383 lines
48 KiB
XML
383 lines
48 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4658242" ID-GBIF-Dataset="71bdd341-af13-4447-ba22-9b2bb8de2f80" ID-GBIF-Taxon="180784332" ID-ISSN="0067-1975" ID-Zenodo-Dep="4658242" checkinTime="1617215540781" checkinUser="felipe" docAuthor="Edgecombe, Gregory D." docDate="1998" docId="03FE7A7C2D46FFAFFA0BFE53FE417B1E" docLanguage="en" docName="RecAustMus.50.3.293-313.pdf.imf" docOrigin="Records of the Australian Museum 50 (3)" docSource="https://journals.australian.museum/edgecombe-1998-rec-aust-mus-503-293313/" docStyle="DocumentStyle:CF64E561BE6B94FA5F31B9132C6D4316.1:RecAustMus.1998.journal_article" docStyleId="CF64E561BE6B94FA5F31B9132C6D4316" docStyleName="RecAustMus.1998.journal_article" docStyleVersion="1" docTitle="Maldybulakia angusi Edgecombe 1998, n.sp." docType="treatment" docVersion="6" lastPageId="17" lastPageNumber="310" masterDocId="FFC702042D4AFFBEFFFFFFE8FFA1714A" masterDocTitle="Early myriapodous arthropods from Australia: Maldybulakia from the Devonian of New South Wales" masterLastPageNumber="313" masterPageNumber="293" pageId="12" pageNumber="305" updateTime="1643595242968" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Early myriapodous arthropods from Australia: Maldybulakia from the Devonian of New South Wales</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Edgecombe, Gregory D.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1998</mods:date>
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<mods:detail type="pubDate">
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<mods:number>1998-11-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>50</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>293</mods:start>
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<mods:end>313</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://journals.australian.museum/edgecombe-1998-rec-aust-mus-503-293313/</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.0067-1975.50.1998.1288</mods:identifier>
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<mods:identifier type="GBIF-Dataset">71bdd341-af13-4447-ba22-9b2bb8de2f80</mods:identifier>
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<mods:identifier type="ISSN">0067-1975</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4653146</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4658242" ID-GBIF-Taxon="180784332" ID-Zenodo-Dep="4658242" LSID="urn:lsid:plazi:treatment:03FE7A7C2D46FFAFFA0BFE53FE417B1E" httpUri="http://treatment.plazi.org/id/03FE7A7C2D46FFAFFA0BFE53FE417B1E" lastPageId="17" lastPageNumber="310" pageId="12" pageNumber="305">
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<subSubSection box="[1524,2010,443,482]" pageId="12" pageNumber="305" type="nomenclature">
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<paragraph blockId="12.[1524,2010,443,482]" box="[1524,2010,443,482]" pageId="12" pageNumber="305">
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<heading box="[1524,2010,443,482]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="305" reason="2">
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[1524,1909,443,482]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="angusi" status="n.sp.">
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<emphasis box="[1524,1909,443,482]" italics="true" pageId="12" pageNumber="305">Maldybulakia angusi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1927,2010,443,482]" pageId="12" pageNumber="305" rank="species">n.sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="17" lastPageNumber="310" pageId="12" pageNumber="305" type="description">
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<paragraph blockId="12.[1677,1863,549,587]" box="[1677,1863,549,587]" pageId="12" pageNumber="305">
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<figureCitation box="[1677,1802,549,587]" captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="12" pageNumber="305">Figs. 9</figureCitation>
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-12
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</paragraph>
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<paragraph blockId="12.[1278,2126,654,688]" box="[1278,2126,654,688]" pageId="12" pageNumber="305">
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[1278,1487,654,688]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="genus" status="new species">
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<emphasis box="[1278,1487,654,688]" italics="true" pageId="12" pageNumber="305">Maldybulakia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1503,1687,654,688]" pageId="12" pageNumber="305" rank="species">new species</taxonomicNameLabel>
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2
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<bibRefCitation author="Edgecombe, G. D." box="[1736,2013,654,688]" pageId="12" pageNumber="305" pagination="172 - 175" refId="ref12512" refString="Edgecombe, G. D., 1998. Devonian terrestrial arthropods from Gondwana. Nature 394: 172 - 175." type="journal article" year="1998">Edgecombe, 1998</bibRefCitation>
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: fig. 2.
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</paragraph>
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<paragraph blockId="12.[1279,1802,756,795]" box="[1279,1802,756,795]" pageId="12" pageNumber="305">
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Etymology. For
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<collectingRegion box="[1566,1673,756,795]" country="United Kingdom" name="Angus" pageId="12" pageNumber="305">Angus</collectingRegion>
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Young.
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</paragraph>
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<paragraph blockId="12.[1279,2261,860,1129]" pageId="12" pageNumber="305">
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Diagnosis.
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<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[1469,1701,860,898]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="genus">
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<emphasis box="[1469,1701,860,898]" italics="true" pageId="12" pageNumber="305">Maldybulakia</emphasis>
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</taxonomicName>
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with considerable serial variation in B-pleurotergites; posteromedian spine present on some tergites of anterior and posterior trunk tagmata; very long, posterolaterally-directed paratergal spines on most bilobate trunk pleurotergites; caudal tergite bearing long median spine; many tergites bearing robust tuberculation.
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</paragraph>
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<paragraph blockId="12.[1278,2262,1196,1466]" pageId="12" pageNumber="305">
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<materialsCitation ID-GBIF-Occurrence="3066851302" collectionCode="AM" county="Sugarloaf Creek Formation" latitude="-35.066666" location="Three Oaks" municipality="Sugarloaf Creek" pageId="12" pageNumber="305" specimenCount="2" stateProvince="New South Wales" typeStatus="holotype">
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Types.
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<typeStatus box="[1399,1581,1196,1234]" pageId="12" pageNumber="305">HOLOTYPE</typeStatus>
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trunk pleurotergite (B-pleurotergite)
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<collectionCode box="[2195,2260,1196,1234]" country="Australia" name="Australian Museum" pageId="12" pageNumber="305" type="Museum">AM</collectionCode>
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F.102565 (Fig. lOa); length
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<quantity box="[1788,1946,1241,1280]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="12" pageNumber="305" unit="mm" value="24.6">24.6 mm</quantity>
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.
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<collectingCounty pageId="12" pageNumber="305">
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<collectingMunicipality box="[1973,2262,1241,1280]" pageId="12" pageNumber="305">Sugarloaf Creek</collectingMunicipality>
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Formation
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</collectingCounty>
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(Lochkovian or Pragian), near
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<location LSID="urn:lsid:plazi:treatment:03FE7A7C2D46FFAFFA0BFE53FE417B1E:8E889DB12D46FFB2F807FAE0F76C7464" box="[2040,2253,1288,1326]" county="Sugarloaf Creek Formation" latitude="-35.066666" municipality="Sugarloaf Creek" name="Three Oaks" pageId="12" pageNumber="305" stateProvince="New South Wales">Three Oaks</location>
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,
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<location LSID="urn:lsid:plazi:treatment:03FE7A7C2D46FFAFFA0BFE53FE417B1E:8E889DB12D46FFB2FB01FADDF9AF7416" box="[1278,1550,1333,1372]" county="Sugarloaf Creek Formation" latitude="-35.066666" municipality="Sugarloaf Creek" name="Taemas district" pageId="12" pageNumber="305" stateProvince="New South Wales">Taemas district</location>
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,
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<collectingRegion box="[1575,1889,1333,1372]" country="Australia" name="New South Wales" pageId="12" pageNumber="305">New South Wales</collectingRegion>
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(
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<geoCoordinate box="[1923,2054,1333,1372]" degrees="35" direction="south" minutes="04" orientation="latitude" pageId="12" pageNumber="305" precision="925" value="-35.066666">35°04'S</geoCoordinate>
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<geoCoordinate box="[2078,2240,1333,1372]" pageId="12" pageNumber="305">148°5l'E</geoCoordinate>
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).
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="3066851303" collectionCode="AM" county="Sugarloaf Creek Formation" latitude="-35.066666" location="Three Oaks" municipality="Sugarloaf Creek" pageId="12" pageNumber="305" specimenCount="2" stateProvince="New South Wales" typeStatus="holotype">
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<typeStatus box="[1278,1467,1379,1418]" pageId="12" pageNumber="305">PARATYPES</typeStatus>
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<collectionCode box="[1480,1547,1379,1418]" country="Australia" name="Australian Museum" pageId="12" pageNumber="305" type="Museum">AM</collectionCode>
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F.l02557-102564, 102566-102580, from type locality
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="12.[1275,2266,1530,3178]" pageId="12" pageNumber="305">
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Description. T-tergites (
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<figureCitation box="[1724,1835,1530,1569]" captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="12" pageNumber="305">Fig. 9</figureCitation>
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a-h) with length 32-39 percent of width. Anterior part of prozonite a flattened, boomerang-shaped surface that slopes forwards and lengthens medially to about 50 percent the length of the prozonite; rear edge of this surface forms a crest-like rise above posteriorpart of prozonite; well preserved specimens show dense, scaly sculpture (
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<figureCitation box="[1804,1931,1806,1845]" captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="12" pageNumber="305">Fig. 9c</figureCitation>
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). Posterior part of prozonite with pair of moderately wide longitudinal furrows varying from faint to distinctly incised. Stricture moderately deep, gently convex forward medially, recurved distally around elliptical or teardrop shaped lateral lobes. Metazonite 70-86 percent length of prozonite. Posteromedian spine absent or short. Posterior margin approximately transverse between lobate processes which vary from broad and rounded to angular to short, slender spines. Tuberculation usually strong, rarely faint, sometimes more prominent on prozonite than on metazonite. Doublure (
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<figureCitation captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="12" pageNumber="305">Fig. 9a</figureCitation>
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) as described for
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<emphasis box="[1630,1854,2312,2351]" italics="true" pageId="12" pageNumber="305">
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<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[1630,1849,2312,2351]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">M. malcolmi</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph blockId="12.[1275,2266,1530,3178]" box="[1324,2027,2358,2396]" pageId="12" pageNumber="305">
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Ring tergite (
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<figureCitation box="[1553,1667,2358,2396]" captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="12" pageNumber="305">Fig. 9i</figureCitation>
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) as for
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<emphasis box="[1805,2027,2358,2396]" italics="true" pageId="12" pageNumber="305">
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<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[1805,2022,2358,2396]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">M. malcolmi</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph blockId="12.[1275,2266,1530,3178]" pageId="12" pageNumber="305">
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B-pleurotergites up to
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<quantity box="[1708,1857,2404,2442]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="12" pageNumber="305" unit="mm" value="24.6">24.6 mm</quantity>
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in length. Anteriormost B-pleurotergite (Fig. lOb,f,h,i) distinguished by relatively short, wide proportions and short, knob-like projections at anterolateral edge of prozonite as seen in this pleurotergite only in
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<emphasis box="[1406,1629,2588,2627]" italics="true" pageId="12" pageNumber="305">
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<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[1406,1624,2588,2627]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="12" pageNumber="305" phylum="Arthropoda" rank="species" species="malcolmi">M. malcolmi</taxonomicName>
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.
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</emphasis>
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Prozonite about 25 percent length of tergite, with cuticular sculpture of dense polygonal depressions amidst a network of low ridges (Fig. lOi). Lateral lobes of metazonite teardrop shaped, outer margin rounded, strongly inflated, with steep lateral and anterior edges encircled by well-impressed furrow. Posteromedian spine on median lobe of metazonite short, conical, and posterodorsally directed or absent. Paratergal spines long, strongly divergent (directed back 40-50 degrees from transverse line), largest known specimen
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<quantity box="[2037,2162,3001,3040]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="12" pageNumber="305" unit="mm" value="88.0">88 mm</quantity>
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wide across spines (extrapolated from one half); inferred width across spines up to approximately
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<quantity box="[1873,2008,3093,3132]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="305" unit="mm" value="115.0">115 mm</quantity>
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(based on size relative to largest known B-pleurotergites); dorsal and
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4653178" ID-Zenodo-Dep="4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="13" pageNumber="306" startId="13.[222,328,3017,3053]" targetBox="[221,2271,344,2984]" targetPageId="13">
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<paragraph blockId="13.[221,2268,3017,3219]" pageId="13" pageNumber="306">
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<emphasis bold="true" box="[222,369,3017,3053]" pageId="13" pageNumber="306">Figure 9.</emphasis>
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<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[385,710,3017,3053]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="13" pageNumber="306" phylum="Arthropoda" rank="species" species="angusi" status="n.sp.">
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<emphasis box="[385,710,3017,3053]" italics="true" pageId="13" pageNumber="306">Maldybulakia angusi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[726,794,3017,3053]" pageId="13" pageNumber="306" rank="species">n.sp.</taxonomicNameLabel>
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Scale bars 10 mm. Latex casts from external moulds except
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<emphasis box="[1771,1913,3017,3053]" italics="true" pageId="13" pageNumber="306">a, d, e, i,</emphasis>
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internal moulds.
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<emphasis box="[2197,2267,3017,3053]" italics="true" pageId="13" pageNumber="306">(a,b)</emphasis>
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dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in
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<emphasis box="[1472,1498,3059,3094]" italics="true" pageId="13" pageNumber="306">a,</emphasis>
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with median flange and expanded posterolateral portion.
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<emphasis box="[361,401,3100,3136]" italics="true" pageId="13" pageNumber="306">(c)</emphasis>
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dorsal view of T-tergite AM F.102558.
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<emphasis box="[1058,1099,3100,3136]" italics="true" pageId="13" pageNumber="306">(d)</emphasis>
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dorsal view of T-tergite AM F.102559.
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<emphasis box="[1758,1798,3100,3136]" italics="true" pageId="13" pageNumber="306">(e)</emphasis>
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dorsa: view of T-tergite AM F.102560.
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<emphasis box="[391,424,3142,3178]" italics="true" pageId="13" pageNumber="306">(j)</emphasis>
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dorsal view of T-tergite AM F.102561.
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<emphasis box="[1072,1113,3142,3178]" italics="true" pageId="13" pageNumber="306">(g)</emphasis>
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dorsal view of T-tergite AM F.102562.
|
|
<emphasis box="[1763,1804,3142,3178]" italics="true" pageId="13" pageNumber="306">(h)</emphasis>
|
|
dorsal view of T-tergite AM F.102563.
|
|
<emphasis box="[388,420,3183,3219]" italics="true" pageId="13" pageNumber="306">(i)</emphasis>
|
|
anterodorsal view of ring tergite AM F.102564.
|
|
</paragraph>
|
|
</caption>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653186" ID-Zenodo-Dep="4653186" httpUri="https://zenodo.org/record/4653186/files/figure.png" pageId="14" pageNumber="307" startId="14.[208,311,2954,2989]" targetBox="[224,2263,375,2901]" targetPageId="14">
|
|
<paragraph blockId="14.[207,2252,2954,3157]" pageId="14" pageNumber="307">
|
|
<emphasis bold="true" box="[208,369,2954,2989]" pageId="14" pageNumber="307">Figure 10.</emphasis>
|
|
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[385,711,2954,2989]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="14" pageNumber="307" phylum="Arthropoda" rank="species" species="angusi" status="n.sp.">
|
|
<emphasis box="[385,711,2954,2989]" italics="true" pageId="14" pageNumber="307">Maldybulakia angusi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[725,792,2954,2989]" pageId="14" pageNumber="307" rank="species">n.sp.</taxonomicNameLabel>
|
|
Scale bars 10 mm, except
|
|
<emphasis box="[1225,1250,2954,2989]" italics="true" pageId="14" pageNumber="307">b,</emphasis>
|
|
5 mm. Latex casts from external moulds except
|
|
<emphasis box="[2023,2122,2954,2989]" italics="true" pageId="14" pageNumber="307">d, f, g,</emphasis>
|
|
internal moulds.
|
|
<emphasis box="[340,381,2996,3031]" italics="true" pageId="14" pageNumber="307">(a)</emphasis>
|
|
dorsal view of holotype B-pleurotergite AM F.102565.
|
|
<emphasis box="[1262,1302,2996,3031]" italics="true" pageId="14" pageNumber="307">(b)</emphasis>
|
|
dorsal view of first B-pleurotergite AM F.102566.
|
|
<emphasis box="[2107,2146,2996,3031]" italics="true" pageId="14" pageNumber="307">(c)</emphasis>
|
|
dorsal view of B-pleurotergite AM F.102567.
|
|
<emphasis box="[838,879,3038,3073]" italics="true" pageId="14" pageNumber="307">(d)</emphasis>
|
|
dorsal view of B-pleurotergite AM F.102568.
|
|
<emphasis box="[1640,1679,3038,3073]" italics="true" pageId="14" pageNumber="307">(e)</emphasis>
|
|
dorsal view of B-pleurotergite AM F.102569.
|
|
<emphasis box="[370,402,3079,3115]" italics="true" pageId="14" pageNumber="307">(j)</emphasis>
|
|
dorsal view of first B-pleurotergite AM F.102570.
|
|
<emphasis box="[1209,1251,3079,3115]" italics="true" pageId="14" pageNumber="307">(g)</emphasis>
|
|
dorsal view of B-pleurotergite AM F.102571.
|
|
<emphasis box="[1985,2046,3079,3115]" italics="true" pageId="14" pageNumber="307">(h,i)</emphasis>
|
|
dorsal views of first B-pleurotergite AM F.102572.
|
|
<emphasis box="[813,845,3121,3157]" italics="true" pageId="14" pageNumber="307">(i)</emphasis>
|
|
detail showing polygonal sculpture on prozonite.
|
|
</paragraph>
|
|
</caption>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653190" ID-Zenodo-Dep="4653190" httpUri="https://zenodo.org/record/4653190/files/figure.png" pageId="15" pageNumber="308" startId="15.[227,330,2817,2869]" targetBox="[232,2257,524,2786]" targetPageId="15">
|
|
<paragraph blockId="15.[222,2265,2817,3070]" pageId="15" pageNumber="308">
|
|
<emphasis bold="true" box="[227,386,2817,2869]" pageId="15" pageNumber="308">Figure 11.</emphasis>
|
|
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[401,724,2817,2869]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="15" pageNumber="308" phylum="Arthropoda" rank="species" species="angusi" status="n.sp.">
|
|
<emphasis box="[401,724,2817,2869]" italics="true" pageId="15" pageNumber="308">Maldybulakia angusi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[737,804,2817,2869]" pageId="15" pageNumber="308" rank="species">n.sp.</taxonomicNameLabel>
|
|
Scale bars 10 mm except
|
|
<emphasis box="[1217,1234,2817,2869]" italics="true" pageId="15" pageNumber="308">i,</emphasis>
|
|
5 mm. Latex casts from external moulds excepta,
|
|
<emphasis box="[2032,2056,2817,2869]" italics="true" pageId="15" pageNumber="308">b,</emphasis>
|
|
g,
|
|
<emphasis box="[2113,2138,2817,2869]" italics="true" pageId="15" pageNumber="308">h,</emphasis>
|
|
internal moulds.
|
|
<emphasis box="[365,407,2870,2910]" italics="true" pageId="15" pageNumber="308">(a)</emphasis>
|
|
dorsal view of posteriormost B-pleurotergite AM F.102573.
|
|
<emphasis box="[1410,1452,2870,2910]" italics="true" pageId="15" pageNumber="308">(b)</emphasis>
|
|
dorsal view of posteriormost B-pleurotergite AM F.102574.
|
|
<emphasis box="[388,426,2911,2950]" italics="true" pageId="15" pageNumber="308">(c)</emphasis>
|
|
dorsal view of B-pleurotergite AM F.102575, with ventral view of another B-pleurotergite.
|
|
<emphasis box="[1873,1914,2911,2950]" italics="true" pageId="15" pageNumber="308">(d)</emphasis>
|
|
dorsal view of caudal pleurotergite AM F.102576.
|
|
<emphasis box="[670,709,2951,2992]" italics="true" pageId="15" pageNumber="308">(e)</emphasis>
|
|
dorsal view of posteriormost B-pleurotergite AM F.102577.
|
|
<emphasis box="[1672,1705,2951,2992]" italics="true" pageId="15" pageNumber="308">(j)</emphasis>
|
|
dorsal view of B-pleurotergite AM F.102578.(g,h)dorsalandrightlateralviewsofcaudalpleurotergiteAMF.102579.(i)dorsalviewofspineAMF.102580,probably
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph blockId="16.[226,1220,409,2839]" pageId="16" pageNumber="309">ventral surfaces of spine bisected by shallow furrow along all but proximalmost part. Small and moderate sized tubercles abundant on lateral and median lobes of metazonite, somewhat stronger on lateral lobes.</paragraph>
|
|
<paragraph blockId="16.[226,1220,409,2839]" pageId="16" pageNumber="309">B-pleurotergite of uncertain position (Fig. lOe,g) with short, posterolaterally directed paratergal spines. Prozonite relatively elongate, about 35 percent length of tergite. Short anterolaterally-directed, conical process at anterolateral corner of metazonite. Posteromedian spine on median lobe of metazonite varying from absent (Fig. 109) to strong (Fig. lOe), variably more posteriorly- or dorsally-directed. Tubercles on lateral lobes of metazonite faint to moderately strong; tubercles weaker on median lobe, sometimes obscure.</paragraph>
|
|
<paragraph blockId="16.[226,1220,409,2839]" pageId="16" pageNumber="309">B-pleurotergites (probably second or third) from middle of tagma (Fig. lOa,c,d) with prozonite about 25 percent length of tergite. Anterolateral corner of pleurotergite extended as a short angulation that gently rises distally. Robust tubercles usually present, abundant on lateral and median lobes of metazonite. Polygonal sculpture grading into transversely elongate, scaly sculpture along anterior margin of prozonite and posterior margin of metazonite and fine tubercles on paratergal spines. Lateral lobes relatively longer than on first B-pleurotergite, less inflated. Paratergal spine long, more posteriorly directed than that on first B-pleurotergite, nearly straight or gently curved for much of length, with slightly stronger posterior curvature distally; spine flattened proximally, more rounded near tip; rounded, ventrally directed process on posterior margin at base of paratergal spine.</paragraph>
|
|
<paragraph blockId="16.[226,1220,409,2839]" pageId="16" pageNumber="309">Posteriormost B-pleurotergite (Fig. lla,b,e) with prozonite up to 35 percent length of tergite. Anterolateral corner of metazonite forming a blunt, gently swollen angulation, separated from lateral lobe by shallow furrow. Median lobe relatively narrow, raised central part set off by pair of shallow longitudinal furrows. Tuberculation of median and lateral lobes subdued in most specimens. Paratergal spine long, approximately straight, weakly directed inwards, markedly flattened in section along entire length; longitudinal furrow on dorsal and ventral surfaces of spine deeper than on other pleurotergites, narrow but well-incised; spines with sculpture of small, crowded tubercles.</paragraph>
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.4653196" ID-Zenodo-Dep="4653196" httpUri="https://zenodo.org/record/4653196/files/figure.png" pageId="16" pageNumber="309" startId="16.[1285,1398,2197,2232]" targetBox="[1313,2221,413,2109]" targetPageId="16">
|
|
<paragraph blockId="16.[1285,2272,2197,2524]" pageId="16" pageNumber="309">
|
|
<emphasis bold="true" box="[1285,1398,2197,2232]" pageId="16" pageNumber="309">Figure</emphasis>
|
|
12.
|
|
<taxonomicName authority="Edgecombe, 1998" authorityName="Edgecombe" authorityYear="1998" box="[1484,1839,2197,2232]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="angusi" status="n.sp.">
|
|
<emphasis box="[1484,1839,2197,2232]" italics="true" pageId="16" pageNumber="309">Maldybulakia angusi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1858,1932,2197,2232]" pageId="16" pageNumber="309" rank="species">n.sp.</taxonomicNameLabel>
|
|
Reconstructions of discrete morphological classes of trunk sclerites, indicating minimal number of tergites.
|
|
<emphasis box="[1750,1792,2281,2316]" italics="true" pageId="16" pageNumber="309">(a)</emphasis>
|
|
T-tergite.
|
|
<emphasis box="[1966,2008,2281,2316]" italics="true" pageId="16" pageNumber="309">(b)</emphasis>
|
|
ring tergite.
|
|
<emphasis box="[2228,2268,2281,2316]" italics="true" pageId="16" pageNumber="309">(c)</emphasis>
|
|
inferred anteriormost B-pleurotergite. Median spine on metazonite sometimes present. (d) B-pleurotergite of uncertain position. Presence of posteromedian spine is variable.
|
|
<emphasis box="[2174,2214,2406,2441]" italics="true" pageId="16" pageNumber="309">(e)</emphasis>
|
|
Bpleurotergite from middle part of tagma. (f) inferred posteriormost B-pleurotergite.
|
|
<emphasis box="[1774,1816,2489,2524]" italics="true" pageId="16" pageNumber="309">(g)</emphasis>
|
|
caudal pleurotergite.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph blockId="16.[226,1220,409,2839]" pageId="16" pageNumber="309">Caudal pleurotergite (Fig. lld,?f,g,h) with teardrop shaped or subtriangular lateral lobes; lobes substantially inflated, tuberculate, set off from median lobe by furrows of similar depth to or shallower than those on other Bpleurotergites; outer edge of lateral lobes steep, set off from more gently sloping outer border by shallow furrow. Lateral margin strongly convex outward. Median spine gently upturned, of about even width along entire known length, bisected by shallow longitudinal furrow.</paragraph>
|
|
<paragraph blockId="16.[226,1220,409,2839]" box="[274,718,2801,2839]" pageId="16" pageNumber="309">Telson sclerites unknown.</paragraph>
|
|
<paragraph blockId="16.[224,1213,2906,3175]" pageId="16" pageNumber="309">
|
|
Discussion.AlthoughMaldybulakia
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[834,945,2906,2945]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="angusi">
|
|
<emphasis box="[834,945,2906,2945]" italics="true" pageId="16" pageNumber="309">angusi</emphasis>
|
|
</taxonomicName>
|
|
is distinguished from its congenerics by numerous autapomorphies, a separate genus cannot be upheld unless reasonable synapomorphies are identified for
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[844,1070,3044,3083]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="malcolmi">
|
|
<emphasis box="[844,1070,3044,3083]" italics="true" pageId="16" pageNumber="309">M. malcolmi</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<emphasis italics="true" pageId="16" pageNumber="309">
|
|
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="mirabilis">M. mirabilis</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
As noted above, most of the similarities between the latter pair of species are probably symplesiomorphies
|
|
</paragraph>
|
|
<paragraph blockId="16.[1283,2275,2631,3175]" pageId="16" pageNumber="309">
|
|
and, if so, restricting
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[1666,1901,2631,2669]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[1666,1901,2631,2669]" italics="true" pageId="16" pageNumber="309">Maldybulakia</emphasis>
|
|
</taxonomicName>
|
|
to these two species would recognise a paraphyletic group.
|
|
<emphasis bold="true" box="[2003,2037,2676,2716]" pageId="16" pageNumber="309">In</emphasis>
|
|
fact, certain characters might instead serve to unite the two Australian species to the exclusion of
|
|
<emphasis box="[1780,2006,2768,2807]" italics="true" pageId="16" pageNumber="309">
|
|
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[1780,2001,2768,2807]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="mirabilis">M. mirabilis</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
The flattened, sloping anterior scarp on the prozonite of the T-tergite is a distinctive modification shared by these two species but appears less developed in
|
|
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[1797,2031,2906,2945]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="mirabilis">
|
|
<emphasis box="[1797,2031,2906,2945]" italics="true" pageId="16" pageNumber="309">M. mirabilis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation author="Tesakov, A. S. & A. S. Alekseev" pageId="16" pageNumber="309" pagination="18 - 23" refId="ref13855" refString="Tesakov, A. S., & A. S. Alekseev, 1992. Myriapod-like arthropods from the Lower Devonian of central Kazakhstan. Paleontological Journal 26: 18 - 23." type="journal article" year="1992">Tesakov & Alekseev, 1992</bibRefCitation>
|
|
: fig. Ig).
|
|
</paragraph>
|
|
<paragraph blockId="16.[1283,2275,2631,3175]" lastBlockId="17.[232,1223,398,2644]" lastPageId="17" lastPageNumber="310" pageId="16" pageNumber="309">
|
|
Considerable variability is observed in the strength of tuberculation on T-tergites as well as B-pleurotergites in
|
|
<emphasis box="[1283,1459,3090,3129]" italics="true" pageId="16" pageNumber="309">
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[1283,1454,3090,3129]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="16" pageNumber="309" phylum="Arthropoda" rank="species" species="angusi">M. angusi</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
Some specimens have only weak expression of tubercles (
|
|
<figureCitation box="[1461,1600,3136,3175]" captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="16" pageNumber="309">Figs. 9b</figureCitation>
|
|
, lOe, lIe), whereas most show them to be more distinct or pronounced. Because of continuous gradation in tuberculation in otherwise similar tergites this is not regarded as an important criterion for distinguishing different pleurotergites. Presence or absence of a median spine on the metazonite is a feature that varies between tergites that are otherwise similar. This is observed on the T-tergites (compare
|
|
<figureCitation box="[579,813,674,712]" captionStart="Figure 9" captionStartId="13.[222,328,3017,3053]" captionTargetBox="[221,2271,344,2984]" captionTargetPageId="13" captionText="Figure 9. Maldybulakia angusi n.sp. Scale bars 10 mm. Latex casts from external moulds except a, d, e, i, internal moulds. (a,b) dorsal views of T-tergite AM F.102557b and AM F.102557a. Note doublure in a, with median flange and expanded posterolateral portion. (c) dorsal view of T-tergite AM F.102558. (d) dorsal view of T-tergite AM F.102559. (e) dorsa: view of T-tergite AM F.102560. (j) dorsal view of T-tergite AM F.102561. (g) dorsal view of T-tergite AM F.102562. (h) dorsal view of T-tergite AM F.102563. (i) anterodorsal view of ring tergite AM F.102564." figureDoi="http://doi.org/10.5281/zenodo.4653178" httpUri="https://zenodo.org/record/4653178/files/figure.png" pageId="17" pageNumber="310">
|
|
Fig. 9b and
|
|
<quantity box="[774,813,674,712]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="17" pageNumber="310" unit="g" value="9.0">9g</quantity>
|
|
</figureCitation>
|
|
) as well as on various Btergites that appear to represent the same position (e.g., Fig. lOb versus lOh; Fig. lOe versus 109). Presence or absence of a median spine is considered to vary between individuals.
|
|
</paragraph>
|
|
<paragraph blockId="17.[232,1223,398,2644]" pageId="17" pageNumber="310">
|
|
The antero-posterior position of B-pleurotergites can be largely determined based on the progressively more posterior direction of the paratergal spines on posterior segments (as observed analogously in other spiny myriapods, such as euphoberiid diplopods). Consistent with this line of evidence are other correspondences with known positions in
|
|
<emphasis box="[457,683,1134,1173]" italics="true" pageId="17" pageNumber="310">
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[457,678,1134,1173]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="malcolmi">M. malcolmi</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
the anterior pleurotergites are shorter relative to their width, and the anteriormost Bpleurotergite is distinguished by its anterolateral processes on the prozonite (Figs. lOf, 12c; compare with
|
|
<figureCitation box="[1072,1199,1273,1311]" captionStart="Figure 6" captionStartId="10.[221,326,3025,3062]" captionTargetBox="[209,2257,404,2977]" captionTargetPageId="10" captionText="Figure 6. Maldybulakia malcolmi n.sp. Scale bars 10 mm. Ca) dorsal view of T-tergite AM F.102549. Cb) ventral view of T-tergite AM F.102550, showing doublure. (c) dorsal view of first B-pleurotergite AM F.102551. (d) dorsal view of first B-pleurotergite AM F.102552. (e) dorsal view of first B-pleurotergite AM F.102553. (j) dorsal view of B-pleurotergite AM F.102554. (g,h) dorsal and left lateral views of second B-pleurotergite AM F.102555. (i,j) right lateral and dorsal views of second B-pleurotergite AM F.102556." figureDoi="http://doi.org/10.5281/zenodo.4653168" httpUri="https://zenodo.org/record/4653168/files/figure.png" pageId="17" pageNumber="310">Fig. 6d</figureCitation>
|
|
). The posteriormost B-pleurotergite (immediately anterior to the caudal pleurotergite) is identified by its posteriorlydirected paratergal spines and relatively long median lobe on the metazonite (
|
|
<figureCitation box="[571,789,1456,1495]" captionStart="Figure 11" captionStartId="15.[227,330,2817,2869]" captionTargetBox="[232,2257,524,2786]" captionTargetPageId="15" captionText="Figure 11. Maldybulakia angusi n.sp. Scale bars 10 mm except i, 5 mm. Latex casts from external moulds excepta, b, g, h, internal moulds. (a) dorsal view of posteriormost B-pleurotergite AM F.102573. (b) dorsal view of posteriormost B-pleurotergite AM F.102574. (c) dorsal view of B-pleurotergite AM F.102575, with ventral view of another B-pleurotergite. (d) dorsal view of caudal pleurotergite AM F.102576. (e) dorsal view of posteriormost B-pleurotergite AM F.102577. (j) dorsal view of B-pleurotergite AM F.102578.(g,h)dorsalandrightlateralviewsofcaudalpleurotergiteAMF.102579.(i)dorsalviewofspineAMF.102580,probably" figureDoi="http://doi.org/10.5281/zenodo.4653190" httpUri="https://zenodo.org/record/4653190/files/figure.png" pageId="17" pageNumber="310">Figs. 11a,b,e</figureCitation>
|
|
, 12f).
|
|
</paragraph>
|
|
<paragraph blockId="17.[232,1223,398,2644]" pageId="17" pageNumber="310">
|
|
Given the above considerations, the minimal number of sclerites in the trunk of
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[675,1030,1548,1587]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="angusi">
|
|
<emphasis box="[675,1030,1548,1587]" italics="true" pageId="17" pageNumber="310">Maldybulakia angusi</emphasis>
|
|
</taxonomicName>
|
|
is depicted in Fig. 12. The B-pleurotergites can be sorted into four morphological groups (Fig. 12c-f)' excluding the caudal pleurotergite (Fig.
|
|
<quantity box="[563,624,1686,1725]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="17" pageNumber="310" unit="g" value="12.0">12g</quantity>
|
|
), thus allowing that the number of pleurotergites may be identical to that in
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[925,1137,1732,1771]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="malcolmi">
|
|
<emphasis box="[925,1137,1732,1771]" italics="true" pageId="17" pageNumber="310">M. malcolmi</emphasis>
|
|
</taxonomicName>
|
|
and, apparently,
|
|
<emphasis box="[472,726,1778,1817]" italics="true" pageId="17" pageNumber="310">
|
|
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[472,720,1778,1817]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="mirabilis">M. mirabilis</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
The position of the Bpleurotergite reconstructed in Fig. 12d is uncertain, constrained only by the likelihood that those shown in Figs. 12c and 12f are the first and last in the series based on their morphological correspondences to
|
|
<emphasis box="[917,1139,1962,2001]" italics="true" pageId="17" pageNumber="310">
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[917,1134,1962,2001]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="malcolmi">M. malcolmi</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
The specimen in
|
|
<figureCitation box="[461,602,2008,2047]" captionStart="Figure 11" captionStartId="15.[227,330,2817,2869]" captionTargetBox="[232,2257,524,2786]" captionTargetPageId="15" captionText="Figure 11. Maldybulakia angusi n.sp. Scale bars 10 mm except i, 5 mm. Latex casts from external moulds excepta, b, g, h, internal moulds. (a) dorsal view of posteriormost B-pleurotergite AM F.102573. (b) dorsal view of posteriormost B-pleurotergite AM F.102574. (c) dorsal view of B-pleurotergite AM F.102575, with ventral view of another B-pleurotergite. (d) dorsal view of caudal pleurotergite AM F.102576. (e) dorsal view of posteriormost B-pleurotergite AM F.102577. (j) dorsal view of B-pleurotergite AM F.102578.(g,h)dorsalandrightlateralviewsofcaudalpleurotergiteAMF.102579.(i)dorsalviewofspineAMF.102580,probably" figureDoi="http://doi.org/10.5281/zenodo.4653190" httpUri="https://zenodo.org/record/4653190/files/figure.png" pageId="17" pageNumber="310">Fig. 11f</figureCitation>
|
|
conforms to a caudal pleurotergite except for the absence of a median spine, although this is possibly due to breakage. Its lensoid outline (i.e., absence of paratergal spines) with tightly curved lateral margins is similar to caudal pleurotergites that have a median spine (
|
|
<figureCitation box="[249,418,2238,2277]" captionStart="Figure 11" captionStartId="15.[227,330,2817,2869]" captionTargetBox="[232,2257,524,2786]" captionTargetPageId="15" captionText="Figure 11. Maldybulakia angusi n.sp. Scale bars 10 mm except i, 5 mm. Latex casts from external moulds excepta, b, g, h, internal moulds. (a) dorsal view of posteriormost B-pleurotergite AM F.102573. (b) dorsal view of posteriormost B-pleurotergite AM F.102574. (c) dorsal view of B-pleurotergite AM F.102575, with ventral view of another B-pleurotergite. (d) dorsal view of caudal pleurotergite AM F.102576. (e) dorsal view of posteriormost B-pleurotergite AM F.102577. (j) dorsal view of B-pleurotergite AM F.102578.(g,h)dorsalandrightlateralviewsofcaudalpleurotergiteAMF.102579.(i)dorsalviewofspineAMF.102580,probably" figureDoi="http://doi.org/10.5281/zenodo.4653190" httpUri="https://zenodo.org/record/4653190/files/figure.png" pageId="17" pageNumber="310">
|
|
Fig.
|
|
<quantity box="[349,418,2238,2277]" metricMagnitude="-2" metricUnit="kg" metricValue="1.1" pageId="17" pageNumber="310" unit="g" value="11.0">11g</quantity>
|
|
</figureCitation>
|
|
).
|
|
<emphasis bold="true" box="[463,499,2238,2277]" pageId="17" pageNumber="310">In</emphasis>
|
|
<emphasis box="[519,770,2238,2277]" italics="true" pageId="17" pageNumber="310">
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[519,763,2238,2277]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="malcolmi">M. malcolmi</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
this specimen is most comparable to a lensoid specimen regarded as a caudal pleurotergite (Fig. 5c).
|
|
</paragraph>
|
|
<paragraph blockId="17.[232,1223,398,2644]" pageId="17" pageNumber="310">
|
|
The best preserved external moulds display a cuticular sculpture of dense polygonal depressions amidst a maze of low ridges (Fig. lOi), as is also seen in
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[998,1217,2467,2506]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="malcolmi">
|
|
M.
|
|
<emphasis box="[1061,1217,2467,2506]" italics="true" pageId="17" pageNumber="310">malcolmi</emphasis>
|
|
</taxonomicName>
|
|
(Fig. 5j,k) and
|
|
<taxonomicName baseAuthorityName="Tesakov & Alekseev" baseAuthorityYear="1992" box="[499,711,2513,2553]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="species" species="mirabilis">
|
|
<emphasis box="[499,711,2513,2553]" italics="true" pageId="17" pageNumber="310">M. mirabilis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation author="Tesakov, A. S. & A. S. Alekseev" box="[740,1209,2513,2553]" pageId="17" pageNumber="310" pagination="18 - 23" refId="ref13855" refString="Tesakov, A. S., & A. S. Alekseev, 1992. Myriapod-like arthropods from the Lower Devonian of central Kazakhstan. Paleontological Journal 26: 18 - 23." type="journal article" year="1992">Tesakov & Alekseev, 1992</bibRefCitation>
|
|
: fig. 1i). This character is, accordingly, general for
|
|
<emphasis box="[232,480,2606,2644]" italics="true" pageId="17" pageNumber="310">
|
|
<taxonomicName authorityName="Edgecombe" authorityYear="1998" box="[232,475,2606,2644]" class="Merostomata" family="Limulidae" genus="Maldybulakia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Xiphosurida" pageId="17" pageNumber="310" phylum="Arthropoda" rank="genus">Maldybulakia</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |