353 lines
43 KiB
XML
353 lines
43 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4658236" ID-GBIF-Dataset="71bdd341-af13-4447-ba22-9b2bb8de2f80" ID-GBIF-Taxon="180784330" ID-ISSN="0067-1975" ID-Zenodo-Dep="4658236" checkinTime="1617215540781" checkinUser="felipe" docAuthor="Edgecombe, Gregory D." docDate="1998" docId="03FE7A7C2D4CFFB2FE45FE48FE7E7D28" docLanguage="en" docName="RecAustMus.50.3.293-313.pdf.imf" docOrigin="Records of the Australian Museum 50 (3)" docSource="https://journals.australian.museum/edgecombe-1998-rec-aust-mus-503-293313/" docStyle="DocumentStyle:CF64E561BE6B94FA5F31B9132C6D4316.1:RecAustMus.1998.journal_article" docStyleId="CF64E561BE6B94FA5F31B9132C6D4316" docStyleName="RecAustMus.1998.journal_article" docStyleVersion="1" docTitle="Maldybulakia malcolmi Edgecombe 1998, n.sp." docType="treatment" docVersion="6" lastPageId="12" lastPageNumber="305" masterDocId="FFC702042D4AFFBEFFFFFFE8FFA1714A" masterDocTitle="Early myriapodous arthropods from Australia: Maldybulakia from the Devonian of New South Wales" masterLastPageNumber="313" masterPageNumber="293" pageId="6" pageNumber="299" updateTime="1643595242968" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early myriapodous arthropods from Australia: Maldybulakia from the Devonian of New South Wales</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Edgecombe, Gregory D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1998</mods:date>
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<mods:detail type="pubDate">
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<mods:number>1998-11-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>50</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>293</mods:start>
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<mods:end>313</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://journals.australian.museum/edgecombe-1998-rec-aust-mus-503-293313/</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.0067-1975.50.1998.1288</mods:identifier>
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<mods:identifier type="GBIF-Dataset">71bdd341-af13-4447-ba22-9b2bb8de2f80</mods:identifier>
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<mods:identifier type="ISSN">0067-1975</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4653146</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4658236" ID-GBIF-Taxon="180784330" ID-Zenodo-Dep="4658236" LSID="urn:lsid:plazi:treatment:03FE7A7C2D4CFFB2FE45FE48FE7E7D28" httpUri="http://treatment.plazi.org/id/03FE7A7C2D4CFFB2FE45FE48FE7E7D28" lastPageId="12" lastPageNumber="305" pageId="6" pageNumber="299">
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<subSubSection pageId="6" pageNumber="299" type="nomenclature">
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<paragraph blockId="6.[442,978,416,456]" box="[442,978,416,456]" pageId="6" pageNumber="299">
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<heading box="[442,978,416,456]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="299" reason="2">
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[442,876,416,456]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="6" pageNumber="299" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
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<emphasis box="[442,876,416,456]" italics="true" pageId="6" pageNumber="299">Maldybulakia malcolmi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[895,978,416,456]" pageId="6" pageNumber="299" rank="species">n.sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph blockId="6.[630,793,523,561]" box="[630,793,523,561]" pageId="6" pageNumber="299">Figs. 3-8</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="305" pageId="6" pageNumber="299" type="description">
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<paragraph blockId="6.[217,1067,627,663]" box="[217,1067,627,663]" pageId="6" pageNumber="299">
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[217,428,627,663]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="6" pageNumber="299" phylum="Arthropoda" rank="genus" status="new species">
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<emphasis box="[217,428,627,663]" italics="true" pageId="6" pageNumber="299">Maldybulakia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[445,629,627,663]" pageId="6" pageNumber="299" rank="species">new species</taxonomicNameLabel>
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1
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<bibRefCitation author="Edgecombe, G. D." box="[678,955,627,663]" pageId="6" pageNumber="299" pagination="172 - 175" refId="ref12512" refString="Edgecombe, G. D., 1998. Devonian terrestrial arthropods from Gondwana. Nature 394: 172 - 175." type="journal article" year="1998">Edgecombe, 1998</bibRefCitation>
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: fig. 1.
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</paragraph>
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<paragraph blockId="6.[218,789,742,781]" box="[218,789,742,781]" pageId="6" pageNumber="299">Etymology. For MaIcolm Young.</paragraph>
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<paragraph blockId="6.[216,1199,847,1116]" pageId="6" pageNumber="299">
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Diagnosis.
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<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[430,677,847,887]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="6" pageNumber="299" phylum="Arthropoda" rank="genus">
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<emphasis box="[430,677,847,887]" italics="true" pageId="6" pageNumber="299">Maldybulakia</emphasis>
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</taxonomicName>
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with relatively minor serial variation in B-pleurotergites; lacking long paratergal spines, posteromedian spines, and tuberculation; caudal pleurotergite with lateral lobes set offby shallow furrows; median projection on caudal pleurotergite blunt, conical, rather than spinose.
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</paragraph>
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<paragraph blockId="6.[214,1199,1184,1591]" pageId="6" pageNumber="299">
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<materialsCitation ID-GBIF-Occurrence="3066851301" collectionCode="AM" collectorName="Z. Iohanson." county="East Boyd State Forest" latitude="-37.15" location="Boyd Volcanic Complex" longLatPrecision="1183" longitude="149.95" municipality="Saltwater Creek" pageId="6" pageNumber="299" specimenCount="2" typeStatus="holotype">
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Types.
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<typeStatus box="[339,521,1184,1223]" pageId="6" pageNumber="299">HOLOTYPE</typeStatus>
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articulated trunkAM F.102533 (Fig. 3a d,f,g), composed of one T-tergite, four B-pleurotergites, a caudal tergite, disarticulated telson sclerites; and possibly associated T-tergite; length from anterior end of T-tergite to posterior end of caudal pleurotergite
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<quantity box="[926,1082,1367,1407]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="6" pageNumber="299" unit="mm" value="31.8">31.8 mm</quantity>
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.
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<location LSID="urn:lsid:plazi:treatment:03FE7A7C2D4CFFB2FE45FE48FE7E7D28:8E889DB12D4CFFB8FBACFABFFD8A74E7" county="East Boyd State Forest" latitude="-37.15" longLatPrecision="1183" longitude="149.95" municipality="Saltwater Creek" name="Boyd Volcanic Complex" pageId="6" pageNumber="299">Boyd Volcanic Complex</location>
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(late Givetian or early Frasnian),
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<location LSID="urn:lsid:plazi:treatment:03FE7A7C2D4CFFB2FE45FE48FE7E7D28:8E889DB12D4CFFB8FF28FA5BFDCF7490" box="[215,622,1459,1498]" county="East Boyd State Forest" latitude="-37.15" longLatPrecision="1183" longitude="149.95" municipality="Saltwater Creek" name="Saltwater Creek Road" pageId="6" pageNumber="299">
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<collectingMunicipality box="[215,508,1459,1498]" pageId="6" pageNumber="299">Saltwater Creek</collectingMunicipality>
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Road
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</location>
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,
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<collectingCounty box="[649,1075,1459,1498]" pageId="6" pageNumber="299">East Boyd State Forest</collectingCounty>
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, NSW (
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<geoCoordinate box="[227,356,1505,1545]" degrees="37" direction="south" minutes="09" orientation="latitude" pageId="6" pageNumber="299" precision="925" value="-37.15">37°09'S</geoCoordinate>
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<geoCoordinate box="[375,529,1505,1545]" degrees="149" direction="east" minutes="57" orientation="longitude" pageId="6" pageNumber="299" precision="925" value="149.95">149°57'E</geoCoordinate>
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). Collected by
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<collectorName box="[784,992,1505,1545]" pageId="6" pageNumber="299">Z. Iohanson.</collectorName>
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="3066851304" collectionCode="AM" collectorName="Z. Iohanson." county="East Boyd State Forest" latitude="-37.15" location="Boyd Volcanic Complex" longLatPrecision="1183" longitude="149.95" municipality="Saltwater Creek" pageId="6" pageNumber="299" specimenCount="2" typeStatus="holotype">
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<typeStatus box="[1007,1196,1505,1545]" pageId="6" pageNumber="299">PARATYPES</typeStatus>
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<collectionCode box="[214,280,1552,1591]" country="Australia" name="Australian Museum" pageId="6" pageNumber="299" type="Museum">AM</collectionCode>
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F.102534-102556, 102581, from type locality
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="6.[205,1197,1656,3167]" pageId="6" pageNumber="299">
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Description. Trunk up to
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<quantity box="[670,807,1656,1696]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="299" unit="mm" value="115.0">115 mm</quantity>
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long, as inferred from size of largest known B-pleurotergite scaled to articulated specimen.
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</paragraph>
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<paragraph blockId="6.[205,1197,1656,3167]" pageId="6" pageNumber="299">
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T-tergite(s) of length up to
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<quantity box="[828,993,1795,1834]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="6" pageNumber="299" unit="mm" value="16.4">16.4 mm</quantity>
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in known specimens, but inferred to be larger based on relative size of B-pleurotergites; smallest disarticulated specimen
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="299" unit="mm" value="5.1">5.1 mm</quantity>
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long medially. Length 38-44 percent of width; of even, moderate convexity (tr.). Anterior part of prozonite a forward-sloping scarp that evenly lengthens medially to about 45 percent the length of the prozonite. Stricture moderately deep and approximately transverse medially, then weakly curving backwards laterally, abruptly shallowing and recurving forward, then inward, such that lateral part of stricture is C-shaped. Metazonite 64-75 percent length (sag.) of prozonite; gently convex (sag.). Lobate posterior projections lie against anterodistal socketlike processes on underlying ring tergite, serving as point of articulation. Posterior margin gently convex forwards between posterior projections. Doublure wide beneath lateral and posterolateral edges ofT-tergite, shortening and scalloped on each side of a flange-like median section that has a transverse course; doublure abruptly truncated anteriorly, with straight anteromedial margin.
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</paragraph>
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<paragraph blockId="6.[205,1197,1656,3167]" lastBlockId="6.[1266,2256,419,3171]" pageId="6" pageNumber="299">Ring tergite (Fig. 5b,d-f) divided by sharp transverse furrow into short, crest-like anterior band of even length and long, elevated posterior band; furrow strongly shallow or effaced near distal margin. Anterior band extended laterally well outside posterior band as slender, pointed process; anterodistal edge folded as a small socket-like process. Posterior band lenticular in outline, convex (sag.), moderately arched transversely. Ring tergite overlapping prozonite of first B-pleurotergite, with distal edge of posterior band extending to inner edge of anterior process on prozonite of B-pleurotergite.</paragraph>
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<paragraph blockId="6.[1266,2256,419,3171]" pageId="6" pageNumber="299">
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Posterior tagma composed of four B-pleurotergites and caudal pleurotergite, widest across first pleurotergite then gently, evening narrowing posteriorly; largest known disarticulated specimen (second B-pleurotergite) of length
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<quantity box="[1274,1427,649,688]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="6" pageNumber="299" unit="mm" value="25.8">25.8 mm</quantity>
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, width
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<quantity box="[1559,1681,649,688]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="299" unit="mm" value="45.0">45 mm</quantity>
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; smallest disarticulated specimen (fourth B-pleurotergite)
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<quantity box="[1700,1835,695,733]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="299" unit="mm" value="7.5">7.5 mm</quantity>
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long, but as short as
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<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="299" unit="mm" value="6.0">6 mm</quantity>
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in articulated specimens. Relative length and width of pleurotergites variable (compare Figs. 3a and 5i for relatively narrow and wide examples, respectively); each pleurotergite longer relative to its width than the preceding one. Furrows separating prozonite and metazonite on all four pleurotergites of similar, moderate depth, long medialIy, narrower but with comparable depth against lateral lobes. In sagittal profile, prozonites gently convex, metazonites flat or weakly convex along most of length; median lobe raised well above lateral lobes. Lateral lobes of all four pleurolergites triangular, with short, blunt posterolateral projections, in some specimens developed as weak spines. Metazonites progressively more strongly arched (tr.) posteriorly in trunk. Diagonal furrows separating lateral and median lobes of equal impression on each B-pleurotergite, gently convex backward.
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</paragraph>
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<paragraph blockId="6.[1266,2256,419,3171]" pageId="6" pageNumber="299">First B-pleurotergite (Figs. 3e, 6c-e) with outer margin of lateral lobe oriented posterolaterally in dorsal view, gently convex outward, this margin posteriorly directed on the succeeding three pleurotergites with a corresponding posterior redirection of the posterolateral projections. Prozonite lenticular, with a strong process at anterodistal edge that is lacking on succeeding pleurotergites. Lateral edge of tergum sharply folded down as vertical scarp; lenticular pleurite separated by sharp, curved pleural furrow; pleural furrow effaced posteriorly; separation of pleuron and tergum not marked by suture on posterior part of pleurotergite; posteroventral edge of pleurotergite turned out as a rounded shelf.</paragraph>
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<paragraph blockId="6.[1266,2256,419,3171]" pageId="6" pageNumber="299">
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Second B-pleurotergite (
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<figureCitation box="[1730,1830,2076,2114]" captionStart="Figure 6" captionStartId="10.[221,326,3025,3062]" captionTargetBox="[209,2257,404,2977]" captionTargetPageId="10" captionText="Figure 6. Maldybulakia malcolmi n.sp. Scale bars 10 mm. Ca) dorsal view of T-tergite AM F.102549. Cb) ventral view of T-tergite AM F.102550, showing doublure. (c) dorsal view of first B-pleurotergite AM F.102551. (d) dorsal view of first B-pleurotergite AM F.102552. (e) dorsal view of first B-pleurotergite AM F.102553. (j) dorsal view of B-pleurotergite AM F.102554. (g,h) dorsal and left lateral views of second B-pleurotergite AM F.102555. (i,j) right lateral and dorsal views of second B-pleurotergite AM F.102556." figureDoi="http://doi.org/10.5281/zenodo.4653168" httpUri="https://zenodo.org/record/4653168/files/figure.png" pageId="6" pageNumber="299">Fig. 6</figureCitation>
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g-j) having anteroventral edge of prozonite forming a narrow, steep rim; anterolateral corner of tergite with ledge-like protrusion markedly distal to pleuron; small ventrally directed pleural process at anterior corner of metazonite, ventral margin of pleuron abruptly flexed ventrally immediately behind this process; pleuron bisected by anterodorsally oriented groove; posterior part of pleuron lensoid in outline, gently sloping outward; pleural furrow indistinct on posterior part of pleurotergite.
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</paragraph>
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<paragraph blockId="6.[1266,2256,419,3171]" pageId="6" pageNumber="299">Third B-pleurotergite (Fig. 3b,c,f) with pleural furrow expanded anteriorly to form a shallow, elongate basin within which a deep, elliptical canal (spiracle) projects inwards; pleurite narrow beneath spiracular region, then abruptly widening; rounded ventral projection on anterior part of pleurite as on preceding diplopleurotergite; sharp, shallow suture between pleurite and pleural furrow anteriorly.</paragraph>
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<paragraph blockId="6.[1266,2256,419,3171]" lastBlockId="11.[269,1259,1354,2862]" lastPageId="11" lastPageNumber="304" pageId="6" pageNumber="299">Fourth B-pleurotergite with lateral lobes of metazonite terminating as posteriorly-directed angulations. Weak nodose swelling usually distinguishable on posteromedian part of median lobe of metazonite (Figs. 3a, 5k), occasionally discernible on other B-pleurotergites. Pleural furrow deep, incised as a V-shaped groove along most of juncture between tergum and pleuron, abruptly shallowing near posterior margin. Pleurite clavate, narrowest anteriorly, projecting only slightly outside lateral edge of tergum.</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4653154" ID-Zenodo-Dep="4653154" httpUri="https://zenodo.org/record/4653154/files/figure.png" pageId="7" pageNumber="300" startId="7.[232,336,2980,3023]" targetBox="[213,2271,419,2948]" targetPageId="7">
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<paragraph blockId="7.[232,2272,2980,3192]" pageId="7" pageNumber="300">
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<emphasis bold="true" box="[232,336,2980,3023]" pageId="7" pageNumber="300">Figure</emphasis>
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3.
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[389,752,2980,3023]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="7" pageNumber="300" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
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<emphasis box="[389,752,2980,3023]" italics="true" pageId="7" pageNumber="300">Maldybulakia malcolmi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[764,833,2980,3023]" pageId="7" pageNumber="300" rank="species">n.sp.</taxonomicNameLabel>
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Scale bars 5 mm except/, 1 mm. Abbreviations: pf, pleural furrow; pI, pleurite; sp, spiracle; t, lateral edge of tergum; tsl, anterior telson sclerite; ts2, posterior telson sclerite.
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<emphasis box="[1502,1629,3024,3064]" italics="true" pageId="7" pageNumber="300">(a-d,J,g)</emphasis>
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holotype articulated trunk AM F.102533.
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<emphasis box="[233,272,3065,3106]" italics="true" pageId="7" pageNumber="300">(a)</emphasis>
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dorsal view of complete specimen.
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<emphasis box="[840,907,3065,3106]" italics="true" pageId="7" pageNumber="300">(b,c)</emphasis>
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lateral views of anterior and posterior B-pleurotergites, respectively. (d) lateral view of complete specimen.
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<emphasis box="[545,577,3107,3148]" italics="true" pageId="7" pageNumber="300">if)</emphasis>
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lateral view of third B-pleurotergite with spiracle.
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<emphasis box="[1369,1410,3107,3148]" italics="true" pageId="7" pageNumber="300">(g)</emphasis>
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dorsal view of caudal pleurotergite and telson sclerites.
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<emphasis box="[232,269,3149,3192]" italics="true" pageId="7" pageNumber="300">(e)</emphasis>
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posterolateral view of first B-pleurotergite AM F.102552 (see Fig. 6d for dorsal view), showing pleural furrow (pf).
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4653158" ID-Zenodo-Dep="4653158" httpUri="https://zenodo.org/record/4653158/files/figure.png" pageId="8" pageNumber="301" startId="8.[214,320,2930,2972]" targetBox="[215,2245,377,2889]" targetPageId="8">
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<paragraph blockId="8.[212,2259,2930,3176]" pageId="8" pageNumber="301">
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<emphasis bold="true" box="[214,320,2930,2972]" pageId="8" pageNumber="301">Figure</emphasis>
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4.
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[376,741,2930,2972]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="8" pageNumber="301" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
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<emphasis box="[376,741,2930,2972]" italics="true" pageId="8" pageNumber="301">Maldybulakia malcolmi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[756,824,2930,2972]" pageId="8" pageNumber="301" rank="species">n.sp.</taxonomicNameLabel>
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Scale bars 10 mm except
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<emphasis box="[1251,1325,2930,2972]" italics="true" pageId="8" pageNumber="301">d,j,g,</emphasis>
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5 mm.
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<emphasis box="[1457,1527,2930,2972]" italics="true" pageId="8" pageNumber="301">(a,b)</emphasis>
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dorsal and left dorsolateral views of T-tergite, ring tergite, and first two B-pleurotergites, AM F.102534. (c) dorsal view of latex cast of articulated trunk AM F.102535a, showing T-tergite, ring tergite, four B-pleurotergites and part of caudal pleurotergite. (d), dorsal view of latex cast of ring tergite and four Bpleurotergites, AM F.l02536.
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<emphasis box="[682,721,3057,3097]" italics="true" pageId="8" pageNumber="301">(e)</emphasis>
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left dorsolateral view of trunk AM F.I02535b (see Fig. 4c for counterpart).
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<emphasis box="[1938,1998,3057,3097]" italics="true" pageId="8" pageNumber="301">If,h)</emphasis>
|
||
dorsal and right lateral views of articulated trunk AM F.102537, showing posterior part of T-tergite, ring tergite (rt), four T-pleurotergites, and part of caudal pleurotergite.
|
||
<emphasis box="[589,630,3140,3176]" italics="true" pageId="8" pageNumber="301">(g)</emphasis>
|
||
dorsal view of telson sclerites AM F.I02538.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653164" ID-Zenodo-Dep="4653164" httpUri="https://zenodo.org/record/4653164/files/figure.png" pageId="9" pageNumber="302" startId="9.[232,336,2913,2949]" targetBox="[216,2278,370,2871]" targetPageId="9">
|
||
<paragraph blockId="9.[231,2272,2913,3199]" pageId="9" pageNumber="302">
|
||
<emphasis bold="true" box="[232,336,2913,2949]" pageId="9" pageNumber="302">Figure</emphasis>
|
||
5.
|
||
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[389,751,2913,2949]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="9" pageNumber="302" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
|
||
<emphasis box="[389,751,2913,2949]" italics="true" pageId="9" pageNumber="302">Maldybulakia malcolmi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[764,833,2913,2949]" pageId="9" pageNumber="302" rank="species">n.sp.</taxonomicNameLabel>
|
||
Scale bars 10 mm except
|
||
<emphasis box="[1246,1269,2913,2949]" italics="true" pageId="9" pageNumber="302">k,</emphasis>
|
||
5 mm.
|
||
<emphasis box="[1395,1435,2913,2949]" italics="true" pageId="9" pageNumber="302">(a)</emphasis>
|
||
dorsal view of ring tergite and three B-pleurotergites, AM E102539.
|
||
<emphasis box="[475,516,2955,2991]" italics="true" pageId="9" pageNumber="302">(b)</emphasis>
|
||
dorsal view of ring tergite AM E102540.
|
||
<emphasis box="[1209,1248,2955,2991]" italics="true" pageId="9" pageNumber="302">(c)</emphasis>
|
||
dorsal view of caudal pleurotergite, AM E102541. The median swelling in this specimen is more dorsally directed than in other specimens.
|
||
<emphasis box="[1421,1461,2997,3032]" italics="true" pageId="9" pageNumber="302">(d)</emphasis>
|
||
dorsal view of ring tergite AM E102542.
|
||
<emphasis box="[2129,2167,2997,3032]" italics="true" pageId="9" pageNumber="302">(e)</emphasis>
|
||
dorsal view of ring tergite (rt) and three B-pleurotergites, AM E102543.
|
||
<emphasis box="[1313,1346,3039,3074]" italics="true" pageId="9" pageNumber="302">(f)</emphasis>
|
||
dorsal view of ring tergite and first B-pleurotergite, AM El02544.
|
||
<emphasis box="[394,462,3080,3116]" italics="true" pageId="9" pageNumber="302">(g,h)</emphasis>
|
||
dorsal and anterior or posterior view of unique sclerite of unknown position, AM El02545. White line in
|
||
<emphasis box="[2110,2127,3080,3116]" italics="true" pageId="9" pageNumber="302">h</emphasis>
|
||
indicates limit of exoskeleton in the embayment. (i) dorsal view of posterior two B-pleurotergites and caudal pleurotergite (cp), AM E102546.
|
||
<emphasis box="[232,290,3164,3199]" italics="true" pageId="9" pageNumber="302">(j,k)</emphasis>
|
||
posterior and dorsal views of fourth B-pleurotergite, AM E102547. (I) dorsal view of caudal pleurotergite AM F.l02548.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653168" ID-Zenodo-Dep="4653168" httpUri="https://zenodo.org/record/4653168/files/figure.png" pageId="10" pageNumber="303" startId="10.[221,326,3025,3062]" targetBox="[209,2257,404,2977]" targetPageId="10">
|
||
<paragraph blockId="10.[221,2270,3025,3186]" pageId="10" pageNumber="303">
|
||
<emphasis bold="true" box="[221,366,3025,3062]" pageId="10" pageNumber="303">Figure 6.</emphasis>
|
||
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[382,748,3025,3062]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="10" pageNumber="303" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
|
||
<emphasis box="[382,748,3025,3062]" italics="true" pageId="10" pageNumber="303">Maldybulakia malcolmi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[763,831,3025,3062]" pageId="10" pageNumber="303" rank="species">n.sp.</taxonomicNameLabel>
|
||
Scale bars 10 mm. Ca) dorsal view of T-tergite AM F.102549. Cb) ventral view of T-tergite AM F.102550, showing doublure.
|
||
<emphasis box="[758,797,3068,3103]" italics="true" pageId="10" pageNumber="303">(c)</emphasis>
|
||
dorsal view of first B-pleurotergite AM F.102551.
|
||
<emphasis box="[1604,1644,3068,3103]" italics="true" pageId="10" pageNumber="303">(d)</emphasis>
|
||
dorsal view of first B-pleurotergite AM F.102552.
|
||
<emphasis box="[387,426,3109,3144]" italics="true" pageId="10" pageNumber="303">(e)</emphasis>
|
||
dorsal view of first B-pleurotergite AM F.102553. (j) dorsal view of B-pleurotergite AM F.102554.
|
||
<emphasis box="[2025,2094,3109,3144]" italics="true" pageId="10" pageNumber="303">(g,h)</emphasis>
|
||
dorsal and left lateral views of second B-pleurotergite AM F.102555.
|
||
<emphasis box="[1123,1176,3151,3186]" italics="true" pageId="10" pageNumber="303">(i,j)</emphasis>
|
||
right lateral and dorsal views of second B-pleurotergite AM F.102556.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653172" ID-Zenodo-Dep="4653172" httpUri="https://zenodo.org/record/4653172/files/figure.png" pageId="11" pageNumber="304" startId="11.[430,539,1176,1211]" targetBox="[283,2318,488,1089]" targetPageId="11">
|
||
<paragraph blockId="11.[430,2156,1176,1253]" pageId="11" pageNumber="304">
|
||
Figure 7.
|
||
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[598,971,1176,1211]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="11" pageNumber="304" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
|
||
<emphasis box="[598,971,1176,1211]" italics="true" pageId="11" pageNumber="304">Maldybulakia malcolmi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[987,1058,1176,1211]" pageId="11" pageNumber="304" rank="species">n.sp.</taxonomicNameLabel>
|
||
Reconstruction of trunk in lateral view, showing serial variation in structures of the pleurotergum. Based mostly on AM F.102533 (Fig. 3b-d).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[269,1259,1354,2862]" pageId="11" pageNumber="304">Caudal tergite (Figs. 3c,g, 5i,l) with prozonite longer than on preceding B-pleurotergites, weakly convex (sag.); furrow between prozonite and metazonite shallow medially, distinctly deeper against lateral lobes; lateral lobes flat, set off by weak furrow; lateral edge of tergum forming a slenderridge that gently bulges beyond otherwise triangular outline of metazonite in dorsal view; pleural furrow not impressed; posteromedian process large, conical, triangular or elliptical in section posteriorly, gently turned up; posterolateral margin vertically inclined, gently convex.</paragraph>
|
||
<paragraph blockId="11.[269,1259,1354,2862]" pageId="11" pageNumber="304">
|
||
Two small telson sclerites (Figs.
|
||
<quantity box="[868,909,2043,2082]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="11" pageNumber="304" unit="g" value="3.0">3g</quantity>
|
||
,
|
||
<quantity box="[929,973,2043,2082]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="11" pageNumber="304" unit="g" value="4.0">4g</quantity>
|
||
). Anterior telson sc1erite divided into a depressed anterior band that resembles an articulating edge and a longer, convex, strongly arched (tr.), posterior band; about 40 percent width of caudal pleurotergite. Posterior telson sc1erite divided by a sharp transverse furrow into short anterior band and flattened, broadly quadrate posterior plate; anterior band overlapped by anterior telson sclerite.
|
||
</paragraph>
|
||
<paragraph blockId="11.[269,1259,1354,2862]" pageId="11" pageNumber="304">
|
||
Dorsal surface of cuticle densely covered with honeycomb-like, shallow, polygonal pits; density of polygons the same on prozonite, stricture, and metazonite. Cuticle replaced with chalcedony, divided into a thin outer layer (
|
||
<quantity box="[392,563,2594,2633]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="304" unit="mm" value="0.04">0.04 mm</quantity>
|
||
) and thicker main layer (
|
||
<quantity box="[1058,1233,2594,2633]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="304" unit="mm" value="0.45">0.45 mm</quantity>
|
||
), measured in thin section; internal fabric of main layer obliterated by chalcedony aggregates. Large, densely arranged canals run through cuticle perpendicular to its surface (Fig. Sa), opening to variable-sized, rounded pits on ventral surface (Fig. Sb).
|
||
</paragraph>
|
||
<paragraph blockId="11.[267,1252,2915,3230]" lastBlockId="11.[1324,2312,1355,3231]" pageId="11" pageNumber="304">
|
||
Discussion. The description of this species regards the trunk as being composed of pleurotergites, the pleurites being fused to the tergal metazonites (
|
||
<figureCitation box="[958,1067,3008,3047]" captionStart="Figure 7" captionStartId="11.[430,539,1176,1211]" captionTargetBox="[283,2318,488,1089]" captionTargetPageId="11" captionText="Figure 7. Maldybulakia malcolmi n.sp. Reconstruction of trunk in lateral view, showing serial variation in structures of the pleurotergum. Based mostly on AM F.102533 (Fig. 3b-d)." figureDoi="http://doi.org/10.5281/zenodo.4653172" httpUri="https://zenodo.org/record/4653172/files/figure.png" pageId="11" pageNumber="304">Fig. 7</figureCitation>
|
||
). The two components are set off by a deep groove, described as a pleural furrow. That the pleuron is actually fused to the tergum is most evident in the posterior part of the first (Fig. 3b,e) and second (
|
||
<figureCitation box="[674,786,3191,3230]" captionStart="Figure 6" captionStartId="10.[221,326,3025,3062]" captionTargetBox="[209,2257,404,2977]" captionTargetPageId="10" captionText="Figure 6. Maldybulakia malcolmi n.sp. Scale bars 10 mm. Ca) dorsal view of T-tergite AM F.102549. Cb) ventral view of T-tergite AM F.102550, showing doublure. (c) dorsal view of first B-pleurotergite AM F.102551. (d) dorsal view of first B-pleurotergite AM F.102552. (e) dorsal view of first B-pleurotergite AM F.102553. (j) dorsal view of B-pleurotergite AM F.102554. (g,h) dorsal and left lateral views of second B-pleurotergite AM F.102555. (i,j) right lateral and dorsal views of second B-pleurotergite AM F.102556." figureDoi="http://doi.org/10.5281/zenodo.4653168" httpUri="https://zenodo.org/record/4653168/files/figure.png" pageId="11" pageNumber="304">Fig. 6i</figureCitation>
|
||
) pleurotergites, where they are in direct continuity, the first pleurotergite without even a trace of a suture behind the abruptly-effaced pleural furrow. The morphology of the pleuron is frequently complicated by crushing, on occasion giving the appearance of several pleurites. In the
|
||
<typeStatus box="[1957,2097,1539,1578]" pageId="11" pageNumber="304">holotype</typeStatus>
|
||
in particular, this appearance is compounded by fracture along the lateral edge of the tergum on the first three pleurotergites (Fig. 3b d). Comparison with other specimens (Figs. 3e, 6h,i) which have an intact down-folded edge of the tergum clarifies the structures (
|
||
<figureCitation box="[1506,1604,1769,1807]" captionStart="Figure 7" captionStartId="11.[430,539,1176,1211]" captionTargetBox="[283,2318,488,1089]" captionTargetPageId="11" captionText="Figure 7. Maldybulakia malcolmi n.sp. Reconstruction of trunk in lateral view, showing serial variation in structures of the pleurotergum. Based mostly on AM F.102533 (Fig. 3b-d)." figureDoi="http://doi.org/10.5281/zenodo.4653172" httpUri="https://zenodo.org/record/4653172/files/figure.png" pageId="11" pageNumber="304">Fig. 7</figureCitation>
|
||
). Every B-pleurotergite (even disarticulated ones) preserves the pleuron in association with the tergites, and they have characteristic morphologies on each of the four B-pleurotergites. Were the pleurites separate, unfused elements (e.g., as in centipedes) this invariant association with the tergum would not be maintained. Whether the pleural component of the pleurotergum may consist of multiple, fused pleurites is not certain. In at least some pleurotergites (Fig. 3c, particularly the fourth Bpleurotergite) the pleuron consists of a simple, elongate band without sutures, and seems likely to be composed of a single pleurite.
|
||
</paragraph>
|
||
<paragraph blockId="11.[1324,2312,1355,3231]" lastBlockId="12.[216,1206,2029,3170]" lastPageId="12" lastPageNumber="305" pageId="11" pageNumber="304">
|
||
Serial variation in the B-pleurotergite series includes a progressive decrease in the height of the lateral edge of the tergum, and an increase in the length of the pleural furrow (short on the first B-pleurotergite versus the entire length of the fourth B-pleurotergite). The pit interpreted as a spiracle is present on a single pleurotergite, and is only distinguishable on the
|
||
<typeStatus box="[1843,1997,2596,2634]" pageId="11" pageNumber="304">holotype</typeStatus>
|
||
(Fig. 3b,f). This opening is not a preservational artifact because it is entirely lined with cuticle, forming a distinct, ovate canal. The position of such a canal at the juncture between the tergum and pleuron resembles that of some tracheates (e.g., pleurostigmophoran chilopods, hexapods). The existence of only one such opening, rather than a series along the trunk, does not negate identity as a spiracle, although it is peculiar given the substantial size of
|
||
<emphasis box="[2046,2308,2962,3001]" italics="true" pageId="11" pageNumber="304">
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[2046,2301,2962,3001]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="11" pageNumber="304" phylum="Arthropoda" rank="genus">Maldybulakia</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
presence of only one or two spiracles in extant myriapods (e.g., Symphyla and some lithobiomorphs) is reasonably regarded as linked to minute body size. To consider alternative interpretations, the tube in
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[2023,2264,3146,3185]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="11" pageNumber="304" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[2023,2264,3146,3185]" italics="true" pageId="11" pageNumber="304">Maldybulakia</emphasis>
|
||
</taxonomicName>
|
||
is situated in the appropriate position for an ozopore, the opening for the repugnatorial glands in helminthomorph diplopods (see
|
||
<bibRefCitation author="Shear" box="[489,696,2075,2114]" pageId="12" pageNumber="305" pagination="309 - 316" refId="ref13547" refString="Shear, w. A., 1993. Myriapodous arthropods from the Visean of East Kirkton, West Lothian, Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences 84: 309 - 316." type="journal article" year="1993">Shear, 1993</bibRefCitation>
|
||
: fig. 2 for a fossil example). Such an identity is considered less likely than a spiracle because an ozopore would be expected to be smaller, whereas myriapod spiracles can be very large (e.g., in scolopendrid chilopods; see lG.E.
|
||
<bibRefCitation author="Lewis, J. G. E. & T. J. Hill & G. E. Wakley" box="[827,1146,2259,2298]" pageId="12" pageNumber="305" pagination="441 - 449" refId="ref12944" refString="Lewis, J. G. E., T. J. Hill & G. E. Wakley, 1996. The structure and possible function of the spiracles of some Scolopendridae (Chilopoda, Scolopendromorpha). In Acta Myriapodologica, eds. J. - I. Geoffroy, J. - P. Mauries & M. Nguyen Duy- Jacquemin, pp. 441 - 449. Paris: Memoires du Museum National d'Histoire Naturelle, Tome 169." type="book chapter" year="1996">
|
||
Lewis
|
||
<emphasis box="[945,1146,2259,2298]" italics="true" pageId="12" pageNumber="305">et al., 1996</emphasis>
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653176" ID-Zenodo-Dep="4653176" httpUri="https://zenodo.org/record/4653176/files/figure.png" pageId="12" pageNumber="305" startId="12.[222,328,1604,1639]" targetBox="[243,1211,441,1546]" targetPageId="12">
|
||
<paragraph blockId="12.[221,1206,1604,1932]" pageId="12" pageNumber="305">
|
||
Figure 8.
|
||
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[384,748,1604,1639]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi" status="n.sp.">
|
||
<emphasis box="[384,748,1604,1639]" italics="true" pageId="12" pageNumber="305">Maldybulakia malcolmi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[763,831,1604,1639]" pageId="12" pageNumber="305" rank="species">n.sp.</taxonomicNameLabel>
|
||
Scale bars 1 mm.
|
||
<emphasis box="[1136,1205,1604,1639]" italics="true" pageId="12" pageNumber="305">(a,b)</emphasis>
|
||
T-tergite AM F.l02581.
|
||
<emphasis box="[649,693,1646,1681]" italics="true" pageId="12" pageNumber="305">(a)</emphasis>
|
||
dorsoventral section through exoskeleton on lateral part of metazonite, ventral surface down, anterior to left, showing large canals filled with clay and silt matrix (light colour, versus darker exoskeleton).
|
||
<emphasis box="[1036,1079,1771,1806]" italics="true" pageId="12" pageNumber="305">(b)</emphasis>
|
||
ventral surface of exoskeleton on lateral part of metazonite, showing openings of canals as large, round pores filled with light-coloured matrix.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="12.[216,1206,2029,3170]" pageId="12" pageNumber="305">
|
||
A lesser degree of serial variation within the two trunk tagmata is a character uniting
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[730,1131,2351,2389]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">
|
||
<emphasis box="[730,1131,2351,2389]" italics="true" pageId="12" pageNumber="305">Maldybulakia malcolmi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[217,435,2396,2435]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis box="[217,435,2396,2435]" italics="true" pageId="12" pageNumber="305">M. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
to the exclusion of
|
||
<emphasis box="[804,990,2396,2435]" italics="true" pageId="12" pageNumber="305">
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[804,985,2396,2435]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="angusi">M. angusi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Suppressed variation is, however, presumed to be a primitive condition (with outgroup comparison to other myriapods), and not indicative of a sister species relationship between
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">
|
||
<emphasis italics="true" pageId="12" pageNumber="305">M. malcolmi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis box="[468,692,2581,2619]" italics="true" pageId="12" pageNumber="305">
|
||
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[468,687,2581,2619]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="mirabilis">M. mirabilis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Given that
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[911,1131,2581,2619]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">
|
||
<emphasis box="[911,1131,2581,2619]" italics="true" pageId="12" pageNumber="305">M. malcolmi</emphasis>
|
||
</taxonomicName>
|
||
has only four B-pleurotergites (not counting the caudal pleurotergite), it is likely that the four articulated Bpleurotergites in the
|
||
<typeStatus box="[611,767,2718,2757]" pageId="12" pageNumber="305">holotype</typeStatus>
|
||
of the very similar
|
||
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis italics="true" pageId="12" pageNumber="305">M. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Tesakov, A. S. & A. S. Alekseev" box="[395,858,2764,2803]" pageId="12" pageNumber="305" pagination="18 - 23" refId="ref13855" refString="Tesakov, A. S., & A. S. Alekseev, 1992. Myriapod-like arthropods from the Lower Devonian of central Kazakhstan. Paleontological Journal 26: 18 - 23." type="journal article" year="1992">Tesakov & Alekseev, 1992</bibRefCitation>
|
||
: fig. la,b) represent the entire B-pleurotergite series.
|
||
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[794,1204,2810,2849]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">
|
||
<emphasis box="[794,1204,2810,2849]" italics="true" pageId="12" pageNumber="305">Maldybulakia malcolmi</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[624,852,2856,2895]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="mirabilis">
|
||
<emphasis box="[624,852,2856,2895]" italics="true" pageId="12" pageNumber="305">M. mirabilis</emphasis>
|
||
</taxonomicName>
|
||
by its pronounced transverse furrow on the T-tergite (as opposed to a pair of chevron-shaped, longitudinally aligned furrows in
|
||
<emphasis italics="true" pageId="12" pageNumber="305">
|
||
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="mirabilis">M. mirabilis</taxonomicName>
|
||
),
|
||
</emphasis>
|
||
much finer granulation on the tergites, shallow (versus deep) furrows setting off the lateral lobes on the caudal pleurotergite, and a less spinose median projection on that sclerite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |