109 lines
14 KiB
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109 lines
14 KiB
XML
<document id="BD69DAA9F86D5B548915FF8C3EB913C8" ID-CLB-Dataset="6611" ID-DOI="10.5281/zenodo.4502132" ID-GBIF-Dataset="60740210-999d-4526-88e3-464fd93143ac" ID-ISSN="2345-7600" ID-Zenodo-Dep="4502132" ID-ZooBank="4D5F3585-04B2-45AA-BA00-8795AEF68A83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612458956132" checkinUser="carolina" docAuthor="Köhler, Frank" docDate="2014" docId="03F3175FEF6B5E40FC733A7EFDF3FE8B" docLanguage="en" docName="RafflesBZool.62.115-123.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Rhagada setzeri subsp. setzeri Maassen 2009" docType="treatment" docVersion="10" lastPageNumber="119" masterDocId="FFCA6F27EF6F5E45FFDE3C2BFFD2FFFE" masterDocTitle="On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands" masterLastPageNumber="123" masterPageNumber="115" pageNumber="119" updateTime="1698926749818" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="469E270C492028B93F6E0B7F4FD9A276">On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands</mods:title>
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<mods:name id="B84958FD692713319A0C27BE6F48CC9D" type="personal">
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<mods:namePart id="31112675ECFBE7DDCACD0D044909FC88">Köhler, Frank</mods:namePart>
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<mods:title id="DF4936E3663781411D8049F267476004">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="E5C91BF55EFA4BA5F8A53232FE2B8F24">2014</mods:date>
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<mods:number id="E2EEE295EE87BA4B752707081FF5281D">2014-03-17</mods:number>
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<mods:number id="60F37ACB77CBBFE66D04370C89711875">62</mods:number>
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<mods:start id="E25A080ECF5E0F2285F18320ABA6F3C9">115</mods:start>
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<treatment id="03F3175FEF6B5E40FC733A7EFDF3FE8B" ID-DOI="http://doi.org/10.5281/zenodo.4774206" ID-GBIF-Taxon="182237997" ID-Zenodo-Dep="4774206" LSID="urn:lsid:plazi:treatment:03F3175FEF6B5E40FC733A7EFDF3FE8B" httpUri="http://treatment.plazi.org/id/03F3175FEF6B5E40FC733A7EFDF3FE8B" lastPageId="5" lastPageNumber="119" pageId="4" pageNumber="119">
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<subSubSection id="C340F5C2EF6B5E41FC733A7EFA8AF993" box="[941,1368,1621,1645]" pageId="4" pageNumber="119" type="nomenclature">
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<paragraph id="8BE5A649EF6B5E41FC733A7EFA8AF993" blockId="4.[941,1368,1621,1645]" box="[941,1368,1621,1645]" pageId="4" pageNumber="119">
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<heading id="D0AD1125EF6B5E41FC733A7EFA8AF993" box="[941,1368,1621,1645]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="119" reason="3">
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<taxonomicName id="4C5ADDCAEF6B5E41FC733A7EFA8AF993" ID-CoL="7V2MG" authority="setzeri Maassen, 2009" authorityName="Maassen" authorityYear="2009" box="[941,1368,1621,1645]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="subSpecies" species="setzeri" subSpecies="setzeri">
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<emphasis id="B92E7A5BEF6B5E41FC733A7EFB7CF993" bold="true" box="[941,1198,1621,1645]" italics="true" pageId="4" pageNumber="119">Rhagada setzeri setzeri</emphasis>
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<bibRefCitation id="EFCBDBB8EF6B5E41FB683A7DFA8AF993" author="Maassen WJM" box="[1206,1368,1622,1645]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340F5C2EF6B5E41FC9D3ABFFBDDF91D" pageId="4" pageNumber="119" type="reference_group">
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<paragraph id="8BE5A649EF6B5E41FC9D3ABFFBDDF91D" blockId="4.[835,1474,1684,1763]" pageId="4" pageNumber="119">
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<treatmentCitationGroup id="AB4A8167EF6B5E41FC9D3ABFFBDDF91D" pageId="4" pageNumber="119">
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<treatmentCitation id="0AFB8058EF6B5E41FC9D3ABFFB1DF957" author="Maassen WJM" box="[835,1231,1684,1705]" page="69" pageId="4" pageNumber="119" year="2009">
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<taxonomicName id="4C5ADDCAEF6B5E41FC9D3ABFFB1DF957" ID-CoL="4RYCN" authority="Maassen, 2009: 66 - 69" authorityName="Maassen" authorityPageNumber="66 - 69" authorityYear="2009" box="[835,1231,1684,1705]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="setzeri">
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<emphasis id="B92E7A5BEF6B5E41FC9D3ABFFC31F957" box="[835,995,1684,1705]" italics="true" pageId="4" pageNumber="119">Rhagada setzeri</emphasis>
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<bibRefCitation id="EFCBDBB8EF6B5E41FC323ABFFB1DF957" author="Maassen WJM" box="[1004,1231,1684,1705]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009: 66–69</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 13–16 (from ‘
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<location id="8E85F092EF6B5E41FA4F3ABFFC74F938" LSID="urn:lsid:plazi:treatment:03F3175FEF6B5E40FC733A7EFDF3FE8B:8E85F092EF6B5E41FA4F3ABFFC74F938" latitude="-8.25" longLatPrecision="21" longitude="124.75" name="Alor Island" pageId="4" pageNumber="119">Alor Island</location>
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, near the airport’,
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<geoCoordinate id="EE6EC08EEF6B5E41FB803A9AFB1FF938" box="[1118,1229,1713,1734]" degrees="8" direction="south" minutes="15" orientation="latitude" pageId="4" pageNumber="119" precision="15" seconds="0" value="-8.25">8° 15' 0" S</geoCoordinate>
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,
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<geoCoordinate id="EE6EC08EEF6B5E41FB073A9AFAB0F938" box="[1241,1378,1713,1734]" degrees="124" direction="east" minutes="45" orientation="longitude" pageId="4" pageNumber="119" precision="15" seconds="0" value="124.75">124° 45' 0" E</geoCoordinate>
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;
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<typeStatus id="54E118EBEF6B5E41FAB03A9AFA10F938" box="[1390,1474,1713,1734]" pageId="4" pageNumber="119" type="holotype">holotype</typeStatus>
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<specimenCode id="DBFC0E32EF6B5E41FCB73AE5FBD6F91D" box="[873,1028,1742,1763]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="119" type="Museum">RMNH 113381</specimenCode>
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).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340F5C2EF6B5E41FC9D3B23FA11F8E3" box="[835,1475,1800,1821]" pageId="4" pageNumber="119" type="materials_examined">
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<paragraph id="8BE5A649EF6B5E41FC9D3B23FA11F8E3" blockId="4.[835,1475,1800,1821]" box="[835,1475,1800,1821]" pageId="4" pageNumber="119">
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<emphasis id="B92E7A5BEF6B5E41FC9D3B23FBD8F8E3" bold="true" box="[835,1034,1800,1821]" pageId="4" pageNumber="119">Material examined.</emphasis>
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<collectionCode id="ED4B3E8CEF6B5E41FBD03B23FBE7F8E3" box="[1038,1077,1800,1821]" country="Australia" name="Australian Museum" pageId="4" pageNumber="119" type="Museum">AM</collectionCode>
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C.471156 (
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<collectingCountry id="F34DE6D9EF6B5E41FB7D3B23FAD6F8E3" box="[1187,1284,1800,1821]" name="Indonesia" pageId="4" pageNumber="119">Indonesia</collectingCountry>
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, Alor Island, 3 dry).
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</paragraph>
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</subSubSection>
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<subSubSection id="C340F5C2EF6B5E40FC9D3B6DFDF3FE8B" lastPageId="5" lastPageNumber="120" pageId="4" pageNumber="119" type="diagnosis">
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<paragraph id="8BE5A649EF6B5E41FC9D3B6DFB01F843" blockId="4.[835,1474,1862,1981]" pageId="4" pageNumber="119">
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<emphasis id="B92E7A5BEF6B5E41FC9D3B6DFC6AF8A3" bold="true" box="[835,952,1862,1885]" pageId="4" pageNumber="119">Diagnosis.</emphasis>
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Shell comparatively large (D =
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<quantity id="4CA20BACEF6B5E41FAD53B6DFA53F8A3" box="[1291,1409,1862,1885]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="4" pageNumber="119" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
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), with usually well-developed axial ribs, umbilicus closed. Penis with long epiphallus, short flagellum, inner penial anatomy unknown (based on
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<bibRefCitation id="EFCBDBB8EF6B5E41FBFF3B8DFB15F843" author="Maassen WJM" box="[1057,1223,1958,1981]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BE5A649EF6B5E40FC9D3BCDFDF3FE8B" blockId="4.[835,1474,2022,2109]" lastBlockId="5.[113,753,190,373]" lastPageId="5" lastPageNumber="120" pageId="4" pageNumber="119">
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<emphasis id="B92E7A5BEF6B5E41FC9D3BCDFB86F803" bold="true" box="[835,1108,2022,2045]" pageId="4" pageNumber="119">Comparative remarks.</emphasis>
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Comparatively large shell with well-developed axial sculpture and closed umbilicus is most characteristic. Well-differentiated from
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<taxonomicName id="4C5ADDCAEF6B5E41FAEB340DFA10F7C3" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[1333,1474,2086,2109]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
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<emphasis id="B92E7A5BEF6B5E41FAEB340DFA10F7C3" box="[1333,1474,2086,2109]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C5ADDCAEF6A5E40FF7D3C95FEDCFF2B" authority="Köhler, 2014" authorityName="Köhler" authorityYear="2014" box="[163,270,190,213]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="120" phylum="Mollusca" rank="species" species="abbasi" status="sp. nov.">
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<emphasis id="B92E7A5BEF6A5E40FF7D3C95FEDCFF2B" box="[163,270,190,213]" italics="true" pageId="5" pageNumber="120">R. abbasi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21DC720EF6A5E40FEC63C95FE84FF2B" box="[280,342,190,213]" pageId="5" pageNumber="120" rank="species">n. sp.</taxonomicNameLabel>
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in a molecular phylogeny based on mitochondrial COI and 16S sequences (
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<figureCitation id="1361BACCEF6A5E40FDFD3CF5FDB4FF0B" box="[547,614,222,245]" captionStart="Fig" captionStartId="5.[113,148,2006,2027]" captionTargetBox="[157,1404,941,1982]" captionTargetId="figure-153@5.[151,1414,931,1991]" captionTargetPageId="5" captionText="Fig. 3. Bayesian consensus phylogram based on analyses of concatenated 16S and COI sequences, adapted from Köhler & Criscione (2013). Lesser Sunda species Rhagada solorensis, R. setzeri setzeri, R. setzeri atauroensis n. subsp., and R. abbasi n. sp. form the sister clade of Rhagada from the Australian Pilbara. Clades from the Australian Kimberley and Pilbara regions collapsed; outgroup pruned from tree; see Köhler & Criscione (2013) for further details." figureDoi="http://doi.org/10.5281/zenodo.4502142" httpUri="https://zenodo.org/record/4502142/files/figure.png" pageId="5" pageNumber="120">Fig. 3</figureCitation>
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). Specimens from Atauro Island differ by consistently smaller shell, weaker sculpture, narrowly open umbilicus but are differentiated by comparatively low genetic distances (
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<tableCitation id="C6D893F2EF6A5E40FDF93D15FDA9FEAB" box="[551,635,318,341]" captionStart="Table 2" captionStartId="6.[136,189,190,211]" captionTargetBox="[147,1457,275,706]" captionText="Table 2. Estimates of evolutionary divergence by means of uncorrected p-distances (base differences per site) within and between species" httpUri="http://table.plazi.org/id/DF25F6C1EF695E43FF563C95FDA8FF0E" pageId="5" pageNumber="120" tableUuid="DF25F6C1EF695E43FF563C95FDA8FF0E">Table 2</tableCitation>
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); therefore being recognised as distinct subspecies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |