142 lines
18 KiB
XML
142 lines
18 KiB
XML
<document id="9ECD8ABA92A3383DE49C269F21B21D0D" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFF97AF3ED1DBDCC3FE1A1131" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Nunoidium Vaz-De-Mello 2008, new genus" docType="treatment" docVersion="3" lastPageNumber="37" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="36" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="81B53AA799CE3172BF8568A7037E8D36">
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<mods:title id="1CE43FF50A7B145E9B4016C87DFD09C1">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
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<mods:name id="AC7874D5F2CD7A14F69AD3C1A892CA3F" type="personal">
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<mods:role id="CFE16F6E05E697A1403F1D0CE61AD07E">
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<mods:roleTerm id="654572464D475BD66BB9E194AD8840A7">Author</mods:roleTerm>
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<mods:namePart id="AF4BC39EABF835596793224D0AA85538">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="0144F6AEDF5D6B5776650174BAFA9497">text</mods:typeOfResource>
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<mods:titleInfo id="3A745BFE4CFD938CD73D9A55E0E92BA9">
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<mods:title id="653012FCD091710771B10D8DDEF24E83">Zootaxa</mods:title>
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<mods:part id="F9B9D012408019F5AEE4BD2BE4C2F7B3">
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<mods:date id="AF75A5CD7C940F99E183D1E048B6A9CF">2008</mods:date>
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<mods:detail id="0A27957DB1A454963D5B68AA4EECCB96" type="pubDate">
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<mods:number id="618492818B67F404EDB0619C81629C6F">2008-12-05</mods:number>
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<mods:detail id="6A7C587E17D50AAE6ED07F6D44F5C314" type="volume">
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<mods:number id="222B74B9A425619EC19CBA221B2024AA">1955</mods:number>
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<mods:start id="C2285DA8CB31BC7143BA5E1924029A3D">1</mods:start>
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<mods:end id="3C61EB5BE48F34241B57DA3FD98A7456">75</mods:end>
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<mods:classification id="A50829BDFFBB7A3F8DDD5DF3AC792117">journal article</mods:classification>
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<mods:identifier id="15DD4EA5DF945D741F4A13C37466F6A1" type="ISSN">11755334</mods:identifier>
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<treatment id="03D287CEFF97AF3ED1DBDCC3FE1A1131" ID-DOI="http://doi.org/10.5281/zenodo.5242027" ID-Zenodo-Dep="5242027" LSID="urn:lsid:plazi:treatment:03D287CEFF97AF3ED1DBDCC3FE1A1131" httpUri="http://treatment.plazi.org/id/03D287CEFF97AF3ED1DBDCC3FE1A1131" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
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<subSubSection id="C3616553FF97AF39D1DBDCC3FE681089" box="[151,462,951,977]" pageId="35" pageNumber="36" type="nomenclature">
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<paragraph id="8BC436D8FF97AF39D1DBDCC3FE681089" blockId="35.[151,462,951,977]" box="[151,462,951,977]" pageId="35" pageNumber="36">
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<heading id="D08C81B4FF97AF39D1DBDCC3FE681089" bold="true" box="[151,462,951,977]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
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<emphasis id="B90FEACAFF97AF39D1DBDCC3FE681089" bold="true" box="[151,462,951,977]" pageId="35" pageNumber="36">
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9.
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<taxonomicName id="4C7B4D5BFF97AF39D1F8DCC3FE981089" ID-CoL="65WB" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[180,318,951,977]" class="Insecta" family="Scarabaeidae" genus="Nunoidium" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90FEACAFF97AF39D1F8DCC3FE981089" bold="true" box="[180,318,951,977]" italics="true" pageId="35" pageNumber="36">Nunoidium</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23C57B1FF97AF39D001DCC3FE681089" box="[333,462,951,977]" pageId="35" pageNumber="36" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF97AF39D1DBDB76FE3717E4" pageId="35" pageNumber="36" type="diagnosis">
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<paragraph id="8BC436D8FF97AF39D1DBDB76FE3717E4" blockId="35.[151,1436,1026,2012]" pageId="35" pageNumber="36">
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<emphasis id="B90FEACAFF97AF39D1DBDB76FEBF1744" bold="true" box="[151,281,1026,1052]" pageId="35" pageNumber="36">Diagnosis:</emphasis>
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Body dark, elongated (
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<figureCitation id="13402A5DFF97AF39D370DB76FD311744" box="[572,663,1026,1052]" captionStart="FIGURES 23–30" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 23–30. Dorsal views. 23. Leotrichillum louzadaorum; 24–25: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="35" pageNumber="36">Fig. 26</figureCitation>
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). Clypeus with two short triangular teeth separated by wide Vshaped emargination. Eyes, in dorsal view, as wide as two-fifths of their length, interocular width seven to eight times eye width (
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<figureCitation id="13402A5DFF97AF39D0D3DB26FE511734" box="[415,503,1106,1132]" captionStart="FIGURES 71–78" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 71–78. 71. Genieridium zanunciorum, parameres; 72–73: Leotrichillum louzadaorum: 72. head; 73. parameres; 74–76: Martinezidium: 74–75: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 77–78: Nunoidium argentinum: 77. head; 78. parameres." pageId="35" pageNumber="36">Fig. 77</figureCitation>
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). Pronotum and hypomeron separated by strong carina, pronotum with posterior bead. Mesofemora and metafemora oval, mesotibiae and metatibiae strongly widened apically. Male claws and tarsi unmodified.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF97AF39D18ADBBEFB0E1484" pageId="35" pageNumber="36" type="description">
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<paragraph id="8BC436D8FF97AF39D18ADBBEFF7F14D4" blockId="35.[151,1436,1026,2012]" pageId="35" pageNumber="36">
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<emphasis id="B90FEACAFF97AF39D18ADBBEFEC617BC" bold="true" box="[198,352,1226,1252]" pageId="35" pageNumber="36">Description:</emphasis>
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Length 3.2–5.0 mm, body elongated, parallel sided (
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<figureCitation id="13402A5DFF97AF39D2ABDBBEFBE717BC" box="[999,1089,1226,1252]" captionStart="FIGURES 23–30" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 23–30. Dorsal views. 23. Leotrichillum louzadaorum; 24–25: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="35" pageNumber="36">Fig. 26</figureCitation>
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), color dark brown to black, shiny, without metallic sheen. Clypeus with two triangular upturned teeth, separated by wide V-shaped emargination, laterally rounded, margin continuous with that of gena. Clypeofrontal and clypeogenal sutures indistinct. Eyes, in dorsal view, as wide as two-fifths of their length; interocular width seven to eight times eye width (
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<figureCitation id="13402A5DFF97AF39D1BDDA1EFEF616DC" box="[241,336,1386,1412]" captionStart="FIGURES 71–78" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 71–78. 71. Genieridium zanunciorum, parameres; 72–73: Leotrichillum louzadaorum: 72. head; 73. parameres; 74–76: Martinezidium: 74–75: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 77–78: Nunoidium argentinum: 77. head; 78. parameres." pageId="35" pageNumber="36">Fig. 77</figureCitation>
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). Pronotum lacking anterior bead, but with uninterrupted posterior one; separated from hypomeron by smooth carina; lateral callosity weakly defined, elongated; disc covered by setose punctures. Hypomeron with strong lateral longitudinal carina; mesoepimera with anterior transverse carina; metasternum covered by simple setose punctures on disc. Elytral discal interstriae (except sutural interstria) with biseriate setose punctures, interstriae flat on disc, mesal ones slightly convex on declivity; striae deeper and wider on declivity than on disc. Pseudoepipleura gradually narrowed posteriorly, with continuous row of punctures. Protibiae with three strong teeth, distributed along apical three-fifths of length, denticulate at base, with ventral scale-like setae on teeth. Mesofemora elongated, oval, as long as three times their medial width; metafemora short, oval, as long as twice its medial width; mesotibiae and metatibiae strongly widened apically; mesotibiae apically as wide as third of its length, metatibiae as long as twice apical width. Pygidium vertical, transverse. Parameres flattened (
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<figureCitation id="13402A5DFF97AF39D351D98EFDDE144C" box="[541,632,1786,1812]" captionStart="FIGURES 71–78" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 71–78. 71. Genieridium zanunciorum, parameres; 72–73: Leotrichillum louzadaorum: 72. head; 73. parameres; 74–76: Martinezidium: 74–75: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 77–78: Nunoidium argentinum: 77. head; 78. parameres." pageId="35" pageNumber="36">Fig. 78</figureCitation>
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), shorter than half length of phallobase, right-angled in relation to phallobase. Internal sac with helicoid pseudoflagellum and three flattened elongated acessory lamellae. Coxites symmetrical, rectangular, mesally toothed. Spermatheca very elongated, C-shaped, apically slightly spiraled.
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</paragraph>
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<paragraph id="8BC436D8FF97AF39D18AD8EEFB0E1484" blockId="35.[151,1436,1026,2012]" pageId="35" pageNumber="36">
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<emphasis id="B90FEACAFF97AF39D18AD8EEFE6114EC" bold="true" box="[198,455,1946,1972]" pageId="35" pageNumber="36">Sexual dimorphism:</emphasis>
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Males with apicomesal tooth in all tibiae; metasternum slightly concave (flat in females), abdominal disc flat (slightly concave in females) and pygidium less transverse.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF90AF3ED18ADFE3FEB61381" pageId="36" pageNumber="37" type="type_taxon">
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<paragraph id="8BC436D8FF90AF3ED18ADFE3FEB61381" blockId="36.[151,1436,151,617]" pageId="36" pageNumber="37">
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<emphasis id="B90FEACAFF90AF3ED18ADFE3FECD13E9" bold="true" box="[198,363,151,177]" pageId="36" pageNumber="37">
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<typeStatus id="54C0887AFF90AF3ED18ADFE3FEA513E9" box="[198,259,151,177]" pageId="36" pageNumber="37">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C7B4D5BFF90AF3ED038DFECFC8E13EA" ID-CoL="768RN" authority="Arrow, 1913" authorityName=": Arrow" authorityYear="1913" box="[372,808,152,178]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="argentinum">
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<emphasis id="B90FEACAFF90AF3ED038DFECFD2A13E9" box="[372,652,152,177]" italics="true" pageId="36" pageNumber="37">Pedaridium argentinum</emphasis>
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<bibRefCitation id="EFEA4B29FF90AF3ED3D8DFECFC8E13EA" author="Arrow, G. J." box="[660,808,152,178]" pageId="36" pageNumber="37" pagination="456 - 466" refId="ref47114" refString="Arrow, G. J. (1913) Some new species of Lamellicorn beetles from Brazil. The Annals and Magazine of Natural History, Series 8, 11, 456 - 466." type="journal article" year="1913">Arrow, 1913</bibRefCitation>
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</taxonomicName>
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(=
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<taxonomicName id="4C7B4D5BFF90AF3ED218DFECFAB113EA" ID-CoL="48469" authority="(Arrow, 1913)" authorityName="Vaz-De-Mello" authorityYear="2008" baseAuthorityName="Vaz-De-Mello" baseAuthorityYear="1913" box="[852,1303,152,178]" class="Insecta" family="Scarabaeidae" genus="Nunoidium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="argentinum" status="comb. nov.">
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<emphasis id="B90FEACAFF90AF3ED218DFECFBC213E9" box="[852,1124,152,177]" italics="true" pageId="36" pageNumber="37">Nunoidium argentinum</emphasis>
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(
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<bibRefCitation id="EFEA4B29FF90AF3ED538DFECFAA813EA" author="Arrow, G. J." box="[1140,1294,152,178]" pageId="36" pageNumber="37" pagination="456 - 466" refId="ref47114" refString="Arrow, G. J. (1913) Some new species of Lamellicorn beetles from Brazil. The Annals and Magazine of Natural History, Series 8, 11, 456 - 466." type="journal article" year="1913">Arrow, 1913</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="B90FEACAFF90AF3ED468DFE3FF591381" bold="true" pageId="36" pageNumber="37">
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<taxonomicNameLabel id="A23C57B1FF90AF3ED468DFE3FF591381" pageId="36" pageNumber="37" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF90AF3ED18ADF93FBD5125A" box="[198,1139,231,258]" pageId="36" pageNumber="37" type="etymology">
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<paragraph id="8BC436D8FF90AF3ED18ADF93FBD5125A" blockId="36.[151,1436,151,617]" box="[198,1139,231,258]" pageId="36" pageNumber="37">
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<emphasis id="B90FEACAFF90AF3ED18ADF93FEF11259" bold="true" box="[198,343,231,257]" pageId="36" pageNumber="37">Etymology:</emphasis>
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Named after my son, Nuno Falqueto Vaz de Mello. Gender neutral.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF90AF3ED18ADE7BFD28120A" pageId="36" pageNumber="37" type="distribution">
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<paragraph id="8BC436D8FF90AF3ED18ADE7BFD28120A" blockId="36.[151,1436,151,617]" pageId="36" pageNumber="37">
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<emphasis id="B90FEACAFF90AF3ED18ADE7BFECF1271" bold="true" box="[198,361,271,297]" pageId="36" pageNumber="37">Distribution:</emphasis>
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Central, northern and northwestern
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<collectingCountry id="F36C7648FF90AF3ED255DE64FC361272" box="[793,912,272,298]" name="Argentina" pageId="36" pageNumber="37">Argentina</collectingCountry>
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, occupying about the Southern
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<collectingRegion id="49BFF83AFF90AF3ED445DE64FF6E120A" country="Argentina" name="Chaco" pageId="36" pageNumber="37">Chaco province</collectingRegion>
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of the Chacoan subgregion (
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<figureCitation id="13402A5DFF90AF3ED356DE4CFD27120A" box="[538,641,312,338]" captionStart="FIGURES 107–108" captionStartId="69.[151,269,1969,1993]" captionTargetBox="[166,1417,182,1945]" captionTargetId="figure-37@69.[166,1420,182,1945]" captionTargetPageId="69" captionText="FIGURES 107–108. Distribution maps. 107. Genieridium medinae (circles) and G. bordoni (triangles); 108. Nunoidium argentinum (circles), Pereiraidium almeidai (squares) and Silvinha unica (triangles)." pageId="36" pageNumber="37">Fig. 108</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF90AF3ED18ADE2BFD4A111A" pageId="36" pageNumber="37" type="discussion">
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<paragraph id="8BC436D8FF90AF3ED18ADE2BFE271142" blockId="36.[151,1436,151,617]" pageId="36" pageNumber="37">
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<emphasis id="B90FEACAFF90AF3ED18ADE2BFEE71221" bold="true" box="[198,321,351,377]" pageId="36" pageNumber="37">Remarks:</emphasis>
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<taxonomicName id="4C7B4D5BFF90AF3ED005DE14FE6D1221" authorityName="Vaz-De-Mello" authorityYear="2008" box="[329,459,352,377]" class="Insecta" family="Scarabaeidae" genus="Nunoidium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFF90AF3ED005DE14FE6D1221" box="[329,459,352,377]" italics="true" pageId="36" pageNumber="37">Nunoidium</emphasis>
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</taxonomicName>
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appears to be closely related to
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<taxonomicName id="4C7B4D5BFF90AF3ED20DDE14FC731221" authorityName="Vaz-De-Mello" authorityYear="2008" box="[833,981,352,377]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFF90AF3ED20DDE14FC731221" box="[833,981,352,377]" italics="true" pageId="36" pageNumber="37">Genieridium</emphasis>
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</taxonomicName>
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, however, the completeness of the carina separating pronotal disc from hypomeron, vertical pygidial position, and presence of posterior pronotal bead, among other characters, will readily separate it. Apomorphies of this genus are the presence of posterior pronotal bead, absence of modification in male claws, elongation of spermatheca, and psammophilous modifications of metalegs.
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</paragraph>
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<paragraph id="8BC436D8FF90AF3ED18ADD53FD4A111A" blockId="36.[151,1436,151,617]" box="[198,748,551,578]" pageId="36" pageNumber="37">
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||
<emphasis id="B90FEACAFF90AF3ED18ADD53FEC91119" bold="true" box="[198,367,551,577]" pageId="36" pageNumber="37">Composition:</emphasis>
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||
Only the
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<typeStatus id="54C0887AFF90AF3ED0A8DD5CFDB0111A" box="[484,534,552,578]" pageId="36" pageNumber="37">type</typeStatus>
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species is known.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FF90AF3ED18ADD3BFE1A1131" box="[198,444,591,617]" pageId="36" pageNumber="37" type="materials_examined">
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<paragraph id="8BC436D8FF90AF3ED18ADD3BFE1A1131" blockId="36.[151,1436,151,617]" box="[198,444,591,617]" pageId="36" pageNumber="37">
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||
<emphasis id="B90FEACAFF90AF3ED18ADD3BFE1A1131" bold="true" box="[198,444,591,617]" pageId="36" pageNumber="37">Material examined:</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |