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<document id="3DC88A65095A8BF9A67EE9E86F2A3994" ID="10.11646/zootaxa.4277.3.1SLASH11404" ID-CLB-Dataset="32824" ID-DOI="10.11646/zootaxa.4277.3.1" ID-GBIF-Dataset="426319f0-f361-4222-bf48-b093743b0b96" ID-ISSN="1175-5326" ID-Zenodo-Dep="810605" ID-ZooBank="70C1663B-F33F-42C1-99C9-FC3A0A4110DA" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497856720584" checkinUser="plazi" docAuthor="Witaliński, Wojciech" docDate="2017" docId="038F8773FFB6FFACFF795FC7958D7AF0" docLanguage="en" docName="zootaxa.4277.3.1SLASH11404.pdf" docOrigin="Zootaxa 4277 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Holoparasitus Witaliński 2017" docType="key" docVersion="7" lastPageNumber="310" masterDocId="FFB6FF0BFFB0FFA5FFEE5B0A90117F5B" masterDocTitle="Key to the world species of Holoparasitus Oudemans, 1936 (Acari: Parasitiformes: Parasitidae)" masterLastPageNumber="351" masterPageNumber="301" pageNumber="307" updateTime="1698447217503" updateUser="ExternalLinkService">
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<mods:title id="5F2E2C581D057109D18A9450A2F83FDE">Key to the world species of Holoparasitus Oudemans, 1936 (Acari: Parasitiformes: Parasitidae)</mods:title>
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<mods:namePart id="C651D21347746DF171F72B62A960392C">Witaliński, Wojciech</mods:namePart>
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<mods:date id="14B9984FF2960D6CF92C20C35BE5315C">2017</mods:date>
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<mods:number id="E0E3C1976DF967639F40675634808C32">4277</mods:number>
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<mods:classification id="EF3C7B30A0AA5F1E6423587CE6A76972">journal article</mods:classification>
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<mods:identifier id="A6C32D00B399DAA5A6248B4B4949C3FD" type="DOI">10.11646/zootaxa.4277.3.1</mods:identifier>
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<treatment id="038F8773FFB6FFACFF795FC7958D7AF0" ID-DOI="http://doi.org/10.5281/zenodo.6004369" ID-GBIF-Taxon="131891889" ID-Zenodo-Dep="6004369" LSID="urn:lsid:plazi:treatment:038F8773FFB6FFACFF795FC7958D7AF0" httpUri="http://treatment.plazi.org/id/038F8773FFB6FFACFF795FC7958D7AF0" lastPageId="9" lastPageNumber="310" pageId="6" pageNumber="307">
<subSubSection id="C33C65EEFFB6FFA3FF795FC7926B7BBC" box="[151,634,1228,1255]" pageId="6" pageNumber="307" type="nomenclature">
<paragraph id="8B993665FFB6FFA3FF795FC7926B7BBC" blockId="6.[151,634,1228,1255]" box="[151,634,1228,1255]" pageId="6" pageNumber="307">
<heading id="D0D18109FFB6FFA3FF795FC7926B7BBC" bold="true" box="[151,634,1228,1255]" fontSize="11" level="1" pageId="6" pageNumber="307" reason="1">
<emphasis id="B952EA77FFB6FFA3FF795FC7926B7BBC" bold="true" box="[151,634,1228,1255]" pageId="6" pageNumber="307">
Key to
<taxonomicName id="4C264DE6FFB6FFA3FF1E5FC6918A7BBD" authorityName="Witaliński" authorityYear="2017" box="[240,411,1228,1254]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B952EA77FFB6FFA3FF1E5FC6918A7BBD" bold="true" box="[240,411,1228,1254]" italics="true" pageId="6" pageNumber="307">Holoparasitus</emphasis>
</taxonomicName>
species (Females)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C65EEFFB6FFACFF795E1E958D7AF0" lastPageId="9" lastPageNumber="310" pageId="6" pageNumber="307" type="key">
<keyStep id="30D22EC0FFB6FFA3FF795E1E958A7A1E" pageId="6" pageNumber="307">
<paragraph id="8B993665FFB6FFA3FF795E1E958D7A72" blockId="6.[151,1436,1299,2020]" box="[151,1436,1299,1321]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795E1E958D7A72" box="[151,1436,1299,1321]" pageId="6" pageNumber="307">
1. Sternum split axially into the left and the right part (
<figureCitation id="131D2AE0FFB6FFA3FD3A5E1E93517A72" box="[724,832,1300,1321]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 11, 14</figureCitation>
)....................... (
<emphasis id="B952EA77FFB6FFA3FB9D5E1995727A72" bold="true" box="[1139,1379,1299,1321]" pageId="6" pageNumber="307">
<taxonomicName id="4C264DE6FFB6FFA3FB9D5E1994C27A72" authorityName="Berlese" authorityYear="1906" box="[1139,1235,1299,1321]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis id="B952EA77FFB6FFA3FB9D5E1994C27A72" bold="true" box="[1139,1235,1299,1321]" italics="true" pageId="6" pageNumber="307">inornatus</emphasis>
</taxonomicName>
species-group
</emphasis>
) … 2
</keyLead>
</paragraph>
<paragraph id="8B993665FFB6FFA3FF795E3A958A7A1E" blockId="6.[151,1436,1299,2020]" box="[151,1435,1328,1349]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795E3A958A7A1E" box="[151,1435,1328,1349]" pageId="6" pageNumber="307">
- Sternum entire (e.g.
<figureCitation id="131D2AE0FFB6FFA3FE4B5E3A91EB7A1E" box="[421,506,1328,1349]" captionStart="FIGURES 1 – 10" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[209,1408,201,1857]" captionTargetId="figure@14.[151,1436,195,1883]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Females. Holoparasitus spp.: 1 ventral side of idiosoma; 2 sternum, presternal plate and lateral presternal platelets; 3 paragynium; 4 epigynium; 5 anterior part of epigynium; 6, 7 endogynium; 8 gnathosoma and gnathotectum, dorsally; 9 gnathosoma, ventrally; 10 pedipalp trochanter." httpUri="https://zenodo.org/record/810607/files/figure.png" pageId="6" pageNumber="307">Figs 1, 2</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FDEA5E3A920C7A1E" box="[516,541,1328,1349]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">33</figureCitation>
)......................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB6FFA3FF795E47958D7AC4" pageId="6" pageNumber="307">
<paragraph id="8B993665FFB6FFA3FF795E47958A7ADB" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795E47958A7ADB" pageId="6" pageNumber="307">
2. Endogynium visible as a transversal thin plate, elliptical or slightly triangular, with central thickening bearing on its dorsal side the centrifugally directed tubercles (
<figureCitation id="131D2AE0FFB6FFA3FDD55E6192B67ADB" box="[571,679,1387,1408]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 12, 13</figureCitation>
)............................
<taxonomicName id="4C264DE6FFB6FFA3FBE65E61950D7ADB" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[1032,1308,1387,1408]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="inornatus">
<emphasis id="B952EA77FFB6FFA3FBE65E6194087ADB" box="[1032,1049,1387,1408]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FBC85E6194937ADB" box="[1062,1154,1387,1408]" italics="true" pageId="6" pageNumber="307">inornatus</emphasis>
(
<bibRefCitation id="EFB74B94FFB6FFA3FB605E6195057ADB" author="Berlese" box="[1166,1300,1387,1408]" pageId="6" pageNumber="307" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFB6FFA3FAC65E6195857ADB" box="[1320,1428,1387,1408]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 11–13</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB6FFA3FF795E80958D7AC4" blockId="6.[151,1436,1299,2020]" box="[151,1436,1417,1439]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795E80958D7AC4" box="[151,1436,1417,1439]" pageId="6" pageNumber="307">
- Endogynium elliptical, axially oriented (
<figureCitation id="131D2AE0FFB6FFA3FD8B5E8092BA7AC4" box="[613,683,1418,1439]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Fig. 14</figureCitation>
).........................
<taxonomicName id="4C264DE6FFB6FFA3FC095E8095557AC4" authority="Schmolzer, 1995" authorityName="Schmolzer" authorityYear="1995" box="[999,1348,1417,1439]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="megacalcaratus">
<emphasis id="B952EA77FFB6FFA3FC095E8093E97AC4" box="[999,1016,1418,1439]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FBEA5E83948C7AC4" box="[1028,1181,1417,1439]" italics="true" pageId="6" pageNumber="307">megacalcaratus</emphasis>
<bibRefCitation id="EFB74B94FFB6FFA3FB4D5E8095557AC4" author="Schmolzer" box="[1187,1348,1418,1439]" pageId="6" pageNumber="307" refString="Schmolzer, K. (1995) Landmilben aus den ostlichen Karawanken (Acarina, Parasitiformes). Carinthia II, 185, 499 - 512." type="journal article" year="1995">Schmölzer, 1995</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB6FFA3FAA15E8095847AC4" box="[1359,1429,1418,1439]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Fig. 14</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB6FFA3FF795EAD958A796A" pageId="6" pageNumber="307">
<paragraph id="8B993665FFB6FFA3FF795EAD958D7AAD" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795EAD958D7AAD" pageId="6" pageNumber="307">
3. Sternum with more or less pronounced axial band, granular or reticulated (e.g.
<figureCitation id="131D2AE0FFB6FFA3FC3E5EAD94717AE7" box="[976,1120,1447,1468]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 33, 46, 50</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FB845EAD94B97AE7" box="[1130,1192,1447,1468]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="6" pageNumber="307">52, 62</figureCitation>
), and/or the anterior margin of presternal plate serrated or coarsely granular (
<figureCitation id="131D2AE0FFB6FFA3FD385ECE93317A82" box="[726,800,1476,1497]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 15</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FCC55ECE93547A82" box="[811,837,1476,1497]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">32</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FCA15ECE93797A82" box="[847,872,1476,1497]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="6" pageNumber="307">52</figureCitation>
) (except
<taxonomicName id="4C264DE6FFB6FFA3FC2C5ECE942D7A82" authorityName="Juvara-Bals" authorityYear="2008" box="[962,1084,1476,1497]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B952EA77FFB6FFA3FC2C5ECE93C27A82" box="[962,979,1476,1497]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FC315ECE942D7A82" box="[991,1084,1476,1497]" italics="true" pageId="6" pageNumber="307">giganteus</emphasis>
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); lateral platelets free........................................................................................ (
<emphasis id="B952EA77FFB6FFA3FB6C5EEB95647AAC" bold="true" box="[1154,1397,1505,1527]" pageId="6" pageNumber="307">
<taxonomicName id="4C264DE6FFB6FFA3FB6C5EEB94F57AAC" authorityName="Juvara-Bals" authorityYear="1975" box="[1154,1252,1505,1527]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="mallorcae">
<emphasis id="B952EA77FFB6FFA3FB6C5EEB94F57AAC" bold="true" box="[1154,1252,1505,1527]" italics="true" pageId="6" pageNumber="307">mallorcae</emphasis>
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species-group
</emphasis>
)...4
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</paragraph>
<paragraph id="8B993665FFB6FFA3FF795EF4958A796A" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795EF4958A796A" pageId="6" pageNumber="307">
- Axial granular or reticulated band on the sternum absent (
<figureCitation id="131D2AE0FFB6FFA3FCE05EF4935B7948" box="[782,842,1534,1555]" captionStart="FIGURES 1 – 10" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[209,1408,201,1857]" captionTargetId="figure@14.[151,1436,195,1883]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Females. Holoparasitus spp.: 1 ventral side of idiosoma; 2 sternum, presternal plate and lateral presternal platelets; 3 paragynium; 4 epigynium; 5 anterior part of epigynium; 6, 7 endogynium; 8 gnathosoma and gnathotectum, dorsally; 9 gnathosoma, ventrally; 10 pedipalp trochanter." httpUri="https://zenodo.org/record/810607/files/figure.png" pageId="6" pageNumber="307">Fig. 2</figureCitation>
); anterior margin of the presternal plate smooth, lateral platelets free or fused to the ends of the presternal plate (
<figureCitation id="131D2AE0FFB6FFA3FD3C5D169301796A" box="[722,784,1564,1585]" captionStart="FIGURES 1 – 10" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[209,1408,201,1857]" captionTargetId="figure@14.[151,1436,195,1883]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Females. Holoparasitus spp.: 1 ventral side of idiosoma; 2 sternum, presternal plate and lateral presternal platelets; 3 paragynium; 4 epigynium; 5 anterior part of epigynium; 6, 7 endogynium; 8 gnathosoma and gnathotectum, dorsally; 9 gnathosoma, ventrally; 10 pedipalp trochanter." httpUri="https://zenodo.org/record/810607/files/figure.png" pageId="6" pageNumber="307">Figs 2</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FCF45D169346796A" box="[794,855,1564,1585]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">16, 17</figureCitation>
)............................................. 23
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<paragraph id="8B993665FFB6FFA3FF795D32958D7931" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
<materialsCitation id="3B4E3C38FFB6FFA3FF795D32958D7931" ID-GBIF-Occurrence="1569232894" location="Central" pageId="6" pageNumber="307" specimenCount="1" stateProvince="Central">
<keyLead id="30D79550FFB6FFA3FF795D32958D7931" pageId="6" pageNumber="307">
4.
<collectingRegion id="49E2F887FFB6FFA3FF0F5D3291397916" box="[225,296,1592,1613]" country="Israel" name="Central" pageId="6" pageNumber="307">Central</collectingRegion>
prong of epigynium slender and considerably elongated (
<figureCitation id="131D2AE0FFB6FFA3FCB75D3293DB7916" box="[857,970,1592,1613]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 18–20</figureCitation>
); endogynium with two teeth, covered ventrally by a dentate lamella (
<figureCitation id="131D2AE0FFB6FFA3FE405D5F920A7931" box="[430,539,1621,1642]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 19–21</figureCitation>
)..........................................
<taxonomicName id="4C264DE6FFB6FFA3FBCE5D5C950A7931" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[1056,1307,1621,1643]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="siculus">
<emphasis id="B952EA77FFB6FFA3FBCE5D5C94207930" box="[1056,1073,1622,1643]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FBD05D5F94907930" box="[1086,1153,1621,1643]" italics="true" pageId="6" pageNumber="307">siculus</emphasis>
(
<bibRefCitation id="EFB74B94FFB6FFA3FB635D5F95027931" author="Berlese" box="[1165,1299,1621,1642]" pageId="6" pageNumber="307" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFB6FFA3FAC95D5F95847931" box="[1319,1429,1621,1642]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 18–21</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB6FFA3FF795D79958D79D3" blockId="6.[151,1436,1299,2020]" box="[151,1436,1651,1672]" pageId="6" pageNumber="307">
<materialsCitation id="3B4E3C38FFB6FFA3FF795D79958D79D3" ID-GBIF-Occurrence="1569232892" box="[151,1436,1651,1672]" location="Central" pageId="6" pageNumber="307" specimenCount="1" stateProvince="Central">
<keyLead id="30D79550FFB6FFA3FF795D79958D79D3" box="[151,1436,1651,1672]" pageId="6" pageNumber="307">
-
<collectingRegion id="49E2F887FFB6FFA3FF0F5D79913979D3" box="[225,296,1651,1672]" country="Israel" name="Central" pageId="6" pageNumber="307">Central</collectingRegion>
prong of epigynium moderately narrow and elongated (e.g.
<figureCitation id="131D2AE0FFB6FFA3FC845D7993C679D3" box="[874,983,1651,1672]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 24, 27</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FC0F5D79945379D3" box="[993,1090,1651,1672]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">34, 41, 48</figureCitation>
)........................... 5
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<keyStep id="30D22EC0FFB6FFA3FF795D9A958D7984" pageId="6" pageNumber="307">
<paragraph id="8B993665FFB6FFA3FF795D9A958A7999" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795D9A958A7999" pageId="6" pageNumber="307">
5.
<emphasis id="B952EA77FFB6FFA3FF0F5D85911879FE" box="[225,265,1679,1701]" italics="true" pageId="6" pageNumber="307">Gv1</emphasis>
glands absent, epigynium with a rounded apex and moderately sharp lateral prongs (
<figureCitation id="131D2AE0FFB6FFA3FBA15D9A94D379FE" box="[1103,1218,1680,1701]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 24, 27</figureCitation>
), endogynium a small sack (
<figureCitation id="131D2AE0FFB6FFA3FEF65DA791BB7999" box="[280,426,1709,1730]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 25, 28, 31</figureCitation>
).................................................................................. 6
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</paragraph>
<paragraph id="8B993665FFB6FFA3FF795DC0958D7984" blockId="6.[151,1436,1299,2020]" box="[151,1436,1737,1759]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795DC0958D7984" box="[151,1436,1737,1759]" pageId="6" pageNumber="307">
-
<emphasis id="B952EA77FFB6FFA3FF0F5DC391187984" box="[225,265,1737,1759]" italics="true" pageId="6" pageNumber="307">Gv1</emphasis>
glands present, epigynium with a triangular apex and salient lateral prongs (
<figureCitation id="131D2AE0FFB6FFA3FC0B5DC094667984" box="[997,1143,1738,1759]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 34, 41, 48</figureCitation>
)...................... 7
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</paragraph>
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<keyStep id="30D22EC0FFB6FFA3FF795DED958D786D" pageId="6" pageNumber="307">
<paragraph id="8B993665FFB6FFA3FF795DED958D79A7" blockId="6.[151,1436,1299,2020]" box="[151,1436,1766,1788]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795DED958D79A7" box="[151,1436,1766,1788]" pageId="6" pageNumber="307">
6. Endogynium—a sack with finely dentate anterior margin (
<figureCitation id="131D2AE0FFB6FFA3FCFE5DED934879A7" box="[784,857,1767,1788]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Fig. 25</figureCitation>
)..........
<taxonomicName id="4C264DE6FFB6FFA3FC365DED950A79A7" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[984,1307,1766,1788]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="algiersensis">
<emphasis id="B952EA77FFB6FFA3FC365DED93F879A7" box="[984,1001,1767,1788]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FC1B5DEC947679A7" box="[1013,1127,1766,1788]" italics="true" pageId="6" pageNumber="307">algiersensis</emphasis>
<bibRefCitation id="EFB74B94FFB6FFA3FB835DED950A79A7" author="Juvara-Bals" box="[1133,1307,1767,1788]" pageId="6" pageNumber="307" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB6FFA3FAC95DED958479A7" box="[1319,1429,1767,1788]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 22–25</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB6FFA3FF795C0E958D786D" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
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- Endogynium—a sack with a round, sclerotised part in its middle (
<figureCitation id="131D2AE0FFB6FFA3FCB95C0E93F97842" box="[855,1000,1796,1817]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 28, 29, 31</figureCitation>
)......................................................................................................
<taxonomicName id="4C264DE6FFB6FFA3FBE75C2B950A786D" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1033,1307,1825,1846]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="eivissa">
<emphasis id="B952EA77FFB6FFA3FBE75C2B940B786D" box="[1033,1050,1825,1846]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FBC85C2B9479786D" box="[1062,1128,1825,1846]" italics="true" pageId="6" pageNumber="307">eivissa</emphasis>
<bibRefCitation id="EFB74B94FFB6FFA3FB835C2B950A786D" author="Juvara-Bals" box="[1133,1307,1825,1846]" pageId="6" pageNumber="307" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB6FFA3FAC95C2B9584786D" box="[1319,1429,1825,1846]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="6" pageNumber="307">Figs 26–31</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB6FFA3FF795C34958A78BF" pageId="6" pageNumber="307">
<paragraph id="8B993665FFB6FFA3FF795C34958D78F1" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
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7. Endogynium—a narrow tube dorso-ventrally oriented and visible as a very small ring (
<figureCitation id="131D2AE0FFB6FFA3FBCE5C3494817808" box="[1056,1168,1854,1875]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 35–39</figureCitation>
), sometimes open on one or both sides, when the tube should be obliquely oriented (
<figureCitation id="131D2AE0FFB6FFA3FCE75C51936D782B" box="[777,892,1883,1904]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 37, 38</figureCitation>
); ventral margin of the endogynium in most specimens bears triangular, asymmetric lamellar protrusion (
<figureCitation id="131D2AE0FFB6FFA3FD535C72933C78D6" box="[701,813,1912,1933]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 35, 36</figureCitation>
); ventrally, delicate endogynial lamella dentate anteriorly covers the endogynium (
<figureCitation id="131D2AE0FFB6FFA3FE675C9F922F78F1" box="[393,574,1941,1962]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 35, 36, 39, 40</figureCitation>
)...................................
<taxonomicName id="4C264DE6FFB6FFA3FC1E5C9F950A78F1" authority="Witalinski, 2016" authorityName="Witalinski" authorityYear="2016" box="[1008,1307,1940,1962]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="calpetanus">
<emphasis id="B952EA77FFB6FFA3FC1E5C9F941078F1" box="[1008,1025,1941,1962]" italics="true" pageId="6" pageNumber="307">H</emphasis>
.
<emphasis id="B952EA77FFB6FFA3FBE05C9E946778F1" box="[1038,1142,1940,1962]" italics="true" pageId="6" pageNumber="307">calpetanus</emphasis>
Witaliński, 2016
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(
<figureCitation id="131D2AE0FFB6FFA3FAC95C9F958478F1" box="[1319,1429,1941,1962]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">Figs 32–40</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB6FFA3FF795CB8958A78BF" blockId="6.[151,1436,1299,2020]" pageId="6" pageNumber="307">
<keyLead id="30D79550FFB6FFA3FF795CB8958A78BF" pageId="6" pageNumber="307">
- Endogynium saccular and large, its diameter similar or larger than the size of epigynium subapical structure (e.g.
<figureCitation id="131D2AE0FFB6FFA3FAC75CB89562789C" box="[1321,1395,1970,1991]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="6" pageNumber="307">
Figs
<date id="FF9810A5FFB6FFA3FAB75CB89562789C" box="[1369,1395,1970,1991]" pageId="6" pageNumber="307" value="1943-04-09">43</date>
</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FA935CB89589789C" box="[1405,1432,1970,1991]" captionStart="FIGURES 1 – 10" captionStartId="14.[151,264,1907,1930]" captionTargetBox="[209,1408,201,1857]" captionTargetId="figure@14.[151,1436,195,1883]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Females. Holoparasitus spp.: 1 ventral side of idiosoma; 2 sternum, presternal plate and lateral presternal platelets; 3 paragynium; 4 epigynium; 5 anterior part of epigynium; 6, 7 endogynium; 8 gnathosoma and gnathotectum, dorsally; 9 gnathosoma, ventrally; 10 pedipalp trochanter." httpUri="https://zenodo.org/record/810607/files/figure.png" pageId="6" pageNumber="307">4 9</figureCitation>
,
<figureCitation id="131D2AE0FFB6FFA3FF0F5CC590EA78BF" box="[225,251,1999,2020]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="6" pageNumber="307">56</figureCitation>
)................................................................................................. 8
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</paragraph>
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<keyStep id="30D22EC0FFB7FFA2FF795B9D958D7E58" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF795B9D95877FBC" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795B9D95877FBC" pageId="7" pageNumber="308">
8. Saccular endogynium with lateral walls forming two large dentated invaginations antero-dorsally directed; stout asymmetric prolongation located in the endogynium midline and protruded anteriorly (
<figureCitation id="131D2AE0FFB7FFA2FC415BBE94317F92" box="[943,1056,180,201]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Figs 43, 44</figureCitation>
); epigynium with broad and blunt apex (
<figureCitation id="131D2AE0FFB7FFA2FF095BD891447FBC" box="[231,341,210,231]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Figs 41, 42</figureCitation>
)..........................................................
<taxonomicName id="4C264DE6FFB7FFA2FBCF5BD895057FBC" authority="Hyatt, 1987" authorityName="Hyatt" authorityYear="1987" box="[1057,1300,209,231]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="B952EA77FFB7FFA2FBCF5BD894237FBC" box="[1057,1074,210,231]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBAE5BDB948F7FBC" box="[1088,1182,209,231]" italics="true" pageId="7" pageNumber="308">lawrencei</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB4D5BD895057FBC" author="Hyatt" box="[1187,1300,210,231]" pageId="7" pageNumber="308" refString="Hyatt, K. H. (1987) Mites of the genus Holoparasitus Oudemans, 1936 (Mesostigmata: Parasitidae) in the British Isles. Bulletin of the British Museum (Natural History), Zoology, 52, 139 - 164." type="journal article" year="1987">Hyatt, 1987</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FACF5BD8959E7FBC" box="[1313,1423,210,231]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Figs 41–44</figureCitation>
)
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<keyLead id="30D79550FFB7FFA2FF735BE4958D7E58" box="[157,1436,238,259]" pageId="7" pageNumber="308">– Endogynium otherwise, with curtain-like structures or protrusions on its posterior margin; epigynium with triangular apex.. 9</keyLead>
</paragraph>
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<keyLead id="30D79550FFB7FFA2FF795A06958A7E7A" box="[151,1435,268,289]" pageId="7" pageNumber="308">
9. Endogynium with two membranous curtain-like structures or two fleshy lobes (
<figureCitation id="131D2AE0FFB7FFA2FC345A0694597E7A" box="[986,1096,268,289]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Figs 49, 51</figureCitation>
)......................... 10
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</paragraph>
<paragraph id="8B993665FFB7FFA2FF795A23958A7E65" blockId="7.[151,1436,151,2033]" box="[151,1435,297,318]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795A23958A7E65" box="[151,1435,297,318]" pageId="7" pageNumber="308">
- Endogynium with one or two protrusions on the posterior margin (e.g.
<figureCitation id="131D2AE0FFB7FFA2FC665A2394087E65" box="[904,1049,297,318]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 56, 60, 66</figureCitation>
,
<figureCitation id="131D2AE0FFB7FFA2FBCD5A23942D7E65" box="[1059,1084,297,318]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">70</figureCitation>
).......................... 11
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<paragraph id="8B993665FFB7FFA2FF795A4C958D7ECE" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795A4C958D7ECE" pageId="7" pageNumber="308">
10. Endogynium with two membranous curtain-like structures (
<figureCitation id="131D2AE0FFB7FFA2FCDE5A4C936B7E00" box="[816,890,326,347]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Fig. 49</figureCitation>
); epigynium with prominent triangular apex; subapical structure anteriorly rounded, with two tiny oval extensions near the apex base (
<figureCitation id="131D2AE0FFB7FFA2FC385A69940C7E23" box="[982,1053,355,376]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Fig. 48</figureCitation>
)................................................................................................
<taxonomicName id="4C264DE6FFB7FFA2FC1D5A8A950B7ECE" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1011,1306,383,405]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="mahnerti">
<emphasis id="B952EA77FFB7FFA2FC1D5A8A94157ECE" box="[1011,1028,384,405]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBFF5A7594767ECE" box="[1041,1127,383,405]" italics="true" pageId="7" pageNumber="308">mahnerti</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB835A8A950B7ECE" author="Juvara-Bals" box="[1133,1306,384,405]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC95A8A95877ECE" box="[1319,1430,384,405]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Figs 45–49</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB7FFA2FF795A97958A7EB7" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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- Endogynium with two arcuate lamellar prolongations (
<figureCitation id="131D2AE0FFB7FFA2FD1C5A97932B7EE9" box="[754,826,413,434]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Fig. 51</figureCitation>
); epigynium with a short, broad tip; subapical structure rectangular, with two membranous triangular extensions near the apex; sternal reticulation shows conspicuous loop-like pattern axially located (
<figureCitation id="131D2AE0FFB7FFA2FEB35ADD91B27EB7" box="[349,419,471,492]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Fig. 50</figureCitation>
).....................................................
<taxonomicName id="4C264DE6FFB7FFA2FBCE5ADD950B7EB7" authority="Hyatt, 1987" authorityName="Hyatt" authorityYear="1987" box="[1056,1306,471,492]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="maritimus">
<emphasis id="B952EA77FFB7FFA2FBCE5ADD94247EB7" box="[1056,1077,471,492]" italics="true" pageId="7" pageNumber="308">H</emphasis>
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<emphasis id="B952EA77FFB7FFA2FBAC5ADD94B27EB7" box="[1090,1187,471,492]" italics="true" pageId="7" pageNumber="308">maritimus</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB475ADD950B7EB7" author="Hyatt" box="[1193,1306,471,492]" pageId="7" pageNumber="308" refString="Hyatt, K. H. (1987) Mites of the genus Holoparasitus Oudemans, 1936 (Mesostigmata: Parasitidae) in the British Isles. Bulletin of the British Museum (Natural History), Zoology, 52, 139 - 164." type="journal article" year="1987">Hyatt, 1987</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC95ADD95847EB7" box="[1319,1429,471,492]" captionStart="FIGURES 32 – 51" captionStartId="16.[151,264,1820,1843]" captionTargetBox="[184,1396,195,1785]" captionTargetId="figure@16.[172,1408,193,1795]" captionTargetPageId="16" captionText="FIGURES 32 – 51. Females. Holoparasitus calpetanus: 32 presternal plate; 33 sternum; 34 paragynium; 35 endogynium covered with endogynial lamella visible between paragynia; 36 endogynium covered with endogynial lamella; 37, 38 endogynium between paragynia, two oblique aspects; 39 endogynium with endogynial lemella; 40 endogynial lamella. Holoparasitus lawrencei: 41 epigynium; 42 central prong of epigynium with subapical structure; 43, 44 endogynium, two aspects. Holoparasitus machnerti: 45 gnathotectum; 46 presternal plate, lateral platelets and sternum; 47 paragynium, posterolateral protrusion; 48 epigynium; 49 endogynium. Holoparasitus maritimus: 50 sternum; 51 endogynium. Scale bars: 32 – 34, 41 – 49 0.1 mm, 35 – 40 0.05 mm." httpUri="https://zenodo.org/record/810611/files/figure.png" pageId="7" pageNumber="308">Figs 50, 51</figureCitation>
)
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<keyLead id="30D79550FFB7FFA2FF795AFE958A7D52" box="[151,1435,500,521]" pageId="7" pageNumber="308">
11. Endogynium with one protrusion on the posterior margin (e.g.
<figureCitation id="131D2AE0FFB7FFA2FCD55AFE93DA7D52" box="[827,971,500,521]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 56, 60, 65</figureCitation>
).................................... 12
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF79591B958A7D7D" blockId="7.[151,1436,151,2033]" box="[151,1435,529,550]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF79591B958A7D7D" box="[151,1435,529,550]" pageId="7" pageNumber="308">
- Endogynium with two protrusions or one deeply split protrusion on the posterior margin (e.g.
<figureCitation id="131D2AE0FFB7FFA2FB8B591B94E47D7D" box="[1125,1269,529,550]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 70, 79, 80</figureCitation>
)........... 14
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</paragraph>
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<keyStep id="30D22EC0FFB7FFA2FF795924958A7D26" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF795924958D7D3B" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795924958D7D3B" pageId="7" pageNumber="308">
12. Endogynium with a single finger-like, occasionally forked out protrusion and 7–8 denticles on its anterior and lateral walls (
<figureCitation id="131D2AE0FFB7FFA2FF065941913F7D3B" box="[232,302,587,608]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Fig. 56</figureCitation>
)...........................................................
<taxonomicName id="4C264DE6FFB7FFA2FBEA5946950B7D3B" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1028,1306,587,609]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="rifensis">
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<emphasis id="B952EA77FFB7FFA2FBCF594194767D3A" box="[1057,1127,587,609]" italics="true" pageId="7" pageNumber="308">rifensis</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB835941950B7D3B" author="Juvara-Bals" box="[1133,1306,587,608]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC9594195847D3B" box="[1319,1429,587,608]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 52–56</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB7FFA2FF795962958A7D26" blockId="7.[151,1436,151,2033]" box="[151,1435,616,637]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795962958A7D26" box="[151,1435,616,637]" pageId="7" pageNumber="308">- Endogynium with a single protrusion only, denticles on its walls absent.......................................... 13</keyLead>
</paragraph>
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<keyStep id="30D22EC0FFB7FFA2FF79598F958D7C77" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF79598F958D7D8E" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF79598F958D7D8E" pageId="7" pageNumber="308">
13. Apex of the finger-like endogynial protrusion sharp (
<figureCitation id="131D2AE0FFB7FFA2FD34598F93597DC1" box="[730,840,645,666]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 60, 61</figureCitation>
); apex of the epigynium mucronate, epigynial subapical structure trapezoidal and well sclerotised, with small hyaline lateral extensions (
<figureCitation id="131D2AE0FFB7FFA2FC5C59A9940E7DE3" box="[946,1055,675,696]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 58, 59</figureCitation>
)...................................................................................................
<taxonomicName id="4C264DE6FFB7FFA2FBFB59CA950B7D8E" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1045,1306,703,725]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="lunae">
<emphasis id="B952EA77FFB7FFA2FBFB59CA94377D8E" box="[1045,1062,704,725]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBDC59B594767D8E" box="[1074,1127,703,725]" italics="true" pageId="7" pageNumber="308">lunae</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB8359CA950B7D8E" author="Juvara-Bals" box="[1133,1306,704,725]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC959CA95847D8E" box="[1319,1429,704,725]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 57–61</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF7959D7958D7C77" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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- Apex of a slender endogynial protrusion usually sharp and split, ending at the anterior margin of the endogynium (
<figureCitation id="131D2AE0FFB7FFA2FACF59D7959F7DA9" box="[1313,1422,733,754]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 65–67</figureCitation>
); apex of the epigynium large and triangular; epigynial subapical structure formed by a sclerotised rectangular line under the apex, and by the two wing-like hyaline lateral extensions (
<figureCitation id="131D2AE0FFB7FFA2FCFE581D93477C77" box="[784,854,791,812]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Fig. 63</figureCitation>
).............
<taxonomicName id="4C264DE6FFB7FFA2FC11581D950B7C77" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1023,1306,790,812]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="malleus">
<emphasis id="B952EA77FFB7FFA2FC11581D94017C77" box="[1023,1040,791,812]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBF3581C94797C77" box="[1053,1128,790,812]" italics="true" pageId="7" pageNumber="308">malleus</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB83581D950B7C77" author="Juvara-Bals" box="[1133,1306,791,812]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC9581D95847C77" box="[1319,1429,791,812]" captionStart="FIGURES 52 – 67" captionStartId="17.[151,264,1738,1761]" captionTargetBox="[200,1411,246,1685]" captionTargetId="figure@17.[151,1436,193,1717]" captionTargetPageId="17" captionText="FIGURES 52 – 67. Females. Holoparasitus rifensis: 52 presternal plate, lateral platelets and sternum; 53 epigynium; 54 epigynium subapical structure under central prong; 55 paragynium; 56 endogynium. Holoparasitus lunae: 57 presternal plate, lateral platelets, sternum and paragynium; 58 epigynium; 59 epigynium subapical structure; 60, 61 endogynium. Holoparasitus malleus: 62 presternal plate, lateral platelets and sternum; 63 epigynium; 64 paragynia and endogynium; 65 – 67 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810613/files/figure.png" pageId="7" pageNumber="308">Figs 62–67</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB7FFA2FF79583E958A7CD8" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF79583E958D7C3D" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF79583E958D7C3D" pageId="7" pageNumber="308">
14. Presternal plate non-serrated (
<figureCitation id="131D2AE0FFB7FFA2FDEA583E925C7C12" box="[516,589,820,841]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Fig. 68</figureCitation>
); endogynium with two short posterior protrusions far apart from each other (
<figureCitation id="131D2AE0FFB7FFA2FAA9583E959C7C12" box="[1351,1421,820,841]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Fig. 70</figureCitation>
); big species........................................................
<taxonomicName id="4C264DE6FFB7FFA2FC02585B950B7C3D" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1004,1306,849,870]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="B952EA77FFB7FFA2FC02585B93EC7C3D" box="[1004,1021,849,870]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBE4585B94767C3D" box="[1034,1127,849,870]" italics="true" pageId="7" pageNumber="308">giganteus</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB83585B950B7C3D" author="Juvara-Bals" box="[1133,1306,849,870]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC9585B95857C3D" box="[1319,1428,849,870]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 68–71</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF795864958A7CD8" blockId="7.[151,1436,151,2033]" box="[151,1435,878,899]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795864958A7CD8" box="[151,1435,878,899]" pageId="7" pageNumber="308">- Presternal plate more or less distinctly serrated; endogynial protrusions of different shape and size.................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB7FFA2FF795881958A7B6A" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF795881958D7CAC" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795881958D7CAC" pageId="7" pageNumber="308">
15. Presternal plate with a few minute denticles (
<figureCitation id="131D2AE0FFB7FFA2FD7E588192C97CFB" box="[656,728,907,928]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Fig. 73</figureCitation>
); gland pore
<emphasis id="B952EA77FFB7FFA2FCBA588193697CFA" box="[852,888,907,929]" italics="true" pageId="7" pageNumber="308">gv1</emphasis>
located on a soft cuticle or on the posterior margin of the sternum; endogynial protrusions involute, unequal in size and distant of each other (
<figureCitation id="131D2AE0FFB7FFA2FBFE58A294487CE6" box="[1040,1113,936,957]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Fig. 76</figureCitation>
); epigynium with triangular apex and a subapical structure with a conspicuous, rectangular sclerotised line continued by the two trapezoidal membranous wings (
<figureCitation id="131D2AE0FFB7FFA2FF0958E891447CAC" box="[231,341,994,1015]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 74, 75</figureCitation>
).......................................................
<taxonomicName id="4C264DE6FFB7FFA2FC1458E9950B7CAC" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1018,1306,994,1016]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="lapalma">
<emphasis id="B952EA77FFB7FFA2FC1458E9941A7CA3" box="[1018,1035,995,1016]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBF758E894767CA3" box="[1049,1127,994,1016]" italics="true" pageId="7" pageNumber="308">lapalma</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB8358E8950B7CAC" author="Juvara-Bals" box="[1133,1306,994,1015]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB7FFA2FAC958E895847CAC" box="[1319,1429,994,1015]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 72–77</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF7958F5958A7B6A" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF7958F5958A7B6A" pageId="7" pageNumber="308">
- Presternal plate distinctly serrated; gland pore
<emphasis id="B952EA77FFB7FFA2FD7358F592D17B4E" box="[669,704,1023,1045]" italics="true" pageId="7" pageNumber="308">gv1</emphasis>
always located on the sternum near the posterior margin; endogynial protrusions well developed; epigynial apex triangular, subapical structure otherwise..................................... 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB7FFA2FF795F30958A7BD2" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF795F30958D7B37" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795F30958D7B37" pageId="7" pageNumber="308">
16. Endogynium—two protrusions of different length, with their bases very close to each other, or one deeply split protrusion (
<figureCitation id="131D2AE0FFB7FFA2FF095F5D91447B37" box="[231,341,1111,1132]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 79–81</figureCitation>
)......................................................
<taxonomicName id="4C264DE6FFB7FFA2FC035F5D950B7B37" authority="Juvara-Bals, 2008" authorityName="Juvara-Bals" authorityYear="2008" box="[1005,1306,1110,1132]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B952EA77FFB7FFA2FC035F5D93EF7B37" box="[1005,1022,1111,1132]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FBE25F5C94797B37" box="[1036,1128,1110,1132]" italics="true" pageId="7" pageNumber="308">variabilis</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB835F5D950B7B37" author="Juvara-Bals" box="[1133,1306,1111,1132]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (2008) New species of Holoparasitus Oudemans, 1936 (Acari, Parasitidae) from Spain, North Africa and Madeira Islands. Revue Suisse de Zoologie, 115, 37 - 84." type="journal article" year="2008">Juvara-Bals, 2008</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB7FFA2FAC95F5D95847B37" box="[1319,1429,1111,1132]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 78–81</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF795F7E958A7BD2" blockId="7.[151,1436,151,2033]" box="[151,1435,1140,1161]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795F7E958A7BD2" box="[151,1435,1140,1161]" pageId="7" pageNumber="308">- Endogynium with two equally sized protrusions............................................................ 17</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8B993665FFB7FFA2FF795F9B958D7B98" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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17. Gnathotectum large, triangular (
<figureCitation id="131D2AE0FFB7FFA2FDF85F9B924F7BFD" box="[534,606,1169,1190]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Fig. 82</figureCitation>
); large cup-like endogynium (
<figureCitation id="131D2AE0FFB7FFA2FC6F5F9B93D97BFD" box="[897,968,1169,1190]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Fig. 84</figureCitation>
) with two protrusions, their apices medially oriented and distant of each other; trochanter IV with ventral protuberance..........
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.
<emphasis id="B952EA77FFB7FFA2FBC25FA494797B98" box="[1068,1128,1198,1219]" italics="true" pageId="7" pageNumber="308">anaga</emphasis>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC95FA495847B98" box="[1319,1429,1198,1219]" captionStart="FIGURES 68 – 84" captionStartId="18.[151,264,1890,1913]" captionTargetBox="[186,1415,194,1864]" captionTargetId="figure@18.[151,1436,177,1897]" captionTargetPageId="18" captionText="FIGURES 68 – 84. Females. Holoparasitus giganteus: 68 presternal plate, lateral platelets and sternum; 69 epigynium; 70 endogynium, laterally; 71 paragynium. Holoparasitus lapalma: 72 pedipalpal trochanter posterolaterally; 73 presternal plate, lateral platelets and sternum; 74 epigynium; 75 epigynium central prong; 76 endogynium; 77 paragynium. Holoparasitus variabilis: 78 epigynium central prong; 79 endogynium; 80 endogynial stipula; 81 paragynium and endogynium. Holoparasitus anaga: 82 gnathotectum; 83 epigynium subapical structure; 84 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810615/files/figure.png" pageId="7" pageNumber="308">Figs 82–84</figureCitation>
)
</keyLead>
</paragraph>
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<keyLead id="30D79550FFB7FFA2FF795FC1958A7BBB" box="[151,1435,1227,1248]" pageId="7" pageNumber="308">- Gnathotectum trispinate; endogynium otherwise; trochanter IV without any protuberance........................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB7FFA2FF795FE2958A7A41" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF795FE2958A7BA6" blockId="7.[151,1436,151,2033]" box="[151,1435,1256,1277]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795FE2958A7BA6" box="[151,1435,1256,1277]" pageId="7" pageNumber="308">
18. Endogynium with the two horn-like or triangular protrusions (
<figureCitation id="131D2AE0FFB7FFA2FCDA5FE293D47BA6" box="[820,965,1256,1277]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 88, 90, 91</figureCitation>
).................................... 19
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF795E0F958A7A41" blockId="7.[151,1436,151,2033]" box="[151,1435,1285,1306]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795E0F958A7A41" box="[151,1435,1285,1306]" pageId="7" pageNumber="308">
- Endogynium with the two finger-like, rather long protrusions (
<figureCitation id="131D2AE0FFB7FFA2FCD65E0F93B67A41" box="[824,935,1285,1306]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 95, 98</figureCitation>
,
<figureCitation id="131D2AE0FFB7FFA2FC5E5E0F94247A41" box="[944,1077,1285,1306]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">101, 102, 104</figureCitation>
)........................... 20
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<keyStep id="30D22EC0FFB7FFA2FF795E28958D7AF0" pageId="7" pageNumber="308">
<paragraph id="8B993665FFB7FFA2FF795E28958D7A2A" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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19. Endogynium with two distant horn-like protrusions (
<figureCitation id="131D2AE0FFB7FFA2FD025E2893267A6C" box="[748,823,1314,1335]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Fig. 88</figureCitation>
); epigynial apex not prominent, subapical structure with oval wings (
<figureCitation id="131D2AE0FFB7FFA2FEC65E3591617A0F" box="[296,368,1343,1364]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Fig. 87</figureCitation>
); palptrochanter seta
<date id="FF9810A5FFB7FFA2FDAE5E3492467A0F" box="[576,599,1342,1364]" pageId="7" pageNumber="308">
<emphasis id="B952EA77FFB7FFA2FDAE5E3492467A0F" box="[576,599,1342,1364]" italics="true" pageId="7" pageNumber="308">v1</emphasis>
</date>
pectinate,
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<emphasis id="B952EA77FFB7FFA2FD2F5E3492C97A0F" box="[705,728,1342,1364]" italics="true" pageId="7" pageNumber="308">v2</emphasis>
</date>
barbed, with a protuberance between them; palpgenu with tubercle close to the anterolateral seta...............................................
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<emphasis id="B952EA77FFB7FFA2FBFA5E5694767A29" box="[1044,1127,1372,1394]" italics="true" pageId="7" pageNumber="308">vaucheri</emphasis>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC95E5695847A2A" box="[1319,1429,1372,1393]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 85–88</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB7FFA2FF795E73958D7AF0" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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- Endogynium with the two triangular protrusions close to each other (
<figureCitation id="131D2AE0FFB7FFA2FC985E7393F77AD5" box="[886,998,1401,1422]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 90, 91</figureCitation>
); epigynial subapical structure extending into the fan-like wings (
<figureCitation id="131D2AE0FFB7FFA2FE755E9C91F07AF0" box="[411,481,1430,1451]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Fig. 92</figureCitation>
)..............................................
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<emphasis id="B952EA77FFB7FFA2FBC15E9C94767AF7" box="[1071,1127,1430,1452]" italics="true" pageId="7" pageNumber="308">franzi</emphasis>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC95E9C95847AF0" box="[1319,1429,1430,1451]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 89–92</figureCitation>
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<keyStep id="30D22EC0FFB7FFA2FF795EBE958A797B" pageId="7" pageNumber="308">
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20. Endogynium with no teeth on either lateral or anterior walls, endogynial protrusions distant, with a gap of 40–45 µm between their bases (
<figureCitation id="131D2AE0FFB7FFA2FEBD5EDA91D07ABE" box="[339,449,1488,1509]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 95, 96</figureCitation>
)...........................................
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<emphasis id="B952EA77FFB7FFA2FC175EDB94767ABD" box="[1017,1127,1489,1510]" italics="true" pageId="7" pageNumber="308">canariensis</emphasis>
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<figureCitation id="131D2AE0FFB7FFA2FAC95EDA95847ABE" box="[1319,1429,1488,1509]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 93–96</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB7FFA2FF795EE7958A797B" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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- Endogynium with teeth on the lateral and anterior walls, long endogynial protrusions straight or sinuous, their tips reaching the anterior margin of the endogynium and their bases close to each other (
<figureCitation id="131D2AE0FFB7FFA2FC645D0193C5797B" box="[906,980,1547,1568]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 98</figureCitation>
,
<figureCitation id="131D2AE0FFB7FFA2FC305D019422797B" box="[990,1075,1547,1568]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">101, 104</figureCitation>
)........................... 21
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21. Lamellar endogynial protrusions straight (
<figureCitation id="131D2AE0FFB7FFA2FD915D2292D87966" box="[639,713,1576,1597]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Fig. 98</figureCitation>
), distance between their bases ca. 10 µm; palptrochanter with simple
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, barbed
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<emphasis id="B952EA77FFB7FFA2FEC65D4E912E7901" box="[296,319,1604,1626]" italics="true" pageId="7" pageNumber="308">v2</emphasis>
</date>
, and a protuberance between them.............................
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.
<emphasis id="B952EA77FFB7FFA2FBE95D4E94767901" box="[1031,1127,1604,1626]" italics="true" pageId="7" pageNumber="308">mallorcae</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FB835D4F950B7901" author="Juvara-Bals" box="[1133,1306,1605,1626]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. (1975) Sur le genre Holoparasitus Oudemans, 1936 et sur certains caracteres morphologiques de la famille Parasitidae Oudem. (Parasitiformes). Acarologia, 17, 384 - 409." type="journal article" year="1975">Juvara-Bals, 1975</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FAC95D4F95847901" box="[1319,1429,1605,1626]" captionStart="FIGURES 85 – 98" captionStartId="19.[151,264,1602,1625]" captionTargetBox="[196,1409,210,1564]" captionTargetId="figure@19.[151,1436,167,1581]" captionTargetPageId="19" captionText="FIGURES 85 – 98. Females. Holoparasitus vaucheri: 85 gnathotectum; 86 paragynium; 87 epigynium; 88 endogynium. Holoparasitus franzi: 89 gnathotectum; 90 paragynium and endogynium; 91 endogynium; 92 epigynium. Holoparasitus canariensis: 93 paragynium; 94 epigynium; 95 endogynium; 96 endogynium under paragynial central prong. Holoparasitus mallorcae: 97 epigynium; 98 endogynium, lateral aspect. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810617/files/figure.png" pageId="7" pageNumber="308">Figs 97, 98</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB7FFA2FF795D68958A79CF" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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- Lamellar endogynial protrusions sinuous (
<figureCitation id="131D2AE0FFB7FFA2FD965D689320792C" box="[632,817,1634,1655]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 101, 102, 104</figureCitation>
), their bases very close to each other; palptrochanter with pectinate
<date id="FF9810A5FFB7FFA2FEE15D74913779CF" box="[271,294,1662,1684]" pageId="7" pageNumber="308">
<emphasis id="B952EA77FFB7FFA2FEE15D74913779CF" box="[271,294,1662,1684]" italics="true" pageId="7" pageNumber="308">v1</emphasis>
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and barbed
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<emphasis id="B952EA77FFB7FFA2FE755D7491A379CF" box="[411,434,1662,1684]" italics="true" pageId="7" pageNumber="308">v2</emphasis>
.
</date>
................................................................................ 22
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<paragraph id="8B993665FFB7FFA2FF795D9695877995" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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22. Endogynium with several (3–7) denticles and two protrusions extending beyond the anterior endogynial margin (
<figureCitation id="131D2AE0FFB7FFA2FAD15D9691177995" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 101, 102</figureCitation>
)..............................................
<taxonomicName id="4C264DE6FFB7FFA2FCD45DB394F47995" authority="Juvara-Bals &amp; Witalinski, 2000" authorityName="Juvara-Bals &amp; Witalinski" authorityYear="2000" box="[826,1253,1720,1742]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="ellipticus">
<emphasis id="B952EA77FFB7FFA2FCD45DB3935A7995" box="[826,843,1721,1742]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FCB65DB293A17995" box="[856,944,1720,1742]" italics="true" pageId="7" pageNumber="308">ellipticus</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FC585DB394F47995" author="Juvara-Bals" box="[950,1253,1721,1742]" pageId="7" pageNumber="308" refString="Juvara-Bals, I. &amp; Witalinski, W. (2000) Description of five new species of Holoparasitus s. str. with redescription of H. apenninorum (Berlese, 1906) and H. cultriger (Berlese, 1906) from Italy and Spain (Acari, Gamasida, Parasitidae). Revue Suisse de Zoologie, 107, 3 - 30." type="journal article" year="2000">Juvara-Bals &amp; Witaliński, 2000</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB7FFA2FB1F5DB3952A7995" box="[1265,1339,1721,1742]" captionStart="FIGURES 11 – 31" captionStartId="15.[151,264,1816,1839]" captionTargetBox="[255,1319,215,1771]" captionTargetId="figure@15.[238,1330,193,1771]" captionTargetPageId="15" captionText="FIGURES 11 – 31. Females. Holoparasitus inornatus: 11 presternal plate, sternum, paragynia; 12 sternum, paragynia, endogynium; 13 endogynium. Holoparasitus megacalcaratus: 14 ventral side of idiosoma. Holoparasitus ellipticus: 15 presternal plate and lateral platelets. Holoparasitus spp.: 16, 17 presternal plate and lateral platelets. Holoparasitus siculus: 18 epigynium; 19, 20 central prong of epigynium and endogynium; 21 endogynium. Holoparasitus algiersensis: 22 presternal plate; 23 presternal plate and sternum; 24 paragynia and epigynium; 25 endogynium. Holoparasitus eivissa: 26 presternal plate and sternum; 27 epigynium; 28, 29 endogynium under paragynia and epigynium; 30 paragynium; 31 endogynium laterally. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810609/files/figure.png" pageId="7" pageNumber="308">Figs 15</figureCitation>
,
<figureCitation id="131D2AE0FFB7FFA2FAA85DB3959E7995" box="[1350,1423,1721,1742]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">99–102</figureCitation>
)
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<paragraph id="8B993665FFB7FFA2FF795DDC958A7853" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795DDC958A7853" pageId="7" pageNumber="308">
- Endogynium (
<figureCitation id="131D2AE0FFB7FFA2FE845DDC91AA79B0" box="[362,443,1750,1771]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Fig. 104</figureCitation>
) with the two lateral denticles, protrusions ending at the anterior endogynial margin......................................................................
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<emphasis id="B952EA77FFB7FFA2FC835DFE93927852" box="[877,899,1780,1801]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FC615DF993DC7852" box="[911,973,1779,1801]" italics="true" pageId="7" pageNumber="308">gibber</emphasis>
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(
<figureCitation id="131D2AE0FFB7FFA2FAE15DF995857853" box="[1295,1428,1779,1800]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 103, 104</figureCitation>
)
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23. Endogynium bubble-shaped (e.g.
<figureCitation id="131D2AE0FFB7FFA2FDCB5C1A92C6787E" box="[549,727,1808,1829]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 109, 113, 119</figureCitation>
)................................... (
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<emphasis id="B952EA77FFB7FFA2FB795C1A94C9787D" bold="true" box="[1175,1240,1808,1830]" italics="true" pageId="7" pageNumber="308">caesus</emphasis>
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species-group)
</emphasis>
...24
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<keyLead id="30D79550FFB7FFA2FF795C27958A7819" box="[151,1435,1837,1858]" pageId="7" pageNumber="308">- Endogynium non-bubble-shaped......................................................................... 27</keyLead>
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<paragraph id="8B993665FFB7FFA2FF795C40958D7827" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
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24. Tube leading to the endogynial sac relatively wide, ca. 1/2 of the sac diameter (
<figureCitation id="131D2AE0FFB7FFA2FC365C4094717804" box="[984,1120,1866,1887]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 107, 109</figureCitation>
).........................................................................................
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<emphasis id="B952EA77FFB7FFA2FC185C6D944F7826" box="[1014,1118,1895,1917]" italics="true" pageId="7" pageNumber="308">ampullaris</emphasis>
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</taxonomicName>
(
<figureCitation id="131D2AE0FFB7FFA2FAE15C6D95847827" box="[1295,1429,1895,1916]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 105–109</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB7FFA2FF795C8E958A78EC" blockId="7.[151,1436,151,2033]" pageId="7" pageNumber="308">
<keyLead id="30D79550FFB7FFA2FF795C8E958A78EC" pageId="7" pageNumber="308">
- Tube leading to the sac narrow, its diameter not exceeding 1/3 of the sac diameter (
<figureCitation id="131D2AE0FFB7FFA2FBFD5C8E94DC78C2" box="[1043,1229,1924,1945]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 113, 116, 119</figureCitation>
); subapical epigynial structure compact (
<figureCitation id="131D2AE0FFB7FFA2FE775CA8925B78EC" box="[409,586,1954,1975]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 111, 115, 118</figureCitation>
).................................................................... 25
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</paragraph>
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<keyStep id="30D22EC0FFB7FFADFF795CB5958A7FF7" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
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<keyLead id="30D79550FFB7FFA2FF795CB5958D78AA" pageId="7" pageNumber="308">
25. Subapical structure of the epigynium with a central rectangular element (
<figureCitation id="131D2AE0FFB7FFA2FC475CB59421788F" box="[937,1072,1983,2004]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 111, 112</figureCitation>
); endogynial sac spherical or slightly ellipsoidal (
<figureCitation id="131D2AE0FFB7FFA2FEBF5CD691C478AA" box="[337,469,2012,2033]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 112, 113</figureCitation>
)..........................................
<taxonomicName id="4C264DE6FFB7FFA2FC345CD6951578AA" authority="Micherdzinski, 1969" authorityName="Micherdzinski" authorityYear="1969" box="[986,1284,2012,2033]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="caesus">
<emphasis id="B952EA77FFB7FFA2FC345CD693FA78AA" box="[986,1003,2012,2033]" italics="true" pageId="7" pageNumber="308">H</emphasis>
.
<emphasis id="B952EA77FFB7FFA2FC165CD6942978AA" box="[1016,1080,2012,2033]" italics="true" pageId="7" pageNumber="308">caesus</emphasis>
<bibRefCitation id="EFB74B94FFB7FFA2FBD05CD6951578AA" author="Micherdzinski" box="[1086,1284,2012,2033]" pageId="7" pageNumber="308" refString="Micherdzinski, W. (1969) Die Familie Parasitidae Oudemans, 1901 (Acarina, Mesostigmata). Panstwowe Wydawnictwo Naukowe, Krakow, 690 pp." type="book" year="1969">Micherdziński, 1969</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB7FFA2FAFF5CD6958478AA" box="[1297,1429,2012,2033]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="7" pageNumber="308">Figs 110–113</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795B9D958A7FF7" blockId="8.[151,1436,151,2035]" box="[151,1435,151,172]" pageId="8" pageNumber="309">
<materialsCitation id="3B4E3C38FFB8FFADFF795B9D958A7FF7" ID-GBIF-Occurrence="1569232893" box="[151,1435,151,172]" location="Central" pageId="8" pageNumber="309" specimenCount="1" stateProvince="Central">
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-
<collectingRegion id="49E2F887FFB8FFADFF0F5B9D91397FF7" box="[225,296,151,172]" country="Israel" name="Central" pageId="8" pageNumber="309">Central</collectingRegion>
rectangular element in the epigynial subapical structure absent.......................................... 26
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</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795BBE958D7FBC" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795BBE958D7F92" blockId="8.[151,1436,151,2035]" box="[151,1436,180,202]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795BBE958D7F92" box="[151,1436,180,202]" pageId="8" pageNumber="309">
26. Endogynial sac ellipsoidal (
<figureCitation id="131D2AE0FFB8FFADFE055BBE922F7F92" box="[491,574,180,201]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="8" pageNumber="309">Fig. 116</figureCitation>
)...............................
<taxonomicName id="4C264DE6FFB8FFADFC535BBF95157F92" authority="Juvara-Bals, 1975" authorityName="Juvara-Bals" authorityYear="1975" box="[957,1284,180,202]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="B952EA77FFB8FFADFC535BBF93DF7F91" box="[957,974,181,202]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC345BBE94407F91" box="[986,1105,180,202]" italics="true" pageId="8" pageNumber="309">tuberculatus</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFBB95BBE95157F92" author="Juvara-Bals" box="[1111,1284,180,201]" pageId="8" pageNumber="309" refString="Juvara-Bals, I. (1975) Sur le genre Holoparasitus Oudemans, 1936 et sur certains caracteres morphologiques de la famille Parasitidae Oudem. (Parasitiformes). Acarologia, 17, 384 - 409." type="journal article" year="1975">Juvara-Bals, 1975</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB8FFADFAFF5BBE95847F92" box="[1297,1429,180,201]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="8" pageNumber="309">Figs 114–116</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB8FFADFF795BD8958D7FBC" blockId="8.[151,1436,151,2035]" box="[151,1436,209,231]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795BD8958D7FBC" box="[151,1436,209,231]" pageId="8" pageNumber="309">
- Endogynial sac posteriorly flattened (
<figureCitation id="131D2AE0FFB8FFADFDA75BD892887FBC" box="[585,665,210,231]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="8" pageNumber="309">Fig. 119</figureCitation>
).........................
<taxonomicName id="4C264DE6FFB8FFADFC3F5BD895157FBC" authority="Juvara-Bals, 1975" authorityName="Juvara-Bals" authorityYear="1975" box="[977,1284,209,231]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="fortunatus">
<emphasis id="B952EA77FFB8FFADFC3F5BD893F37FBC" box="[977,994,210,231]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC015BDB94407FBC" box="[1007,1105,209,231]" italics="true" pageId="8" pageNumber="309">fortunatus</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFBB95BD895157FBC" author="Juvara-Bals" box="[1111,1284,210,231]" pageId="8" pageNumber="309" refString="Juvara-Bals, I. (1975) Sur le genre Holoparasitus Oudemans, 1936 et sur certains caracteres morphologiques de la famille Parasitidae Oudem. (Parasitiformes). Acarologia, 17, 384 - 409." type="journal article" year="1975">Juvara-Bals, 1975</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB8FFADFAFF5BD895847FBC" box="[1297,1429,210,231]" captionStart="FIGURES 99 – 119" captionStartId="20.[151,264,1810,1833]" captionTargetBox="[192,1380,195,1770]" captionTargetId="figure@20.[172,1400,167,1778]" captionTargetPageId="20" captionText="FIGURES 99 – 119. Females. Holoparasitus ellipticus: 99 presternal plate, lateral platelets and sternum; 100 epigynium, anterior part; 101, 102 endogynium. Holoparasitus gibber: 103 epigynium, anterior part; 104 endogynium. Holoparasitus ampullaris: 105 ventral side, genital region showing endogynium under paragynia and epigynium; 106 epigynium, anterior part; 107 – 109 endogynium, different aspects. Holoparasitus caesus: 110 gnathotectum; 111 epigynium central prong with subapical structure; 112 endogynium under subapical epigynial structure; 113 endogynium. Holoparasitus tuberculatus: 114 gnathotectum; 115 epigynial central prong with subapical structure; 116 endogynium under genital opening. Holoparasitus fortunatus: 117 gnathotectum; 118 epigynial central prong with subapical structure; 119 endogynium. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/810619/files/figure.png" pageId="8" pageNumber="309">Figs 117–119</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795BE4958A7E7A" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795BE4958A7E58" blockId="8.[151,1436,151,2035]" box="[151,1435,238,259]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795BE4958A7E58" box="[151,1435,238,259]" pageId="8" pageNumber="309">
27. Endogynium cup-shaped, circular (e.g.
<figureCitation id="131D2AE0FFB8FFADFDB55BE493017E58" box="[603,784,238,259]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 121, 125, 133</figureCitation>
,
<figureCitation id="131D2AE0FFB8FFADFCF45BE4932E7E58" box="[794,831,238,259]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">158</figureCitation>
) or rhombic (
<figureCitation id="131D2AE0FFB8FFADFC2D5BE494597E58" box="[963,1096,238,259]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 144, 146</figureCitation>
) in the ventral perspective..... 28
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795A06958A7E7A" blockId="8.[151,1436,151,2035]" box="[151,1435,268,289]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795A06958A7E7A" box="[151,1435,268,289]" pageId="8" pageNumber="309">
- Endogynium non-cup-shaped, neither circular nor rhombic in its outline (e.g.
<figureCitation id="131D2AE0FFB8FFADFC275A06945E7E7A" box="[969,1103,268,289]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="8" pageNumber="309">Figs 201, 208</figureCitation>
,
<figureCitation id="131D2AE0FFB8FFADFBB75A0694CD7E7A" box="[1113,1244,268,289]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="8" pageNumber="309">211, 216, 225</figureCitation>
)............. 45
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<paragraph id="8B993665FFB8FFADFF795A23958A7E00" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
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28. Endogynium without any thorns, teeth or lamellar protrusions, except a long band-like structure (endogynial lamella?) located posteriorly (
<figureCitation id="131D2AE0FFB8FFADFEB85A4C91B67E00" box="[342,423,326,347]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Fig. 121</figureCitation>
)...................................................
<taxonomicName id="4C264DE6FFB8FFADFBFF5A4C95127E00" authority="Vinnik, 1994" authorityName="Vinnik" authorityYear="1994" box="[1041,1283,326,347]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="inventus">
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<emphasis id="B952EA77FFB8FFADFBDE5A4C946E7E00" box="[1072,1151,326,347]" italics="true" pageId="8" pageNumber="309">inventus</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFB6B5A4C95127E00" author="Vinnik" box="[1157,1283,326,347]" pageId="8" pageNumber="309" refString="Vinnik, O. M. (1994) A new mite species of the genus Holoparasitus (Mesostigmata, Parasitidae) from the Crimea. Uestnik zoologii, 2, 82 - 84. [in Russian]" type="journal article" year="1994">Vinnik, 1994</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB8FFADFAE15A4C95847E00" box="[1295,1429,326,347]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 120, 121</figureCitation>
)
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<paragraph id="8B993665FFB8FFADFF795A69958A7E23" blockId="8.[151,1436,151,2035]" box="[151,1435,355,376]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795A69958A7E23" box="[151,1435,355,376]" pageId="8" pageNumber="309">- Endogynium with thorns, teeth and/or lamellar protrusions.................................................... 29</keyLead>
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<paragraph id="8B993665FFB8FFADFF795A8A958D7EE9" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
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29. Subapical structure of epigynium with a distinct, &quot;inverted–T&quot; thickening located axially (
<figureCitation id="131D2AE0FFB8FFADFBAE5A8A94D77ECE" box="[1088,1222,384,405]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 123, 127</figureCitation>
)............................................................................................. (
<emphasis id="B952EA77FFB8FFADFB665A9695787EE9" bold="true" box="[1160,1385,412,434]" pageId="8" pageNumber="309">
<taxonomicName id="4C264DE6FFB8FFADFB665A9694C97EE9" authorityName="Berlese" authorityYear="1906" box="[1160,1240,412,434]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="peraltus">
<emphasis id="B952EA77FFB8FFADFB665A9694C97EE9" bold="true" box="[1160,1240,412,434]" italics="true" pageId="8" pageNumber="309">peraltus</emphasis>
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species-group
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)...30
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795AB0958A7E94" blockId="8.[151,1436,151,2035]" box="[151,1435,442,463]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795AB0958A7E94" box="[151,1435,442,463]" pageId="8" pageNumber="309">- &quot;Inverted-T&quot; thickening absent.......................................................................... 31</keyLead>
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<keyStep id="30D22EC0FFB8FFADFF795ADD958D7D26" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795ADD958D7D7D" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795ADD958D7D7D" pageId="8" pageNumber="309">
30. Endogynium with narrow lamellar protrusions sometimes branched distally; endogynial sac bears many dents or thorns (
<figureCitation id="131D2AE0FFB8FFADFA9F5ADD91287D52" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 125, 126</figureCitation>
); the anterior thickenings of epigynial subapical structure well pronounced, the posterior thickenings oriented parallelly to each other (
<figureCitation id="131D2AE0FFB8FFADFE74591B920E7D7D" box="[410,543,529,550]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 122, 123</figureCitation>
)...........................
<taxonomicName id="4C264DE6FFB8FFADFC80591895127D7D" authority="Witalinski &amp; Skorupski, 2003" authorityName="Witalinski &amp; Skorupski" authorityYear="2003" box="[878,1283,529,551]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="nonaltus">
<emphasis id="B952EA77FFB8FFADFC805918936E7D7C" box="[878,895,530,551]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC65591B93CF7D7C" box="[907,990,529,551]" italics="true" pageId="8" pageNumber="309">nonaltus</emphasis>
Witaliński &amp; Skorupski, 2003
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(
<figureCitation id="131D2AE0FFB8FFADFAE1591B95847D7D" box="[1295,1429,529,550]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 122–126</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795924958D7D26" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795924958D7D26" pageId="8" pageNumber="309">
- Endogynium with broad lamellar protrusions indented terminally; endogynial sac with reticulation, but dents or thorns absent (
<figureCitation id="131D2AE0FFB8FFADFF065941912D7D3B" box="[232,316,587,608]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Fig. 129</figureCitation>
); the anterior thickenings of subapical epigynial structure not discernible, the posterior thickenings run convergently (
<figureCitation id="131D2AE0FFB8FFADFF065962912B7D26" box="[232,314,616,637]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Fig. 127</figureCitation>
)..........................................................
<taxonomicName id="4C264DE6FFB8FFADFC12596395127D26" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[1020,1283,616,638]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="peraltus">
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<emphasis id="B952EA77FFB8FFADFBF4596294797D25" box="[1050,1128,616,638]" italics="true" pageId="8" pageNumber="309">peraltus</emphasis>
(
<bibRefCitation id="EFB74B94FFB8FFADFB9B596294EA7D26" author="Berlese" box="[1141,1275,616,637]" pageId="8" pageNumber="309" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB8FFADFAE1596295847D26" box="[1295,1429,616,637]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 127–129</figureCitation>
)
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31. Endogynium with thorns gathered in the two lateral (left and right) groups (
<figureCitation id="131D2AE0FFB8FFADFC5B598F94857DC1" box="[949,1172,645,666]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 133,137, 141, 146</figureCitation>
)................... 32
</keyLead>
</paragraph>
<paragraph id="8B993665FFB8FFADFF7959A9958A7D8E" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
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- Endogynium with the two large thorns located posteriorly, and other thorns or denticles; the latter, if numerous, not arranged in lateral groups but distributed more regularly around the endogynial sac entrance (e.g.
<figureCitation id="131D2AE0FFB8FFADFBF959CA948D7D8E" box="[1047,1180,704,725]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 154, 164</figureCitation>
,
<figureCitation id="131D2AE0FFB8FFADFB4859CA94ED7D8E" box="[1190,1276,704,725]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">167, 187</figureCitation>
,
<figureCitation id="131D2AE0FFB8FFADFAE859CA954A7D8E" box="[1286,1371,704,725]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="8" pageNumber="309">195, 197</figureCitation>
).. 35
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32. Posterior endogynial margin regular, without two lamellae protruding anteriorly (
<figureCitation id="131D2AE0FFB8FFADFC0159D794667DA9" box="[1007,1143,733,754]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 133, 137</figureCitation>
); posterior paragynial lobes in close proximity, or moderately distant of each other......................................................... 33
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<keyLead id="30D79550FFB8FFADFF79581D958D7C3D" pageId="8" pageNumber="309">
- Posterior endogynial margin forms the two broad, partially overlapped elliptical or leaf-shaped lamellae covering most of the endogynial entrance (
<figureCitation id="131D2AE0FFB8FFADFE40583E92247C12" box="[430,565,820,841]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 141, 144</figureCitation>
); posterior paragynial lobes (and the posterior subapical thickenings of the epigynium) distant (
<figureCitation id="131D2AE0FFB8FFADFEFC585B91D67C3D" box="[274,455,849,870]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 138, 139, 143</figureCitation>
).................................................. (
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)...34
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33. Endogynial thorns directed anteriorly; posterior paragynial lobes in close proximity; distal margin of Tr IV with lamellar process ending in one or two prongs (
<figureCitation id="131D2AE0FFB8FFADFDC7588192A07CFB" box="[553,689,907,928]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 130, 131</figureCitation>
); endogynium with a pair of double lateral teeth located posteriorly and a pair of smaller teeth located more anteriorly, spherular organ occupies the posterior half of endogynium, whereas the endogynial sac is in the anterior half (
<figureCitation id="131D2AE0FFB8FFADFE3458CF923D7C81" box="[474,556,965,986]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Fig. 133</figureCitation>
)........................
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<emphasis id="B952EA77FFB8FFADFC9458CF93DF7C81" box="[890,974,965,986]" italics="true" pageId="8" pageNumber="309">cornutus</emphasis>
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(
<figureCitation id="131D2AE0FFB8FFADFAE158CF95857C81" box="[1295,1428,965,986]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 130–133</figureCitation>
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<paragraph id="8B993665FFB8FFADFF7958E8958D7B4F" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
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- Endogynial thorns directed posteriorly (
<figureCitation id="131D2AE0FFB8FFADFDB058E892A07CAC" box="[606,689,994,1015]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Fig. 137</figureCitation>
); posterior paragynial lobes more distant; lamellar process at the distal margin of Tr IV absent.................................................
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.
<emphasis id="B952EA77FFB8FFADFC3258F5945C7B4E" box="[988,1101,1023,1045]" italics="true" pageId="8" pageNumber="309">intermedius</emphasis>
(
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(
<figureCitation id="131D2AE0FFB8FFADFAE158F595847B4F" box="[1295,1429,1023,1044]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 134–137</figureCitation>
)
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34. Endogynium roundish in its outline (
<figureCitation id="131D2AE0FFB8FFADFDAF5F1692827B6A" box="[577,659,1052,1073]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Fig. 141</figureCitation>
)......................
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<emphasis id="B952EA77FFB8FFADFC4A5F1793A47B69" box="[932,949,1053,1074]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC2F5F1694467B69" box="[961,1111,1052,1074]" italics="true" pageId="8" pageNumber="309">hemisphaericus</emphasis>
(
<bibRefCitation id="EFB74B94FFB8FFADFB8A5F1694EA7B6A" author="Vitzthum" box="[1124,1275,1052,1073]" pageId="8" pageNumber="309" refString="Vitzthum, H. v. (1923) Acarologische Beobachtungen. 7. Reihe. Archiv fur Naturgeschichte, Berlin, 89 A, 97 - 181." type="journal article" year="1923">Vitzthum, 1923</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB8FFADFAE15F1695857B6A" box="[1295,1428,1052,1073]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 138–142</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795F30958D7B14" blockId="8.[151,1436,151,2035]" box="[151,1436,1081,1103]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795F30958D7B14" box="[151,1436,1081,1103]" pageId="8" pageNumber="309">
- Endogynium rhombic (
<figureCitation id="131D2AE0FFB8FFADFE515F3092547B14" box="[447,581,1082,1103]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 144, 146</figureCitation>
)............................
<taxonomicName id="4C264DE6FFB8FFADFC445F3095127B14" authority="Witalinski, 2006" authorityName="Witalinski" authorityYear="2006" box="[938,1283,1081,1103]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="rhombogynialis">
<emphasis id="B952EA77FFB8FFADFC445F3093AA7B14" box="[938,955,1082,1103]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC265F33944F7B14" box="[968,1118,1081,1103]" italics="true" pageId="8" pageNumber="309">rhombogynialis</emphasis>
Witaliński, 2006
</taxonomicName>
(
<figureCitation id="131D2AE0FFB8FFADFAE15F3095847B14" box="[1295,1429,1082,1103]" captionStart="FIGURES 120 – 146" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[231,1306,193,1748]" captionTargetId="figure@21.[231,1308,193,1748]" captionTargetPageId="21" captionText="FIGURES 120 – 146. Females. Holoparasitus inventus: 120 idiosoma ventral side; 121 endogynium. Holoparasitus nonaltus: 122 endogynium under paragynia and epigynial central prong; 123 epigynium, anterior part; 124 paragynia; 125, 126 endogynium. Holoparasitus peraltus: 127 epigynium, anterior part; 128 paragynia; 129 endogynium. Holoparasitus cornutus: 130 trochanter IV with lamellar process ending in two prongs; 131 ibid., distal end of segment; 132 epigynium; 133 endogynium. Holoparasitus intermedius: 134, 135 gnathotectum; 136 epigynium; 137 paragynia and endogynium. Holoparasitus hemisphaericus: 138 endogynium under paragynia and epigynium; 139, 140 epigynium; 141 endogynium; 142 endogynial lamella. Holoparasitus rhombogynialis: 143 epigynium; 144 endogynium with endogynial lamella outlined; 145 endogynial lamella; 146 endogynium, another aspect. Scale bars: 130 – 132, 139 – 146 0.1 mm, 133 0.05 mm." httpUri="https://zenodo.org/record/810621/files/figure.png" pageId="8" pageNumber="309">Figs 143–146</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795F5D958A7BFD" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795F5D958A7B37" blockId="8.[151,1436,151,2035]" box="[151,1435,1110,1132]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795F5D958A7B37" box="[151,1435,1110,1132]" pageId="8" pageNumber="309">
35. Postcoxal
<emphasis id="B952EA77FFB8FFADFEAA5F5C91777B37" box="[324,358,1110,1132]" italics="true" pageId="8" pageNumber="309">gv2</emphasis>
glands empty into the large foveolae formed by a thickened cuticle (
<figureCitation id="131D2AE0FFB8FFADFC175F5D94CF7B37" box="[1017,1246,1111,1132]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 147, 151, 152, 156</figureCitation>
)............ 35
</keyLead>
</paragraph>
<paragraph id="8B993665FFB8FFADFF795F7E958A7BFD" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795F7E958A7BFD" pageId="8" pageNumber="309">
- Evident foveolae absent (
<figureCitation id="131D2AE0FFB8FFADFE3C5F7E92327BD2" box="[466,547,1140,1161]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Fig. 162</figureCitation>
) (but a cuticle surrounding
<emphasis id="B952EA77FFB8FFADFCC65F79935A7BD2" box="[808,843,1139,1161]" italics="true" pageId="8" pageNumber="309">gv2</emphasis>
pores in
<taxonomicName id="4C264DE6FFB8FFADFC4F5F7E94747BD2" authorityName="Berlese" authorityYear="1906" box="[929,1125,1139,1161]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="pseudoperforatus">
<emphasis id="B952EA77FFB8FFADFC4F5F7E93A37BD2" box="[929,946,1140,1161]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC535F7994747BD2" box="[957,1125,1139,1161]" italics="true" pageId="8" pageNumber="309">pseudoperforatus</emphasis>
</taxonomicName>
may occasionally be thickened)................................................................................................... 38
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795FA4958D7A6C" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795FA495877BA6" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795FA495877BA6" pageId="8" pageNumber="309">
36. Hyaline endogynial lamella with a thickened, sinuous anterior edge (
<figureCitation id="131D2AE0FFB8FFADFC945FA494127B98" box="[890,1027,1198,1219]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 149, 150</figureCitation>
); epigynial subapical structure and central prong relatively large; roundish endogynial sac visible as a distinct structure smaller than the endogynium (
<figureCitation id="131D2AE0FFB8FFADFB0B5FC195267BBB" box="[1253,1335,1227,1248]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Fig. 149</figureCitation>
).........................................................................
<taxonomicName id="4C264DE6FFB8FFADFC125FE294ED7BA6" authority="Witalinski, 1994" authorityName="Witalinski" authorityYear="1994" box="[1020,1276,1255,1277]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="dallaii">
<emphasis id="B952EA77FFB8FFADFC125FE2941C7BA6" box="[1020,1037,1256,1277]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFBF75FED94497BA6" box="[1049,1112,1255,1277]" italics="true" pageId="8" pageNumber="309">dallaii</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFBB05FE294ED7BA6" author="Witalinski" box="[1118,1276,1256,1277]" pageId="8" pageNumber="309" refString="Witalinski, W. (1994) Two new Holoparasitus species from Italy and Poland (Acari: Gamasida: Pergamasidae). Genus, 5, 215 - 222." type="journal article" year="1994">Witaliński, 1994</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB8FFADFAE75FE2959E7BA6" box="[1289,1423,1256,1277]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 147–150</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB8FFADFF795E0F958D7A6C" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795E0F958D7A6C" pageId="8" pageNumber="309">
- The edge of endogynial lamella not thickened, frequently poorly visible (
<figureCitation id="131D2AE0FFB8FFADFC775E0F945E7A41" box="[921,1103,1285,1306]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 154, 155, 160</figureCitation>
); epigynial subapical structure and central prong of normal size; endogynial sac as large as the endogynium, not discernible as a smaller distinct structure.... 37
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795E35958D7AD5" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795E35958D7A2A" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795E35958D7A2A" pageId="8" pageNumber="309">
37. Idiosoma strikingly spherical; anterior epigynial margins on the sides of the central prong deeply concaved (
<figureCitation id="131D2AE0FFB8FFADFB005E3595517A0F" box="[1262,1344,1343,1364]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Fig. 153</figureCitation>
)........................................................................
<taxonomicName id="4C264DE6FFB8FFADFC055E5695127A2A" authority="Witalinski, 1994" authorityName="Witalinski" authorityYear="1994" box="[1003,1283,1372,1394]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B952EA77FFB8FFADFC055E5693ED7A2A" box="[1003,1020,1372,1393]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFBE65E56944C7A29" box="[1032,1117,1372,1394]" italics="true" pageId="8" pageNumber="309">globosus</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFB8A5E5695127A2A" author="Witalinski" box="[1124,1283,1372,1393]" pageId="8" pageNumber="309" refString="Witalinski, W. (1994) Two new Holoparasitus species from Italy and Poland (Acari: Gamasida: Pergamasidae). Genus, 5, 215 - 222." type="journal article" year="1994">Witaliński, 1994</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB8FFADFAE15E5695857A2A" box="[1295,1428,1372,1393]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 151–155</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB8FFADFF795E73958D7AD5" blockId="8.[151,1436,151,2035]" box="[151,1436,1401,1423]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795E73958D7AD5" box="[151,1436,1401,1423]" pageId="8" pageNumber="309">
- Idiosoma elongated; anterior epigynial margins straight (
<figureCitation id="131D2AE0FFB8FFADFD105E7393417AD5" box="[766,848,1401,1422]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Fig. 157</figureCitation>
) …
<taxonomicName id="4C264DE6FFB8FFADFC945E7095157AD5" authority="Witalinski &amp; Skorupski, 2007" authorityName="Witalinski &amp; Skorupski" authorityYear="2007" box="[890,1284,1401,1423]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="fovealis">
<emphasis id="B952EA77FFB8FFADFC945E70939A7AD4" box="[890,907,1402,1423]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC785E7393F07AD4" box="[918,993,1401,1423]" italics="true" pageId="8" pageNumber="309">fovealis</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFC085E7395157AD5" author="Witalinski" box="[998,1284,1401,1422]" pageId="8" pageNumber="309" refString="Witalinski, W. &amp; Skorupski, M. (2007) Two new species of Holoparasitus Oudemans, 1936 from Italy - Tuscany and Trentino (Acari: Gamasida: Parasitidae). Genus, 18, 125 - 138." type="journal article" year="2007">Witaliński &amp; Skorupski, 2007</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB8FFADFAFE5E7395847AD5" box="[1296,1429,1401,1422]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 156–160</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795E9C958D7922" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795E9C958D7979" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795E9C958D7979" pageId="8" pageNumber="309">
38. Endogynium relatively large, circular (
<figureCitation id="131D2AE0FFB8FFADFDBB5E9C92CB7AF0" box="[597,730,1430,1451]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Fig. 164, 165</figureCitation>
); margins of endogynium and endogynial sac distinctly visible as the two concentric circles; both posterior main thorns protrude more or less centripetally at ca. 120o from sac margin; anterior main thorn absent; two to four additional denticles can be distributed bilaterally at the sac entrance, occasionally one or two small denticles, located randomly and much deeper in the sac, are encountered; epigynial central prong weakly sclerotised and less pigmented apically (
<figureCitation id="131D2AE0FFB8FFADFE4A5D07923B7979" box="[420,554,1549,1570]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 162, 163</figureCitation>
)...........................
<taxonomicName id="4C264DE6FFB8FFADFC9A5D0795127979" authority="Witalinski &amp; Skorupski, 2007" authorityName="Witalinski &amp; Skorupski" authorityYear="2007" box="[884,1283,1549,1570]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="amiatus">
<emphasis id="B952EA77FFB8FFADFC9A5D0793947979" box="[884,901,1549,1570]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFC7C5D0793CF7979" box="[914,990,1549,1570]" italics="true" pageId="8" pageNumber="309">amiatus</emphasis>
<bibRefCitation id="EFB74B94FFB8FFADFC0A5D0795127979" author="Witalinski" box="[996,1283,1549,1570]" pageId="8" pageNumber="309" refString="Witalinski, W. &amp; Skorupski, M. (2007) Two new species of Holoparasitus Oudemans, 1936 from Italy - Tuscany and Trentino (Acari: Gamasida: Parasitidae). Genus, 18, 125 - 138." type="journal article" year="2007">Witaliński &amp; Skorupski, 2007</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB8FFADFAE15D0795857979" box="[1295,1428,1549,1570]" captionStart="FIGURES 147 – 165" captionStartId="22.[151,264,1850,1873]" captionTargetBox="[163,1418,197,1826]" captionTargetId="figure@22.[151,1436,163,1844]" captionTargetPageId="22" captionText="FIGURES 147 – 165. Females. Holoparasitus dallaii: 147 gv 2 pore in thicken cuticular foveola; 148 epigynium; 149, 150 endogynium. Holoparasitus globosus: 151 gv 2 pore in foveola behind coxa IV opening; 152 region of genital opening with endogynium under paragynia and epigynium; 153 epigynium; 154 endogynium; 155 endogynial lamella. Holoparasitus fovealis: 156 region of genital opening with endogynium under paragynia and epigynium; 157 epigynium; 158 – 160 endogynium. Holoparasitus amiatus: 161 gnathotectum; 162 epigynium; 163 epigynium central prong; 164, 165 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810623/files/figure.png" pageId="8" pageNumber="309">Figs 161–165</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795D20958D7922" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
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- Endogynium at least with the two posterior and one anterior main thorns; the anterior main thorn or dent usually smaller, located axially and directed backward; additional denticles usually present and distributed regularly on both sides, between posterior and anterior main thorns (e.g.
<figureCitation id="131D2AE0FFB8FFADFDB35D6E93027922" box="[605,787,1636,1657]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 167, 173, 186</figureCitation>
,
<figureCitation id="131D2AE0FFB8FFADFCF25D6E93607922" box="[796,881,1636,1657]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="8" pageNumber="309">195, 197</figureCitation>
)................... (
<emphasis id="B952EA77FFB8FFADFB9C5D6E95787921" bold="true" box="[1138,1385,1636,1658]" pageId="8" pageNumber="309">
<taxonomicName id="4C264DE6FFB8FFADFB9C5D6E94C97921" authorityName="Koch" authorityYear="Koch" box="[1138,1240,1636,1658]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="B952EA77FFB8FFADFB9C5D6E94C97921" bold="true" box="[1138,1240,1636,1658]" italics="true" pageId="8" pageNumber="309">calcaratus</emphasis>
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species-group
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)...39
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<keyStep id="30D22EC0FFB8FFADFF795D8B958A798A" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795D8B958D79E8" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
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39. Endogynium with the three main thorns, endogynial sac visible as an elliptical, distinct structure with many long centripetal thorns protruding from the wall (
<figureCitation id="131D2AE0FFB8FFADFDF25D9492B079E8" box="[540,673,1694,1715]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 167–169</figureCitation>
)..........................
<taxonomicName id="4C264DE6FFB8FFADFC0B5D95951279E8" authority="Willmann, 1940" authorityName="Willmann" authorityYear="1940" box="[997,1283,1694,1716]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="rotulifer">
<emphasis id="B952EA77FFB8FFADFC0B5D9593E779EF" box="[997,1014,1695,1716]" italics="true" pageId="8" pageNumber="309">H</emphasis>
.
<emphasis id="B952EA77FFB8FFADFBED5D94944379EF" box="[1027,1106,1694,1716]" italics="true" pageId="8" pageNumber="309">rotulifer</emphasis>
(
<bibRefCitation id="EFB74B94FFB8FFADFBB05D9494EA79E8" author="Willmann" box="[1118,1275,1694,1715]" pageId="8" pageNumber="309" refString="Willmann, C. (1940) Neue Milben aus Hohlen der Balkanhalbinsel, gesammelt von Prof. Dr. K. Absolon, Brunn (2. Mitteilung). Zoologischer Anzeiger, 130, 209 - 218." type="journal article" year="1940">Willmann, 1940</bibRefCitation>
)
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(
<figureCitation id="131D2AE0FFB8FFADFAE15D94958479E8" box="[1295,1429,1694,1715]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 166–169</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB8FFADFF795DB6958A798A" blockId="8.[151,1436,151,2035]" box="[151,1435,1724,1745]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795DB6958A798A" box="[151,1435,1724,1745]" pageId="8" pageNumber="309">- The wall of the endogynial sac without any thorns or dents.................................................... 40</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB8FFADFF795DD2958A7873" pageId="8" pageNumber="309">
<paragraph id="8B993665FFB8FFADFF795DD2958A7851" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795DD2958A7851" pageId="8" pageNumber="309">
40. Endogynium with distinct spherular organ protruding from the posterior surface of the endogynium into the endogynial lumen (e.g.
<figureCitation id="131D2AE0FFB8FFADFEFF5DFF91E77851" box="[273,502,1781,1802]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 173–175, 184–187</figureCitation>
)........................................................................... 41
</keyLead>
</paragraph>
<paragraph id="8B993665FFB8FFADFF795C19958A7873" blockId="8.[151,1436,151,2035]" box="[151,1435,1811,1832]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795C19958A7873" box="[151,1435,1811,1832]" pageId="8" pageNumber="309">
- Distinct spherular organ absent (e.g.
<figureCitation id="131D2AE0FFB8FFADFDAC5C1993387873" box="[578,809,1811,1832]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="8" pageNumber="309">Figs 189, 193, 195, 197</figureCitation>
).................................................. 42
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B993665FFB8FFADFF795C3A958D78C7" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyStep id="30D22EC0FFB8FFADFF795C3A958D78C7" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795C3A958D78C7" pageId="8" pageNumber="309">
41. Anterior part of endogynium perimeter not covered from its ventral side with any lamella protruding from the endogynial margin; spherular organ triangular or subtriangular, frequently more or less pointed anteriad and protruding into the endogynial lumen (
<figureCitation id="131D2AE0FFB8FFADFEC75C6091A37824" box="[297,434,1898,1919]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 173–175</figureCitation>
); the apex of epigynial plate blunt, tetrangular or rounded (
<figureCitation id="131D2AE0FFB8FFADFC095C60947C7824" box="[999,1133,1898,1919]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 170–172</figureCitation>
)...................................................................................
<taxonomicName id="4C264DE6FFB8FFADFC4C5C8D951278C7" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[930,1283,1926,1948]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="pseudoperforatus">
<emphasis id="B952EA77FFB8FFADFC4C5C8D93A278C7" box="[930,947,1927,1948]" italics="true" pageId="8" pageNumber="309">H</emphasis>
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<emphasis id="B952EA77FFB8FFADFC2F5C8C947878C7" box="[961,1129,1926,1948]" italics="true" pageId="8" pageNumber="309">pseudoperforatus</emphasis>
(
<bibRefCitation id="EFB74B94FFB8FFADFB9B5C8D94ED78C7" author="Berlese" box="[1141,1276,1927,1948]" pageId="8" pageNumber="309" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB8FFADFAE15C8D958578C7" box="[1295,1428,1927,1948]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 170–175</figureCitation>
)
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</keyStep>
</paragraph>
<paragraph id="8B993665FFB8FFADFF795CAE958D78A8" blockId="8.[151,1436,151,2035]" pageId="8" pageNumber="309">
<keyLead id="30D79550FFB8FFADFF795CAE958D78A8" pageId="8" pageNumber="309">
- The anterior margin of the endogynium forms a thin bilobate lamella partly covering the endogynium entrance (
<figureCitation id="131D2AE0FFB8FFADFAE35CAE954E78E2" box="[1293,1375,1956,1977]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Fig. 183</figureCitation>
), with the lobes separated by a distinct incision (usually with an additional triangular protrusion); spherular organ noticeable, its anterior margin arcuate in its central part and straight laterally (
<figureCitation id="131D2AE0FFB8FFADFCF65CD493B078A8" box="[792,929,2014,2035]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="8" pageNumber="309">Figs 184–187</figureCitation>
); the central prong of epigynium present, but located
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</paragraph>
<paragraph id="8B993665FFB9FFACFF0F5B9D958D7E00" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
ventrally over the anterior, triangular part of the subapical epigynial thickening, thus poorly discernible (
<figureCitation id="131D2AE0FFB9FFACFB0C5B9D957A7FF7" box="[1250,1387,151,172]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="9" pageNumber="310">Figs 179–182</figureCitation>
); the lamellar extensions of the subapical epigynial structure projecting laterally beyond the epigynial apex, occassionally also anteriorly (
<figureCitation id="131D2AE0FFB9FFACFECD5BD891B97FBC" box="[291,424,210,231]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="9" pageNumber="310">Figs 179, 182</figureCitation>
).............................................
<taxonomicName id="4C264DE6FFB9FFACFC0E5BD895127FBC" authority="Berlese, 1906" authorityName="Berlese" authorityYear="Berlese" box="[992,1283,209,231]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="excipuliger">
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<emphasis id="B952EA77FFB9FFACFC135BDB94787FBC" box="[1021,1129,209,231]" italics="true" pageId="9" pageNumber="310">excipuliger</emphasis>
(
<bibRefCitation id="EFB74B94FFB9FFACFB9B5BD894EA7FBC" author="Berlese" box="[1141,1275,210,231]" pageId="9" pageNumber="310" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB9FFACFAE15BD895847FBC" box="[1295,1429,210,231]" captionStart="FIGURES 166 – 187" captionStartId="23.[151,264,1790,1813]" captionTargetBox="[159,1429,202,1740]" captionTargetId="figure@23.[151,1436,171,1789]" captionTargetPageId="23" captionText="FIGURES 166 – 187. Females. Holoparasitus rotulifer: 166 epigynium, central and lateral prongs; 167 – 169 endogynium. Holoparasitus pseudoperforatus: 170 epigynium; 171 paragynium; 172 epigynium subapical structure; 173, 174 endogynium, ventrally; 175 endogynium in posteroventral perspective. Holoparasitus excipuliger: 176 – 178 gnathotectum; 179 epigynium, anterior part; 180 – 182 epigynium central prong with subapical structure; 183 – 187 endogynium, different aspects. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810625/files/figure.png" pageId="9" pageNumber="310">Figs 176–187</figureCitation>
)
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42. Endogynium with the three main thorns, large and of a similar size; additional 2–3 dents are located on each side between the anterior and posterior main thorns (
<figureCitation id="131D2AE0FFB9FFACFDD55A0692D77E7A" box="[571,710,268,289]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 189, 190</figureCitation>
); central epigynial prong long and narrow, lateral protrusions of subapical epigynial structure wing-like and elongated, but practically not extending beyond the epigynial margin (
<figureCitation id="131D2AE0FFB9FFACFB285A2395087E65" box="[1222,1305,297,318]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 188</figureCitation>
)..............................................................
<taxonomicName id="4C264DE6FFB9FFACFCB95A4C95127E00" authority="Witalinski &amp; Skorupski, 2002" authorityName="Witalinski &amp; Skorupski" authorityYear="2002" box="[855,1283,325,347]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="kerkirensis">
<emphasis id="B952EA77FFB9FFACFCB95A4C93797E00" box="[855,872,326,347]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFC9B5A4F93CC7E00" box="[885,989,325,347]" italics="true" pageId="9" pageNumber="310">kerkirensis</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFC0A5A4C95127E00" author="Witalinski" box="[996,1283,326,347]" pageId="9" pageNumber="310" refString="Witalinski, W. &amp; Skorupski, M. (2002) Genus Holoparasitus Oudemans, 1936 in Berlese Acaroteca (Acari: Gamasida: Parasitidae). Part I. Redia, 85, 37 - 60." type="journal article" year="2002">Witaliński &amp; Skorupski, 2002</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB9FFACFAE15A4C95847E00" box="[1295,1429,326,347]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 188–191</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB9FFACFF795A69958A7ECE" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795A69958A7ECE" pageId="9" pageNumber="310">- Endogynium with the posterior main thorns evidently larger than the anterior one; additional lateral dents (1–2 on each side) may also be encountered............................................................................... 43</keyLead>
</paragraph>
<keyStep id="30D22EC0FFB9FFACFF795A97958A7D3B" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF795A97958D7D52" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
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43. Epigynium (
<figureCitation id="131D2AE0FFB9FFACFEB45A9791BF7EE9" box="[346,430,413,434]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 192</figureCitation>
) with the central prong short; lateral hyaline protrusions of the subapical epigynial structure wing-like and elongated, rounded terminally and extending far beyond the epigynium margin; the thickening of the subapical epigynial structure forms the four sides of a regular pentagon, with the anterior sides straight (
<figureCitation id="131D2AE0FFB9FFACFBE25ADD94717EB7" box="[1036,1120,471,492]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 192</figureCitation>
); endogynium irregularly roundish (
<figureCitation id="131D2AE0FFB9FFACFEE45AFE914D7D52" box="[266,348,500,521]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 193</figureCitation>
)....................................................
<taxonomicName id="4C264DE6FFB9FFACFC375AFE95127D52" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[985,1283,500,522]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="pollicipatus">
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<emphasis id="B952EA77FFB9FFACFC165AFE94787D51" box="[1016,1129,500,522]" italics="true" pageId="9" pageNumber="310">pollicipatus</emphasis>
(
<bibRefCitation id="EFB74B94FFB9FFACFB9B5AFE94EA7D52" author="Berlese" box="[1141,1275,500,521]" pageId="9" pageNumber="310" refString="Berlese, A. (1903) Acari nuovi. Manipulus Ius. Redia, 1, 235 - 252." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB9FFACFAE15AFE95847D52" box="[1295,1429,500,521]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 192, 193</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB9FFACFF79591B958A7D3B" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
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- The thickening of the subapical epigynial structure with the anterior parts evidently wavy (
<figureCitation id="131D2AE0FFB9FFACFBAB591B94DD7D7D" box="[1093,1228,529,550]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 194, 196</figureCitation>
), lamellar lateral protrusions only moderately extending beyond the epigynium margin, three-sided, with the posterior sides concave (
<figureCitation id="131D2AE0FFB9FFACFAD1592491197D3B" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 194, 196</figureCitation>
); endogynium regularly roundish..................................................................... 44
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<paragraph id="8B993665FFB9FFACFF795962958D7DC1" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
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44. The posterior main thorns of endogynium curved centripetally and settled on a joint, broad base (
<figureCitation id="131D2AE0FFB9FFACFB6A596294C97D26" box="[1156,1240,616,637]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 195</figureCitation>
)..................................................................................
<taxonomicName id="4C264DE6FFB9FFACFC14598C95127DC1" authority="Koch, 1839" authorityName="Koch" authorityYear="Koch" box="[1018,1283,645,667]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="calcaratus">
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<emphasis id="B952EA77FFB9FFACFBF9598F946D7DC0" box="[1047,1148,645,667]" italics="true" pageId="9" pageNumber="310">calcaratus</emphasis>
(
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)
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(
<figureCitation id="131D2AE0FFB9FFACFAE1598F95847DC1" box="[1295,1429,645,666]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 194, 195</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB9FFACFF7959A9958D7D8E" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF7959A9958D7D8E" pageId="9" pageNumber="310">
- The posterior main thorns of the endogynium straight and settled independently; endogynial sac visible as a distinct smaller circle (
<figureCitation id="131D2AE0FFB9FFACFECD59CA91647D8E" box="[291,373,704,725]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 197</figureCitation>
).....................................
<taxonomicName id="4C264DE6FFB9FFACFCD659CA95127D8E" authority="Witalinski &amp; Skorupski, 2003b" authorityName="Witalinski &amp; Skorupski" authorityYear="2003" box="[824,1283,703,725]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="paradisiacus">
<emphasis id="B952EA77FFB9FFACFCD659CA93587D8E" box="[824,841,704,725]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFCB859B593C37D8E" box="[854,978,703,725]" italics="true" pageId="9" pageNumber="310">paradisiacus</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFC3659CA95127D8E" author="Witalinski" box="[984,1283,704,725]" pageId="9" pageNumber="310" refString="Witalinski, W. &amp; Skorupski, M. (2003 b) A new species of Holoparasitus Oudemans, 1936 from North Italy (Acari: Gamasida: Parasitidae). Genus, 14, 431 - 438." type="journal article" year="2003" yearSuffix="b">Witaliński &amp; Skorupski, 2003b</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB9FFACFAE159CA95847D8E" box="[1295,1429,704,725]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 196, 197</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB9FFACFF7959D7958A7C12" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF7959D7958A7C77" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF7959D7958A7C77" pageId="9" pageNumber="310">
45. The anterior margin of the sternum between
<emphasis id="B952EA77FFB9FFACFD6259D692B97DA9" box="[652,680,732,754]" italics="true" pageId="9" pageNumber="310">st1</emphasis>
setae better sclerotised, partly overlaps a presternal plate (
<figureCitation id="131D2AE0FFB9FFACFB3659D795717DA9" box="[1240,1376,733,754]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 198, 202</figureCitation>
); endogynium visible as an arcuate sclerotised band with an axially located opening (
<figureCitation id="131D2AE0FFB9FFACFC3059F094D57C54" box="[990,1220,762,783]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 200, 201, 204, 205</figureCitation>
).............................................................................................. (
<emphasis id="B952EA77FFB9FFACFB69581C95787C77" bold="true" box="[1159,1385,790,812]" pageId="9" pageNumber="310">
<taxonomicName id="4C264DE6FFB9FFACFB69581C94C97C77" authorityName="Juvara-Bals &amp; Witalinski" authorityYear="2006" box="[1159,1240,790,812]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="annulus">
<emphasis id="B952EA77FFB9FFACFB69581C94C97C77" bold="true" box="[1159,1240,790,812]" italics="true" pageId="9" pageNumber="310">annulus</emphasis>
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species-group
</emphasis>
)...46
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</paragraph>
<paragraph id="8B993665FFB9FFACFF79583E958A7C12" blockId="9.[151,1436,151,1452]" box="[151,1435,820,841]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF79583E958A7C12" box="[151,1435,820,841]" pageId="9" pageNumber="310">- Sternum anterior margin developed normally; endogynium different from the above................................ 47</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB9FFACFF79585B958D7CAC" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF79585B958D7CFB" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF79585B958D7CFB" pageId="9" pageNumber="310">
46. Gland pores
<emphasis id="B952EA77FFB9FFACFEB3585A91917C3D" box="[349,384,848,870]" italics="true" pageId="9" pageNumber="310">gv1</emphasis>
present; opening in the cuticular band of endogynium with a subcircular anterior margin, a dentate lamellar structure (endogynial lamella?) located laterally to the opening not discernible............................................................................................
<taxonomicName id="4C264DE6FFB9FFACFC8E588695137CFB" authority="Juvara-Bals &amp; Witalinski, 2006" authorityName="Juvara-Bals &amp; Witalinski" authorityYear="2006" box="[864,1282,907,929]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="annulus">
<emphasis id="B952EA77FFB9FFACFC8E588693647CFA" box="[864,885,908,929]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFC6C588193DF7CFA" box="[898,974,907,929]" italics="true" pageId="9" pageNumber="310">annulus</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFC3A588195137CFB" author="Juvara-Bals" box="[980,1282,907,928]" pageId="9" pageNumber="310" refString="Juvara-Bals, I. &amp; Witalinski, W. (2006) Two new species of the genus Holoparasitus Oudemans from the Mediterranean Basin - Algeria and Sardinia (Acari: Gamasida: Parasitidae). Genus, 17, 437 - 448." type="journal article" year="2006">Juvara-Bals &amp; Witaliński, 2006</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB9FFACFAE1588195857CFB" box="[1295,1428,907,928]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 198–201</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB9FFACFF7958A2958D7CAC" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF7958A2958D7CAC" pageId="9" pageNumber="310">
- Gland pores
<emphasis id="B952EA77FFB9FFACFEB558A2916F7CE5" box="[347,382,936,958]" italics="true" pageId="9" pageNumber="310">gv1</emphasis>
absent; opening in the cuticular band of endogynium usually with an undulate anterior margin, dentate lamellar structure (endogynial lamella?) can be discernible laterally to the opening...........................................................................................
<taxonomicName id="4C264DE6FFB9FFACFCB858E995137CAC" authority="Juvara-Bals &amp; Witalinski, 2006" authorityName="Juvara-Bals &amp; Witalinski" authorityYear="2006" box="[854,1282,994,1016]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="sardensis">
<emphasis id="B952EA77FFB9FFACFCB858E993767CA3" box="[854,871,995,1016]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFC9A58E893DF7CA3" box="[884,974,994,1016]" italics="true" pageId="9" pageNumber="310">sardensis</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFC3A58E895137CAC" author="Juvara-Bals" box="[980,1282,994,1015]" pageId="9" pageNumber="310" refString="Juvara-Bals, I. &amp; Witalinski, W. (2006) Two new species of the genus Holoparasitus Oudemans from the Mediterranean Basin - Algeria and Sardinia (Acari: Gamasida: Parasitidae). Genus, 17, 437 - 448." type="journal article" year="2006">Juvara-Bals &amp; Witaliński, 2006</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB9FFACFAE158E895847CAC" box="[1295,1429,994,1015]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 202–205</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB9FFACFF7958F5958A7B37" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF7958F5958D7B6A" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF7958F5958D7B6A" pageId="9" pageNumber="310">
47. Endogynium large, irregularly semicircular, with a split lamellar protrusion, anteriorly pointed and dentate (
<figureCitation id="131D2AE0FFB9FFACFB1758F5955E7B4F" box="[1273,1359,1023,1044]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 208</figureCitation>
); posterior paragynial lobes distant of each other (
<figureCitation id="131D2AE0FFB9FFACFD935F1692DE7B6A" box="[637,719,1052,1073]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Fig. 206</figureCitation>
).........................
<taxonomicName id="4C264DE6FFB9FFACFBE85F1795127B6A" authority="Berlese, 1906" authorityName="Berlese" authorityYear="1906" box="[1030,1283,1052,1074]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="excisus">
<emphasis id="B952EA77FFB9FFACFBE85F1794067B69" box="[1030,1047,1053,1074]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFBCA5F1794787B69" box="[1060,1129,1053,1074]" italics="true" pageId="9" pageNumber="310">excisus</emphasis>
(
<bibRefCitation id="EFB74B94FFB9FFACFB9B5F1694ED7B6A" author="Berlese" box="[1141,1276,1052,1073]" pageId="9" pageNumber="310" refString="Berlese, A. (1906) Monografia del genere Gamasus Latr. Redia, 3, 66 - 304." type="journal article" year="1906">Berlese, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131D2AE0FFB9FFACFAE15F1695847B6A" box="[1295,1429,1052,1073]" captionStart="FIGURES 188 – 208" captionStartId="24.[151,264,1807,1830]" captionTargetBox="[238,1346,194,1761]" captionTargetId="figure@24.[207,1375,155,1803]" captionTargetPageId="24" captionText="FIGURES 188 – 208. Females. Holoparasitus kerkirensis: 188 epigynium, central and lateral prongs; 189, 190 endogynium; 191 gnathotectum. Holoparasitus pollicipatus: 192 epigynium, central and lateral prongs; 193 endogynium. Holoparasitus calcaratus: 194 epigynium, central and lateral prongs; 195 endogynium. Holoparasitus paradisiacus: 196 epigynium, central and lateral prongs; 197 endogynium. Holoparasitus annulus: 198 presternal plate, lateral platelets and sternum anterior margin; 199 epigynium, anterior part; 200, 201 endogynium in situ. Holoparasitus sardensis: 202 presternal plate, lateral platelets and sternum anterior margin; 203 epigynium, anterior part; 204, 205 endogynium in situ. Holoparasitus excisus: 206 endogynium under paragynia and epigynium central prong; 207 epigynium, anterior part; 208 endogynium. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810627/files/figure.png" pageId="9" pageNumber="310">Figs 206–208</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB9FFACFF795F30958A7B37" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795F30958A7B37" pageId="9" pageNumber="310">
- Endogynium very small (e.g.
<figureCitation id="131D2AE0FFB9FFACFE105F3092F57B14" box="[510,740,1082,1103]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 209, 215, 219, 223</figureCitation>
); posterior paragynial lobes in close proximity of each other or distant.................................................................................................... 48
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</paragraph>
</keyStep>
<keyStep id="30D22EC0FFB9FFACFF795F7E958D7BBB" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF795F7E958D7BFD" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795F7E958D7BFD" pageId="9" pageNumber="310">
48. Posterior paragynial lobes distant; central epigynial prong large and sharp, the thickening of subapical epigynial structure not discernible (
<figureCitation id="131D2AE0FFB9FFACFEB95F9B91BD7BFD" box="[343,428,1169,1190]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Fig. 210</figureCitation>
); endogynium subcircular with two &quot;orifices&quot; (
<figureCitation id="131D2AE0FFB9FFACFC805F9B93AF7BFD" box="[878,958,1169,1190]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Fig. 211</figureCitation>
)...
<taxonomicName id="4C264DE6FFB9FFACFC1A5F9B95127BFD" authority="Juvara-Bals, 1995" authorityName="Juvara-Bals" authorityYear="1995" box="[1012,1283,1168,1190]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="vasilei">
<emphasis id="B952EA77FFB9FFACFC1A5F9B94147BFD" box="[1012,1029,1169,1190]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFBFC5F9A94417BFD" box="[1042,1104,1168,1190]" italics="true" pageId="9" pageNumber="310">vasilei</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFBB85F9B95127BFD" author="Juvara-Bals" box="[1110,1283,1169,1190]" pageId="9" pageNumber="310" refString="Juvara-Bals, I. (1995) Holoparasitus (Holoparasitus) vasilei n. sp. from Israel (Acari, Gamasida: Parasitidae). In: Nitzu, E. (Ed.), Soil Fauna of Israel. Editura Academiei Romane, Bucuresti, pp. 47 - 52." type="book chapter" year="1995">Juvara-Bals, 1995</bibRefCitation>
</taxonomicName>
(
<figureCitation id="131D2AE0FFB9FFACFAFE5F9B95847BFD" box="[1296,1429,1169,1190]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 209–211</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B993665FFB9FFACFF795FA4958D7BBB" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795FA4958D7BBB" pageId="9" pageNumber="310">
- Posterior paragynial lobes in close proximity of each other (
<figureCitation id="131D2AE0FFB9FFACFCCB5FA493BC7B98" box="[805,941,1198,1219]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 214, 223</figureCitation>
); central epigynial prong wider than the thickening of the subapical epigynial structure, which is narrow and compact, anteriorly tripartite or rounded (e.g.
<figureCitation id="131D2AE0FFB9FFACFB505FC195557BBB" box="[1214,1348,1227,1248]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 213, 218</figureCitation>
)..... 49
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<keyStep id="30D22EC0FFB9FFACFF795FE2958A7A6C" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF795FE2958A7A41" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795FE2958A7A41" pageId="9" pageNumber="310">
49. The thickening of the subapical epigynial structure—anteriorly tripartite (
<figureCitation id="131D2AE0FFB9FFACFC4A5FE2943D7BA6" box="[932,1068,1256,1277]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 212–214</figureCitation>
), lateral hyaline protrusions extending a little beyond the epigynial margin.......................................
<taxonomicName id="4C264DE6FFB9FFACFBE65E0F95127A41" authority="Witalinski, 2016" authorityName="Witalinski" authorityYear="2016" box="[1032,1283,1284,1306]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="fanes">
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.
<emphasis id="B952EA77FFB9FFACFBC55E0E944C7A41" box="[1067,1117,1284,1306]" italics="true" pageId="9" pageNumber="310">fanes</emphasis>
Witaliński, 2016
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(
<figureCitation id="131D2AE0FFB9FFACFAE15E0F95847A41" box="[1295,1429,1285,1306]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 212–216</figureCitation>
)
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</paragraph>
<paragraph id="8B993665FFB9FFACFF795E28958A7A6C" blockId="9.[151,1436,151,1452]" box="[151,1435,1314,1336]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795E28958A7A6C" box="[151,1435,1314,1336]" pageId="9" pageNumber="310">
- The thickening of the subapical epigynial structure—anteriorly rounded (
<figureCitation id="131D2AE0FFB9FFACFC705E2894357A6C" box="[926,1060,1314,1335]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 218, 221</figureCitation>
) … (
<emphasis id="B952EA77FFB9FFACFBB95E2895767A63" bold="true" box="[1111,1383,1314,1336]" pageId="9" pageNumber="310">
<taxonomicName id="4C264DE6FFB9FFACFBB95E2894C67A63" authorityName="Juvara-Bals &amp; Witalinski" authorityYear="2000" box="[1111,1239,1314,1336]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="crassisetosus">
<emphasis id="B952EA77FFB9FFACFBB95E2894C67A63" bold="true" box="[1111,1239,1314,1336]" italics="true" pageId="9" pageNumber="310">crassisetosus</emphasis>
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species-group
</emphasis>
). 50
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<keyStep id="30D22EC0FFB9FFACFF795E35958D7AF0" pageId="9" pageNumber="310">
<paragraph id="8B993665FFB9FFACFF795E35958A7A2A" blockId="9.[151,1436,151,1452]" pageId="9" pageNumber="310">
<keyLead id="30D79550FFB9FFACFF795E35958A7A2A" pageId="9" pageNumber="310">
50. Ti IV with the thickened seta
<emphasis id="B952EA77FFB9FFACFDE15E3492207A0F" box="[527,561,1342,1364]" italics="true" pageId="9" pageNumber="310">pv1</emphasis>
(
<figureCitation id="131D2AE0FFB9FFACFDAC5E35928B7A0F" box="[578,666,1343,1364]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Fig. 217</figureCitation>
); lateral hyaline protrusions of the subapical epigynial structure roundish and extending a little beyond the epigynial margin (
<figureCitation id="131D2AE0FFB9FFACFD495E5692E87A2A" box="[679,761,1372,1393]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Fig. 218</figureCitation>
)...
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<emphasis id="B952EA77FFB9FFACFCDC5E5693527A2A" box="[818,835,1372,1393]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFCA15E5693DC7A2A" box="[847,973,1372,1393]" italics="true" pageId="9" pageNumber="310">crassisetosus</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFC3A5E5695137A2A" author="Juvara-Bals" box="[980,1282,1372,1393]" pageId="9" pageNumber="310" refString="Juvara-Bals, I. &amp; Witalinski, W. (2000) Description of five new species of Holoparasitus s. str. with redescription of H. apenninorum (Berlese, 1906) and H. cultriger (Berlese, 1906) from Italy and Spain (Acari, Gamasida, Parasitidae). Revue Suisse de Zoologie, 107, 3 - 30." type="journal article" year="2000">Juvara-Bals &amp; Witaliński, 2000</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB9FFACFAE15E5695847A2A" box="[1295,1429,1372,1393]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 217–220</figureCitation>
)
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- Ti IV with normal setae
<emphasis id="B952EA77FFB9FFACFE205E7391E17AD4" box="[462,496,1401,1423]" italics="true" pageId="9" pageNumber="310">pv1</emphasis>
; prominent lateral hyaline protrusions of the subapical epigynial structure wing-like and directed either laterally, or anterolaterally (
<figureCitation id="131D2AE0FFB9FFACFDC95E9C926C7AF0" box="[551,637,1430,1451]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 221</figureCitation>
)................
<taxonomicName id="4C264DE6FFB9FFACFCD35E9D95137AF0" authority="Juvara-Bals &amp; Witalinski, 2000" authorityName="Juvara-Bals &amp; Witalinski" authorityYear="2000" box="[829,1282,1430,1452]" class="Arachnida" family="Parasitidae" genus="Holoparasitus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="digitiformis">
<emphasis id="B952EA77FFB9FFACFCD35E9D93407AF7" box="[829,849,1431,1452]" italics="true" pageId="9" pageNumber="310">H</emphasis>
.
<emphasis id="B952EA77FFB9FFACFCB05E9C93DF7AF7" box="[862,974,1430,1452]" italics="true" pageId="9" pageNumber="310">digitiformis</emphasis>
<bibRefCitation id="EFB74B94FFB9FFACFC3A5E9C95137AF0" author="Juvara-Bals" box="[980,1282,1430,1451]" pageId="9" pageNumber="310" refString="Juvara-Bals, I. &amp; Witalinski, W. (2000) Description of five new species of Holoparasitus s. str. with redescription of H. apenninorum (Berlese, 1906) and H. cultriger (Berlese, 1906) from Italy and Spain (Acari, Gamasida, Parasitidae). Revue Suisse de Zoologie, 107, 3 - 30." type="journal article" year="2000">Juvara-Bals &amp; Witaliński, 2000</bibRefCitation>
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(
<figureCitation id="131D2AE0FFB9FFACFAE15E9C95847AF0" box="[1295,1429,1430,1451]" captionStart="FIGURES 209 – 227" captionStartId="25.[151,264,1822,1845]" captionTargetBox="[267,1326,198,1789]" captionTargetId="figure@25.[249,1335,193,1794]" captionTargetPageId="25" captionText="FIGURES 209 – 227. Females. Holoparasitus vasilei: 209 paragynia and endogynium; 210 anterior part of epigynium and central epigynial prong with subapical structure (courtesy of Dr. I. Juvara-Bals, Nov. 2016); 211 endogynium. Holoparasitus fanes: 212, 213 epigynium central prong; 214 paragynia and epigynium central prong; 215 endogynium between paragynia; 216 endogynium. Holoparasitus crassisetosus: 217 tibia IV with thickened seta pv 1; 218 epigynium, anterior part; 219 paragynium and endogynium; 220 endogynium. Holoparasitus digitiformis: 221 epigynium, central and lateral prongs; 222, 223 paragynium and endogynium; 224 – 226 endogynium, different perspectives; 227 endogynium and endogynial lamella. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/810629/files/figure.png" pageId="9" pageNumber="310">Figs 221–227</figureCitation>
)
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</treatment>
</document>
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