="124" unit="m" value="97.5" valueMax="125.0" valueMin="70.0">
|
||
70–125
|
||
<emphasis id="B952EA83C107435DFB1EEAE6FB1BFF2F" box="[1253,1269,189,212]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m in diameter, with
|
||
<quantity id="4CDE9B74C107435DFF2BEA84FEC0FF03" box="[208,302,223,248]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="23" pageNumber="124" unit="m" value="17.5" valueMax="25.0" valueMin="10.0">
|
||
10–25
|
||
<emphasis id="B952EA83C107435DFEE5EABAFEC0FF03" box="[286,302,225,248]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m diameter central perforation, and 4–15 (
|
||
<quantity id="4CDE9B74C107435DFCF4EA84FCD5FF03" box="[783,827,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="23" pageNumber="124" unit="in" value="8.0">8 in</quantity>
|
||
the most symmetrical tables) holes arranged in a ring (
|
||
<figureCitation id="131D2A14C107435DFF64EB5FFF10FEE6" box="[159,254,260,285]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13d</figureCitation>
|
||
) or dispersed more irregularly in more highly perforated discs (
|
||
<figureCitation id="131D2A14C107435DFC2FEB5FFBDEFEE6" box="[980,1072,260,285]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13f</figureCitation>
|
||
), disc margin smooth (
|
||
<figureCitation id="131D2A14C107435DFACEEB5FFA7DFEE6" box="[1333,1427,260,285]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13e</figureCitation>
|
||
) to undulating (
|
||
<figureCitation id="131D2A14C107435DFEB2EB7CFE4BFEBB" box="[329,421,295,320]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13f</figureCitation>
|
||
), to partially (
|
||
<figureCitation id="131D2A14C107435DFDA3EB7CFD59FEBB" box="[600,695,295,320]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13a</figureCitation>
|
||
,
|
||
<figureCitation id="131D2A14C107435DFD31EB7CFCDEFEBB" box="[714,816,295,320]" captionStart="FIGURE 14" captionStartId="21.[151,249,1909,1931]" captionTargetBox="[196,1390,199,1881]" captionTargetId="figure-59@21.[189,1397,193,1887]" captionTargetPageId="21" captionText="FIGURE 14. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert." figureDoi="http://doi.org/10.5281/zenodo.4908376" httpUri="https://zenodo.org/record/4908376/files/figure.png" pageId="23" pageNumber="124">
|
||
Fig.
|
||
<quantity id="4CDE9B74C107435DFD06EB7CFCDEFEBB" box="[765,816,295,320]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="23" pageNumber="124" unit="m" value="14.0">14m</quantity>
|
||
</figureCitation>
|
||
) or completely serrated, barely raised; spire absent (
|
||
<figureCitation id="131D2A14C107435DFF64EB17FEF3FE9E" box="[159,285,332,357]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">
|
||
Fig.
|
||
<quantity id="4CDE9B74C107435DFF2FEB17FF12FE9E" box="[212,252,332,357]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="23" pageNumber="124" unit="g" value="13.0">13g</quantity>
|
||
–h
|
||
</figureCitation>
|
||
), to partially (
|
||
<figureCitation id="131D2A14C107435DFE2BEB17FDA7FE9E" box="[464,585,332,357]" captionStart="FIGURE 14" captionStartId="21.[151,249,1909,1931]" captionTargetBox="[196,1390,199,1881]" captionTargetId="figure-59@21.[189,1397,193,1887]" captionTargetPageId="21" captionText="FIGURE 14. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert." figureDoi="http://doi.org/10.5281/zenodo.4908376" httpUri="https://zenodo.org/record/4908376/files/figure.png" pageId="23" pageNumber="124">Fig. 14a–e</figureCitation>
|
||
) or well developed (
|
||
<figureCitation id="131D2A14C107435DFCB6EB17FC42FE9E" box="[845,940,332,357]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13a</figureCitation>
|
||
), when complete, formed by four pillars connected by a single cross-beam half way along spire; crown variable in diameter when present, forming a spiny ring when well developed (
|
||
<figureCitation id="131D2A14C107435DFE2AEBCFFDAAFE56" box="[465,580,404,429]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13a,c</figureCitation>
|
||
). Podia with perforated plates,
|
||
<quantity id="4CDE9B74C107435DFC57EBCFFBC9FE57" box="[940,1063,404,429]" metricMagnitude="2" metricUnit="m" metricValue="1.575" metricValueMax="1.85" metricValueMin="1.3" pageId="23" pageNumber="124" unit="m" value="157.5" valueMax="185.0" valueMin="130.0">
|
||
130–185
|
||
<emphasis id="B952EA83C107435DFBECEBCEFBC9FE57" box="[1047,1063,405,428]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m long,
|
||
<quantity id="4CDE9B74C107435DFB73EBCFFB1BFE57" box="[1160,1269,404,429]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="23" pageNumber="124" unit="m" value="85.0" valueMax="100.0" valueMin="70.0">
|
||
70–100
|
||
<emphasis id="B952EA83C107435DFB1EEBCEFB1BFE57" box="[1253,1269,405,428]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m wide; wider and not as elongated as in
|
||
<taxonomicName id="4C264D12C107435DFE46EBE2FDC4FE2B" box="[445,554,439,464]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="fragilis">
|
||
<emphasis id="B952EA83C107435DFE46EBE2FDC4FE2B" box="[445,554,439,464]" italics="true" pageId="23" pageNumber="124">P. fragilis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C264D12C107435DFDB4EBE2FD0DFE2B" baseAuthorityName="Clark" baseAuthorityYear="1938" box="[591,739,439,464]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="thyonoides">
|
||
<emphasis id="B952EA83C107435DFDB4EBE2FD0DFE2B" box="[591,739,439,464]" italics="true" pageId="23" pageNumber="124">P. thyonoides</emphasis>
|
||
</taxonomicName>
|
||
; with large holes in center and smaller toward periphery; with large well-defined serrations, along at least part of the more convex margin (
|
||
<figureCitation id="131D2A14C107435DFC06EB87FB9BFE0E" box="[1021,1141,476,501]" captionStart="FIGURE 13" captionStartId="20.[151,249,1777,1799]" captionTargetBox="[195,1389,196,1753]" captionTargetId="figure-54@20.[192,1394,193,1756]" captionTargetPageId="20" captionText="FIGURE 13. Ossicle assemblage of Phyrella mookiei UF 10336 (Holotype). a–k: tables (a–h) and perforated plates (i–k) from mid-dorsal body wall; l–o: perforated plates (l–n) and tube foot end plate (o) from ventral body wall." figureDoi="http://doi.org/10.5281/zenodo.4908372" httpUri="https://zenodo.org/record/4908372/files/figure.png" pageId="23" pageNumber="124">Fig. 13l–o</figureCitation>
|
||
;
|
||
<figureCitation id="131D2A14C107435DFB79EB87FAC3FE0E" box="[1154,1325,476,501]" captionStart="FIGURE 14" captionStartId="21.[151,249,1909,1931]" captionTargetBox="[196,1390,199,1881]" captionTargetId="figure-59@21.[189,1397,193,1887]" captionTargetPageId="21" captionText="FIGURE 14. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert." figureDoi="http://doi.org/10.5281/zenodo.4908376" httpUri="https://zenodo.org/record/4908376/files/figure.png" pageId="23" pageNumber="124">Fig. 14f–i, o–p</figureCitation>
|
||
).
|
||
<emphasis id="B952EA83C107435DFAB9EB87FEE0FDE3" bold="true" pageId="23" pageNumber="124">Cloacal body wall</emphasis>
|
||
with tables and abundant rosettes. Tables of variable form ranging from similar to body wall tables (
|
||
<figureCitation id="131D2A14C107435DFF64E87FFF11FDC6" box="[159,255,548,573]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">
|
||
Fig.
|
||
<quantity id="4CDE9B74C107435DFF2FE87FFF11FDC6" box="[212,255,548,573]" metricMagnitude="-2" metricUnit="kg" metricValue="1.5" pageId="23" pageNumber="124" unit="g" value="15.0">15g</quantity>
|
||
</figureCitation>
|
||
), to others that show a much greater number of disc perforations (
|
||
<figureCitation id="131D2A14C107435DFC15E87FFBBEFDC6" box="[1006,1104,548,573]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">Fig. 15k</figureCitation>
|
||
); disc
|
||
<quantity id="4CDE9B74C107435DFB61E87FFB16FDC7" box="[1178,1272,548,573]" metricMagnitude="1" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="23" pageNumber="124" unit="m" value="80.0" valueMax="85.0" valueMin="75.0">
|
||
75–85
|
||
<emphasis id="B952EA83C107435DFB13E87EFB16FDC7" box="[1256,1272,549,572]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m in diameter, rim typically more serrated in tables of cloacal region than elsewhere in body wall; spire absent (
|
||
<figureCitation id="131D2A14C107435DFAD4E81CFA60FD9B" box="[1327,1422,583,608]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">Fig. 15e</figureCitation>
|
||
), incomplete (
|
||
<figureCitation id="131D2A14C107435DFEDCE837FE61FD7E" box="[295,399,620,645]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">
|
||
Fig.
|
||
<quantity id="4CDE9B74C107435DFEA0E837FE61FD7E" box="[347,399,620,645]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="124" unit="m" value="15.0">15m</quantity>
|
||
</figureCitation>
|
||
), or complete, ending in a crown with teeth or lateral projections (
|
||
<figureCitation id="131D2A14C107435DFB5CE837FAE7FD7E" box="[1191,1289,620,645]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">Fig. 15k</figureCitation>
|
||
). Podia near cloaca with abundant rosettes (
|
||
<figureCitation id="131D2A14C107435DFE0BE8D4FDBAFD53" box="[496,596,655,680]" captionStart="FIGURE 12" captionStartId="19.[151,249,1978,2000]" captionTargetBox="[153,1398,1644,1956]" captionTargetId="figure-74@19.[150,1436,1644,1956]" captionTargetPageId="19" captionText="FIGURE 12. Rosettes of Phyrella mookiei. A: UF 10336 (Holotype), mid-dorsal body wall; B: UF 4770 (Paratype), introvert; C: UF 4770 (Paratype), cloacal region. Scale bar, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4908368" httpUri="https://zenodo.org/record/4908368/files/figure.png" pageId="23" pageNumber="124">Fig. 12C</figureCitation>
|
||
) and perforated plates; latter smaller (
|
||
<quantity id="4CDE9B74C107435DFC06E8D4FB87FD53" box="[1021,1129,655,680]" metricMagnitude="1" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="23" pageNumber="124" unit="m" value="90.0" valueMax="100.0" valueMin="80.0">
|
||
80–100
|
||
<emphasis id="B952EA83C107435DFBA2E8CAFB87FD53" box="[1113,1129,657,680]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m long) and often narrower (
|
||
<quantity id="4CDE9B74C107435DFF65E8EFFF10FD37" box="[158,254,692,717]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="23" pageNumber="124" unit="m" value="50.0" valueMax="60.0" valueMin="40.0">
|
||
40–60
|
||
<emphasis id="B952EA83C107435DFF15E8EEFF10FD37" box="[238,254,693,716]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m wide) than those elsewhere in body wall (
|
||
<figureCitation id="131D2A14C107435DFD0EE8EFFC87FD36" box="[757,873,692,717]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">Fig. 15c,d</figureCitation>
|
||
), some with unusual lateral extensions (
|
||
<figureCitation id="131D2A14C107435DFAD4E8EFFA61FD36" box="[1327,1423,692,717]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">Fig. 15a</figureCitation>
|
||
). Podia that flank anal teeth with rods and abundant rosettes; rods often with marked curvature; ends with 2–6 holes and sometimes small spines (
|
||
<figureCitation id="131D2A14C107435DFE19E8A7FDB0FCEE" box="[482,606,764,789]" captionStart="FIGURE 16" captionStartId="22.[151,249,1941,1963]" captionTargetBox="[388,1201,1020,1915]" captionTargetId="figure-64@22.[381,1206,1012,1920]" captionTargetPageId="22" captionText="FIGURE 16. Ossicle assemblage of Phyrella mookiei UF 11359 (Paratype). a–d: rods in podia that flank anal teeth." figureDoi="http://doi.org/10.5281/zenodo.4908384" httpUri="https://zenodo.org/record/4908384/files/figure.png" pageId="23" pageNumber="124">Fig. 16a–d</figureCitation>
|
||
).
|
||
<emphasis id="B952EA83C107435DFD88E8A7FD0FFCEE" bold="true" box="[627,737,764,789]" pageId="23" pageNumber="124">Introvert</emphasis>
|
||
with similar ossicle assemblage to cloacal region; with tables and abundant rosettes. Tables
|
||
<quantity id="4CDE9B74C107435DFDF9E944FD9AFCC3" box="[514,628,799,824]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.05" metricValueMin="0.95" pageId="23" pageNumber="124" unit="m" value="100.0" valueMax="105.0" valueMin="95.0">
|
||
95–105
|
||
<emphasis id="B952EA83C107435DFD9FE97AFD9AFCC3" box="[612,628,801,824]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m in diameter. Podia with abundant rosettes, and perforated plates. Perforated plates variable, ranging from similar to those from the body wall (
|
||
<figureCitation id="131D2A14C107435DFBF3E91FFB4EFCA6" box="[1032,1184,836,861]" captionStart="FIGURE 14" captionStartId="21.[151,249,1909,1931]" captionTargetBox="[196,1390,199,1881]" captionTargetId="figure-59@21.[189,1397,193,1887]" captionTargetPageId="21" captionText="FIGURE 14. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert." figureDoi="http://doi.org/10.5281/zenodo.4908376" httpUri="https://zenodo.org/record/4908376/files/figure.png" pageId="23" pageNumber="124">Fig. 14u–v, x</figureCitation>
|
||
), to much smaller (
|
||
<quantity id="4CDE9B74C107435DFA85E91FFF49FC7B" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="124" unit="m" value="80.0">
|
||
80
|
||
<emphasis id="B952EA83C107435DFF6CE932FF49FC7B" box="[151,167,873,896]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m long), almost circular, with small perforations (
|
||
<figureCitation id="131D2A14C107435DFD27E93CFCB9FC7B" box="[732,855,871,896]" captionStart="FIGURE 14" captionStartId="21.[151,249,1909,1931]" captionTargetBox="[196,1390,199,1881]" captionTargetId="figure-59@21.[189,1397,193,1887]" captionTargetPageId="21" captionText="FIGURE 14. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–i: tables (a–e) and perforated plates (f–i) from mid-dorsal body wall; j–p: tables (j–n) and perforated plates (o–p) from mid-ventral body wall; q–z: tables (q–t) and perforated plates (u–z) from introvert." figureDoi="http://doi.org/10.5281/zenodo.4908376" httpUri="https://zenodo.org/record/4908376/files/figure.png" pageId="23" pageNumber="124">Fig. 14w,y</figureCitation>
|
||
).
|
||
<emphasis id="B952EA83C107435DFC97E93CFC35FC7B" bold="true" box="[876,987,871,896]" pageId="23" pageNumber="124">Tentacles</emphasis>
|
||
with rods,
|
||
<quantity id="4CDE9B74C107435DFBA6E93CFB55FC7B" box="[1117,1211,871,896]" metricMagnitude="1" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="23" pageNumber="124" unit="m" value="57.5" valueMax="65.0" valueMin="50.0">
|
||
50–65
|
||
<emphasis id="B952EA83C107435DFB50E932FB55FC7B" box="[1195,1211,873,896]" italics="true" pageId="23" pageNumber="124">µ</emphasis>
|
||
</quantity>
|
||
m long, with one or no perforation at ends (
|
||
<figureCitation id="131D2A14C107435DFE55E9D7FDE1FC5E" box="[430,527,908,933]" captionStart="FIGURE 15" captionStartId="22.[151,249,909,931]" captionTargetBox="[168,1418,196,886]" captionTargetId="figure-60@22.[164,1423,193,888]" captionTargetPageId="22" captionText="FIGURE 15. Ossicle assemblage of Phyrella mookiei UF 4770 (Paratype). a–m: perforated plates (a–e) and tables (f–m) from cloacal region; n: rods from tentacles." figureDoi="http://doi.org/10.5281/zenodo.4908382" httpUri="https://zenodo.org/record/4908382/files/figure.png" pageId="23" pageNumber="124">Fig. 15n</figureCitation>
|
||
). No ossicles observed in cloacal, longitudinal and retractor muscles; cloaca; intestine; gonads; and respiratory tree.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C651AC107435DFF3CE98FFF1BFBCE" pageId="23" pageNumber="124" type="etymology">
|
||
<paragraph id="8B993691C107435DFF3CE98FFF1BFBCE" blockId="23.[151,1436,151,1365]" pageId="23" pageNumber="124">
|
||
<emphasis id="B952EA83C107435DFF3CE98FFEA0FC16" bold="true" box="[199,334,980,1005]" inLineHeading="true" pageId="23" pageNumber="124" reason="1">Etymology.</emphasis>
|
||
Named after Mookie, the dog of our collection assistant Ms. Mandy Bemis, because the “woolly” appearance and color of this species is similar to the soft coat of wheaten terriers, the breed to which Mookie belongs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C651AC107435DFF3CEE64FB76FAF0" pageId="23" pageNumber="124" type="discussion">
|
||
<paragraph id="8B993691C107435DFF3CEE64FB76FAF0" blockId="23.[151,1436,151,1365]" pageId="23" pageNumber="124">
|
||
<emphasis id="B952EA83C107435DFF3CEE64FED5FBA3" bold="true" box="[199,315,1087,1112]" inLineHeading="true" pageId="23" pageNumber="124" reason="1">Remarks.</emphasis>
|
||
<taxonomicName id="4C264D12C107435DFEBCEE64FDE5FBA3" box="[327,523,1087,1112]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C107435DFEBCEE64FDE5FBA3" box="[327,523,1087,1112]" italics="true" pageId="23" pageNumber="124">Phyrella mookiei</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4C264D12C107435DFD68EE1AFCEAFBA3" box="[659,772,1087,1112]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="fragilis">
|
||
<emphasis id="B952EA83C107435DFD68EE1AFCEAFBA3" box="[659,772,1087,1112]" italics="true" pageId="23" pageNumber="124">P. fragilis</emphasis>
|
||
</taxonomicName>
|
||
externally, but the ossicle assemblage and color pattern distinguish the two. In
|
||
<emphasis id="B952EA83C107435DFE5BEE3EFDF3FB86" box="[416,541,1124,1149]" italics="true" pageId="23" pageNumber="124">
|
||
<taxonomicName id="4C264D12C107435DFE5BEE3EFDF6FB86" box="[416,536,1124,1149]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="mookiei">P. mookiei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holes in the disc of the tables are smaller, the margins of the disc are often at least partially serrated, the crowns are often more complete and wider; the perforated plates of the podia are shorter and wider; and the irregular tables in cloacal region have many holes in their rim. Rosettes occur throughout the body wall in
|
||
<taxonomicName id="4C264D12C107435DFEC9EE8AFE47FB13" box="[306,425,1231,1256]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C107435DFEC9EE8AFE47FB13" box="[306,425,1231,1256]" italics="true" pageId="23" pageNumber="124">P. mookiei</emphasis>
|
||
</taxonomicName>
|
||
but are restricted to introvert and near-cloacal body wall in other
|
||
<taxonomicName id="4C264D12C107435DFB57EE94FAE5FB13" authorityName="Heding & Panning" authorityYear="1954" box="[1196,1291,1231,1256]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B952EA83C107435DFB57EE94FAE5FB13" box="[1196,1291,1231,1256]" italics="true" pageId="23" pageNumber="124">Phyrella</emphasis>
|
||
</taxonomicName>
|
||
. While both species have a beige-tan base color, the well-defined dark spotting distinguishes
|
||
<taxonomicName id="4C264D12C107435DFBE6EEAEFB7CFAF6" box="[1053,1170,1268,1293]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C107435DFBE6EEAEFB7CFAF6" box="[1053,1170,1268,1293]" italics="true" pageId="23" pageNumber="124">P. mookiei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C651AC107435DFF3CEF4CFAB5FAA8" pageId="23" pageNumber="124" type="biology_ecology">
|
||
<paragraph id="8B993691C107435DFF3CEF4CFAB5FAA8" blockId="23.[151,1436,151,1365]" pageId="23" pageNumber="124">
|
||
<emphasis id="B952EA83C107435DFF3CEF4CFE30FACB" bold="true" box="[199,478,1303,1328]" pageId="23" pageNumber="124">Ecology & distribution.</emphasis>
|
||
The specimens were found in shallow waters (<
|
||
<quantity id="4CDE9B74C107435DFC05EF4CFBC3FACB" box="[1022,1069,1303,1328]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="124" unit="m" value="2.0">2 m</quantity>
|
||
), under rocks, in coarse coralline sediments in areas of high flow.
|
||
<taxonomicName id="4C264D12C107435DFDF9EF67FD2FFAAE" box="[514,705,1340,1365]" class="Holothuroidea" family="Phyllophoridae" genus="Phyrella" kingdom="Animalia" order="Dendrochirotida" pageId="23" pageNumber="124" phylum="Echinodermata" rank="species" species="mookiei">
|
||
<emphasis id="B952EA83C107435DFDF9EF67FD2FFAAE" box="[514,705,1340,1365]" italics="true" pageId="23" pageNumber="124">Phyrella mookiei</emphasis>
|
||
</taxonomicName>
|
||
is currently only known from
|
||
<collectingCountry id="F3317601C107435DFBECEF67FBB0FAAE" box="[1047,1118,1340,1365]" name="Guam" pageId="23" pageNumber="124">Guam</collectingCountry>
|
||
, where it is fairly rare.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |