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<document id="BACCDFDC580FFB3ECBEC973DEEE97128" ID="10.11646/zootaxa.4277.3.1SLASH11404" ID-CLB-Dataset="32824" ID-DOI="10.11646/zootaxa.4277.3.1" ID-GBIF-Dataset="426319f0-f361-4222-bf48-b093743b0b96" ID-ISSN="1175-5326" ID-Zenodo-Dep="810605" ID-ZooBank="70C1663B-F33F-42C1-99C9-FC3A0A4110DA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497856720584" checkinUser="plazi" docAuthor="Witaliński, Wojciech" docDate="2017" docId="038F8773FFB9FFAFFF795EFF9332788B" docLanguage="en" docName="zootaxa.4277.3.1SLASH11404.pdf" docOrigin="Zootaxa 4277 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holoparasitus Witaliński 2017" docType="treatment" docVersion="7" lastPageNumber="310" masterDocId="FFB6FF0BFFB0FFA5FFEE5B0A90117F5B" masterDocTitle="Key to the world species of Holoparasitus Oudemans, 1936 (Acari: Parasitiformes: Parasitidae)" masterLastPageNumber="351" masterPageNumber="301" pageNumber="310" updateTime="1698447217503" updateUser="ExternalLinkService">
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<mods:title id="751141DB23FEE3F565009ECF4EC8B0A7">Key to the world species of Holoparasitus Oudemans, 1936 (Acari: Parasitiformes: Parasitidae)</mods:title>
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<mods:date id="CF166E3E461C89A20E1126973EEC99AC">2017</mods:date>
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<treatment id="038F8773FFB9FFAFFF795EFF9332788B" ID-DOI="http://doi.org/10.5281/zenodo.6004373" ID-GBIF-Taxon="131891888" ID-Zenodo-Dep="6004373" LSID="urn:lsid:plazi:treatment:038F8773FFB9FFAFFF795EFF9332788B" httpUri="http://treatment.plazi.org/id/038F8773FFB9FFAFFF795EFF9332788B" lastPageId="10" lastPageNumber="310" pageId="9" pageNumber="310">
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Key to
<taxonomicName id="4C264DE6FFB9FFACFF1E5EFE918A7955" authorityName="Witaliński" authorityYear="2017" box="[240,411,1524,1550]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="B952EA77FFB9FFACFF1E5EFE918A7955" bold="true" box="[240,411,1524,1550]" italics="true" pageId="9" pageNumber="310">Holoparasitus</emphasis>
</taxonomicName>
species (Males)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<keyStep id="30D22EC0FFB9FFACFF795D34958A79D6" pageId="9" pageNumber="310">
<subSubSection id="C33C65EEFFB9FFACFF795D34958A792B" pageId="9" pageNumber="310" type="key">
<paragraph id="8B993665FFB9FFACFF795D34958A792B" blockId="9.[151,1436,1595,1971]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795D34958A792B" pageId="9" pageNumber="310">
1. Sternum with an excipulum (e.g.
<figureCitation id="131D2AE0FFB9FFACFDC55D3492A77908" box="[555,694,1598,1619]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="9" pageNumber="310">Figs 229, 230</figureCitation>
); if excipulum is not encountered, a distinct, not-reticulated elliptic area1 is found on the sternum at the coxae II level (
<figureCitation id="131D2AE0FFB9FFACFD935D5192DE792B" box="[637,719,1627,1648]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="9" pageNumber="310">Fig. 231</figureCitation>
).......................................................... 2
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</paragraph>
</subSubSection>
<paragraph id="8B993665FFB9FFACFF795D72958A79D6" blockId="9.[151,1436,1595,1971]" box="[151,1435,1656,1677]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795D72958A79D6" box="[151,1435,1656,1677]" pageId="9" pageNumber="310">
- Excipulum or its equivalent, a non-reticulated area on the sternum absent (e.g.
<figureCitation id="131D2AE0FFB9FFACFC3A5D72943479D6" box="[980,1061,1656,1677]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="9" pageNumber="310">Fig. 228</figureCitation>
)............................ 15
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</paragraph>
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<keyStep id="30D22EC0FFB9FFACFF795D9C958D7866" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF795D9C958A7993" blockId="9.[151,1436,1595,1971]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795D9C958A7993" pageId="9" pageNumber="310">
2. Gnathotectum with the central prong well pronounced, acute, flanked by the two lateral prongs, sometimes minute or altogether absent (e.g.
<figureCitation id="131D2AE0FFB9FFACFE785DB9920A7993" box="[406,539,1715,1736]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="9" pageNumber="310">Figs 234, 239</figureCitation>
)......................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8B993665FFB9FFACFF795DDA958D7866" blockId="9.[151,1436,1595,1971]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795DDA958D7866" pageId="9" pageNumber="310">
- Gnathotectum central prong lobate (
<figureCitation id="131D2AE0FFB9FFACFDD05DDA928279BE" box="[574,659,1744,1765]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="9" pageNumber="310">Fig. 235</figureCitation>
), rounded or obtuse anteriorly, either spineless, or with minute spines (13) apically (e.g.
<figureCitation id="131D2AE0FFB9FFACFEAB5DE4918B7858" box="[325,410,1774,1795]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="9" pageNumber="310">Figs 281</figureCitation>
,
<figureCitation id="131D2AE0FFB9FFACFE4A5DE491DD7858" box="[420,460,1774,1795]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="9" pageNumber="310">310</figureCitation>
); lateral small prongs present; a mobile digit of chelicera much thicker in the basal half (e.g.
<figureCitation id="131D2AE0FFB9FFACFAAC5DE495897858" box="[1346,1432,1774,1795]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="9" pageNumber="310">Figs 282</figureCitation>
,
<figureCitation id="131D2AE0FFB9FFACFF0F5C019116787B" box="[225,263,1803,1824]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="9" pageNumber="310">288</figureCitation>
,
<figureCitation id="131D2AE0FFB9FFACFEFC5C019127787B" box="[274,310,1803,1824]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="9" pageNumber="310">311</figureCitation>
) (in
<taxonomicName id="4C264DE6FFB9FFACFE895C069298787B" authority="Willmann, 1940" authorityName="Willmann" authorityYear="1940" box="[359,649,1803,1825]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="rotulifer">
<emphasis id="B952EA77FFB9FFACFE895C069169787A" box="[359,376,1804,1825]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFE685C0191C4787A" box="[390,469,1803,1825]" italics="true" pageId="9" pageNumber="310">rotulifer</emphasis>
(
<bibRefCitation id="EFB74B94FFB9FFACFE0D5C019293787B" author="Willmann" box="[483,642,1803,1824]" pageId="9" pageNumber="310" refString="Willmann, C. (1940) Neue Milben aus Hohlen der Balkanhalbinsel, gesammelt von Prof. Dr. K. Absolon, Brunn (2. Mitteilung). Zoologischer Anzeiger, 130, 209 - 218." type="journal article" year="1940">Willmann, 1940</bibRefCitation>
)
</taxonomicName>
gnathotectum with a single central prong only, and a movable digit of chelicera with a tubercle on the external (ventral) margin)..................................... (
<emphasis id="B952EA77FFB9FFACFB905C2295647865" bold="true" box="[1150,1397,1832,1854]" pageId="9" pageNumber="310">
<taxonomicName id="4C264DE6FFB9FFACFB905C2294F57865" authorityName="Koch" authorityYear="Koch" box="[1150,1252,1832,1854]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="B952EA77FFB9FFACFB905C2294F57865" bold="true" box="[1150,1252,1832,1854]" italics="true" pageId="9" pageNumber="310">calcaratus</emphasis>
</taxonomicName>
species-group
</emphasis>
)...9
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<keyStep id="30D22EC0FFB9FFACFF795C4C958A7823" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF795C4C958D7800" blockId="9.[151,1436,1595,1971]" box="[151,1436,1861,1883]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795C4C958D7800" box="[151,1436,1861,1883]" pageId="9" pageNumber="310">
3. Postcoxal glands
<emphasis id="B952EA77FFB9FFACFE675C4F91BA7800" box="[393,427,1861,1883]" italics="true" pageId="9" pageNumber="310">gv2</emphasis>
empty into the cuticular foveolae (
<figureCitation id="131D2AE0FFB9FFACFD155C4C93407800" box="[763,849,1862,1883]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="9" pageNumber="310">Figs 231</figureCitation>
,
<figureCitation id="131D2AE0FFB9FFACFCB55C4C93907800" box="[859,897,1862,1883]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="9" pageNumber="310">243</figureCitation>
)........................................... 4
</keyLead>
</paragraph>
<paragraph id="8B993665FFB9FFACFF795C69958A7823" blockId="9.[151,1436,1595,1971]" box="[151,1435,1891,1912]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795C69958A7823" box="[151,1435,1891,1912]" pageId="9" pageNumber="310">- Cuticular foveolae behind coxae IV—not encountered........................................................ 5</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B993665FFB9FFACFF795C8A958D78E8" blockId="9.[151,1436,1595,1971]" pageId="9" pageNumber="310">
<keyStep id="30D22EC0FFB9FFACFF795C8A958D78E8" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795C8A958D78E8" pageId="9" pageNumber="310">
4. Sternum with a well-developed excipulum (
<figureCitation id="131D2AE0FFB9FFACFD6C5C8A92C678CE" box="[642,727,1920,1941]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="9" pageNumber="310">Fig. 243</figureCitation>
); corniculi slightly indented; dorsal edge of fixed digit of chelicera normal (
<figureCitation id="131D2AE0FFB9FFACFF065C94912B78E8" box="[232,314,1950,1971]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="9" pageNumber="310">Fig. 244</figureCitation>
)...........................................................
<taxonomicName id="4C264DE6FFB9FFACFC125C94951278E8" authority="Witalinski, 1994" authorityName="Witalinski" authorityYear="1994" box="[1020,1283,1949,1971]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="dallaii">
<emphasis id="B952EA77FFB9FFACFC125C94940378E8" box="[1020,1042,1950,1971]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFBF15C97944F78E8" box="[1055,1118,1949,1971]" italics="true" pageId="9" pageNumber="310">dallaii</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFB8A5C94951278E8" author="Witalinski" box="[1124,1283,1950,1971]" pageId="9" pageNumber="310" refString="Witalinski, W. (1994) Two new Holoparasitus species from Italy and Poland (Acari: Gamasida: Pergamasidae). Genus, 5, 215 - 222." type="journal article" year="1994">Witaliński, 1994</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB9FFACFAE15C94958478E8" box="[1295,1429,1950,1971]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="9" pageNumber="310">Figs 243245</figureCitation>
)
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B993665FFB9FFAFFF795CD69332788B" blockId="9.[151,1166,2011,2033]" lastBlockId="10.[151,1051,1978,2033]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
<keyStep id="30D22EC0FFB9FFACFF795CD6949F78AA" box="[151,1166,2011,2033]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795CD6949F78AA" box="[151,1166,2011,2033]" pageId="9" pageNumber="310">
1. In
<taxonomicName id="4C264DE6FFB9FFACFF095CD6914878AA" authorityName="Witalinski" authorityYear="1994" box="[231,345,2011,2033]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B952EA77FFB9FFACFF095CD690E978AA" box="[231,248,2012,2033]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFEEA5CD1914878AA" box="[260,345,2011,2033]" italics="true" pageId="9" pageNumber="310">globosus</emphasis>
</taxonomicName>
excipulum is either weakly visible or only not-reticulated area in that place is present
</keyLead>
</keyStep>
<subSection id="E2A92F8EFFBAFFAFFF795B9D9332788B" pageId="10" pageNumber="311" type="multiple">
- Sternum with a non-reticulated area (
<figureCitation id="131D2AE0FFBAFFAFFDBA5B9D92BA7FF7" box="[596,683,151,172]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="10" pageNumber="311">Fig. 231</figureCitation>
); in some specimens the traces of excipulum can be detected; corniculi not indented; dorsal edge of fixed digit of chelicera with an elevation (
<figureCitation id="131D2AE0FFBAFFAFFCB15BBF93A07F91" box="[863,945,181,202]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 246</figureCitation>
).....................................................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC545BD894C27FBC" authority="Witalinski, 1994" authorityName="Witalinski" authorityYear="1994" box="[954,1235,209,231]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B952EA77FFBAFFAFFC545BD893DA7FBC" box="[954,971,210,231]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC375BDB943F7FBC" box="[985,1070,209,231]" italics="true" pageId="10" pageNumber="311">globosus</emphasis>
<bibRefCitation id="EFB74B94FFBAFFAFFBDA5BD894C27FBC" author="Witalinski" box="[1076,1235,210,231]" pageId="10" pageNumber="311" refString="Witalinski, W. (1994) Two new Holoparasitus species from Italy and Poland (Acari: Gamasida: Pergamasidae). Genus, 5, 215 - 222." type="journal article" year="1994">Witaliński, 1994</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFB315BD895277FBC" box="[1247,1334,210,231]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="10" pageNumber="311">Figs 231</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFAAE5BD895847FBC" box="[1344,1429,210,231]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">246, 247</figureCitation>
) 5. Gnathotectum with all prongs similarly well pronounced (
<figureCitation id="131D2AE0FFBAFFAFFCF75BE593617E5F" box="[793,880,239,260]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 248</figureCitation>
); Ti II with additional tubercle dorsally (
<figureCitation id="131D2AE0FFBAFFAFFAEF5BE595497E5F" box="[1281,1368,239,260]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 249</figureCitation>
); fixed digit of chelicera knife-shaped and toothless (
<figureCitation id="131D2AE0FFBAFFAFFD785A0792F97E79" box="[662,744,269,290]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 250</figureCitation>
)......................
<taxonomicName id="4C264DE6FFBAFFAFFC195A0795127E79" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[1015,1283,268,290]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="cultriger">
<emphasis id="B952EA77FFBAFFAFFC195A0794197E79" box="[1015,1032,269,290]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFBFB5A0694787E79" box="[1045,1129,268,290]" italics="true" pageId="10" pageNumber="311">cultriger</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFB9B5A0794EA7E79" author="Berlese" box="[1141,1275,269,290]" pageId="10" pageNumber="311" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15A0795847E79" box="[1295,1429,269,290]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Figs 248250</figureCitation>
) - Gnathotectum with the central prong well pronounced, whereas the lateral ones are either small or altogether absent (
<figureCitation id="131D2AE0FFBAFFAFFAD25A20959C7E64" box="[1340,1421,298,319]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 251</figureCitation>
); Ti II with a regular prong only; fixed digit of chelicera either toothless or with several dents.......................... 6 6. Excipulum horseshoe-shaped, its anterior thickened margin complete (
<figureCitation id="131D2AE0FFBAFFAFFC495A6F94137E21" box="[935,1026,357,378]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Figs 252</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFBFF5A6F94287E21" box="[1041,1081,357,378]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">259</figureCitation>
); sternal reticulation unremarkable; hypostomatic setae not on a separate piece of cuticle (
<figureCitation id="131D2AE0FFBAFFAFFD0B5A8893287ECC" box="[741,825,386,407]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 253</figureCitation>
); fixed digit of chelicera practically toothless............................................................................................ (
<emphasis id="B952EA77FFBAFFAFFB7A5A9595647EEE" bold="true" box="[1172,1397,415,437]" pageId="10" pageNumber="311">
<taxonomicName id="4C264DE6FFBAFFAFFB7A5A9594F57EEE" authorityName="Berlese" authorityYear="1906" box="[1172,1252,415,437]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="peraltus">
<emphasis id="B952EA77FFBAFFAFFB7A5A9594F57EEE" bold="true" box="[1172,1252,415,437]" italics="true" pageId="10" pageNumber="311">peraltus</emphasis>
</taxonomicName>
species-group
</emphasis>
)...7 - Excipulum “open” at both anterior and posterior ends and flanked by two very prominent lines of reticulation running parallel to the excipulum anterolateral margins (
<figureCitation id="131D2AE0FFBAFFAFFD865AD092D17EB4" box="[616,704,474,495]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">Figs 265</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFD225AD092E57EB4" box="[716,756,474,495]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">270</figureCitation>
); hypostomatic setae on a separate piece of cuticle (like in
<figureCitation id="131D2AE0FFBAFFAFFAD55AD095807EB4" box="[1339,1425,474,495]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="10" pageNumber="311">Fig. 232</figureCitation>
); fixed digit of chelicera with a row of many tiny denticles......................... (
<emphasis id="B952EA77FFBAFFAFFBA75AFD95647D56" bold="true" box="[1097,1397,503,525]" pageId="10" pageNumber="311">
<taxonomicName id="4C264DE6FFBAFFAFFBA75AFD94F57D56" authorityName="Vitzthum" authorityYear="1923" box="[1097,1252,503,525]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="hemisphaericus">
<emphasis id="B952EA77FFBAFFAFFBA75AFD94F57D56" bold="true" box="[1097,1252,503,525]" italics="true" pageId="10" pageNumber="311">hemisphaericus</emphasis>
</taxonomicName>
species-group
</emphasis>
)...8 7. Spur on Ti II singular, long and low (
<figureCitation id="131D2AE0FFBAFFAFFDA5591F92C77D71" box="[587,726,533,554]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Figs 256, 257</figureCitation>
); pedipalp trochanter with seta
<date id="FF9810A5FFBAFFAFFBE8591E940C7D71" box="[1030,1053,532,554]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFBE8591E940C7D71" box="[1030,1053,532,554]" italics="true" pageId="10" pageNumber="311">v1</emphasis>
</date>
simple,
<date id="FF9810A5FFBAFFAFFB9E591E94967D71" box="[1136,1159,532,554]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFB9E591E94967D71" box="[1136,1159,532,554]" italics="true" pageId="10" pageNumber="311">v2</emphasis>
</date>
stout and terminally brushy, settled on eminence not far from
<date id="FF9810A5FFBAFFAFFDF1593B92277D1C" box="[543,566,561,583]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFDF1593B92277D1C" box="[543,566,561,583]" italics="true" pageId="10" pageNumber="311">v1</emphasis>
</date>
(
<figureCitation id="131D2AE0FFBAFFAFFDAD593892847D1C" box="[579,661,562,583]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Fig. 254</figureCitation>
).............................
<taxonomicName id="4C264DE6FFBAFFAFFC12593895127D1C" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[1020,1283,561,583]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="peraltus">
<emphasis id="B952EA77FFBAFFAFFC125938941C7D1C" box="[1020,1037,562,583]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFBF4593B94797D1C" box="[1050,1128,561,583]" italics="true" pageId="10" pageNumber="311">peraltus</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFB9B593894ED7D1C" author="Berlese" box="[1141,1276,562,583]" pageId="10" pageNumber="311" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE1593895847D1C" box="[1295,1429,562,583]" captionStart="FIGURES 243 258" captionStartId="27.[151,264,1844,1867]" captionTargetBox="[265,1309,199,1796]" captionTargetId="figure@27.[253,1321,193,1804]" captionTargetPageId="27" captionText="FIGURES 243 258. Males. Holoparasitus dallaii: 243 idiosoma ventral side; 244 chelicera, antiaxially; 245 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus globosus: 246 chelicera, adaxially; 247 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus cultriger: 248 gnathotectum; 249 Fe II, Ge II and Ti II, posterolaterally; 250 chelicera. Holoparasitus peraltus: 251 gnathotectum; 252 excipulum; 253 hypostome; 254 pedipalp trochanter, ventrally; 255 chelicera, adaxially; 256 Fe II, Ge II and Ti II, posterolaterally; 257 spurs on Fe II, Ge II and Ti II; 258 main spur and axillary process in ventral perspective (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810633/files/figure.png" pageId="10" pageNumber="311">Figs 251258</figureCitation>
) - Spur on Ti II bipartite (
<figureCitation id="131D2AE0FFBAFFAFFE2D5945925C7D3F" box="[451,589,591,612]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">Figs 262, 263</figureCitation>
): proximal part with a sigmoidal edge, distal one forming a tooth-like structure; seta
<date id="FF9810A5FFBAFFAFFA6B5945958D7D3E" box="[1413,1436,591,613]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFA6B5945958D7D3E" box="[1413,1436,591,613]" italics="true" pageId="10" pageNumber="311">v2</emphasis>
</date>
pectinate distally, neither stout nor located on eminence close to
<date id="FF9810A5FFBAFFAFFCA2596693607DD9" box="[844,881,620,642]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFCA2596693757DD9" box="[844,868,620,642]" italics="true" pageId="10" pageNumber="311">v1</emphasis>
.
</date>
...................................................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC8C598095137DC4" authority="Witalinski &amp; Skorupski, 2003a" authorityName="Witalinski &amp; Skorupski" authorityYear="2003" box="[866,1282,649,671]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="nonaltus">
<emphasis id="B952EA77FFBAFFAFFC8C598093627DC4" box="[866,883,650,671]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC6F598393C27DC4" box="[897,979,649,671]" italics="true" pageId="10" pageNumber="311">nonaltus</emphasis>
<bibRefCitation id="EFB74B94FFBAFFAFFC37598095137DC4" author="Witalinski" box="[985,1282,650,671]" pageId="10" pageNumber="311" refString="Witalinski, W. &amp; Skorupski, M. (2003 a) Genus Holoparasitus Oudemans, 1936 in Berlese Acaroteca (Acari: Gamasida: Parasitidae). Part II. Redia, 86, 17 - 22." type="journal article" year="2003" yearSuffix="a">Witaliński &amp; Skorupski, 2003a</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE1598095877DC4" box="[1295,1430,650,671]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">Figs 259264</figureCitation>
) 8. Spur on Ti II (
<figureCitation id="131D2AE0FFBAFFAFFE8759AD91FF7DE7" box="[361,494,679,700]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">Figs 267269</figureCitation>
) ending distally at the segment margin and visible as an arcuate tape from both the ventral and the lateral sides; the cuticle located behind the spur (posteriorly) is weakly pigmented (
<figureCitation id="131D2AE0FFBAFFAFFC1059CF94927D81" box="[1022,1155,709,730]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">Figs 268, 269</figureCitation>
).................................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC4759E895127DAC" authority="Witalinski, 2006" authorityName="Witalinski" authorityYear="2006" box="[937,1283,737,759]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="rhombogynialis">
<emphasis id="B952EA77FFBAFFAFFC4759E893AB7DAC" box="[937,954,738,759]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC2659EB944F7DAC" box="[968,1118,737,759]" italics="true" pageId="10" pageNumber="311">rhombogynialis</emphasis>
Witaliński, 2006
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE159E895877DAC" box="[1295,1430,738,759]" captionStart="FIGURES 259 269" captionStartId="28.[151,264,1649,1672]" captionTargetBox="[177,1394,205,1615]" captionTargetId="figure@28.[151,1436,193,1654]" captionTargetPageId="28" captionText="FIGURES 259 269. Males. Holoparasitus nonaltus: 259 genital lamina and sternogenital shield with excipulum; 260 chelicera, adaxially; 261 movable digit of chelicera; 262 Fe II, Ge II and Ti II, anterolaterally; 263 spur on Ti II, posterolaterally; 264 spur on Ge II, posterolaterally. Holoparasitus rhombogynialis: 265 sternogenital shield with excipulum; 266 chelicera, adaxially; 267 Fe II, Ge II and Ti II, anterolaterally; 268 ibid., ventral aspect; 269 Ti II, ventrally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810635/files/figure.png" pageId="10" pageNumber="311">Figs 265269</figureCitation>
) - Spur on Ti II ending far behind the distal margin of the segment (
<figureCitation id="131D2AE0FFBAFFAFFCBC59F593C67C4F" box="[850,983,767,788]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Figs 272, 273</figureCitation>
)................................................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC4D581795137C69" authority="Vitzthum, 1923" authorityName="Vitzthum" authorityYear="1923" box="[931,1282,796,818]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="hemisphaericus">
<emphasis id="B952EA77FFBAFFAFFC4D581793A57C69" box="[931,948,797,818]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC2F581694467C69" box="[961,1111,796,818]" italics="true" pageId="10" pageNumber="311">hemisphaericus</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFB8A581794EA7C69" author="Vitzthum" box="[1124,1275,797,818]" pageId="10" pageNumber="311" refString="Vitzthum, H. v. (1923) Acarologische Beobachtungen. 7. Reihe. Archiv fur Naturgeschichte, Berlin, 89 A, 97 - 181." type="journal article" year="1923">Vitzthum, 1923</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE1581795847C69" box="[1295,1429,797,818]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Figs 270273</figureCitation>
) 9. Excipulum strong, well pronounced, with the lateral margins partially overlapped by a thin cuticular lamellae (e.g.
<figureCitation id="131D2AE0FFBAFFAFFAAF583091167C37" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Figs 274, 280</figureCitation>
)............................................................................................... 10 - Excipulum weakly to moderately pronounced, its lateral margins not overlapped by the cuticular lamellae (e.g.
<figureCitation id="131D2AE0FFBAFFAFFAE4587F95717CD1" box="[1290,1376,885,906]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 298</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFA87587F959F7CD1" box="[1385,1422,885,906]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">308</figureCitation>
)................................................................................................... 11 10. Gnathotectum central lobe pointed; spurs on Ge II large and high, the ones on Ti II smaller and low (
<figureCitation id="131D2AE0FFBAFFAFFB5058A595057C9F" box="[1214,1300,943,964]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Fig. 278</figureCitation>
); very narrow movable digit of chelicera with semicircular tubercle on the external (ventral) edge (
<figureCitation id="131D2AE0FFBAFFAFFBE758C7944A7CB9" box="[1033,1115,973,994]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Fig. 276</figureCitation>
)..........................................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC0B58E095127CA4" authority="Willmann, 1940" authorityName="Willmann" authorityYear="1940" box="[997,1283,1001,1023]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="rotulifer">
<emphasis id="B952EA77FFBAFFAFFC0B58E093E77CA4" box="[997,1014,1002,1023]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFBED58E394437CA4" box="[1027,1106,1001,1023]" italics="true" pageId="10" pageNumber="311">rotulifer</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFBB058E094ED7CA4" author="Willmann" box="[1118,1276,1002,1023]" pageId="10" pageNumber="311" refString="Willmann, C. (1940) Neue Milben aus Hohlen der Balkanhalbinsel, gesammelt von Prof. Dr. K. Absolon, Brunn (2. Mitteilung). Zoologischer Anzeiger, 130, 209 - 218." type="journal article" year="1940">Willmann, 1940</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE158E095847CA4" box="[1295,1429,1002,1023]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Figs 274278</figureCitation>
) - Gnathotectum central lobe obtuse (
<figureCitation id="131D2AE0FFBAFFAFFDDB5F0D929B7B47" box="[565,650,1031,1052]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Fig. 281</figureCitation>
); fixed digit of chelicera with one-fourth to one-third distal part—narrower (
<figureCitation id="131D2AE0FFBAFFAFFA9B5F0D91197B61" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Fig. 282</figureCitation>
), in dorso-ventral perspective a tubercular to tooth-like protrusion directed laterally at
<emphasis id="B952EA77FFBAFFAFFBBB5F2E94C37B61" box="[1109,1234,1060,1082]" italics="true" pageId="10" pageNumber="311">pilus dentilis</emphasis>
level is visible (
<figureCitation id="131D2AE0FFBAFFAFFA9B5F2F91187B0C" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">Fig. 283</figureCitation>
); movable digit without any tubercle on the external (ventral) edge...
<taxonomicName id="4C264DE6FFBAFFAFFC5E5F4894C37B0C" authority="Berlese, 1906" authorityName="Berlese" authorityYear="Berlese" box="[944,1234,1089,1111]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="excipuliger">
<emphasis id="B952EA77FFBAFFAFFC5E5F4893D07B0C" box="[944,961,1090,1111]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC235F4B94287B0C" box="[973,1081,1089,1111]" italics="true" pageId="10" pageNumber="311">excipuliger</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFBAB5F4894DA7B0C" author="Berlese" box="[1093,1227,1090,1111]" pageId="10" pageNumber="311" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFB315F4895277B0C" box="[1247,1334,1090,1111]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="10" pageNumber="311">Figs 229</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFAD15F4895847B0C" box="[1343,1429,1090,1111]" captionStart="FIGURES 270 287" captionStartId="29.[151,264,1862,1885]" captionTargetBox="[282,1317,219,1823]" captionTargetId="figure@29.[258,1343,193,1853]" captionTargetPageId="29" captionText="FIGURES 270 287. Males. Holoparasitus hemisphaericus: 270 sternogenital shield with excipulum; 271 chelicera, antiaxially; 272 Fe II, Ge II and Ti II, anterolaterally; 273 ibid., ventrally (arrow anterolateral side). Holoparasitus rotulifer: 274, 275 excipulum on sternogenital shield; 276 movable digit of chelicera; 277 fixed digit of chelicera; 278 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus excipuliger: 279, 280 excipulum on sternogenital shield; 281 gnathotectum; 282 chelicera, antiaxially; 283 apical part of chelicera in ventral perspective; 284 Fe II, Ge II and Ti II, anterolaterally; 285, 286 Ge II and Ti II, ventrally (arrow anterolateral side); 287 Fe II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810637/files/figure.png" pageId="10" pageNumber="311">279287</figureCitation>
) 11. The pores of postcoxal glands
<emphasis id="B952EA77FFBAFFAFFDE85F5592387B2E" box="[518,553,1119,1141]" italics="true" pageId="10" pageNumber="311">gv2</emphasis>
surrounded by a thickened cuticle forming foveolae (
<figureCitation id="131D2AE0FFBAFFAFFBC55F55946C7B2F" box="[1067,1149,1119,1140]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Fig. 293</figureCitation>
)..................... 12 - Cuticular foveolae around the
<emphasis id="B952EA77FFBAFFAFFDEA5F7692377BC9" box="[516,550,1148,1170]" italics="true" pageId="10" pageNumber="311">gv2</emphasis>
gland pores absent....................................................... 13 12. Axillary process on Fe II asymmetric and bent towards the segment (
<figureCitation id="131D2AE0FFBAFFAFFC955F9094127BF4" box="[891,1027,1178,1199]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 290, 291</figureCitation>
); cheliceral fixed digit with a lateral triangular projection (
<figureCitation id="131D2AE0FFBAFFAFFE685FBD91CB7B97" box="[390,474,1207,1228]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Fig. 289</figureCitation>
), seta
<date id="FF9810A5FFBAFFAFFDF85FBD923C7B96" box="[534,557,1207,1229]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFDF85FBD923C7B96" box="[534,557,1207,1229]" italics="true" pageId="10" pageNumber="311">v2</emphasis>
</date>
on pedipalp trochanter pectinate distally on both sides and thicker than
<date id="FF9810A5FFBAFFAFFB305FBD94ED7B97" box="[1246,1276,1207,1229]" pageId="10" pageNumber="311">
<emphasis id="B952EA77FFBAFFAFFB305FBD94E47B96" box="[1246,1269,1207,1229]" italics="true" pageId="10" pageNumber="311">v1</emphasis>
.
</date>
........................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC4C5FDF95127BB1" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[930,1283,1236,1258]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="pseudoperforatus">
<emphasis id="B952EA77FFBAFFAFFC4C5FDF93A27BB1" box="[930,947,1237,1258]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC2F5FDE94787BB1" box="[961,1129,1236,1258]" italics="true" pageId="10" pageNumber="311">pseudoperforatus</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFB9B5FDF94EA7BB1" author="Berlese" box="[1141,1275,1237,1258]" pageId="10" pageNumber="311" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15FDF95847BB1" box="[1295,1429,1237,1258]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 288292</figureCitation>
) - Axillary process on Fe II symmetric (
<figureCitation id="131D2AE0FFBAFFAFFDA65FF892C17A5C" box="[584,720,1266,1287]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 295, 296</figureCitation>
); cheliceral fixed digit without a lateral projection (
<figureCitation id="131D2AE0FFBAFFAFFB535FF8951E7A5C" box="[1213,1295,1266,1287]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Fig. 294</figureCitation>
)..........................................................................
<taxonomicName id="4C264DE6FFBAFFAFFC375E1A95127A7F" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[985,1283,1295,1317]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="pollicipatus">
<emphasis id="B952EA77FFBAFFAFFC375E1A93FB7A7E" box="[985,1002,1296,1317]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC165E0594787A7E" box="[1016,1129,1295,1317]" italics="true" pageId="10" pageNumber="311">pollicipatus</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFB9B5E0594ED7A7F" author="Berlese" box="[1141,1276,1295,1316]" pageId="10" pageNumber="311" refString="Berlese, A. (1903) Acari nuovi. Manipulus Ius. Redia, 1, 235 - 252." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15E0595877A7F" box="[1295,1430,1295,1316]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 293297</figureCitation>
) 13. Spurs on Ge II and Ti II very similar, low and ridgelike (
<figureCitation id="131D2AE0FFBAFFAFFD115E2793957A19" box="[767,900,1325,1346]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 300, 301</figureCitation>
)................................................................................................
<taxonomicName id="4C264DE6FFBAFFAFFCB95E4095127A04" authority="Witalinski &amp; Skorupski, 2002" authorityName="Witalinski &amp; Skorupski" authorityYear="2002" box="[855,1283,1353,1375]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="kerkirensis">
<emphasis id="B952EA77FFBAFFAFFCB95E4093797A04" box="[855,872,1354,1375]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC9B5E4393CC7A04" box="[885,989,1353,1375]" italics="true" pageId="10" pageNumber="311">kerkirensis</emphasis>
<bibRefCitation id="EFB74B94FFBAFFAFFC0A5E4095127A04" author="Witalinski" box="[996,1283,1354,1375]" pageId="10" pageNumber="311" refString="Witalinski, W. &amp; Skorupski, M. (2002) Genus Holoparasitus Oudemans, 1936 in Berlese Acaroteca (Acari: Gamasida: Parasitidae). Part I. Redia, 85, 37 - 60." type="journal article" year="2002">Witaliński &amp; Skorupski, 2002</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15E4095847A04" box="[1295,1429,1354,1375]" captionStart="FIGURES 288 301" captionStartId="30.[151,264,1870,1893]" captionTargetBox="[271,1305,217,1796]" captionTargetId="figure@30.[245,1329,193,1853]" captionTargetPageId="30" captionText="FIGURES 288 301. Males. Holoparasitus pseudoperforatus: 288, 289 chelicera, two aspects; 290 spurs on Fe II, Ge II and Ti II; 291 Fe II, Ge II and Ti II, anterolaterally; 292 Fe II, ventrally (arrow anterolateral side). Holoparasitus pollicipatus: 293 gv 2 pore in foveola behind opening for coxa IV; 294 chelicera, antiaxially; 295 Fe II, Ge II and Ti II, posterolaterally; 296 ibid, anterolaterally; 297 Fe II and Ge II, ventrally (arrow anterolateral side). Holoparasitus kerkirensis: 298 excipulum on sternogenital shield; 299 Fe II, posterolaterally; 300, 301 Ge II and Ti II, respectively, posterolateral view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810639/files/figure.png" pageId="10" pageNumber="311">Figs 298301</figureCitation>
) - Spurs on Ge II and Ti II different........................................................................ 14 14. Corniculi slender with an adaxial border only slightly sinuous; movable digit of chelicera with one large proximal tooth followed by 46 minute distal teeth (
<figureCitation id="131D2AE0FFBAFFAFFDC95EA8926D7AEC" box="[551,636,1442,1463]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">Fig. 302</figureCitation>
); main spur on the femur with an apical segment blunt terminally and quadrilateral, if observed from a ventral perspective (
<figureCitation id="131D2AE0FFBAFFAFFDB75EB592CE7A8F" box="[601,735,1471,1492]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">Figs 303, 306</figureCitation>
)...................
<taxonomicName id="4C264DE6FFBAFFAFFC245ECA94C27A8F" authority="Koch, 1839" authorityName="Koch" authorityYear="Koch" box="[970,1235,1471,1493]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="B952EA77FFBAFFAFFC245ECA93CA7A8E" box="[970,987,1472,1493]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC065EB5945C7A8E" box="[1000,1101,1471,1493]" italics="true" pageId="10" pageNumber="311">calcaratus</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFBB45EB594DD7A8F" author="Koch" box="[1114,1228,1471,1492]" pageId="10" pageNumber="311" refString="Koch, C. L. (1839) Deutschlands Crustaceen, Myriapoden und Arachniden. Heft 26. No. 6. Bei Friedrich Pustet, Regensburg, 156 pp." type="book" year="1839">Koch, 1839</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFB315EB595277A8F" box="[1247,1334,1471,1492]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="10" pageNumber="311">Figs 230</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFAAE5EB595847A8F" box="[1344,1429,1471,1492]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">302307</figureCitation>
) - Corniculi with a small protuberance on the adaxial border (
<figureCitation id="131D2AE0FFBAFFAFFCF65ED7937C7AA9" box="[792,877,1501,1522]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">Fig. 309</figureCitation>
); movable digit of chelicera with one large proximal tooth followed by 2 smaller distal teeth (
<figureCitation id="131D2AE0FFBAFFAFFDC05EF0926E7954" box="[558,639,1530,1551]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">Fig. 311</figureCitation>
)...............
<taxonomicName id="4C264DE6FFBAFFAFFCD65EF095127954" authority="Witalinski &amp; Skorupski, 2003b" authorityName="Witalinski &amp; Skorupski" authorityYear="2003" box="[824,1283,1529,1551]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="paradisiacus">
<emphasis id="B952EA77FFBAFFAFFCD65EF093587954" box="[824,841,1530,1551]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFCB85EF393C37954" box="[854,978,1529,1551]" italics="true" pageId="10" pageNumber="311">paradisiacus</emphasis>
<bibRefCitation id="EFB74B94FFBAFFAFFC365EF095127954" author="Witalinski" box="[984,1283,1530,1551]" pageId="10" pageNumber="311" refString="Witalinski, W. &amp; Skorupski, M. (2003 b) A new species of Holoparasitus Oudemans, 1936 from North Italy (Acari: Gamasida: Parasitidae). Genus, 14, 431 - 438." type="journal article" year="2003" yearSuffix="b">Witaliński &amp; Skorupski, 2003b</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15EF095877954" box="[1295,1430,1530,1551]" captionStart="FIGURES 302 314" captionStartId="31.[151,264,1696,1719]" captionTargetBox="[175,1416,211,1631]" captionTargetId="figure@31.[151,1436,193,1675]" captionTargetPageId="31" captionText="FIGURES 302 314. Males. Holoparasitus calcaratus: 302 chelicera, adaxially; 303 main spur and axillary process on Fe II, ventral perspective; 304 spurs on Fe II, Ge II and Ti II, posterolaterally; 305 ibid., anterolaterally; 306 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side); 307 Ge II and Ti II, posterolaterally. Holoparasitus paradisiacus: 308 genital lamina and sternogenital shield with excipulum; 309 gnathosoma, ventrally; 310 gnathotectum; 311 chelicera, adaxially; 312 314 Fe II, Ge II and Ti II, respectively, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810641/files/figure.png" pageId="10" pageNumber="311">Figs 308314</figureCitation>
) 15. Ti II with several spurs, the two ventral ones huge (
<figureCitation id="131D2AE0FFBAFFAFFD235D1D93307977" box="[717,801,1559,1580]" captionStart="FIGURES 315 320" captionStartId="32.[151,264,1601,1624]" captionTargetBox="[205,1411,222,1562]" captionTargetId="figure@32.[151,1436,193,1580]" captionTargetPageId="32" captionText="FIGURES 315 320. Males. Holoparasitus karawankianus: 315 gnathosoma, ventrally; 316 leg II, posterolaterally. Holoparasitus fovealis: 317 genital lamina and sternogenital shield; 318 gnathotectum; 319 chelicera, adaxially; 320 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810643/files/figure.png" pageId="10" pageNumber="311">Fig. 316</figureCitation>
).........
<taxonomicName id="4C264DE6FFBAFFAFFC755D1295137977" authority="Schmolzer, 1991" authorityName="Schmolzer" authorityYear="1991" box="[923,1282,1559,1581]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="karawankianus">
<emphasis id="B952EA77FFBAFFAFFC755D1293BD7976" box="[923,940,1560,1581]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC575D1D945D7976" box="[953,1100,1559,1581]" italics="true" pageId="10" pageNumber="311">karawankianus</emphasis>
(
<bibRefCitation id="EFB74B94FFBAFFAFFBB95D1D94ED7977" author="Schmolzer" box="[1111,1276,1559,1580]" pageId="10" pageNumber="311" refString="Schmolzer, K. (1991) Landmilben aus Karnten I. (Acarina, Parasitiformes). Carinthia II, 181, 343 - 358." type="journal article" year="1991">Schmölzer, 1991</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15D1D95847977" box="[1295,1429,1559,1580]" captionStart="FIGURES 315 320" captionStartId="32.[151,264,1601,1624]" captionTargetBox="[205,1411,222,1562]" captionTargetId="figure@32.[151,1436,193,1580]" captionTargetPageId="32" captionText="FIGURES 315 320. Males. Holoparasitus karawankianus: 315 gnathosoma, ventrally; 316 leg II, posterolaterally. Holoparasitus fovealis: 317 genital lamina and sternogenital shield; 318 gnathotectum; 319 chelicera, adaxially; 320 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810643/files/figure.png" pageId="10" pageNumber="311">Figs 315, 316</figureCitation>
) - The two huge spurs on Ti II—not encountered.............................................................. 16 16. Hypostomatic setae on a piece of cuticle separated laterally and posteriorly with narrow incisions................... 17 - Incisions in a cuticle laterally to and behind the hypostomatic setae absent...................................... 46 17. Postcoxal
<emphasis id="B952EA77FFBAFFAFFEAA5D86917779F9" box="[324,358,1676,1698]" italics="true" pageId="10" pageNumber="311">gv2</emphasis>
glands empty into the cuticular foveolae (
<figureCitation id="131D2AE0FFBAFFAFFD155D87935C79F9" box="[763,845,1677,1698]" captionStart="FIGURES 315 320" captionStartId="32.[151,264,1601,1624]" captionTargetBox="[205,1411,222,1562]" captionTargetId="figure@32.[151,1436,193,1580]" captionTargetPageId="32" captionText="FIGURES 315 320. Males. Holoparasitus karawankianus: 315 gnathosoma, ventrally; 316 leg II, posterolaterally. Holoparasitus fovealis: 317 genital lamina and sternogenital shield; 318 gnathotectum; 319 chelicera, adaxially; 320 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810643/files/figure.png" pageId="10" pageNumber="311">Fig. 317</figureCitation>
)..
<taxonomicName id="4C264DE6FFBAFFAFFC985D87951279F9" authority="Witalinski &amp; Skorupski, 2006" authorityName="Witalinski &amp; Skorupski" authorityYear="2006" box="[886,1283,1676,1698]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="fovealis">
<emphasis id="B952EA77FFBAFFAFFC985D87939679F9" box="[886,903,1677,1698]" italics="true" pageId="10" pageNumber="311">H</emphasis>
.
<emphasis id="B952EA77FFBAFFAFFC7D5D8693CF79F9" box="[915,990,1676,1698]" italics="true" pageId="10" pageNumber="311">fovealis</emphasis>
Witaliński &amp; Skorupski, 2006
</taxonomicName>
(
<figureCitation id="131D2AE0FFBAFFAFFAE15D87958779F9" box="[1295,1430,1677,1698]" captionStart="FIGURES 315 320" captionStartId="32.[151,264,1601,1624]" captionTargetBox="[205,1411,222,1562]" captionTargetId="figure@32.[151,1436,193,1580]" captionTargetPageId="32" captionText="FIGURES 315 320. Males. Holoparasitus karawankianus: 315 gnathosoma, ventrally; 316 leg II, posterolaterally. Holoparasitus fovealis: 317 genital lamina and sternogenital shield; 318 gnathotectum; 319 chelicera, adaxially; 320 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810643/files/figure.png" pageId="10" pageNumber="311">Figs 317320</figureCitation>
) - Cuticular foveolae behind coxae IV absent............................................................... 18 18. Hypostome with a central part distinctly sclerotised, tongue-like (e.g.
<figureCitation id="131D2AE0FFBAFFAFFC905DC093C27984" box="[894,979,1738,1759]" captionStart="FIGURES 336 345" captionStartId="34.[151,264,1759,1782]" captionTargetBox="[187,1391,199,1707]" captionTargetId="figure@34.[151,1436,173,1738]" captionTargetPageId="34" captionText="FIGURES 336 345. Males. Holoparasitus franzi: 336 pedipalp trochanter; 337 gnathotectum; 338 chelicera, antiaxially; 339 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus lawrencei: 340 gnathosoma and pedipalp trochanters, ventrally; 341 gnathotectum; 342 chelicera, adaxially; 343 movable digit of chelicera, antiaxially; 344 spurs on Fe II, Ge II and Ti II; 345 Fe II, Ge II and Ti II, posterolaterally. Scale bars: 1, 4 10 0.1 mm, 2, 3 0.05 mm." httpUri="https://zenodo.org/record/810647/files/figure.png" pageId="10" pageNumber="311">Figs 340</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFC335DC094237984" box="[989,1074,1738,1759]" captionStart="FIGURES 391 401" captionStartId="38.[151,264,1871,1894]" captionTargetBox="[183,1417,192,1826]" captionTargetId="figure@38.[176,1420,183,1837]" captionTargetPageId="38" captionText="FIGURES 391 401. Males. Holoparasitus calpetanus: 391 gnathosoma, pedipalp trochanter and femur in ventral perspective; 392 gnathotectum; 393 chelicera, antiaxially; 394 genital lamina; 395 Fe II, Ge II and Ti II, anterolaterally; 396 Ti II, posterolaterally; 397 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side). Holoparasitus mallorcae: 398 gnathosoma and pedipalp trochanter, ventrally; 399 gnathotectum; 400 chelicera, antiaxially; 401 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810657/files/figure.png" pageId="10" pageNumber="311">391, 398</figureCitation>
) or extended anteriorly2 (
<figureCitation id="131D2AE0FFBAFFAFFAC85DC095697984" box="[1318,1400,1738,1759]" captionStart="FIGURES 346 360" captionStartId="35.[151,264,1856,1879]" captionTargetBox="[243,1358,195,1810]" captionTargetId="figure@35.[217,1378,171,1821]" captionTargetPageId="35" captionText="FIGURES 346 360. Males. Holoparasitus gibber: 346 chelicera, adaxially; 347 ibid., antiaxially; 348 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus ellipticus: 349 gnathosoma and pedipalp trochanter, ventrally; 350 gnathotectum; 351 chelicera, adaxially; 352 ibid., antiaxially; 353 Fe II, Ge II and Ti II, anterolaterally; 354 Ge II and Ti II, ventrally (arrow anterolateral side). Holoparasitus anaga: 355 pedipalp trochanter and corniculus; 356 gnathotectum; 357 chelicera, adaxially; 358 ibid., antiaxially; 359 main spur and axillary process on Fe II, anterolaterally; 360 Ge II and Ti II, anterolaterally. Scale bars: 34 6 354, 359, 360 0.1 mm, 355 358 0.05 mm." httpUri="https://zenodo.org/record/810651/files/figure.png" pageId="10" pageNumber="311">Fig. 349</figureCitation>
)............................................................................ (
<emphasis id="B952EA77FFBAFFAFFB8D5DE09546785B" bold="true" box="[1123,1367,1770,1792]" pageId="10" pageNumber="311">
<taxonomicName id="4C264DE6FFBAFFAFFB8D5DE094D7785B" authorityName="Juvara-Bals" authorityYear="1975" box="[1123,1222,1770,1792]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="mallorcae">
<emphasis id="B952EA77FFBAFFAFFB8D5DE094D7785B" bold="true" box="[1123,1222,1770,1792]" italics="true" pageId="10" pageNumber="311">mallorcae</emphasis>
</taxonomicName>
species-group
</emphasis>
) 3...19 -
<collectingRegion id="49E2F887FFBAFFAFFF0F5C0291397846" box="[225,296,1800,1821]" country="Israel" name="Central" pageId="10" pageNumber="311">Central</collectingRegion>
part of the hypostome triangular and weakly sclerotised (e.g.
<figureCitation id="131D2AE0FFBAFFAFFC975C0293EF7846" box="[889,1022,1800,1821]" captionStart="FIGURES 228 242" captionStartId="26.[151,264,1847,1870]" captionTargetBox="[219,1355,227,1789]" captionTargetId="figure@26.[151,1436,171,1844]" captionTargetPageId="26" captionText="FIGURES 228 242. Males. 228 Holoparasitus cornutus, genital lamina and sternogenital shield, II-IV openings for coxae II to IV, iv 1 iv 3 sternal pores, st 1 st 4 sternal setae, gv 1 opening of gland gv 1; 229 Holoparasitus excipuliger, genital lamina and sternogenital shield, gv 2 pore of gland gv 2; 230 Holoparasitus calcaratus, genital lamina and sternogenital shield; 231 Holoparasitus globosus, sternogenital shield; 232 Holoparasitus sp., pedipalp trochanter and gnathosoma, ventrally; 233 Holoparasitus crassisetosus, pedipalp trochanter, femur and hypostome; 234 240 Holoparasitus spp., gnathotectum, different shapes; 241 genital lamina; 242 femur, genu and tibia of leg II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810631/files/figure.png" pageId="10" pageNumber="311">Figs 232, 233</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFBE65C02943F7846" box="[1032,1070,1800,1821]" captionStart="FIGURES 443 454" captionStartId="42.[151,264,1866,1889]" captionTargetBox="[162,1432,208,1832]" captionTargetId="figure@42.[151,1436,193,1845]" captionTargetPageId="42" captionText="FIGURES 443 454. Males. Holoparasitus excisus: 443 gnathotectum; 444 chelicera, adaxially; 445 pedipalp trochanter and femur, ventrally; 446 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus apenninorum: 447 chelicera, adaxially; 448 chelicera, antiaxially; 449 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus amiatus: 450 genital lamina and anterior part of sternogenital shield; 451 pedipalp trochanter and gnathosoma, ventrally; 452 chelicera, antiaxially; 453 spurs on Fe II, Ge II and Ti II in anteroventral perspective (arrow anterolateral side); 454 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810665/files/figure.png" pageId="10" pageNumber="311">451</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFBD65C0294707846" box="[1080,1121,1800,1821]" captionStart="FIGURES 465 475" captionStartId="44.[151,264,1798,1821]" captionTargetBox="[161,1428,203,1766]" captionTargetId="figure@44.[151,1436,193,1778]" captionTargetPageId="44" captionText="FIGURES 465 475. Males. Holoparasitus crassisetosus: 465 gnathosoma with pedipalp trochanters and femurs, ventrally; 466 corniculus; 467 Ti IV, thickened seta; 468 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus annulus: 469 gnathotectum; 470 chelicera, antiaxially; 471 pedipalp trochanter, corniculi and hypognathal groove; 472 corniculus; 473 genital lamina; 474 Fe II, Ge II and Ti II, anterolaterally; 475 ibid., ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810669/files/figure.png" pageId="10" pageNumber="311">465</figureCitation>
)........................ 39 19. Glands
<emphasis id="B952EA77FFBAFFAFFEC55C2E915F7861" box="[299,334,1828,1850]" italics="true" pageId="10" pageNumber="311">gv1</emphasis>
absent; genital lamina rounded, heavily sclerotised on the inner face (
<figureCitation id="131D2AE0FFBAFFAFFC005C2F94C67861" box="[1006,1239,1829,1850]" captionStart="FIGURES 321 335" captionStartId="33.[151,264,1888,1911]" captionTargetBox="[215,1361,190,1863]" captionTargetId="figure@33.[204,1381,177,1878]" captionTargetPageId="33" captionText="FIGURES 321 335. Males. Holoparasitus singularis: 321 genital lamina; 322 gnathotectum and corniculu; 323 chelicera, antiaxially; 324 ibid., adaxially; 325 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus algiersensis: 326, 327 genital lamina; 328 chelicera, antiaxially; 329 movable digit of chelicera; 330 corniculus; 331 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus eivissa: 332 genital lamina; 333 chelicera, antiaxially; 334 ibid., adaxially; 335 Fe II, Ge II and Ti II, ventrally (arrow anterolateral side). Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810645/files/figure.png" pageId="10" pageNumber="311">Figs 321, 326, 327, 332</figureCitation>
); microsclerite bearing tritosternum large and trapezoidal..................................................................... 20 - Gland
<emphasis id="B952EA77FFBAFFAFFECF5C559155782E" box="[289,324,1887,1909]" italics="true" pageId="10" pageNumber="311">gv1</emphasis>
present; genital lamina of another shape and rather weakly sclerotised, largely transparent (e.g.
<figureCitation id="131D2AE0FFBAFFAFFB355C6A9521782E" box="[1243,1328,1888,1909]" captionStart="FIGURES 361 375" captionStartId="36.[151,264,1851,1874]" captionTargetBox="[209,1390,196,1813]" captionTargetId="figure@36.[183,1403,169,1864]" captionTargetPageId="36" captionText="FIGURES 361 375. Males. Holoparasitus giganteus: 361 pedipalp trochanter and corniculus; 362 gnathotectum; 363 chelicera, antiaxially; 364 Fe II, Ge II and Ti II, posterolaterally. Holoparasitus lapalma: 365 pedipalp trochanter and corniculus; 366 gnathotectum; 367 chelicera, antiaxially; 368 Fe II, posterolaterally; 369 Ge II and Ti II, posterolaterally; 3 70 Ti II, ventrally (arrow anterolateral side). Holoparasitus canariensis: 371 pedipalp trochanter and corniculus; 372 gnathotectum, different aspects; 373 genital lamina; 374 Fe II, Ge II and Ti II, anterolaterally; 375 Ge II and Ti II, ventrally (arrow anterolateral side). Scale bars: 361, 362, 364 366, 368 375 0.1 mm, 363, 367 0.05 mm." httpUri="https://zenodo.org/record/810653/files/figure.png" pageId="10" pageNumber="311">Figs 373</figureCitation>
,
<figureCitation id="131D2AE0FFBAFFAFFAD45C6A9580782E" box="[1338,1425,1888,1909]" captionStart="FIGURES 402 413" captionStartId="39.[151,264,1690,1713]" captionTargetBox="[155,1432,212,1652]" captionTargetId="figure@39.[151,1436,193,1669]" captionTargetPageId="39" captionText="FIGURES 402 413. Males. Holoparasitus variabilis: 402 gnathotectum; 403 corniculus; 404 pedipalp trochanter and femur, ventrally; 405 genital lamina; 406 chelicera, antiaxially; 407 Fe II, Ge II and Ti II, anterolaterally. Holoparasitus malleus: 408 gnathotectum; 409 corniculus; 410 pedipalp trochanter and femur, ventrally; 411 genital lamina; 412 chelicera, antiaxially; 413 Fe II, Ge II and Ti II, anterolaterally. Scale bars: 402, 404, 405, 407, 410, 411, 413 0.1 mm, 403, 406, 408, 409, 412 0.05 mm." httpUri="https://zenodo.org/record/810659/files/figure.png" pageId="10" pageNumber="311">405, 411</figureCitation>
); 2. Similar in appearance to those in the
<taxonomicName id="4C264DE6FFBAFFAFFDA95CB092F3788B" authorityName="C. Athias-Henriot" authorityYear="1981" box="[583,738,1978,2000]" class="Arachnida" family="Parasitidae" genus="Rhabdocarpais" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="B952EA77FFBAFFAFFDA95CB092F3788B" box="[583,738,1978,2000]" italics="true" pageId="10" pageNumber="311">Rhabdocarpais</emphasis>
</taxonomicName>
males
</subSection>
</paragraph>
</treatment>
</document>