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<document id="C718A31F432714F089351772AF8CF58C" ID-CLB-Dataset="3608" ID-DOI="10.11646/zootaxa.5067.2.7" ID-GBIF-Dataset="59e0113f-90f0-4cc7-a332-7a8c7269435b" ID-ISSN="1175-5326" ID-Zenodo-Dep="5677973" ID-ZooBank="FE8F2CB8-72EC-4A12-95AF-260F3C2941AB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636527369303" checkinUser="plazi" docAuthor="Xu, Yifan, Zheng, Bingyu, Fang, Hui, Shih, Chungkun, Ren, Dong &amp; Wang, Yongjie" docDate="2021" docId="F70987CA151EFFB4A5B936DBFD68FD13" docLanguage="en" docName="zootaxa.5067.2.7.pdf" docOrigin="Zootaxa 5067 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fuscopolystoechotes reticulatus Xu, Zheng, Shih &amp; Wang 2021, sp. nov." docType="treatment" docUuid="EC1A65B9-8258-442B-896D-D62C415A225D" docUuidSource="ZooBank" docVersion="8" lastPageNumber="262" masterDocId="0B30FFB2151CFFB7A52E3725FF88FFDE" masterDocTitle="New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)" masterLastPageNumber="266" masterPageNumber="259" pageNumber="261" updateTime="1699468132269" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DBA2AB173E8EA3AF24EF0D64EFB1C591">New fossil giant lacewings from the Middle Jurassic Jiulongshan Formation of China (Neuroptera, Ithonidae)</mods:title>
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<mods:namePart id="906B410EDF22D80EC686F3218DBC3AE1">Xu, Yifan</mods:namePart>
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<mods:namePart id="3EAF395FD251359623A583409EB5B7C9">Zheng, Bingyu</mods:namePart>
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<mods:namePart id="362F31BAEA60B4BE3C4E2F48712DE20C">Fang, Hui</mods:namePart>
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<mods:namePart id="93DE4BF5B428E72DB9F18DBB988AC660">Shih, Chungkun</mods:namePart>
<mods:affiliation id="13ECCE059CB32F271E14095B2FA9D07C">Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC, 20013 - 7012, USA Corresponding author. wangyongjie @ cnu. edu. cn</mods:affiliation>
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<treatment id="F70987CA151EFFB4A5B936DBFD68FD13" ID-DOI="http://doi.org/10.5281/zenodo.5698939" ID-GBIF-Taxon="190302571" ID-Zenodo-Dep="5698939" LSID="urn:lsid:zoobank.org:act:EC1A65B9-8258-442B-896D-D62C415A225D" httpUri="http://treatment.plazi.org/id/F70987CA151EFFB4A5B936DBFD68FD13" lastPageId="3" lastPageNumber="262" pageId="2" pageNumber="261">
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<paragraph id="7F1F36DC151EFFB5A5B936DBFC40FDC7" blockId="2.[151,970,510,609]" box="[151,968,510,537]" pageId="2" pageNumber="261">
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<taxonomicName id="B8A04D5F151EFFB5A5B936DBFCD5FDC7" authority="Xu, Zheng, Shih &amp; Wang" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[151,861,510,537]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="4DD4EACE151EFFB5A5B936DBFD91FDC7" bold="true" box="[151,537,510,537]" italics="true" pageId="2" pageNumber="261">Fuscopolystoechotes reticulatus</emphasis>
Xu, Zheng, Shih &amp; Wang
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,
<taxonomicNameLabel id="56E757B5151EFFB5A64436DAFC40FDC7" box="[874,968,511,537]" pageId="2" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="37BA6557151EFFB5A5B93562FF63FDBF" box="[151,235,583,609]" pageId="2" pageNumber="261" type="description">
<paragraph id="7F1F36DC151EFFB5A5B93562FF63FDBF" blockId="2.[151,970,510,609]" box="[151,235,583,609]" pageId="2" pageNumber="261">
(
<figureCitation id="E79B2A59151EFFB5A58E3562FF68FDBF" box="[160,224,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1950]" captionTargetBox="[151,1436,630,1900]" captionTargetId="figure-161@2.[151,1436,630,1900]" captionTargetPageId="2" captionText="FIGURE 1. Fuscopolystoechotes reticulatus Xu, Zheng, Shih &amp; Wang, gen. et sp. nov. (AD). (A) Holotype of F. reticulatus, CNUNEUNN2016005. (B) Overlay drawing of holotype of F. reticulatus, with forewing pattern. (C) Photos of paratype of F. reticulatus, CNUNEUNN2016008P (D), Photos of paratype of F. reticulatus. Scale bars: 5 mm in AD." figureDoi="http://doi.org/10.5281/zenodo.5677977" httpUri="https://zenodo.org/record/5677977/files/figure.png" pageId="2" pageNumber="261">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="2BDF6654151EFFB5A5B930A3FB45F839" ID-DOI="http://doi.org/10.5281/zenodo.5677977" ID-Zenodo-Dep="5677977" httpUri="https://zenodo.org/record/5677977/files/figure.png" pageId="2" pageNumber="261" startId="2.[151,250,1926,1950]" targetBox="[151,1436,630,1900]" targetPageId="2">
<paragraph id="7F1F36DC151EFFB5A5B930A3FB45F839" blockId="2.[151,1437,1926,2023]" pageId="2" pageNumber="261">
<emphasis id="4DD4EACE151EFFB5A5B930A3FE9BF840" bold="true" box="[151,275,1926,1950]" pageId="2" pageNumber="261">FIGURE 1.</emphasis>
<taxonomicName id="B8A04D5F151EFFB5A43430A3FCE3F841" authority="Xu, Zheng, Shih &amp; Wang" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[282,875,1926,1951]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="reticulatus" status="gen. et sp. nov.">
<emphasis id="4DD4EACE151EFFB5A43430A3FDD6F840" box="[282,606,1926,1950]" italics="true" pageId="2" pageNumber="261">Fuscopolystoechotes reticulatus</emphasis>
Xu, Zheng, Shih &amp; Wang
</taxonomicName>
,
<emphasis id="4DD4EACE151EFFB5A65B30A2FB99F841" bold="true" box="[885,1041,1927,1951]" pageId="2" pageNumber="261">
<taxonomicNameLabel id="56E757B5151EFFB5A65B30A2FB99F841" box="[885,1041,1927,1951]" pageId="2" pageNumber="261" rank="species">
gen.
<emphasis id="4DD4EACE151EFFB5A68630A2FC32F840" bold="true" box="[936,954,1927,1950]" italics="true" pageId="2" pageNumber="261">et</emphasis>
sp. nov.
</taxonomicNameLabel>
</emphasis>
(AD). (A) Holotype of
<taxonomicName id="B8A04D5F151EFFB5A03F30A2FA1DF840" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[1297,1429,1926,1950]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="4DD4EACE151EFFB5A03F30A2FA1DF840" box="[1297,1429,1926,1950]" italics="true" pageId="2" pageNumber="261">F. reticulatus</emphasis>
</taxonomicName>
, CNUNEUNN2016005. (B) Overlay drawing of holotype of
<taxonomicName id="B8A04D5F151EFFB5A63F308EFC1CF81C" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[785,916,1962,1986]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="4DD4EACE151EFFB5A63F308EFC1CF81C" box="[785,916,1962,1986]" italics="true" pageId="2" pageNumber="261">F. reticulatus</emphasis>
</taxonomicName>
, with forewing pattern. (C) Photos of paratype of
<taxonomicName id="B8A04D5F151EFFB5A0A4308EFE8AF838" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="4DD4EACE151EFFB5A0A4308EFE8AF838" italics="true" pageId="2" pageNumber="261">F. reticulatus</emphasis>
</taxonomicName>
, CNUNEUNN2016008P (D), Photos of paratype of
<taxonomicName id="B8A04D5F151EFFB5A61830EAFC32F838" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[822,954,1998,2022]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="261" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="4DD4EACE151EFFB5A61830EAFC32F838" box="[822,954,1998,2022]" italics="true" pageId="2" pageNumber="261">F. reticulatus</emphasis>
</taxonomicName>
. Scale bars: 5 mm in AD.
</paragraph>
</caption>
<subSubSection id="37BA6557151FFFB4A5B937B2FDACFF0B" pageId="3" pageNumber="262" type="etymology">
<paragraph id="7F1F36DC151FFFB4A5B937B2FDACFF0B" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="262">
<emphasis id="4DD4EACE151FFFB4A5B937B2FE96FF6F" bold="true" box="[151,286,151,177]" pageId="3" pageNumber="262">Etymology.</emphasis>
The specific name
<taxonomicName id="B8A04D5F151FFFB4A4D837B2FDE5FF6F" authorityName="Xu, Zheng, Shih &amp; Wang" authorityYear="2021" box="[502,621,151,177]" class="Insecta" family="Ithonidae" genus="Fuscopolystoechotes" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="262" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="4DD4EACE151FFFB4A4D837B2FDE5FF6F" box="[502,621,151,177]" italics="true" pageId="3" pageNumber="262">reticulatus</emphasis>
</taxonomicName>
is a Latin word meaning “reticulated”, referring to the reticulated pattern present on forewing of this species.
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151FFFB4A5E937FBFC7CFEBB" pageId="3" pageNumber="262" type="materials_examined">
<paragraph id="7F1F36DC151FFFB4A5E937FBFDA4FEC3" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="262">
<emphasis id="4DD4EACE151FFFB4A5E937FBFEBEFF26" bold="true" box="[199,310,222,248]" pageId="3" pageNumber="262">Material.</emphasis>
<typeStatus id="A01B887E151FFFB4A41337FAFE21FF27" box="[317,425,223,249]" pageId="3" pageNumber="262" type="holotype">Holotype</typeStatus>
. CNUNEUNN2016005 (
<figureCitation id="E79B2A59151FFFB4A7CF37FAFCD4FF27" box="[737,860,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1950]" captionTargetBox="[151,1436,630,1900]" captionTargetId="figure-161@2.[151,1436,630,1900]" captionTargetPageId="2" captionText="FIGURE 1. Fuscopolystoechotes reticulatus Xu, Zheng, Shih &amp; Wang, gen. et sp. nov. (AD). (A) Holotype of F. reticulatus, CNUNEUNN2016005. (B) Overlay drawing of holotype of F. reticulatus, with forewing pattern. (C) Photos of paratype of F. reticulatus, CNUNEUNN2016008P (D), Photos of paratype of F. reticulatus. Scale bars: 5 mm in AD." figureDoi="http://doi.org/10.5281/zenodo.5677977" httpUri="https://zenodo.org/record/5677977/files/figure.png" pageId="3" pageNumber="262">Fig. 1AB</figureCitation>
);
<typeStatus id="A01B887E151FFFB4A65E37FAFC5EFF27" box="[880,982,223,249]" pageId="3" pageNumber="262" type="paratype">Paratype</typeStatus>
. CNUNEUNN2016008P/C (
<figureCitation id="E79B2A59151FFFB4A01937FAFA05FF27" box="[1335,1421,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1950]" captionTargetBox="[151,1436,630,1900]" captionTargetId="figure-161@2.[151,1436,630,1900]" captionTargetPageId="2" captionText="FIGURE 1. Fuscopolystoechotes reticulatus Xu, Zheng, Shih &amp; Wang, gen. et sp. nov. (AD). (A) Holotype of F. reticulatus, CNUNEUNN2016005. (B) Overlay drawing of holotype of F. reticulatus, with forewing pattern. (C) Photos of paratype of F. reticulatus, CNUNEUNN2016008P (D), Photos of paratype of F. reticulatus. Scale bars: 5 mm in AD." figureDoi="http://doi.org/10.5281/zenodo.5677977" httpUri="https://zenodo.org/record/5677977/files/figure.png" pageId="3" pageNumber="262">Fig. 1C</figureCitation>
), CNUNEUNN2016009 (
<figureCitation id="E79B2A59151FFFB4A4EB3626FD96FEC3" box="[453,542,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1926,1950]" captionTargetBox="[151,1436,630,1900]" captionTargetId="figure-161@2.[151,1436,630,1900]" captionTargetPageId="2" captionText="FIGURE 1. Fuscopolystoechotes reticulatus Xu, Zheng, Shih &amp; Wang, gen. et sp. nov. (AD). (A) Holotype of F. reticulatus, CNUNEUNN2016005. (B) Overlay drawing of holotype of F. reticulatus, with forewing pattern. (C) Photos of paratype of F. reticulatus, CNUNEUNN2016008P (D), Photos of paratype of F. reticulatus. Scale bars: 5 mm in AD." figureDoi="http://doi.org/10.5281/zenodo.5677977" httpUri="https://zenodo.org/record/5677977/files/figure.png" pageId="3" pageNumber="262">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="7F1F36DC151FFFB4A5E93602FC7CFEBB" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="262">
<materialsCitation id="CFC83C81151FFFB4A5E93602FC78FEBB" ID-GBIF-Occurrence="3400676302" country="China" county="Ningcheng County" location="Daohugou Village" municipality="Middle Jurassic" pageId="3" pageNumber="262" specimenCount="1" stateProvince="Nei Mongol" typeStatus="holotype">
<emphasis id="4DD4EACE151FFFB4A5E93602FE74FE9E" bold="true" box="[199,508,294,321]" pageId="3" pageNumber="262">
<typeStatus id="A01B887E151FFFB4A5E93602FE8AFE9F" box="[199,258,295,321]" pageId="3" pageNumber="262">Type</typeStatus>
locality and horizon.
</emphasis>
<location id="7A7F6007151FFFB4A72C3602FD58FE9F" LSID="urn:lsid:plazi:treatment:F70987CA151EFFB4A5B936DBFD68FD13:7A7F6007151FFFB4A72C3602FD58FE9F" box="[514,720,295,321]" country="China" county="Ningcheng County" municipality="Middle Jurassic" name="Daohugou Village" pageId="3" pageNumber="262" stateProvince="Nei Mongol">Daohugou Village</location>
,
<location id="7A7F6007151FFFB4A7F43602FC25FE9F" LSID="urn:lsid:plazi:treatment:F70987CA151EFFB4A5B936DBFD68FD13:7A7F6007151FFFB4A7F43602FC25FE9F" box="[730,941,295,321]" country="China" county="Ningcheng County" municipality="Middle Jurassic" name="Shantou Township" pageId="3" pageNumber="262" stateProvince="Nei Mongol">Shantou Township</location>
,
<collectingCounty id="967E4E50151FFFB4A6993602FB04FE9F" box="[951,1164,295,321]" pageId="3" pageNumber="262">Ningcheng County</collectingCounty>
,
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,
<collectingCountry id="07B7764C151FFFB4A07C3602FA10FE9F" box="[1362,1432,295,321]" name="China" pageId="3" pageNumber="262">China</collectingCountry>
;
<location id="7A7F6007151FFFB4A5B9366EFE14FEBB" LSID="urn:lsid:plazi:treatment:F70987CA151EFFB4A5B936DBFD68FD13:7A7F6007151FFFB4A5B9366EFE14FEBB" box="[151,412,331,357]" country="China" county="Ningcheng County" municipality="Middle Jurassic" name="Jiulongshan Formation" pageId="3" pageNumber="262" stateProvince="Nei Mongol">Jiulongshan Formation</location>
,
<location id="7A7F6007151FFFB4A489366EFD18FEBB" LSID="urn:lsid:plazi:treatment:F70987CA151EFFB4A5B936DBFD68FD13:7A7F6007151FFFB4A489366EFD18FEBB" box="[423,656,331,357]" country="China" county="Ningcheng County" municipality="Middle Jurassic" name="Callovian-Oxfordian" pageId="3" pageNumber="262" stateProvince="Nei Mongol">Callovian-Oxfordian</location>
boundary, late
<collectingMunicipality id="9F7BACA6151FFFB4A612366EFC78FEBB" box="[828,1008,331,357]" pageId="3" pageNumber="262">Middle Jurassic</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151FFFB4A5E9364BFD42FE57" box="[199,714,366,393]" pageId="3" pageNumber="262" type="diagnosis">
<paragraph id="7F1F36DC151FFFB4A5E9364BFD42FE57" blockId="3.[151,1437,151,717]" box="[199,714,366,393]" pageId="3" pageNumber="262">
<emphasis id="4DD4EACE151FFFB4A5E9364BFECAFE56" bold="true" box="[199,322,366,392]" pageId="3" pageNumber="262">Diagnosis.</emphasis>
As for the genus due to monotypy.
</paragraph>
</subSubSection>
<subSubSection id="37BA6557151FFFB4A5E936B7FD68FD13" pageId="3" pageNumber="262" type="description">
<paragraph id="7F1F36DC151FFFB4A5E936B7FC42FD77" blockId="3.[151,1437,151,717]" pageId="3" pageNumber="262">
<emphasis id="4DD4EACE151FFFB4A5E936B7FED1FE72" bold="true" box="[199,345,402,428]" pageId="3" pageNumber="262">Description.</emphasis>
Forewing elongated,
<emphasis id="4DD4EACE151FFFB4A76436B6FDECFE72" box="[586,612,403,428]" italics="true" pageId="3" pageNumber="262">ca</emphasis>
.
<quantity id="B8589B39151FFFB4A75C36B6FD48FE73" box="[626,704,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="262" unit="mm" value="39.0">39 mm</quantity>
long and
<quantity id="B8589B39151FFFB4A60336B6FCF3FE73" box="[813,891,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="262" unit="mm" value="13.0">13 mm</quantity>
wide; membrane dark, with the distinct transparent patches along anterior and posterior margins; trichosors well-defined in most margin except for the proximal half of anterior margin; humeral vein recurrent, producing 45 short crossveins; costal crossveins with the distal forks, and absence of interlinked crossveins in between; space between Sc and RA relatively broad, sc-sa crossvein absent; ra-rp crossveins numerous; RP with 18 close-set branches, crossveins in RP area numerous and irregularly arranged, only one outer gradate series present; M forked close to the divergence of RP1 from RP; MA with 4 distal branches, MP with 6 distal branches; Cu proximally branched; CuA with about 9 oblique pectinate branches; CuP dichotomously branched; A1A3 multi-dichotomously branched distally.
</paragraph>
<paragraph id="7F1F36DC151FFFB4A5E93596FD68FD13" blockId="3.[151,1437,151,717]" box="[199,736,691,717]" pageId="3" pageNumber="262">Only forewing present, body features unknown.</paragraph>
</subSubSection>
</treatment>
</document>