280 lines
41 KiB
XML
280 lines
41 KiB
XML
<document id="1B9742E8CFFDC308E3C50D24A76708AD" ID-DOI="10.11646/zootaxa.3931.3.3" ID-GBIF-Dataset="f8048196-a12e-40d0-8349-c830479322db" ID-ISSN="1175-5326" ID-Zenodo-Dep="244956" ID-ZooBank="13BA1A35-C31A-4910-8EAC-1D840C5CAD18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461010774241" checkinUser="plazi" docAuthor="Lina, Almeida-Silva, Charles, Griswold & Antonio, Brescovit" docDate="2015" docId="03D287E3FFD37F5DFF28FAE14847FB3B" docLanguage="en" docName="zt03931p400.pdf" docOrigin="Zootaxa 3931 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Emmenomma Simon" docType="treatment" docVersion="7" lastPageNumber="390" masterDocId="FFEBFF9BFFD27F5EFFBFFFD34B64FFF6" masterDocTitle="Revision of Emmenomma Simon (Amaurobiidae, Macrobuninae)" masterLastPageNumber="400" masterPageNumber="387" pageNumber="388" updateTime="1698567196116" updateUser="plazi">
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<mods:titleInfo id="8D0CDAD029645A4461DC683273805FF3">
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<mods:title id="76E47D5E525F1B6B5FCAAF5F5EA77678">Revision of Emmenomma Simon (Amaurobiidae, Macrobuninae)</mods:title>
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<mods:namePart id="2967317B05D88505AD18C2D32922F176">Lina, Almeida-Silva</mods:namePart>
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<mods:namePart id="3B4A239668C839EBD9075D7353D66425">Charles, Griswold</mods:namePart>
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<mods:name id="10C06302CEA9495ED8AAE906DB4BE87E" type="personal">
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<mods:roleTerm id="2C6B0ABC9D3A651DB167AEF793955436">Author</mods:roleTerm>
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<mods:namePart id="D07D546F723CCC50CA216D76AF95ADCA">Antonio, Brescovit</mods:namePart>
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<mods:typeOfResource id="3F2DAED80E0AD6BDEFAF5C1FCBF88258">text</mods:typeOfResource>
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<mods:title id="59D8AC12602B113619ECF05FC8DB4643">Zootaxa</mods:title>
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<mods:part id="93C967D09236622D48D86DF6A7F8DBCA">
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<mods:date id="9C46001112FB2C8C888D4E2834FA21C4">2015</mods:date>
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<mods:detail id="53B98F7F7656F24C6D2CFFEF66DAD3E7" type="volume">
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<mods:number id="A34DBDB32AB04C259BB2D2F814A532D9">3931</mods:number>
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<mods:classification id="6E97F82DE65B485F489B2D98DF8EAF7C">journal article</mods:classification>
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<mods:identifier id="7FB076E3A29243BE3A7F6337BCC87D84" type="DOI">10.11646/zootaxa.3931.3.3</mods:identifier>
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<treatment id="03D287E3FFD37F5DFF28FAE14847FB3B" ID-DOI="http://doi.org/10.5281/zenodo.6115507" ID-GBIF-Taxon="119600146" ID-Zenodo-Dep="6115507" LSID="urn:lsid:plazi:treatment:03D287E3FFD37F5DFF28FAE14847FB3B" httpUri="http://treatment.plazi.org/id/03D287E3FFD37F5DFF28FAE14847FB3B" lastPageId="3" lastPageNumber="390" pageId="1" pageNumber="388">
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<subSubSection id="C361657EFFD37F5FFF28FAE14AF3FABB" box="[151,407,1330,1357]" pageId="1" pageNumber="388" type="nomenclature">
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<paragraph id="8BC436F5FFD37F5FFF28FAE14AF3FABB" blockId="1.[151,407,1330,1357]" box="[151,407,1330,1357]" pageId="1" pageNumber="388">
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<heading id="D08C8199FFD37F5FFF28FAE14AF3FABB" bold="true" box="[151,407,1330,1357]" fontSize="11" level="1" pageId="1" pageNumber="388" reason="1">
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<taxonomicName id="4C7B4D76FFD37F5FFF28FAE14AF3FABB" ID-CoL="62RTH" authority="Simon" authorityName="Simon" box="[151,407,1330,1357]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFF28FAE14AF3FABB" bold="true" box="[151,407,1330,1357]" pageId="1" pageNumber="388">
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<emphasis id="B90FEAE7FFD37F5FFF28FAE14A24FABA" bold="true" box="[151,320,1330,1356]" italics="true" pageId="1" pageNumber="388">Emmenomma</emphasis>
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Simon
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C361657EFFD37F5FFF28FAA94A8DFA58" pageId="1" pageNumber="388" type="reference_group">
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<paragraph id="8BC436F5FFD37F5FFF28FAA94A8DFA58" blockId="1.[151,1436,1401,1454]" pageId="1" pageNumber="388">
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<treatmentCitationGroup id="AB6B11DBFFD37F5FFF28FAA94A8DFA58" pageId="1" pageNumber="388">
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<taxonomicName id="4C7B4D76FFD37F5FFF28FAA94A8FFA79" ID-CoL="62RTH" authority="Simon, 1884: 126" authorityName="Simon" authorityPageNumber="126" authorityYear="1884" box="[151,491,1401,1423]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFF28FAA94A4DFA79" bold="true" box="[151,297,1402,1423]" italics="true" pageId="1" pageNumber="388">Emmenomma</emphasis>
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<treatmentCitation id="0ADA10E4FFD37F5FFE8EFAAA4A8FFA79" author="Simon" box="[305,491,1401,1423]" page="126" pageId="1" pageNumber="388" year="1884">
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<bibRefCitation id="EFEA4B04FFD37F5FFE8EFAAA4AD3FA79" author="Simon" box="[305,439,1401,1423]" pageId="1" pageNumber="388" refString="Simon, E. (1884) Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France, 9, 117 - 144." type="journal article" year="1884">Simon, 1884</bibRefCitation>
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: 126
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</treatmentCitation>
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</taxonomicName>
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, pl. 3, figs. 8–11 (type-species by monotypy
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<taxonomicName id="4C7B4D76FFD37F5FFC01FAA94FCAFA79" ID-CoL="39H5J" box="[958,1198,1401,1423]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="oculatum">
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<emphasis id="B90FEAE7FFD37F5FFC01FAA94FCAFA79" box="[958,1198,1401,1423]" italics="true" pageId="1" pageNumber="388">Emmenomma oculatum</emphasis>
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</taxonomicName>
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);
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<treatmentCitation id="0ADA10E4FFD37F5FFB7AFAAA4EF3FA79" author="Lehtinen" box="[1221,1431,1401,1423]" page="232" pageId="1" pageNumber="388" year="1967">
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<bibRefCitation id="EFEA4B04FFD37F5FFB7AFAAA4E07FA79" author="Lehtinen" box="[1221,1379,1401,1423]" pageId="1" pageNumber="388" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
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: 232
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</treatmentCitation>
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; World Spider Catalog, 2014.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C361657EFFD37F5FFF28FA094E1FF919" pageId="1" pageNumber="388" type="diagnosis">
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<paragraph id="8BC436F5FFD37F5FFF28FA094E1FF919" blockId="1.[151,1437,1498,2027]" pageId="1" pageNumber="388">
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<emphasis id="B90FEAE7FFD37F5FFF28FA094A75FA05" bold="true" box="[151,273,1498,1523]" pageId="1" pageNumber="388">Diagnosis.</emphasis>
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<taxonomicName id="4C7B4D76FFD37F5FFEA1FA084AD3FA04" box="[286,439,1499,1522]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFEA1FA084AD3FA04" box="[286,439,1499,1522]" italics="true" pageId="1" pageNumber="388">Emmenomma</emphasis>
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</taxonomicName>
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differs from the non-macrobunine amaurobiids by the presence of denticles on the proximal end of the retromarginal row of teeth of the chelicerae (
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<figureCitation id="13402A70FFD37F5FFCCCFA2D48D2F9E1" box="[883,950,1534,1559]" captionStart="FIGURE 2" captionStartId="3.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1048]" captionTargetId="figure@3.[151,1436,193,1049]" captionTargetPageId="3" captionText="FIGURE 2. Emmenomma oculatum. A. Eyes, ♀; B. Eyes detail, AME and ALE, ♀; C. Tarsal organ, ♂ palp; D. Right endite, dorsal, ♀. E. Right chelicerae teeth, retromargin at top; F. Palpal tarsal claw, ♀." httpUri="https://zenodo.org/record/244958/files/figure.png" pageId="1" pageNumber="388">Fig. 2</figureCitation>
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E).
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<taxonomicName id="4C7B4D76FFD37F5FFC61FA2C4F12F9E0" box="[990,1142,1535,1558]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFC61FA2C4F12F9E0" box="[990,1142,1535,1558]" italics="true" pageId="1" pageNumber="388">Emmenomma</emphasis>
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</taxonomicName>
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can be recognized among other species of Macrobuninae by the recurved posterior eye row (
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<figureCitation id="13402A70FFD37F5FFC23F9F148B9F9CD" box="[924,989,1570,1595]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1989]" captionTargetBox="[204,1382,879,1942]" captionTargetId="figure@2.[204,1383,879,1942]" captionTargetPageId="2" captionText="FIGURE 1. Emmenomma oculatum. A – C: ♂ (recently molted); D: Immature. A. Carapace frontal view; B. Habitus, dorsal view; C. Habitus ventral view; D. Tapetum removed from ALE." httpUri="https://zenodo.org/record/244957/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
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A), enlarged ALE and PLE (
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<figureCitation id="13402A70FFD37F5FFA8FF9F14E19F9CD" box="[1328,1405,1570,1595]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1989]" captionTargetBox="[204,1382,879,1942]" captionTargetId="figure@2.[204,1383,879,1942]" captionTargetPageId="2" captionText="FIGURE 1. Emmenomma oculatum. A – C: ♂ (recently molted); D: Immature. A. Carapace frontal view; B. Habitus, dorsal view; C. Habitus ventral view; D. Tapetum removed from ALE." httpUri="https://zenodo.org/record/244957/files/figure.png" pageId="1" pageNumber="388">Figs. 1</figureCitation>
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A; 2A–B), and grate-shaped tapetum in all indirect eyes (
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<figureCitation id="13402A70FFD37F5FFD47F995485CF9A9" box="[760,824,1606,1631]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1989]" captionTargetBox="[204,1382,879,1942]" captionTargetId="figure@2.[204,1383,879,1942]" captionTargetPageId="2" captionText="FIGURE 1. Emmenomma oculatum. A – C: ♂ (recently molted); D: Immature. A. Carapace frontal view; B. Habitus, dorsal view; C. Habitus ventral view; D. Tapetum removed from ALE." httpUri="https://zenodo.org/record/244957/files/figure.png" pageId="1" pageNumber="388">Fig. 1</figureCitation>
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D). The males resemble
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<taxonomicName id="4C7B4D76FFD37F5FFBF0F9944FD9F9A8" box="[1103,1213,1607,1630]" class="Arachnida" family="Amaurobiidae" genus="Anisacate" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFBF0F9944FD9F9A8" box="[1103,1213,1607,1630]" italics="true" pageId="1" pageNumber="388">Anisacate</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7B4D76FFD37F5FFB4BF9944E2AF9A8" box="[1268,1358,1607,1630]" class="Arachnida" family="Amaurobiidae" genus="Naevius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFB4BF9944E2AF9A8" box="[1268,1358,1607,1630]" italics="true" pageId="1" pageNumber="388">Naevius</emphasis>
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</taxonomicName>
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by the presence of a gland in the male palpal tibia which discharges through an opening on the DTA (
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<figureCitation id="13402A70FFD37F5FFB64F9B94E4DF975" box="[1243,1321,1642,1667]" captionStart="FIGURE 3" captionStartId="4.[151,250,1749,1771]" captionTargetBox="[151,1426,447,1725]" captionTargetId="figure@4.[151,1436,447,1727]" captionTargetPageId="4" captionText="FIGURE 3. Emmenomma oculatum right ♂ palp. A. Bulb, prolateral view; B. Median apophysis detail; C. Bulb, ventral view, including DTA tip (Arrow pointing to gland pore); D. Bulb, apical view, including DTA tip (Arrow pointing to gland pore). Abbrev.: DTP—distal tegular process; E—embolus; MA—median apophysis; TA—tegular apophysis; VTA—ventral tibial Apophysis." httpUri="https://zenodo.org/record/244959/files/figure.png" pageId="1" pageNumber="388">Figs. 3</figureCitation>
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C–D), but differ from
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<taxonomicName id="4C7B4D76FFD37F5FFE9FF95C4AEAF950" box="[288,398,1679,1702]" class="Arachnida" family="Amaurobiidae" genus="Anisacate" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFE9FF95C4AEAF950" box="[288,398,1679,1702]" italics="true" pageId="1" pageNumber="388">Anisacate</emphasis>
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</taxonomicName>
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by having the DTA flexibly attached and mobile (
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<figureCitation id="13402A70FFD37F5FFC5EF95D4F56F951" box="[993,1074,1678,1703]" captionStart="FIGURE 4" captionStartId="5.[151,250,1862,1884]" captionTargetBox="[244,1337,193,1835]" captionTargetId="figure@5.[244,1345,193,1842]" captionTargetPageId="5" captionText="FIGURE 4. Emmenomma oculatum ♂. A. Right palp dorsal view. B. Left palp, retrolateral view showing RTA and DTA. C. Left palp, retrolateral view, iRTA. D. Left palp, expanded, retrolateral view, bulb. E. Left palp, expanded, prolateral view, bulb. F. Left palp, expanded, ventro-prolateral view. Abbrev.: DTA—dorsal tibial apophysis; DTP—distal tegular process; E—embolus; iRTA—internal branch of the retrolateral tibial apophysis; MA—median apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; TA—tegular apophysis TR—tegular rim; ST—subtegulum; VTA—ventral tibial apophysis." httpUri="https://zenodo.org/record/244960/files/figure.png" pageId="1" pageNumber="388">Figs. 4</figureCitation>
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A–B) and from
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<taxonomicName id="4C7B4D76FFD37F5FFB51F95C4E2CF950" box="[1262,1352,1679,1702]" class="Arachnida" family="Amaurobiidae" genus="Naevius" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAE7FFD37F5FFB51F95C4E2CF950" box="[1262,1352,1679,1702]" italics="true" pageId="1" pageNumber="388">Naevius</emphasis>
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</taxonomicName>
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by the presence of a bifid MA (
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<figureCitation id="13402A70FFD37F5FFE11F9614A95F93D" box="[430,497,1714,1739]" captionStart="FIGURE 3" captionStartId="4.[151,250,1749,1771]" captionTargetBox="[151,1426,447,1725]" captionTargetId="figure@4.[151,1436,447,1727]" captionTargetPageId="4" captionText="FIGURE 3. Emmenomma oculatum right ♂ palp. A. Bulb, prolateral view; B. Median apophysis detail; C. Bulb, ventral view, including DTA tip (Arrow pointing to gland pore); D. Bulb, apical view, including DTA tip (Arrow pointing to gland pore). Abbrev.: DTP—distal tegular process; E—embolus; MA—median apophysis; TA—tegular apophysis; VTA—ventral tibial Apophysis." httpUri="https://zenodo.org/record/244959/files/figure.png" pageId="1" pageNumber="388">Fig. 3</figureCitation>
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B), by the elbowed DTA and RTA not ventrally displaced (
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<figureCitation id="13402A70FFD37F5FFB34F9614FB4F93D" box="[1163,1232,1714,1739]" captionStart="FIGURE 4" captionStartId="5.[151,250,1862,1884]" captionTargetBox="[244,1337,193,1835]" captionTargetId="figure@5.[244,1345,193,1842]" captionTargetPageId="5" captionText="FIGURE 4. Emmenomma oculatum ♂. A. Right palp dorsal view. B. Left palp, retrolateral view showing RTA and DTA. C. Left palp, retrolateral view, iRTA. D. Left palp, expanded, retrolateral view, bulb. E. Left palp, expanded, prolateral view, bulb. F. Left palp, expanded, ventro-prolateral view. Abbrev.: DTA—dorsal tibial apophysis; DTP—distal tegular process; E—embolus; iRTA—internal branch of the retrolateral tibial apophysis; MA—median apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; TA—tegular apophysis TR—tegular rim; ST—subtegulum; VTA—ventral tibial apophysis." httpUri="https://zenodo.org/record/244960/files/figure.png" pageId="1" pageNumber="388">Fig. 4</figureCitation>
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B). Females differ from all Macrobuninae by having an ET at the posterior end of lateral lobe of the epigynum (
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<figureCitation id="13402A70FFD37F5FFB16F9054F93F919" box="[1193,1271,1750,1775]" captionStart="FIGURE 5" captionStartId="6.[151,250,933,955]" captionTargetBox="[151,1436,193,911]" captionTargetId="figure@6.[151,1436,193,912]" captionTargetPageId="6" captionText="FIGURE 5. Emmenomma oculatum ♀. A. Epigynum, ventral view. B. Epigynum, ventro-posterior view. C. Epigynum, posterior view; D. Vulva, left spermatheca. E. Vulva, dorsal view. F. Vulva, anterior view. Abbrev.: BG—Bennetts’s gland; CD—copulatory duct; LL—lateral lobe of the epigynum; MF—epigynum middle-field; S—spermathecae; ET- epigynum tooth; FD—fertilization duct." httpUri="https://zenodo.org/record/244961/files/figure.png" pageId="1" pageNumber="388">Figs. 5</figureCitation>
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B; 8A; 9D).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C361657EFFD37F5CFF78F92949E1FD96" lastPageId="2" lastPageNumber="389" pageId="1" pageNumber="388" type="description">
|
||
<paragraph id="8BC436F5FFD37F5CFF78F92949E1FD96" blockId="1.[151,1437,1498,2027]" lastBlockId="2.[151,1437,151,824]" lastPageId="2" lastPageNumber="389" pageId="1" pageNumber="388">
|
||
<emphasis id="B90FEAE7FFD37F5FFF78F9294A3DF8E5" bold="true" box="[199,345,1786,1811]" pageId="1" pageNumber="388">Description.</emphasis>
|
||
Total length: 4.90–15.00. Anterior eye row slightly procurved and posterior eye row recurved; ALE enlarged, approximately twice the size of PME; AME reduced, 1/2 size of PME; PLE and PME about same size. All indirect eyes with grate-shaped tapetum. Thoracic fovea long and shallow in
|
||
<taxonomicName id="4C7B4D76FFD37F5FFB12F8904E24F8AD" box="[1197,1344,1859,1883]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="obscurum">
|
||
<emphasis id="B90FEAE7FFD37F5FFB12F8904E24F8AD" box="[1197,1344,1859,1883]" italics="true" pageId="1" pageNumber="388">E. obscurum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D76FFD37F5FFA3BF8904A70F888" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="joshuabelli" status="sp. nov.">
|
||
<emphasis id="B90FEAE7FFD37F5FFA3BF8904A70F888" italics="true" pageId="1" pageNumber="388">E. joshuabelli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAE7FFD37F5FFEA2F8B54A1EF889" bold="true" box="[285,378,1894,1919]" pageId="1" pageNumber="388">
|
||
<taxonomicNameLabel id="A23C579CFFD37F5FFEA2F8B54A1EF889" box="[285,378,1894,1919]" pageId="1" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and deep in
|
||
<taxonomicName id="4C7B4D76FFD37F5FFDAAF8B449FBF888" box="[533,671,1894,1918]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD37F5FFDAAF8B449FBF888" box="[533,671,1894,1918]" italics="true" pageId="1" pageNumber="388">E. oculatum</emphasis>
|
||
</taxonomicName>
|
||
. Coloration variable between pale and dark brown, with scattered guanine deposits in the abdomen, legs usually maculate (
|
||
<figureCitation id="13402A70FFD37F5FFC89F85948E7F855" box="[822,899,1930,1955]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1989]" captionTargetBox="[204,1382,879,1942]" captionTargetId="figure@2.[204,1383,879,1942]" captionTargetPageId="2" captionText="FIGURE 1. Emmenomma oculatum. A – C: ♂ (recently molted); D: Immature. A. Carapace frontal view; B. Habitus, dorsal view; C. Habitus ventral view; D. Tapetum removed from ALE." httpUri="https://zenodo.org/record/244957/files/figure.png" pageId="1" pageNumber="388">Figs. 1</figureCitation>
|
||
A–C). Chilum absent except in females of
|
||
<taxonomicName id="4C7B4D76FFD37F5FFA3BF8584B9BF830" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD37F5FFA3BF8584B9BF830" italics="true" pageId="1" pageNumber="388">E. oculatum</emphasis>
|
||
</taxonomicName>
|
||
where it is divided. Clypeal hood absent in
|
||
<taxonomicName id="4C7B4D76FFD37F5FFCA5F87C48C2F830" box="[794,934,1966,1990]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD37F5FFCA5F87C48C2F830" box="[794,934,1966,1990]" italics="true" pageId="1" pageNumber="388">E. oculatum</emphasis>
|
||
</taxonomicName>
|
||
, present in
|
||
<taxonomicName id="4C7B4D76FFD37F5FFB87F87C4FBDF830" box="[1080,1241,1966,1990]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="joshuabelli" status="sp. nov.">
|
||
<emphasis id="B90FEAE7FFD37F5FFB87F87C4FBDF830" box="[1080,1241,1966,1990]" italics="true" pageId="1" pageNumber="388">E. joshuabelli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAE7FFD37F5FFB5BF87D4E20F831" bold="true" box="[1252,1348,1966,1991]" pageId="1" pageNumber="388">
|
||
<taxonomicNameLabel id="A23C579CFFD37F5FFB5BF87D4E20F831" box="[1252,1348,1966,1991]" pageId="1" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7B4D76FFD37F5FFA3BF87C4A61F81D" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="obscurum">
|
||
<emphasis id="B90FEAE7FFD37F5FFA3BF87C4A61F81D" italics="true" pageId="1" pageNumber="388">E. obscurum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13402A70FFD37F5FFEA8F8014A11F81D" box="[279,373,2002,2027]" captionStart="FIGURE 10" captionStartId="11.[151,249,1296,1318]" captionTargetBox="[151,1436,157,1274]" captionTargetId="figure@11.[151,1436,157,1274]" captionTargetPageId="11" captionText="FIGURE 10. Emmenomma obscurum and E. joshuabelli sp. nov. A, D, G—E. obscurum, ♀; B, E, H—E. joshuabelli, ♀; C, F—E. joshuabelli, ♂. A – C. Carapace frontal view showing clypeal hood. D – F. Chelicerae ectal view; G – H. Habitus, dorsal view." httpUri="https://zenodo.org/record/244966/files/figure.png" pageId="1" pageNumber="388">Figs. 10</figureCitation>
|
||
A–F). Chelicerae geniculate only in
|
||
<taxonomicName id="4C7B4D76FFD37F5FFCA5F80048CEF81D" box="[794,938,2003,2027]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="obscurum">
|
||
<emphasis id="B90FEAE7FFD37F5FFCA5F80048CEF81D" box="[794,938,2003,2027]" italics="true" pageId="1" pageNumber="388">E. obscurum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13402A70FFD37F5FFC02F8014F74F81D" box="[957,1040,2002,2027]" captionStart="FIGURE 10" captionStartId="11.[151,249,1296,1318]" captionTargetBox="[151,1436,157,1274]" captionTargetId="figure@11.[151,1436,157,1274]" captionTargetPageId="11" captionText="FIGURE 10. Emmenomma obscurum and E. joshuabelli sp. nov. A, D, G—E. obscurum, ♀; B, E, H—E. joshuabelli, ♀; C, F—E. joshuabelli, ♂. A – C. Carapace frontal view showing clypeal hood. D – F. Chelicerae ectal view; G – H. Habitus, dorsal view." httpUri="https://zenodo.org/record/244966/files/figure.png" pageId="1" pageNumber="388">Fig. 10</figureCitation>
|
||
A, D) and
|
||
<taxonomicName id="4C7B4D76FFD37F5FFB31F8004E7CF81D" box="[1166,1304,2003,2027]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD37F5FFB31F8004E7CF81D" box="[1166,1304,2003,2027]" italics="true" pageId="1" pageNumber="388">E. oculatum</emphasis>
|
||
</taxonomicName>
|
||
. Cheliceral promargin with four teeth, retromargin with variable number of teeth and denticles (
|
||
<figureCitation id="13402A70FFD07F5CFBF8FF444FEFFF46" box="[1095,1163,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1048]" captionTargetId="figure@3.[151,1436,193,1049]" captionTargetPageId="3" captionText="FIGURE 2. Emmenomma oculatum. A. Eyes, ♀; B. Eyes detail, AME and ALE, ♀; C. Tarsal organ, ♂ palp; D. Right endite, dorsal, ♀. E. Right chelicerae teeth, retromargin at top; F. Palpal tarsal claw, ♀." httpUri="https://zenodo.org/record/244958/files/figure.png" pageId="2" pageNumber="389">Fig. 2</figureCitation>
|
||
E), see details in species description. Labium slightly wider than long. Sternum longer than wide, projected between legs IV or not. Trochanter shallowly notched. Tracheal spiracle approximately as wide as colulus. Median tracheae branched and laterals simple. Ecribellate, colulus entire, wider than long, width approximately 1/4 of the distance between the outer margin of ALS (
|
||
<figureCitation id="13402A70FFD07F5CFE25FEF44A84FEB6" box="[410,480,295,320]" captionStart="FIGURE 7" captionStartId="8.[151,250,1903,1925]" captionTargetBox="[151,1436,589,1881]" captionTargetId="figure@8.[151,1436,589,1882]" captionTargetPageId="8" captionText="FIGURE 7. Emmenomma oculatum, spinnerets. A – D. ♂. A. Overview. B. Left ALS; C. Left PMS; D. Right PLS. E – I. ♀. E. Spinnerets overview (colulus indicated by arrow). F. Left ALS. G. PMS overview; H. Right PMS. I. Right PLS. Abbrev.: AC—aciniform gland spigots; Cy—cylindrical gland spigot; MAP—major ampullate gland spigot; Pi—piriform gland spigot." httpUri="https://zenodo.org/record/244963/files/figure.png" pageId="2" pageNumber="389">Fig. 7</figureCitation>
|
||
E). PLS slightly smaller than ALS but longer than PMS.
|
||
<collectingCountry id="F36C7665FFD07F5CFBC8FEF44FD5FEB6" box="[1143,1201,295,320]" name="Malta" pageId="2" pageNumber="389">Male</collectingCountry>
|
||
palp with triangular cymbium, longer than wide, cymbial apophysis present only in
|
||
<taxonomicName id="4C7B4D76FFD07F5CFCD7FE9E4F61FE92" box="[872,1029,332,356]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="joshuabelli" status="sp. nov.">
|
||
<emphasis id="B90FEAE7FFD07F5CFCD7FE9E4F61FE92" box="[872,1029,332,356]" italics="true" pageId="2" pageNumber="389">E. joshuabelli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAE7FFD07F5CFBB3FE9F4F03FE93" bold="true" box="[1036,1127,332,357]" pageId="2" pageNumber="389">
|
||
<taxonomicNameLabel id="A23C579CFFD07F5CFBB3FE9F4F03FE93" box="[1036,1127,332,357]" pageId="2" pageNumber="389" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
MA bifid with a triangular branch and a flat, almost rectangular, concave branch. E rising at prolateral side of bulb. TA present, located between DTP and MA. DTP membranous. DTA elbowed and retrolaterally projected with an opening for the tibial gland. RTA reduced, laminar, longer than wide and strongly sclerotized. iRTA bifid, with an external triangular and an internal laminate branch (
|
||
<figureCitation id="13402A70FFD07F5CFE68FE0F4941FE03" box="[471,549,476,501]" captionStart="FIGURE 6" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[158,1428,633,1578]" captionTargetId="figure@7.[151,1436,625,1588]" captionTargetPageId="7" captionText="FIGURE 6. Emmenomma oculatum. A – C: Left palp, ♂: A. Retrolateral view; B. Ventral view; C. Dorsal view. D – F: ♀ genitalia: D. Epigynum ventral view; E. Vulva dorsal view. Abbrev.: CD—copulatory ducts; DTA—dorsal tibial apophysis; DTP—distal tegular process; E—embolus; ET—epigynum tooth; FD—fertilization ducts; iRTA—internal branch of the RTA; MA—median apophysis; S—spermatheca." httpUri="https://zenodo.org/record/244962/files/figure.png" pageId="2" pageNumber="389">Figs. 6</figureCitation>
|
||
B; 9B–C). VTA distally rounded. PTA small and subtriangular, interacting with a small cymbial, prolateral projection (
|
||
<figureCitation id="13402A70FFD07F5CFDF8FE2C49F2FDEE" box="[583,662,511,536]" captionStart="FIGURE 4" captionStartId="5.[151,250,1862,1884]" captionTargetBox="[244,1337,193,1835]" captionTargetId="figure@5.[244,1345,193,1842]" captionTargetPageId="5" captionText="FIGURE 4. Emmenomma oculatum ♂. A. Right palp dorsal view. B. Left palp, retrolateral view showing RTA and DTA. C. Left palp, retrolateral view, iRTA. D. Left palp, expanded, retrolateral view, bulb. E. Left palp, expanded, prolateral view, bulb. F. Left palp, expanded, ventro-prolateral view. Abbrev.: DTA—dorsal tibial apophysis; DTP—distal tegular process; E—embolus; iRTA—internal branch of the retrolateral tibial apophysis; MA—median apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; TA—tegular apophysis TR—tegular rim; ST—subtegulum; VTA—ventral tibial apophysis." httpUri="https://zenodo.org/record/244960/files/figure.png" pageId="2" pageNumber="389">Figs. 4</figureCitation>
|
||
A; 8A–C). Female epigynum with teeth and well-developed posterior plate (
|
||
<figureCitation id="13402A70FFD07F5CFF5EFDF74A56FDCB" box="[225,306,548,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,933,955]" captionTargetBox="[151,1436,193,911]" captionTargetId="figure@6.[151,1436,193,912]" captionTargetPageId="6" captionText="FIGURE 5. Emmenomma oculatum ♀. A. Epigynum, ventral view. B. Epigynum, ventro-posterior view. C. Epigynum, posterior view; D. Vulva, left spermatheca. E. Vulva, dorsal view. F. Vulva, anterior view. Abbrev.: BG—Bennetts’s gland; CD—copulatory duct; LL—lateral lobe of the epigynum; MF—epigynum middle-field; S—spermathecae; ET- epigynum tooth; FD—fertilization duct." httpUri="https://zenodo.org/record/244961/files/figure.png" pageId="2" pageNumber="389">Figs. 5</figureCitation>
|
||
B–C; 6D; 8C; 9F). CD wall heavily sclerotized (
|
||
<figureCitation id="13402A70FFD07F5CFCC9FDF748D8FDCB" box="[886,956,548,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,933,955]" captionTargetBox="[151,1436,193,911]" captionTargetId="figure@6.[151,1436,193,912]" captionTargetPageId="6" captionText="FIGURE 5. Emmenomma oculatum ♀. A. Epigynum, ventral view. B. Epigynum, ventro-posterior view. C. Epigynum, posterior view; D. Vulva, left spermatheca. E. Vulva, dorsal view. F. Vulva, anterior view. Abbrev.: BG—Bennetts’s gland; CD—copulatory duct; LL—lateral lobe of the epigynum; MF—epigynum middle-field; S—spermathecae; ET- epigynum tooth; FD—fertilization duct." httpUri="https://zenodo.org/record/244961/files/figure.png" pageId="2" pageNumber="389">Fig. 5</figureCitation>
|
||
D–F; 6E; 8B; 9F). Spermathecae oval to mushroom-shaped (
|
||
<figureCitation id="13402A70FFD07F5CFECAFD944AD2FD96" box="[373,438,583,608]" captionStart="FIGURE 5" captionStartId="6.[151,250,933,955]" captionTargetBox="[151,1436,193,911]" captionTargetId="figure@6.[151,1436,193,912]" captionTargetPageId="6" captionText="FIGURE 5. Emmenomma oculatum ♀. A. Epigynum, ventral view. B. Epigynum, ventro-posterior view. C. Epigynum, posterior view; D. Vulva, left spermatheca. E. Vulva, dorsal view. F. Vulva, anterior view. Abbrev.: BG—Bennetts’s gland; CD—copulatory duct; LL—lateral lobe of the epigynum; MF—epigynum middle-field; S—spermathecae; ET- epigynum tooth; FD—fertilization duct." httpUri="https://zenodo.org/record/244961/files/figure.png" pageId="2" pageNumber="389">Fig. 5</figureCitation>
|
||
D–F; 6E; 8B; 9E).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361657EFFD07F5CFF78FDBF4F33FD06" pageId="2" pageNumber="389" type="discussion">
|
||
<paragraph id="8BC436F5FFD07F5CFF78FDBF4F33FD06" blockId="2.[151,1437,151,824]" pageId="2" pageNumber="389">
|
||
<emphasis id="B90FEAE7FFD07F5CFF78FDBF4A5FFD72" bold="true" box="[199,315,620,645]" pageId="2" pageNumber="389">Remarks.</emphasis>
|
||
A circular region of closely spaced pores, slightly everted, can be found on the distal end of the copulatory duct (
|
||
<figureCitation id="13402A70FFD07F5CFEE1FD5C4AC0FD5E" box="[350,420,655,680]" captionStart="FIGURE 5" captionStartId="6.[151,250,933,955]" captionTargetBox="[151,1436,193,911]" captionTargetId="figure@6.[151,1436,193,912]" captionTargetPageId="6" captionText="FIGURE 5. Emmenomma oculatum ♀. A. Epigynum, ventral view. B. Epigynum, ventro-posterior view. C. Epigynum, posterior view; D. Vulva, left spermatheca. E. Vulva, dorsal view. F. Vulva, anterior view. Abbrev.: BG—Bennetts’s gland; CD—copulatory duct; LL—lateral lobe of the epigynum; MF—epigynum middle-field; S—spermathecae; ET- epigynum tooth; FD—fertilization duct." httpUri="https://zenodo.org/record/244961/files/figure.png" pageId="2" pageNumber="389">Fig. 5</figureCitation>
|
||
D) of
|
||
<taxonomicName id="4C7B4D76FFD07F5CFE4EFD424919FD5E" box="[497,637,656,680]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD07F5CFE4EFD424919FD5E" box="[497,637,656,680]" italics="true" pageId="2" pageNumber="389">E. oculatum</emphasis>
|
||
</taxonomicName>
|
||
and seems to be compatible with what
|
||
<bibRefCitation id="EFEA4B04FFD07F5CFBE4FD5C4E2AFD5E" author="Ramirez" box="[1115,1358,655,680]" pageId="2" pageNumber="389" refString="Ramirez, M. J. (2014) The morphology and phylogeny of dionychan spiders (Araneae, Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramirez (2014: 270)</bibRefCitation>
|
||
called Bennett’s gland (BG). Although there is not enough evidence to corroborate or refute the homology between those features we decided to call this porous area a BG so it will be noted in further studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361657EFFD07F5DFF78FD2F4847FB3B" lastPageId="3" lastPageNumber="390" pageId="2" pageNumber="389" type="distribution">
|
||
<paragraph id="8BC436F5FFD07F5CFF78FD2F492FFCCE" blockId="2.[151,1437,151,824]" pageId="2" pageNumber="389">
|
||
<emphasis id="B90FEAE7FFD07F5CFF78FD2F4A04FCE2" bold="true" box="[199,352,764,789]" pageId="2" pageNumber="389">Distribution.</emphasis>
|
||
<collectingCountry id="F36C7665FFD07F5CFED6FD2F4947FCE3" box="[361,547,764,789]" name="Falkland Islands" pageId="2" pageNumber="389">Falkland Islands</collectingCountry>
|
||
(
|
||
<collectingCountry id="F36C7665FFD07F5CFD8CFD2F49C5FCE3" box="[563,673,764,789]" name="Falkland Islands" pageId="2" pageNumber="389">Malvinas</collectingCountry>
|
||
), southern
|
||
<collectingCountry id="F36C7665FFD07F5CFCA0FD2F4838FCE3" box="[799,860,764,789]" name="Chile" pageId="2" pageNumber="389">Chile</collectingCountry>
|
||
and
|
||
<collectingCountry id="F36C7665FFD07F5CFC29FD2F4F61FCE3" box="[918,1029,764,789]" name="Argentina" pageId="2" pageNumber="389">Argentina</collectingCountry>
|
||
including islands around Tierra del Fuego—see distribution map (
|
||
<figureCitation id="13402A70FFD07F5CFE54FCCC4959FCCE" box="[491,573,799,824]" captionStart="FIGURE 11" captionStartId="12.[151,250,1663,1685]" captionTargetBox="[155,1431,781,1592]" captionTargetId="figure@12.[151,1436,733,1642]" captionTargetPageId="12" captionText="FIGURE 11. Distribution map for species of Emmenomma. E. obscurum (black star); E. joshuabelli (black triangle) and E. oculatum (white squares)." httpUri="https://zenodo.org/record/244967/files/figure.png" pageId="2" pageNumber="389">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF04667DFFD07F5CFF28F87C487BF815" httpUri="https://zenodo.org/record/244957/files/figure.png" pageId="2" pageNumber="389" targetBox="[204,1382,879,1942]" targetPageId="2">
|
||
<paragraph id="8BC436F5FFD07F5CFF28F87C487BF815" blockId="2.[151,1436,1967,2020]" pageId="2" pageNumber="389">
|
||
<emphasis id="B90FEAE7FFD07F5CFF28F87C4A71F832" bold="true" box="[151,277,1967,1989]" pageId="2" pageNumber="389">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C7B4D76FFD07F5CFEA2F863496AF833" box="[285,526,1967,1989]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD07F5CFEA2F863496AF833" box="[285,526,1967,1989]" italics="true" pageId="2" pageNumber="389">Emmenomma oculatum</emphasis>
|
||
</taxonomicName>
|
||
. A–C: ♂ (recently molted); D: Immature. A. Carapace frontal view; B. Habitus, dorsal view; C. Habitus ventral view; D. Tapetum removed from ALE.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF04667DFFD17F5DFF28FBFC48DDFB95" httpUri="https://zenodo.org/record/244958/files/figure.png" pageId="3" pageNumber="390" targetBox="[151,1436,193,1048]" targetPageId="3">
|
||
<paragraph id="8BC436F5FFD17F5DFF28FBFC48DDFB95" blockId="3.[151,1436,1071,1123]" pageId="3" pageNumber="390">
|
||
<emphasis id="B90FEAE7FFD17F5DFF28FBFC4A77FBB2" bold="true" box="[151,275,1071,1093]" pageId="3" pageNumber="390">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7B4D76FFD17F5DFEA5FBE3496DFBB3" box="[282,521,1071,1093]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD17F5DFEA5FBE3496DFBB3" box="[282,521,1071,1093]" italics="true" pageId="3" pageNumber="390">Emmenomma oculatum</emphasis>
|
||
</taxonomicName>
|
||
. A. Eyes, ♀; B. Eyes detail, AME and ALE, ♀; C. Tarsal organ, ♂ palp; D. Right endite, dorsal, ♀. E. Right chelicerae teeth, retromargin at top; F. Palpal tarsal claw, ♀.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC436F5FFD17F5DFF78FB434847FB3B" blockId="3.[151,1436,1168,1229]" pageId="3" pageNumber="390">
|
||
<emphasis id="B90FEAE7FFD17F5DFF78FB434A02FB5E" bold="true" box="[199,358,1168,1193]" pageId="3" pageNumber="390">Composition.</emphasis>
|
||
<taxonomicName id="4C7B4D76FFD17F5DFECCFB424846FB5E" authority="Simon, 1884" authorityName="Simon" authorityYear="1884" box="[371,802,1168,1193]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="oculatum">
|
||
<emphasis id="B90FEAE7FFD17F5DFECCFB4249E4FB5E" box="[371,640,1168,1192]" italics="true" pageId="3" pageNumber="390">Emmenomma oculatum</emphasis>
|
||
<bibRefCitation id="EFEA4B04FFD17F5DFD32FB434846FB5E" author="Simon" box="[653,802,1168,1193]" pageId="3" pageNumber="390" refString="Simon, E. (1884) Arachnides recueillis par la Mission du Cap Horn en 1882 - 1883. Bulletin de la Societe Zoologique de France, 9, 117 - 144." type="journal article" year="1884">Simon, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C7B4D76FFD17F5DFC87FB424F09FB5F" authority="Simon, 1905" authorityName="Simon" authorityYear="1905" box="[824,1133,1168,1193]" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="obscurum">
|
||
<emphasis id="B90FEAE7FFD17F5DFC87FB4248AFFB5E" box="[824,971,1168,1192]" italics="true" pageId="3" pageNumber="390">E. obscurum</emphasis>
|
||
<bibRefCitation id="EFEA4B04FFD17F5DFC67FB434F09FB5F" author="Simon" box="[984,1133,1168,1193]" pageId="3" pageNumber="390" refString="Simon, E. (1905) Etude sur les arachnides recueillis en Patagonie par le Dr. Fillipo Silvestri. Bollettino dei Musei di Zoologia e di Anatomia Comparata della Reale Universita di Torino, 20 (511), 1 - 17." type="journal article" year="1905">Simon, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
sp. reval.,
|
||
<emphasis id="B90FEAE7FFD17F5DFB46FB434E0BFB5F" bold="true" box="[1273,1391,1168,1193]" pageId="3" pageNumber="390">new rank</emphasis>
|
||
;
|
||
<taxonomicName id="4C7B4D76FFD17F5DFA3BFB4249A7FB3B" authority="Almeida-Silva, Griswold & Brescovit" authorityName="Almeida-Silva, Griswold & Brescovit" class="Arachnida" family="Amaurobiidae" genus="Emmenomma" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="joshuabelli" status="sp. nov.">
|
||
<emphasis id="B90FEAE7FFD17F5DFA3BFB424A70FB3A" italics="true" pageId="3" pageNumber="390">E. joshuabelli</emphasis>
|
||
Almeida-Silva, Griswold & Brescovit
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAE7FFD17F5DFD75FB674847FB3B" bold="true" box="[714,803,1204,1229]" pageId="3" pageNumber="390">
|
||
<taxonomicNameLabel id="A23C579CFFD17F5DFD75FB674847FB3B" box="[714,803,1204,1229]" pageId="3" pageNumber="390" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |