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<mods:title id="FE29374DF32F42F6EF34DA095C9993A3">Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species</mods:title>
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<treatment id="03D287A2FD183715FEF7FF22FAB1F996" ID-DOI="http://doi.org/10.5281/zenodo.5630040" ID-GBIF-Taxon="127668907" ID-Zenodo-Dep="5630040" LSID="urn:lsid:plazi:treatment:03D287A2FD183715FEF7FF22FAB1F996" httpUri="http://treatment.plazi.org/id/03D287A2FD183715FEF7FF22FAB1F996" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="C361653FFD183715FEF7FF22FDB2FF16" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BC436B4FD183715FEF7FF22FE63FF77" blockId="17.[151,558,151,211]" box="[151,511,151,178]" pageId="17" pageNumber="18">
<heading id="D08C81D8FD183715FEF7FF22FE63FF77" bold="true" box="[151,511,151,178]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<taxonomicName id="4C7B4D37FD183715FEF7FF22FE06FF74" box="[151,410,151,177]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="mazzai" status="sp. nov.">
<emphasis id="B90FEAA6FD183715FEF7FF22FE06FF74" bold="true" box="[151,410,151,177]" italics="true" pageId="17" pageNumber="18">Scontromeryx mazzai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57DDFD183715FFC1FF2DFE63FF77" box="[417,511,152,178]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC436B4FD183715FEF7FF0FFDB2FF16" blockId="17.[151,558,151,211]" box="[151,558,186,211]" pageId="17" pageNumber="18">
<figureCitation id="13402A31FD183715FEF7FF0FFF4BFF16" box="[151,215,186,211]" captionStart="FIGURE 6" captionStartId="12.[151,250,1824,1846]" captionTargetBox="[205,1378,205,1796]" captionTargetId="figure@12.[196,1392,193,1803]" captionTargetPageId="12" captionText="FIGURE 6. Postcranial elements of Hoplitomeryx. A, left tibia (RGM 335.882 + 425.328; San Giovannino): medial view; B, right tibia (RGM 425.144; Chiro D 1): anterior view. Astragals of different size in anterior (dorsal) view: C, left astragal (RGM 425.385; Falcone 2 B); D, left astragal (RGM 260.863; San Giovannino); E, right astragal (RGM 425.421; Fina N). F, left astragal (RGM 263.951; Gargano, unknown fissure): lateral view; G, left calcaneum (RGM 260.901; San Giovannino): anterior view; H, same specimen: posterior view; I, same specimen: medial view; J,; K, detail of distal right metatarsal (RGM 178.518; San Giovannino) showing the proximal extension of the dorsal gully into the (fused) cubonavicular." httpUri="https://zenodo.org/record/286821/files/figure.png" pageId="17" pageNumber="18">Fig. 6</figureCitation>
H in
<bibRefCitation id="EFEA4B45FD183715FF70FF0EFDB2FF16" author="Mazza" box="[272,558,186,211]" pageId="17" pageNumber="18" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni (2011)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD183715FEF7FEB7FE48FE77" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BC436B4FD183715FEF7FEB7FD4DFEF2" blockId="17.[151,1436,258,434]" pageId="17" pageNumber="18">
<taxonomicName id="4C7B4D37FD183715FEF7FEB7FEBEFEDD" box="[151,290,258,280]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B90FEAA6FD183715FEF7FEB7FEBEFEDD" box="[151,290,258,280]" italics="true" pageId="17" pageNumber="18">Hoplitomeryx</emphasis>
</taxonomicName>
sp.
<bibRefCitation id="EFEA4B45FD183715FF2FFEB7FE75FEDD" author="Leinders" box="[335,489,258,280]" pageId="17" pageNumber="18" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
<bibRefCitation id="EFEA4B45FD183715FF9EFEB7FD52FEDD" author="Rustioni" box="[510,718,258,280]" pageId="17" pageNumber="18" refString="Rustioni, M., Mazza, P., Azzaroli, A., Boscagli, G., Cozzini, F., Di Vito, E., Masseti, M. &amp; Pisano, A. (1992) Miocene Vertebrate remains from Scontrone, National Park of Abruzzi, Central Italy. Rendiconti della Classe Scienze Fisiche, Matematiche e Naturali dell'Accademia dei Lincei, 9, 227 - 237." type="journal article" year="1992">
Rustioni
<emphasis id="B90FEAA6FD183715FC3CFEB6FD16FEDD" box="[604,650,258,280]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. 1992
</bibRefCitation>
: fig. 3 [uncatalogued skull fragment];
<bibRefCitation id="EFEA4B45FD183715FA3DFEB6FAC5FEDD" author="Mazza" box="[1117,1369,258,280]" pageId="17" pageNumber="18" refString="Mazza, P. &amp; Rustioni, M. (1996) The Turolian fossil Artiodactyls from Scontrone (Abruzzo, Central Italy) and their paleoecological and paleoenvironmental implications. Bollettino della Societa Paleontologica Italiana, 35, 93 - 106." type="journal article" year="1996">Mazza &amp; Rustioni 1996</bibRefCitation>
: fig. 1 [SCT17, redrawn after
<bibRefCitation id="EFEA4B45FD183715FFD1FE94FD18FEF2" author="Rustioni" box="[433,644,289,311]" pageId="17" pageNumber="18" refString="Rustioni, M., Mazza, P., Azzaroli, A., Boscagli, G., Cozzini, F., Di Vito, E., Masseti, M. &amp; Pisano, A. (1992) Miocene Vertebrate remains from Scontrone, National Park of Abruzzi, Central Italy. Rendiconti della Classe Scienze Fisiche, Matematiche e Naturali dell'Accademia dei Lincei, 9, 227 - 237." type="journal article" year="1992">
Rustioni
<emphasis id="B90FEAA6FD183715FC6EFE97FDA6FEF2" box="[526,570,289,311]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., 1992
</bibRefCitation>
: fig. 1].
</paragraph>
<paragraph id="8BC436B4FD183715FEF7FEF4FCF9FE93" blockId="17.[151,1436,258,434]" box="[151,869,320,342]" pageId="17" pageNumber="18">
Taxon C—Mazza &amp;
<bibRefCitation id="EFEA4B45FD183715FF0AFEF5FE65FE93" author="Rustioni" box="[362,505,320,342]" pageId="17" pageNumber="18" refString="Mazza, P. &amp; Rustioni, M. (1999) Dental characters of the Turolian fossil artiodactyls from Scontrone (Abruzzo, Central Italy). In: Mayhall, J. &amp; Heikkinen, T. (Eds.), Proceedings of the 11 th International Symposium on Dental Morphology, 26 - 30 August 1998, Oulu (Finland). Oulu University, Oulu, pp. 300 - 309." type="journal article" year="1999">Rustioni 1999</bibRefCitation>
(partim): p. 307 [STC 72, STC 83].
</paragraph>
<paragraph id="8BC436B4FD183715FEF7FEEBFC44FEB1" blockId="17.[151,1436,258,434]" box="[151,984,350,372]" pageId="17" pageNumber="18">
<taxonomicName id="4C7B4D37FD183715FEF7FEEBFE22FEB1" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,446,350,372]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B90FEAA6FD183715FEF7FEEBFEBEFEB1" box="[151,290,350,372]" italics="true" pageId="17" pageNumber="18">Hoplitomeryx</emphasis>
<bibRefCitation id="EFEA4B45FD183715FF48FEEBFE22FEB1" author="Leinders" box="[296,446,350,372]" pageId="17" pageNumber="18" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
</taxonomicName>
<bibRefCitation id="EFEA4B45FD183715FFB9FEEBFD50FEB1" author="Mazza" box="[473,716,350,372]" pageId="17" pageNumber="18" refString="Mazza, P. &amp; Rustioni, M. (1999) Dental characters of the Turolian fossil artiodactyls from Scontrone (Abruzzo, Central Italy). In: Mayhall, J. &amp; Heikkinen, T. (Eds.), Proceedings of the 11 th International Symposium on Dental Morphology, 26 - 30 August 1998, Oulu (Finland). Oulu University, Oulu, pp. 300 - 309." type="journal article" year="1999">Mazza &amp; Rustioni 1999</bibRefCitation>
(partim): p. 305 [SCT80].
</paragraph>
<paragraph id="8BC436B4FD183715FEF7FEC8FE48FE77" blockId="17.[151,1436,258,434]" pageId="17" pageNumber="18">
<taxonomicName id="4C7B4D37FD183715FEF7FEC8FD87FE56" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,539,381,403]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="matthei">
<emphasis id="B90FEAA6FD183715FEF7FEC8FEE4FE56" box="[151,376,381,403]" italics="true" pageId="17" pageNumber="18">Hoplitomeryx matthei</emphasis>
<bibRefCitation id="EFEA4B45FD183715FFE2FEC8FD87FE56" author="Leinders" box="[386,539,381,403]" pageId="17" pageNumber="18" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders, 1984</bibRefCitation>
</taxonomicName>
<bibRefCitation id="EFEA4B45FD183715FC56FEC8FCAEFE56" author="Mazza" box="[566,818,381,403]" pageId="17" pageNumber="18" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni 2011</bibRefCitation>
(partim): p. 1320, 1321, 1330, fig. 6, tables 1, 2 [SCT17, SCT72, SC T 80, SCT83].
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD183715FEF7FE6BFCC9FDDE" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BC436B4FD183715FEF7FE6BFD8BFE32" blockId="17.[151,1437,478,1619]" box="[151,535,478,503]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEF7FE6BFE97FE32" bold="true" box="[151,267,478,503]" pageId="17" pageNumber="18">
<typeStatus id="54C08816FD183715FEF7FE6BFE9AFE32" box="[151,262,478,503]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
Skull fragment SCT17.
</paragraph>
<paragraph id="8BC436B4FD183715FEA7FDB7FCC9FDDE" blockId="17.[151,1437,478,1619]" box="[199,853,514,539]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FDB7FEDAFDDE" bold="true" box="[199,326,514,539]" pageId="17" pageNumber="18">
<typeStatus id="54C08816FD183715FEA7FDB7FEDDFDDE" box="[199,321,514,539]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Mandible fragments SC T 80, SCT83, SCT72.
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD183715FEA7FD93FA9FFCFE" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BC436B4FD183715FEA7FD93FA06FDA6" blockId="17.[151,1437,478,1619]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FD93FEDDFDFA" bold="true" box="[199,321,550,575]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
A hoplitomerycid with two pairs of orbital horns. Mandible markedly pachyostotic with brachyodont dentition. The lower m2 and m3 lack ectostylids, a cingulum and cementum and have rugose enamel.
</paragraph>
<paragraph id="8BC436B4FD183715FEA7FDDBFC38FD36" blockId="17.[151,1437,478,1619]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FDDBFE4CFD42" bold="true" box="[199,464,622,647]" pageId="17" pageNumber="18">Differential diagnosis.</emphasis>
Differs from
<taxonomicName id="4C7B4D37FD183715FC11FDDAFD74FD43" box="[625,744,623,646]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B90FEAA6FD183715FC11FDDAFD74FD43" box="[625,744,623,646]" italics="true" pageId="17" pageNumber="18">S. minutus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B4D37FD183715FC98FDDAFCE0FD43" box="[760,892,622,646]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="falcidens">
<emphasis id="B90FEAA6FD183715FC98FDDAFCE0FD43" box="[760,892,622,646]" italics="true" pageId="17" pageNumber="18">S. falcidens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D37FD183715FDD7FDDAFBA8FD43" box="[951,1076,622,646]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="apulicus">
<emphasis id="B90FEAA6FD183715FDD7FDDAFBA8FD43" box="[951,1076,622,646]" italics="true" pageId="17" pageNumber="18">S. apulicus</emphasis>
</taxonomicName>
by pachyostosis of mandibular ramus and presence of orbital horns, differs from
<taxonomicName id="4C7B4D37FD183715FCBDFD26FCFDFD6E" box="[733,865,659,683]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="falcidens">
<emphasis id="B90FEAA6FD183715FCBDFD26FCFDFD6E" box="[733,865,659,683]" italics="true" pageId="17" pageNumber="18">S. falcidens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D37FD183715FDFEFD26FB81FD6E" box="[926,1053,659,683]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="apulicus">
<emphasis id="B90FEAA6FD183715FDFEFD26FB81FD6E" box="[926,1053,659,683]" italics="true" pageId="17" pageNumber="18">S. apulicus</emphasis>
</taxonomicName>
by rugose enamel on its molars, differs from
<taxonomicName id="4C7B4D37FD183715FF46FD02FE34FD0B" box="[294,424,694,718]" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="falcidens">
<emphasis id="B90FEAA6FD183715FF46FD02FE34FD0B" box="[294,424,694,718]" italics="true" pageId="17" pageNumber="18">S. falcidens</emphasis>
</taxonomicName>
by absence of ectostylid in m2 and m3 and by having brachyodont teeth. Differs from
<taxonomicName id="4C7B4D37FD183715FBE7FD02FF6CFD37" class="Mammalia" family="Hoplitomerycidae" genus="Scontromeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="minutus">
<emphasis id="B90FEAA6FD183715FBE7FD02FF6CFD37" italics="true" pageId="17" pageNumber="18">S. minutus</emphasis>
</taxonomicName>
by absence of a cingulum and by rugose lingual enamel wall.
</paragraph>
<paragraph id="8BC436B4FD183715FEA7FD4BFAF3FCD2" blockId="17.[151,1437,478,1619]" box="[199,1391,766,791]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FD4BFE28FCD3" bold="true" box="[199,436,766,791]" pageId="17" pageNumber="18">Derivation of name.</emphasis>
Named in honour of Paul Mazza for his research on the artiodactyls from Scontrone.
</paragraph>
<paragraph id="8BC436B4FD183715FEA7FC97FA9FFCFE" blockId="17.[151,1437,478,1619]" box="[199,1283,802,827]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FC97FD80FCFE" bold="true" box="[199,540,802,827]" pageId="17" pageNumber="18">Preservation and deposition.</emphasis>
Soprintendenza Archeologica dellAbruzzo (Chieti, central
<collectingCountry id="F36C7624FD183715FADFFC97FB6BFCFE" box="[1215,1271,802,827]" name="Italy" pageId="17" pageNumber="18">Italy</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD183715FEA7FCF3FE09FC62" pageId="17" pageNumber="18" type="materials_examined">
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<typeStatus id="54C08816FD183715FEA7FCF3FE9EFC9A" box="[199,258,838,863]" pageId="17" pageNumber="18">Type</typeStatus>
locality and horizon.
</emphasis>
Tortonian Scontrone Member of the Lithothamnium Limestone (
<bibRefCitation id="EFEA4B45FD183715FA99FCF2FF4FFC46" author="Patacca" pageId="17" pageNumber="18" refString="Patacca, E., Scandone, P. &amp; Mazza, P. (2008) Oligocene migration path for Apulia macromammals: the Central-Adriatic bridge. Bollettino della Societa Geologica Italiana, 127, 337 - 355." type="journal article" year="2008">
Patacca
<emphasis id="B90FEAA6FD183715FB3AFCF2FA09FC9B" box="[1370,1429,838,862]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
, 2008
</bibRefCitation>
;
<geoCoordinate id="EE4F5073FD183715FE80FCDFFE14FC46" box="[224,392,874,899]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="41.75432">41°45'15.54''N</geoCoordinate>
,
<geoCoordinate id="EE4F5073FD183715FFF6FCDEFDDAFC46" box="[406,582,875,899]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="14.0369835">14°02' 13.14''E</geoCoordinate>
), outskirts of Scontrone, southern border of the National Park of Abruzzi, LAquila, central
<collectingCountry id="F36C7624FD183715FF3CFC3BFE0DFC62" box="[348,401,910,935]" name="Italy" pageId="17" pageNumber="18">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD183715FEA7FC07FD91FB2E" pageId="17" pageNumber="18" type="description">
<paragraph id="8BC436B4FD183715FEA7FC07FD0FFB02" blockId="17.[151,1437,478,1619]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FC07FEC5FC0E" bold="true" box="[199,345,946,971]" pageId="17" pageNumber="18">Description.</emphasis>
See
<bibRefCitation id="EFEA4B45FD183715FFF0FC06FD2FFC0E" author="Mazza" box="[400,691,946,971]" pageId="17" pageNumber="18" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni (2011)</bibRefCitation>
. Additional characters shown by
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revised after
<bibRefCitation id="EFEA4B45FD183715FB54FC06FED6FC2A" author="Mazza" pageId="17" pageNumber="18" refString="Mazza, P. &amp; Rustioni, M. (1999) Dental characters of the Turolian fossil artiodactyls from Scontrone (Abruzzo, Central Italy). In: Mayhall, J. &amp; Heikkinen, T. (Eds.), Proceedings of the 11 th International Symposium on Dental Morphology, 26 - 30 August 1998, Oulu (Finland). Oulu University, Oulu, pp. 300 - 309." type="journal article" year="1999">Mazza &amp; Rustioni (1999)</bibRefCitation>
. The
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SCT80 (m3), SCT83 (incomplete m2, badly preserved m3) and SCT72 (incomplete m2, m3, alveoles p2m1) are fragmentary mandibles. The description here is after
<bibRefCitation id="EFEA4B45FD183715FA2EFC4EFAEEFBD6" author="Mazza" box="[1102,1394,1018,1043]" pageId="17" pageNumber="18" refString="Mazza, P. &amp; Rustioni, M. (1999) Dental characters of the Turolian fossil artiodactyls from Scontrone (Abruzzo, Central Italy). In: Mayhall, J. &amp; Heikkinen, T. (Eds.), Proceedings of the 11 th International Symposium on Dental Morphology, 26 - 30 August 1998, Oulu (Finland). Oulu University, Oulu, pp. 300 - 309." type="journal article" year="1999">Mazza &amp; Rustioni (1999)</bibRefCitation>
for SCT72 and SCT83. Horizontal ramus with marked pachyostosis. m2-m3. Brachyodont without ectostylids. The ribs and the styles are slightly prominent on the lingual enamel walls. The rear wings of the protoconids are fused to the anterior wings of the entoconids. The labial crescents are obliquely arranged backwards and have a V-shaped outline in occlusal view. The posterior lobe is bicuspid with the labial lobe exceeding the lingual lobe in size. No cingula, no cementum. The enamel is rugose.
</paragraph>
<paragraph id="8BC436B4FD183715FEA7FB67FD91FB2E" blockId="17.[151,1437,478,1619]" box="[199,525,1234,1259]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FB67FEE1FB2E" bold="true" box="[199,381,1234,1259]" pageId="17" pageNumber="18">Measurements.</emphasis>
See
<tableCitation id="C6F9030FFD183715FFD3FB67FD95FB2E" box="[435,521,1234,1259]" captionStart="TABLE 2" captionStartId="15.[151,239,152,175]" captionTargetBox="[159,1391,298,1424]" captionTargetPageId="15" captionText="TABLE 2. Dental measurements (L = length, W = width; in mm) of the mandibular (above) and maxillar fragments (below) from Scontrone, mandibular premolar / molar ratio and body mass estimation (BM in kg) based on lower molar row length after Damuth (1990) for selenodont browsers. * alveolar measurements. The specimens are sorted according to increasing size. Dental measurements from Mazza &amp; Rustioni (2011, 1999, 1996)." httpUri="http://table.plazi.org/id/DF04663CFD06370BFEF7FF2DFBBDFECA" pageId="17" pageNumber="18" tableUuid="DF04663CFD06370BFEF7FF2DFBBDFECA">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361653FFD183715FEA7FB43FAB1F996" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BC436B4FD183715FEA7FB43FAB1F996" blockId="17.[151,1437,478,1619]" pageId="17" pageNumber="18">
<emphasis id="B90FEAA6FD183715FEA7FB43FEA8FACA" bold="true" box="[199,308,1270,1295]" pageId="17" pageNumber="18">Remarks</emphasis>
.
<bibRefCitation id="EFEA4B45FD183715FF23FB42FDF8FACA" author="Mazza" box="[323,612,1270,1295]" pageId="17" pageNumber="18" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni (2011)</bibRefCitation>
attributed skull fragment SCT17 and three mandible fragments (SCT80, SCT83, SCT72) from Scontrone to
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<emphasis id="B90FEAA6FD183715FC4FFAAEFCBAFAF6" box="[559,806,1307,1331]" italics="true" pageId="17" pageNumber="18">Hoplitomeryx matthei</emphasis>
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. However, these specimens cannot be attributed to this species. According to
<bibRefCitation id="EFEA4B45FD183715FFEFFA8AFD30FA92" author="Mazza" box="[399,684,1342,1367]" pageId="17" pageNumber="18" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza &amp; Rustioni (2011)</bibRefCitation>
there is no evidence of the possible presence of the nasal horn core in skull fragment SCT17, yet a nasal horn core is diagnostic for
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<emphasis id="B90FEAA6FD183715FDACFAD6FB55FABE" box="[972,1225,1379,1403]" italics="true" pageId="17" pageNumber="18">Hoplitomeryx matthei</emphasis>
(
<bibRefCitation id="EFEA4B45FD183715FA80FAD7FA0CFABE" author="Leinders" box="[1248,1424,1378,1403]" pageId="17" pageNumber="18" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." type="journal article" year="1984">Leinders 1984</bibRefCitation>
)
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. Furthermore, the orbital horn cores of SCT17 show a combination of characters of Leinders (1984) horn core
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I and II and constitutes thus a new horn core
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, unlike any of Leinders (1984)
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.
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far as the mandibles are concerned, there is no evidence for the loss of the second premolar. This means that the presence of
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<emphasis id="B90FEAA6FD183715FEF7FA46FE12F9CE" box="[151,398,1523,1547]" italics="true" pageId="17" pageNumber="18">Hoplitomeryx matthei</emphasis>
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cannot be confirmed among the material from Scontrone. In order to avoid confusion, the four Scontrone specimens which are excluded here from the
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<emphasis id="B90FEAA6FD183715FD59F9A2FC32F9EB" box="[825,942,1558,1582]" italics="true" pageId="17" pageNumber="18">H. matthei</emphasis>
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material are attributed here to a new species as they cannot be attributed to any of the other Scontrone species defined by
<bibRefCitation id="EFEA4B45FD183715FD94F98EFAB4F996" author="Mazza" box="[1012,1320,1594,1619]" pageId="17" pageNumber="18" refString="Mazza, P. P. A. &amp; Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00737. x" type="journal article" year="2011">Mazza and Rustioni (2011)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
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