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<document id="B1967275D4FF8FD4CF80AA63095370DE" ID-CLB-Dataset="53513" ID-DOI="10.11646/zootaxa.5296.4.6" ID-GBIF-Dataset="278cb4ab-18d4-4abe-a154-ce6d9cccec35" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984426" ID-ZooBank="A667A27C-CBB4-432C-8901-8399950699B3" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685428816830" checkinUser="plazi" docAuthor="Makbun, Noppadon" docDate="2023" docId="03F0827BFFB0503DFF6CFDE36F11FF5B" docLanguage="en" docName="zootaxa.5296.4.6.pdf" docOrigin="Zootaxa 5296 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sympetrum thailandensis Makbun 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="578" masterDocId="FFC9FA03FFB55037FFFBFF896A4FFF8E" masterDocTitle="The genus Sympetrum Newman, 1833 in Thailand, with description of S. thailandensis sp. nov. (Odonata: Libellulidae)" masterLastPageNumber="581" masterPageNumber="569" pageNumber="574" updateTime="1699511272782" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0F1CB36B3E3425969ABB417C4BFAC5A0">The genus Sympetrum Newman, 1833 in Thailand, with description of S. thailandensis sp. nov. (Odonata: Libellulidae)</mods:title>
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<mods:date id="463F8E7231CCFB483AD49363DA4F5D9D">2023</mods:date>
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<treatment id="03F0827BFFB0503DFF6CFDE36F11FF5B" ID-DOI="http://doi.org/10.5281/zenodo.7990838" ID-GBIF-Taxon="209663931" ID-Zenodo-Dep="7990838" LSID="urn:lsid:plazi:treatment:03F0827BFFB0503DFF6CFDE36F11FF5B" httpUri="http://treatment.plazi.org/id/03F0827BFFB0503DFF6CFDE36F11FF5B" lastPageId="10" lastPageNumber="578" pageId="5" pageNumber="574">
<subSubSection id="C34360E6FFB05032FF6CFDE36860FD0A" box="[151,559,618,645]" pageId="5" pageNumber="574" type="nomenclature">
<paragraph id="8BE6336DFFB05032FF6CFDE36860FD0A" blockId="5.[151,559,618,680]" box="[151,559,618,645]" pageId="5" pageNumber="574">
<heading id="D0AE8401FFB05032FF6CFDE36860FD0A" bold="true" box="[151,559,618,645]" fontSize="11" level="1" pageId="5" pageNumber="574" reason="1">
<emphasis id="B92DEF7FFFB05032FF6CFDE36860FD0A" bold="true" box="[151,559,618,645]" pageId="5" pageNumber="574">
<taxonomicName id="4C5948EEFFB05032FF6CFDE36B85FD0B" ID-CoL="BHZMK" authority="Makbun, 2023" authorityName="Makbun" authorityYear="2023" box="[151,458,618,645]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="574" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B92DEF7FFFB05032FF6CFDE36B85FD0B" bold="true" box="[151,458,618,645]" italics="true" pageId="5" pageNumber="574">Sympetrum thailandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E5204FFB05032FE29FDE36860FD0A" box="[466,559,618,644]" pageId="5" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34360E6FFB05032FF6CFD076B61FD26" box="[151,302,654,680]" pageId="5" pageNumber="574" type="description">
<paragraph id="8BE6336DFFB05032FF6CFD076B61FD26" blockId="5.[151,559,618,680]" box="[151,302,654,680]" pageId="5" pageNumber="574">
(
<figureCitation id="13622FE8FFB05032FF64FD076B44FD26" box="[159,267,654,680]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,1917,1941]" captionStartId-1="6.[151,250,1489,1513]" captionStartId-2="7.[151,250,1887,1911]" captionTargetBox-0="[151,1436,1028,1893]" captionTargetBox-1="[151,1436,181,1440]" captionTargetBox-2="[151,1435,619,1849]" captionTargetId-0="figure-312@5.[151,1436,1028,1893]" captionTargetId-1="figure-18@6.[151,1436,181,1465]" captionTargetId-2="figure-192@7.[151,1436,597,1864]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 4. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) right wing in dorsal view." captionText-1="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." captionText-2="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7984434" figureDoi-1="http://doi.org/10.5281/zenodo.7984436" figureDoi-2="http://doi.org/10.5281/zenodo.7984438" httpUri-0="https://zenodo.org/record/7984434/files/figure.png" httpUri-1="https://zenodo.org/record/7984436/files/figure.png" httpUri-2="https://zenodo.org/record/7984438/files/figure.png" pageId="5" pageNumber="574">Figs. 46</figureCitation>
,
<figureCitation id="13622FE8FFB05032FEECFD076B6AFD26" box="[279,293,654,680]" captionStart="FIGURE 8" captionStartId="9.[151,250,1728,1752]" captionTargetBox="[151,1436,849,1705]" captionTargetId="figure-310@9.[151,1436,849,1705]" captionTargetPageId="9" captionText="FIGURE 8. Sympetrum thailandensis sp. nov. in life, males (ab), females (cd); (a) Hin Tung, Muang, Nakhon Nayok province, Thailand, 3-xi-2018 (photo by Noppadon Makbun); (b) same locality, 24-x-2018 (photo by Dennis Farrell); (c) same locality, 24-x-2018 (photo by Dennis Farrell); (d) same locality, 3-xi-2018 (photo by Noppadon Makbun)." figureDoi="http://doi.org/10.5281/zenodo.7984442" httpUri="https://zenodo.org/record/7984442/files/figure.png" pageId="5" pageNumber="574">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34360E6FFB05032FF6CFD5F6B54FC2B" pageId="5" pageNumber="574" type="materials_examined">
<paragraph id="8BE6336DFFB05032FF6CFD5F6FD2FD7E" blockId="5.[151,1437,726,1004]" box="[151,1437,726,755]" pageId="5" pageNumber="574">
<materialsCitation id="3B313930FFB05032FF6CFD5F6FD2FD7E" ID-GBIF-Occurrence="4121715306" box="[151,1437,726,755]" collectingDate="2018-11-03" collectorName="N. Makbun" country="Thailand" elevation="1235" location="Hin Tang" municipality="Mueng" pageId="5" pageNumber="574" specimenCount="1" stateProvince="Nakhon Nayok" typeStatus="holotype">
<emphasis id="B92DEF7FFFB05032FF6CFD5F6B44FD7E" bold="true" box="[151,267,726,752]" pageId="5" pageNumber="574">
<typeStatus id="54E28DCFFFB05032FF6CFD5F6B49FD7E" box="[151,262,726,752]" pageId="5" pageNumber="574" type="holotype">Holotype</typeStatus>
.
</emphasis>
1 ♁,
<collectingCountry id="F34E73FDFFB05032FEABFD5F6BF9FD7E" box="[336,438,726,752]" name="Thailand" pageId="5" pageNumber="574">Thailand</collectingCountry>
,
<collectingRegion id="499DFD8FFFB05032FE3FFD5F6890FD7E" box="[452,735,726,752]" country="Thailand" name="Nakhon Nayok" pageId="5" pageNumber="574">Nakhon Nayok province</collectingRegion>
,
<collectingMunicipality id="6B82A917FFB05032FD17FD5E6971FD7F" box="[748,830,727,753]" pageId="5" pageNumber="574">Mueng</collectingMunicipality>
,
<location id="8E8665B6FFB05032FCB7FD5F69F8FD7F" LSID="urn:lsid:plazi:treatment:03F0827BFFB0503DFF6CFDE36F11FF5B:8E8665B6FFB05032FCB7FD5F69F8FD7F" box="[844,951,726,753]" country="Thailand" municipality="Mueng" name="Hin Tang" pageId="5" pageNumber="574" stateProvince="Nakhon Nayok">Hin Tang</location>
(
<elevation id="0074D45EFFB05032FC33FD5F6E1DFD7E" box="[968,1106,726,753]" metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" pageId="5" pageNumber="574" unit="m" value="1235.0">
<quantity id="4CA19E88FFB05032FC33FD5F6E68FD7F" box="[968,1063,726,753]" metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" pageId="5" pageNumber="574" unit="m" value="1235.0">1,235 m</quantity>
asl
</elevation>
),
<date id="FFE715ADFFB05032FB92FD5F6E95FD7E" box="[1129,1242,726,752]" pageId="5" pageNumber="574" value="2018-11-03">
<collectingDate id="EFA3EC45FFB05032FB92FD5F6E95FD7E" box="[1129,1242,726,752]" pageId="5" pageNumber="574" value="2018-11-03">3-xi-2018</collectingDate>
</date>
,
<collectorName id="26AC56BBFFB05032FB13FD5E6F26FD7E" box="[1256,1385,726,753]" pageId="5" pageNumber="574">N. Makbun</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="8BE6336DFFB05032FF6CFD7369BBFC9A" blockId="5.[151,1437,726,1004]" box="[151,1012,762,791]" pageId="5" pageNumber="574">
<emphasis id="B92DEF7FFFB05032FF6CFD736B59FC9A" bold="true" box="[151,278,762,788]" pageId="5" pageNumber="574">
<typeStatus id="54E28DCFFFB05032FF6CFD736B5EFC9A" box="[151,273,762,788]" pageId="5" pageNumber="574" type="paratype">Paratypes</typeStatus>
.
</emphasis>
3 ♁,
<specimenCount id="9D5FF8E4FFB05032FEAEFD736B30FC9B" box="[341,383,762,789]" pageId="5" pageNumber="574" type="female">1 ♀</specimenCount>
, locality and date same as for
<typeStatus id="54E28DCFFFB05032FD2FFD736976FC9A" box="[724,825,762,788]" pageId="5" pageNumber="574" type="holotype">holotype</typeStatus>
, N. Makbun leg.
</paragraph>
<paragraph id="8BE6336DFFB05032FF3CFC976B54FC2B" blockId="5.[151,1437,726,1004]" pageId="5" pageNumber="574">
<emphasis id="B92DEF7FFFB05032FF3CFC9768C4FCB6" bold="true" box="[199,651,798,824]" pageId="5" pageNumber="574">Additional photographic observations</emphasis>
.
<materialsCitation id="3B313930FFB05032FD63FC9769FAFCD2" ID-GBIF-Occurrence="4121715310" collectingDate="2018-10-09" collectingDateMax="2018-10-25" collectingDateMin="2018-10-09" country="Thailand" location="Hin Tang" municipality="Mueng" pageId="5" pageNumber="574" specimenCount="1" stateProvince="Nakhon Nayok">
1 ♁,
<collectingCountry id="F34E73FDFFB05032FD35FC97697BFCB6" box="[718,820,798,824]" name="Thailand" pageId="5" pageNumber="574">Thailand</collectingCountry>
,
<collectingRegion id="499DFD8FFFB05032FCC5FC976E1CFCB6" box="[830,1107,798,824]" country="Thailand" name="Nakhon Nayok" pageId="5" pageNumber="574">Nakhon Nayok province</collectingRegion>
,
<collectingMunicipality id="6B82A917FFB05032FBA6FC966EE0FCB7" box="[1117,1199,799,825]" pageId="5" pageNumber="574">Mueng</collectingMunicipality>
,
<location id="8E8665B6FFB05032FB41FC976F6BFCB7" LSID="urn:lsid:plazi:treatment:03F0827BFFB0503DFF6CFDE36F11FF5B:8E8665B6FFB05032FB41FC976F6BFCB7" box="[1210,1316,798,825]" country="Thailand" municipality="Mueng" name="Hin Tang" pageId="5" pageNumber="574" stateProvince="Nakhon Nayok">Hin Tang</location>
,
<date id="FFE715ADFFB05032FAD4FC976FD7FCB6" box="[1327,1432,798,824]" pageId="5" pageNumber="574" value="2018-10-09">
<collectingDate id="EFA3EC45FFB05032FAD4FC976FD7FCB6" box="[1327,1432,798,824]" pageId="5" pageNumber="574" value="2018-10-09">9-x-2018</collectingDate>
</date>
, Reinthong Ruangrong; 1 ♁, same locality,
<date id="FFE715ADFFB05032FD73FCCB694FFCD3" box="[648,768,834,861]" pageId="5" pageNumber="574" value="2018-10-25">
<collectingDate id="EFA3EC45FFB05032FD73FCCB694FFCD3" box="[648,768,834,861]" pageId="5" pageNumber="574" value="2018-10-25">25-x-2018</collectingDate>
</date>
, Dennis Farrell
</materialsCitation>
;
<materialsCitation id="3B313930FFB05032FC3FFCCB6B19FC0E" ID-GBIF-Occurrence="4121715302" collectingDate="2020-07-26" collectingDateMax="2018-10-25" collectingDateMin="2018-10-09" country="Thailand" location="Hin Tang" municipality="Mueng" pageId="5" pageNumber="574" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Nayok province">
<specimenCount id="9D5FF8E4FFB05032FC3FFCCB69BEFCD3" box="[964,1009,834,861]" pageId="5" pageNumber="574" type="female">1 ♀</specimenCount>
(teneral), same locality,
<date id="FFE715ADFFB05032FAEAFCCB6FD7FCD2" box="[1297,1432,834,860]" pageId="5" pageNumber="574" value="2020-07-26">
<collectingDate id="EFA3EC45FFB05032FAEAFCCB6FD7FCD2" box="[1297,1432,834,860]" pageId="5" pageNumber="574" value="2020-07-26">26-vii-2020</collectingDate>
</date>
, Dennis Farrell; 1
</materialsCitation>
♁,
<materialsCitation id="3B313930FFB05032FE79FCEF6B58FC2B" ID-GBIF-Occurrence="4121715312" collectingDate="2020-10-03" collectingDateMax="2021-08-23" collectingDateMin="2020-10-03" country="Thailand" location="Hin Tang" municipality="Mueng" pageId="5" pageNumber="574" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Nayok province">
<specimenCount id="9D5FF8E4FFB05032FE79FCEF6BE2FC0F" box="[386,429,870,897]" pageId="5" pageNumber="574" type="female">1 ♀</specimenCount>
(copula), same locality,
<date id="FFE715ADFFB05032FD38FCEF6963FC0E" box="[707,812,870,896]" pageId="5" pageNumber="574" value="2020-10-03">
<collectingDate id="EFA3EC45FFB05032FD38FCEF6963FC0E" box="[707,812,870,896]" pageId="5" pageNumber="574" value="2020-10-03">3-x-2020</collectingDate>
</date>
, John Sim, 1 ♁, same locality,
<date id="FFE715ADFFB05032FB76FCEF6F53FC0E" box="[1165,1308,870,896]" pageId="5" pageNumber="574" value="2021-08-23">
<collectingDate id="EFA3EC45FFB05032FB76FCEF6F53FC0E" box="[1165,1308,870,896]" pageId="5" pageNumber="574" value="2021-08-23">23-viii-2021</collectingDate>
</date>
, Reinthong Ruangrong
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34360E6FFB05032FF3CFC276B57FC62" pageId="5" pageNumber="574" type="etymology">
<paragraph id="8BE6336DFFB05032FF3CFC276B57FC62" blockId="5.[151,1437,726,1004]" pageId="5" pageNumber="574">
<emphasis id="B92DEF7FFFB05032FF3CFC276B01FC46" bold="true" box="[199,334,942,968]" pageId="5" pageNumber="574">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C5948EEFFB05032FDB7FC2768AEFC46" authorityName="Makbun" authorityYear="2023" box="[588,737,942,968]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="574" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B92DEF7FFFB05032FDB7FC2768AEFC46" box="[588,737,942,968]" italics="true" pageId="5" pageNumber="574">thailandensis</emphasis>
</taxonomicName>
” refers to the country,
<collectingCountry id="F34E73FDFFB05032FC16FC276E1CFC46" box="[1005,1107,942,968]" name="Thailand" pageId="5" pageNumber="574">Thailand</collectingCountry>
, where this new species was discovered.
</paragraph>
</subSubSection>
<caption id="DF2663E5FFB05032FF6CF8F46966F837" ID-DOI="http://doi.org/10.5281/zenodo.7984434" ID-Zenodo-Dep="7984434" httpUri="https://zenodo.org/record/7984434/files/figure.png" pageId="5" pageNumber="574" startId="5.[151,250,1917,1941]" targetBox="[151,1436,1028,1893]" targetPageId="5" targetType="figure">
<paragraph id="8BE6336DFFB05032FF6CF8F46966F837" blockId="5.[151,1436,1917,1977]" pageId="5" pageNumber="574">
<emphasis id="B92DEF7FFFB05032FF6CF8F46B5DF81B" bold="true" box="[151,274,1917,1941]" pageId="5" pageNumber="574">FIGURE 4.</emphasis>
General morphology of
<taxonomicName id="4C5948EEFFB05032FDF3F8F46949F81B" authority="Makbun, 2023" authorityName="Makbun" authorityYear="2023" box="[520,774,1917,1941]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="574" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B92DEF7FFFB05032FDF3F8F46949F81B" box="[520,774,1917,1941]" italics="true" pageId="5" pageNumber="574">Sympetrum thailandensis</emphasis>
</taxonomicName>
<emphasis id="B92DEF7FFFB05032FCF0F8F46915F81B" bold="true" box="[779,858,1917,1941]" pageId="5" pageNumber="574">
<taxonomicNameLabel id="A21E5204FFB05032FCF0F8F46915F81B" box="[779,858,1917,1941]" pageId="5" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♁; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) right wing in dorsal view.
</paragraph>
</caption>
<caption id="DF2663E5FFB35031FF6CFA5868D1F9BC" ID-DOI="http://doi.org/10.5281/zenodo.7984436" ID-Zenodo-Dep="7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="6" pageNumber="575" startId="6.[151,250,1489,1513]" targetBox="[151,1436,181,1440]" targetPageId="6" targetType="figure">
<paragraph id="8BE6336DFFB35031FF6CFA5868D1F9BC" blockId="6.[151,1436,1489,1586]" pageId="6" pageNumber="575">
<emphasis id="B92DEF7FFFB35031FF6CFA586B5CFA64" bold="true" box="[151,275,1489,1514]" pageId="6" pageNumber="575">FIGURE 5.</emphasis>
General morphology of
<taxonomicName id="4C5948EEFFB35031FDF5FA5B6941FA64" authority="Makbun, 2023" authorityName="Makbun" authorityYear="2023" box="[526,782,1490,1514]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="575" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B92DEF7FFFB35031FDF5FA5B6941FA64" box="[526,782,1490,1514]" italics="true" pageId="6" pageNumber="575">Sympetrum thailandensis</emphasis>
</taxonomicName>
<emphasis id="B92DEF7FFFB35031FCEEFA5B692AFA64" bold="true" box="[789,869,1490,1514]" pageId="6" pageNumber="575">
<taxonomicNameLabel id="A21E5204FFB35031FCEEFA5B692AFA64" box="[789,869,1490,1514]" pageId="6" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view.
</paragraph>
</caption>
<subSubSection id="C34360E6FFB3503EFF6CF9E96F6EFE93" lastPageId="9" lastPageNumber="578" pageId="6" pageNumber="575" type="description">
<paragraph id="8BE6336DFFB35031FF6CF9E96860F9F5" blockId="6.[151,1437,1632,2018]" box="[151,559,1632,1659]" pageId="6" pageNumber="575">
<emphasis id="B92DEF7FFFB35031FF6CF9E96BE1F9F4" bold="true" box="[151,430,1632,1658]" pageId="6" pageNumber="575">
Description of
<typeStatus id="54E28DCFFFB35031FEBCF9E96BE1F9F4" box="[327,430,1632,1658]" pageId="6" pageNumber="575" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="13622FE8FFB35031FE46F9E96867F9F5" box="[445,552,1632,1659]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="5.[151,250,1917,1941]" captionStartId-1="6.[151,250,1489,1513]" captionTargetBox-0="[151,1436,1028,1893]" captionTargetBox-1="[151,1436,181,1440]" captionTargetId-0="figure-312@5.[151,1436,1028,1893]" captionTargetId-1="figure-18@6.[151,1436,181,1465]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 4. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) right wing in dorsal view." captionText-1="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.7984434" figureDoi-1="http://doi.org/10.5281/zenodo.7984436" httpUri-0="https://zenodo.org/record/7984434/files/figure.png" httpUri-1="https://zenodo.org/record/7984436/files/figure.png" pageId="6" pageNumber="575">Figs. 45</figureCitation>
)
</paragraph>
<paragraph id="8BE6336DFFB35031FF3CF90D6808F8A0" blockId="6.[151,1437,1632,2018]" pageId="6" pageNumber="575">
<emphasis id="B92DEF7FFFB35031FF3CF90D6B4BF910" box="[199,260,1668,1694]" italics="true" pageId="6" pageNumber="575">Head</emphasis>
covered with black setae (
<figureCitation id="13622FE8FFB35031FDCFF90D68FFF910" box="[564,688,1668,1694]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1941]" captionTargetBox="[151,1436,1028,1893]" captionTargetId="figure-312@5.[151,1436,1028,1893]" captionTargetPageId="5" captionText="FIGURE 4. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) right wing in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7984434" httpUri="https://zenodo.org/record/7984434/files/figure.png" pageId="6" pageNumber="575">Figs. 4ab</figureCitation>
). Compound eyes brownish yellow (upper part red and lower part brown in life). Occipital triangle reddish. Vertex black with reddish yellow dorsally. Antenna entirely black. Postfrons reddish yellow with a black basal band, but the margin of the band ambiguous. Antefrons and postclypeus reddish yellow. Anteclypeus, gena, mandible and labrum yellow. In the labium, the middle lobe yellow with narrow black inner margin in the lateral lobes.
</paragraph>
<paragraph id="8BE6336DFFB35030FF3CF8B169ABFF5B" blockId="6.[151,1437,1632,2018]" lastBlockId="7.[151,1437,151,573]" lastPageId="7" lastPageNumber="576" pageId="6" pageNumber="575">
<emphasis id="B92DEF7FFFB35031FF3CF8B16B57F8DC" box="[199,280,1848,1874]" italics="true" pageId="6" pageNumber="575">Thorax</emphasis>
(
<figureCitation id="13622FE8FFB35031FED3F8B16B34F8DC" box="[296,379,1848,1874]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1941]" captionTargetBox="[151,1436,1028,1893]" captionTargetId="figure-312@5.[151,1436,1028,1893]" captionTargetPageId="5" captionText="FIGURE 4. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) right wing in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7984434" httpUri="https://zenodo.org/record/7984434/files/figure.png" pageId="6" pageNumber="575">Fig. 4c</figureCitation>
). Prothorax black, posterior lobe red dorsally with a fringe of long brown setae on upper margin. Synthorax covered with long setae. Mesepisternum and antelar sinus brown (reddish brown in life). Humeral stripe black and thick. Mesepimeron and metepisterum red with black at lower part, connecting with the humeral stripe. Black triangular stripe covered interpleural suture and metastigma, reaching about half of its length. Metapleural suture covered with black narrow stripe. The area between the interpleural and metapleural stripes blackish brown. Metepimeron entirely red. Leg black except coxa and trochanter reddish brown; inner sides of pro-femur yellowish; femur with short robust spines; tibia with numerous long spines.
</paragraph>
<paragraph id="8BE6336DFFB25030FF3CFF566AA9FE07" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="576">
<emphasis id="B92DEF7FFFB25030FF3CFF566B44FF76" box="[199,267,223,248]" italics="true" pageId="7" pageNumber="576">Wings</emphasis>
brown with small yellow basal patch (
<figureCitation id="13622FE8FFB25030FD35FF56696BFF77" box="[718,804,223,249]" captionStart="FIGURE 4" captionStartId="5.[151,250,1917,1941]" captionTargetBox="[151,1436,1028,1893]" captionTargetId="figure-312@5.[151,1436,1028,1893]" captionTargetPageId="5" captionText="FIGURE 4. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) right wing in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7984434" httpUri="https://zenodo.org/record/7984434/files/figure.png" pageId="7" pageNumber="576">Fig. 4d</figureCitation>
). Pt brown with darken costal margin, covering 2 cells below. Ax
<quantity id="4CA19E88FFB25030FEF5FE8B6B02FE93" box="[270,333,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" pageId="7" pageNumber="576" unit="in" value="8.5">8.5 in</quantity>
FW,
<quantity id="4CA19E88FFB25030FE73FE8A6BFDFE93" box="[392,434,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="576" unit="in" value="6.0">6 in</quantity>
HW. Px 7 (right)8 (left) in FW, 7 (left)8 (right) in HW. Arc arched and situated between the first and the second antenodal crossveins. Median space entire in both wings. Cubital space with 1 crossvein in both wings. Triangle with one crossvein in FW and entire in HW. Anal loop 18 (left)19 (right) in HW. Membranule brown.
</paragraph>
<paragraph id="8BE6336DFFB25030FF3CFE1A6B0FFE7B" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="576">
<emphasis id="B92DEF7FFFB25030FF3CFE1A6B7EFE23" box="[199,305,403,429]" italics="true" pageId="7" pageNumber="576">Abdomen</emphasis>
typical for the genus, bright red in color. S1 reddish brown. S2 red with black basally. S37 entirely red and unmarked. S89 red with black middorsal marking and apico-lateral band. S10 black with two red lateral irregular spots.
</paragraph>
<paragraph id="8BE6336DFFB25030FF3CFE766F14FDB3" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="576">
<emphasis id="B92DEF7FFFB25030FF3CFE766BE7FD97" box="[199,424,511,537]" italics="true" pageId="7" pageNumber="576">Secondary genitalia</emphasis>
(
<figureCitation id="13622FE8FFB25030FE4DFE76684AFD97" box="[438,517,510,537]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="7" pageNumber="576">Fig. 5a</figureCitation>
) entire black with yellow long setae. Genital lobe erect, semicircular. Hamule long triangular, blunt tip, directing posteroventrally, with pale triangular inner hook. Anterior lamina semicircular.
</paragraph>
<caption id="DF2663E5FFB25030FF6CF8D66B9EF831" ID-DOI="http://doi.org/10.5281/zenodo.7984438" ID-Zenodo-Dep="7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="7" pageNumber="576" startId="7.[151,250,1887,1911]" targetBox="[151,1435,619,1849]" targetPageId="7" targetType="figure">
<paragraph id="8BE6336DFFB25030FF6CF8D66B9EF831" blockId="7.[151,1437,1887,1983]" pageId="7" pageNumber="576">
<emphasis id="B92DEF7FFFB25030FF6CF8D66B5DF8F9" bold="true" box="[151,274,1887,1911]" pageId="7" pageNumber="576">FIGURE 6.</emphasis>
General morphology of
<taxonomicName id="4C5948EEFFB25030FDF3F8E96949F8F9" authority="Makbun, 2023" authorityName="Makbun" authorityYear="2023" box="[520,774,1887,1911]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="576" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B92DEF7FFFB25030FDF3F8E96949F8F9" box="[520,774,1887,1911]" italics="true" pageId="7" pageNumber="576">Sympetrum thailandensis</emphasis>
</taxonomicName>
<emphasis id="B92DEF7FFFB25030FCF7F8E96914F8F6" bold="true" box="[780,859,1888,1912]" pageId="7" pageNumber="576">
<taxonomicNameLabel id="A21E5204FFB25030FCF7F8E96914F8F6" box="[780,859,1888,1912]" pageId="7" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view.
</paragraph>
</caption>
<caption id="DF2663E5FFBD503FFF6CFBAC6E19FB0B" ID-DOI="http://doi.org/10.5281/zenodo.7984440" ID-Zenodo-Dep="7984440" httpUri="https://zenodo.org/record/7984440/files/figure.png" pageId="8" pageNumber="577" startId="8.[151,250,1061,1085]" targetBox="[151,1436,181,1037]" targetPageId="8" targetType="figure">
<paragraph id="8BE6336DFFBD503FFF6CFBAC6E19FB0B" blockId="8.[151,1437,1061,1157]" pageId="8" pageNumber="577">
<emphasis id="B92DEF7FFFBD503FFF6CFBAC6B5BFBB3" bold="true" box="[151,276,1061,1085]" pageId="8" pageNumber="577">FIGURE 7.</emphasis>
<taxonomicName id="4C5948EEFFBD503FFEE7FBAF684EFBB3" baseAuthorityName="Selys" baseAuthorityYear="1884" box="[284,513,1061,1085]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="577" phylum="Arthropoda" rank="species" species="hypomelas">
<emphasis id="B92DEF7FFFBD503FFEE7FBAF684EFBB3" box="[284,513,1061,1085]" italics="true" pageId="8" pageNumber="577">Sympetrum hypomelas</emphasis>
</taxonomicName>
in life, males (ab), females (cd); (a) Hin Tung, Muang, Nakhon Nayok province, Thailand, 26vii2020 (photo by Dennis Farrell); (b) same locality, 3xi2018 (photo by Noppadon Makbun); (c) same locality, 24x2018 (photo by Dennis Farrell); (d) same locality, 24x2018 (photo by Dennis Farrell).
</paragraph>
</caption>
<paragraph id="8BE6336DFFBD503FFF3CFB3D6E77FA0C" blockId="8.[151,1437,1203,1770]" pageId="8" pageNumber="577">
<emphasis id="B92DEF7FFFBD503FFF3CFB3D6BCAFB43" box="[199,389,1203,1229]" italics="true" pageId="8" pageNumber="577">Vesica spermalis</emphasis>
(
<figureCitation id="13622FE8FFBD503FFE6CFB3A685FFB40" box="[407,528,1203,1230]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Figs. 5bc</figureCitation>
). V1 black, large, rather semicircular in lateral view, covered with yellow lone setae. V2 pale, tubular with wrinkled surface, slightly expanded apically. V3 dark, short, and curved posteroventally. V4 dark, spoon-shaped; medial lobe well-developed, curved, covered most of cornua except its apical part (
<figureCitation id="13622FE8FFBD503FFAC6FB726FC0FA98" box="[1341,1423,1275,1302]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Fig. 5b</figureCitation>
). The visible apical part of cornua filamentous with pointed tip, curved downward in lateral view. In ventral view, V4 semicircular with ventral crest (or medial nodule) at middle; medial lobe triangular covering cornua except its apical part; cornua bending sideway with exposed apical part curved downward (
<figureCitation id="13622FE8FFBD503FFC22FAEE6E64FA0C" box="[985,1067,1383,1410]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Fig. 5c</figureCitation>
).
</paragraph>
<paragraph id="8BE6336DFFBD503FFF3CFA026E8BF9B8" blockId="8.[151,1437,1203,1770]" pageId="8" pageNumber="577">
<emphasis id="B92DEF7FFFBD503FFF3CFA026BC6FA2B" box="[199,393,1419,1445]" italics="true" pageId="8" pageNumber="577">Anal appendages</emphasis>
(
<figureCitation id="13622FE8FFBD503FFE63FA02685FFA28" box="[408,528,1419,1446]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Figs. 5df</figureCitation>
). Cerci, about 3.5 times as long as S10. In lateral view, cerci curved with pointed tip, the ventral margin possess a row of 6 small blunt subapical teeth (
<figureCitation id="13622FE8FFBD503FFC52FA2669B2FA44" box="[937,1021,1455,1482]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Fig. 5d</figureCitation>
). In dorsal view, cerci black with red basally, straight and slightly convergent, broad basally, with pointed tip, directed outward (
<figureCitation id="13622FE8FFBD503FFB6EFA5A6EA8FA60" box="[1173,1255,1491,1518]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Fig. 5e</figureCitation>
). Paraprocts 0.8 times as long as cerci. In lateral view, paraproct curved upward at middle, tapered to its hooked tip. In ventral view paraprocts triangular, broad at base, concaved subbasally and tapered to its blunt apex (
<figureCitation id="13622FE8FFBD503FFB93F9926EF7F9B8" box="[1128,1208,1563,1590]" captionStart="FIGURE 5" captionStartId="6.[151,250,1489,1513]" captionTargetBox="[151,1436,181,1440]" captionTargetId="figure-18@6.[151,1436,181,1465]" captionTargetPageId="6" captionText="FIGURE 5. General morphology of Sympetrum thailandensis sp. nov., holotype ♁; (a) secondary genitalia in lateral view; (b) vesica spermalis in lateral view; (c) vesica spermalis in ventral view; (d) anal appendages in lateral view; (e) anal appendages in dorsal view; (f) anal appendages in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984436" httpUri="https://zenodo.org/record/7984436/files/figure.png" pageId="8" pageNumber="577">Fig. 5f</figureCitation>
).
</paragraph>
<paragraph id="8BE6336DFFBD503FFF3CF9C96F6EF9D4" blockId="8.[151,1437,1203,1770]" box="[199,1313,1599,1626]" pageId="8" pageNumber="577">
<emphasis id="B92DEF7FFFBD503FFF3CF9C96B39F9D4" bold="true" box="[199,374,1600,1626]" pageId="8" pageNumber="577">Measurements</emphasis>
(in mm). HW 26.64, abdomen including appendages 22.38, total body length 35.04
</paragraph>
<paragraph id="8BE6336DFFBD503FFF3CF9EA697CF967" blockId="8.[151,1437,1203,1770]" pageId="8" pageNumber="577">
<emphasis id="B92DEF7FFFBD503FFF3CF9EA6865F9F0" bold="true" box="[199,554,1635,1662]" pageId="8" pageNumber="577">
Variations in male
<typeStatus id="54E28DCFFFBD503FFE56F9ED686AF9F0" box="[429,549,1636,1662]" pageId="8" pageNumber="577" type="paratype">paratypes</typeStatus>
.
</emphasis>
Male
<typeStatus id="54E28DCFFFBD503FFD8EF9ED68AFF9F0" box="[629,736,1636,1662]" pageId="8" pageNumber="577" type="paratype">paratypes</typeStatus>
are larger in size, HW, length of abdomen including anal appendages and total body length (27.5028.16, 22.7423.40, and 35.1437.33, respectively). Ax and Px in HW are more than the
<typeStatus id="54E28DCFFFBD503FFEC1F9226BD3F94B" box="[314,412,1707,1733]" pageId="8" pageNumber="577" type="holotype">holotype</typeStatus>
(7 and 89, respectively). The number of cells in anal loop of
<specimenCount id="9D5FF8E4FFBD503FFB9BF9256EBFF948" box="[1120,1264,1708,1734]" pageId="8" pageNumber="577" type="generic" typeStatus="paratype">one paratype</specimenCount>
is less (1718) and the other one is more (1920) than that of the
<typeStatus id="54E28DCFFFBD503FFD32F9466960F967" box="[713,815,1743,1769]" pageId="8" pageNumber="577" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BE6336DFFBD503FFF6CF89E682BF8BF" blockId="8.[151,1437,1815,2022]" box="[151,612,1815,1842]" pageId="8" pageNumber="577">
<emphasis id="B92DEF7FFFBD503FFF6CF89E6847F8BF" bold="true" box="[151,520,1815,1842]" pageId="8" pageNumber="577">
Description of
<typeStatus id="54E28DCFFFBD503FFEBCF8916BFDF8BC" box="[327,434,1816,1842]" pageId="8" pageNumber="577" type="paratype">paratype</typeStatus>
female
</emphasis>
(
<figureCitation id="13622FE8FFBD503FFDE3F89E6814F8BF" box="[536,603,1815,1841]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="8" pageNumber="577">Fig. 6</figureCitation>
)
</paragraph>
<paragraph id="8BE6336DFFBD503EFF3CF8B26937FF77" blockId="8.[151,1437,1815,2022]" lastBlockId="9.[151,1436,151,825]" lastPageId="9" lastPageNumber="578" pageId="8" pageNumber="577">
Similar to
<typeStatus id="54E28DCFFFBD503FFEC4F8B26BEEF8DB" box="[319,417,1851,1877]" pageId="8" pageNumber="577" type="holotype">holotype</typeStatus>
male unless otherwise stated (
<figureCitation id="13622FE8FFBD503FFD00F8B26901F8DB" box="[763,846,1851,1877]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="8" pageNumber="577">Fig. 6a</figureCitation>
). Postfrons, antefrons and postclypeus yellow (
<figureCitation id="13622FE8FFBD503FFA9FF8B26AF9F8F7" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="8" pageNumber="577">Figs. 6b</figureCitation>
). Side of synthorax pale yellow covered with a black complete stripe on humeral suture and two incomplete stripes covered separately on metapleural and interpleural sutures. The area between the interpleural and metapleural stripes blackish brown (
<figureCitation id="13622FE8FFBD503FFE3FF82E685AF84F" box="[452,533,1959,1985]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="8" pageNumber="577">Fig. 6c</figureCitation>
). Wings brownish with yellow patch at the base, Px
<quantity id="4CA19E88FFBD503FFBA3F8216ECDF84F" box="[1112,1154,1959,1986]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="577" unit="in" value="8.0">8 in</quantity>
FW and 7 (right)8 (left) in HW. Anal loop 19 (right) and 21 (left) in HW. Abdomen cylindrical, yellow with red dorsally; S7 with black apico-lateral spot; S89 with black apico-lateral marking and middorsal spots; S10 black with reddish yellow lateral spot (
<figureCitation id="13622FE8FFBC503EFF2DFF326B65FF5B" box="[214,298,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="9" pageNumber="578">Fig. 6d</figureCitation>
). Cerci black, conical with tip pointed sideway, about 1.8 times as long as S10, in top view (
<figureCitation id="13622FE8FFBC503EFAC6FF326FC0FF5B" box="[1341,1423,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="9" pageNumber="578">Fig. 6e</figureCitation>
). Valvula vulvae black, short, bluntly triangular-tip bilobe (
<figureCitation id="13622FE8FFBC503EFCE7FF566924FF77" box="[796,875,223,249]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[151,1435,619,1849]" captionTargetId="figure-192@7.[151,1436,597,1864]" captionTargetPageId="7" captionText="FIGURE 6. General morphology of Sympetrum thailandensis sp. nov., paratype ♀; (a) habitus in lateral view; (b) head in dorsal view; (c) synthorax in lateral view; (d) abdominal S610 in lateral view; (e) anal appendages in dorsal view; (f) valvula vulvae (white outline) in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7984438" httpUri="https://zenodo.org/record/7984438/files/figure.png" pageId="9" pageNumber="578">Fig. 6f</figureCitation>
).
</paragraph>
<paragraph id="8BE6336DFFBC503EFF3CFE8A6F6EFE93" blockId="9.[151,1436,151,825]" box="[199,1313,258,285]" pageId="9" pageNumber="578">
<emphasis id="B92DEF7FFFBC503EFF3CFE8A6B39FE93" bold="true" box="[199,374,259,285]" pageId="9" pageNumber="578">Measurements</emphasis>
(in mm). HW 29.51, abdomen including appendages 23.10, total body length 36.13
</paragraph>
</subSubSection>
<subSubSection id="C34360E6FFBC503DFF3CFEAF6F11FF5B" lastPageId="10" lastPageNumber="579" pageId="9" pageNumber="578" type="diagnosis">
<paragraph id="8BE6336DFFBC503EFF3CFEAF6EEAFE5F" blockId="9.[151,1436,151,825]" pageId="9" pageNumber="578">
<emphasis id="B92DEF7FFFBC503EFF3CFEAF6B82FECE" bold="true" box="[199,461,294,320]" pageId="9" pageNumber="578">Differential diagnosis.</emphasis>
Following
<bibRefCitation id="EFC84E9CFFBC503EFDB5FEAE69D3FECF" author="Pilgrim, E. M. &amp; von Dohlen, C. D." box="[590,924,295,321]" pageId="9" pageNumber="578" pagination="281 - 295" refId="ref7118" refString="Pilgrim, E. M. &amp; von Dohlen, C. D. (2012) Phylogeny of the dragonfly genus Sympetrum (Odonata: Libellulidae). Organisms, Diversity and Evolution, 12 (3), 281 - 295. https: // doi. org / 10.1007 / s 13127 - 012 - 0081 - 7" type="journal article" year="2012">Pilgrim &amp; von Dohlen (2012)</bibRefCitation>
, who categorised
<taxonomicName id="4C5948EEFFBC503EFB9FFEAE6EADFECF" authorityName="Newman" authorityYear="1833" box="[1124,1250,295,321]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEF7FFFBC503EFB9FFEAE6EADFECF" box="[1124,1250,295,321]" italics="true" pageId="9" pageNumber="578">Sympetrum</emphasis>
</taxonomicName>
into six speciesgroups based on penile morphology,
<taxonomicName id="4C5948EEFFBC503EFDC7FEC268A0FEEB" authorityName="Makbun" authorityYear="2023" box="[572,751,331,357]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B92DEF7FFFBC503EFDC7FEC268A0FEEB" box="[572,751,331,357]" italics="true" pageId="9" pageNumber="578">S. thailandensis</emphasis>
</taxonomicName>
belongs to
<taxonomicName id="4C5948EEFFBC503EFC8DFEC269BEFEEB" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[886,1009,331,357]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="infuscatum">
<emphasis id="B92DEF7FFFBC503EFC8DFEC269BEFEEB" box="[886,1009,331,357]" italics="true" pageId="9" pageNumber="578">infuscatum</emphasis>
</taxonomicName>
-group. With five more species added to the group as
<bibRefCitation id="EFC84E9CFFBC503EFEBFFEE66842FE07" author="Tang, H. - C. &amp; Yeh, W. - C. &amp; Chen, S. - L." box="[324,525,367,393]" pageId="9" pageNumber="578" pagination="351 - 357" refId="ref7367" refString="Tang, H. - C., Yeh, W. - C. &amp; Chen, S. - L. (2013) Description of an endemic and endangered new Sympetrum species (Odonata: Libellulidae) from the subtropical area of Taiwan. Zootaxa, 3693 (3), 351 - 357. https: // doi. org / 10.11646 / zootaxa. 3693.3.5" type="journal article" year="2013">
Tang
<emphasis id="B92DEF7FFFBC503EFE78FEE66BF3FE07" box="[387,444,367,393]" italics="true" pageId="9" pageNumber="578">et al.</emphasis>
(2013)
</bibRefCitation>
suggested, the
<taxonomicName id="4C5948EEFFBC503EFD41FEE6697AFE07" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[698,821,367,393]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="infuscatum">
<emphasis id="B92DEF7FFFBC503EFD41FEE6697AFE07" box="[698,821,367,393]" italics="true" pageId="9" pageNumber="578">infuscatum</emphasis>
</taxonomicName>
-group so far comprises 10 species, viz.
<taxonomicName id="4C5948EEFFBC503EFB02FEE66F36FE07" authorityName="Zhu" authorityYear="1999" box="[1273,1401,367,393]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92DEF7FFFBC503EFB02FEE66F36FE07" box="[1273,1401,367,393]" italics="true" pageId="9" pageNumber="578">S. daliensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFA7CFEE66B63FE23" baseAuthorityName="Selys" baseAuthorityYear="1883" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="darwinianum">
<emphasis id="B92DEF7FFFBC503EFA7CFEE66B63FE23" italics="true" pageId="9" pageNumber="578">S. darwinianum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFEC0FE1A6804FE23" authority="Oguma, 1915" authorityName="Oguma" authorityYear="1915" box="[315,587,402,429]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="gracile">
<emphasis id="B92DEF7FFFBC503EFEC0FE1A6BE8FE23" box="[315,423,403,429]" italics="true" pageId="9" pageNumber="578">S. gracile</emphasis>
<bibRefCitation id="EFC84E9CFFBC503EFE54FE1A6804FE23" author="Oguma, K." box="[431,587,402,429]" pageId="9" pageNumber="578" pagination="142 - 145" refId="ref6935" refString="Oguma, K. (1915) A new species of Sympetrum. Entomological Magazine, Kyoto, 1 (4), 142 - 145." type="journal article" year="1915">Oguma, 1915</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFDACFE1A68A3FE23" baseAuthorityName="Selys" baseAuthorityYear="1884" box="[599,748,403,429]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="hypomelas">
<emphasis id="B92DEF7FFFBC503EFDACFE1A68A3FE23" box="[599,748,403,429]" italics="true" pageId="9" pageNumber="578">S. hypomelas</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFD01FE1A6E7DFE23" authority="(Selys, 1883)" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[762,1074,403,429]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="infuscatum">
<emphasis id="B92DEF7FFFBC503EFD01FE1A69DDFE23" box="[762,914,403,429]" italics="true" pageId="9" pageNumber="578">S. infuscatum</emphasis>
(
<bibRefCitation id="EFC84E9CFFBC503EFC59FE1A6E66FE23" author="Selys, E." box="[930,1065,403,429]" pageId="9" pageNumber="578" pagination="82 - 143" refId="ref7278" refString="Selys, E. (1883) Les odonates du Japon. Annales de la Societe Entomologique de Belgique, 27, 82 - 143." type="journal article" year="1883">Selys, 1883</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFBC5FE1A6F34FE23" authority="Oguma, 1922" authorityName="Oguma" authorityYear="1922" box="[1086,1403,403,429]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B92DEF7FFFBC503EFBC5FE1A6E98FE23" box="[1086,1239,403,429]" italics="true" pageId="9" pageNumber="578">S. maculatum</emphasis>
<bibRefCitation id="EFC84E9CFFBC503EFB24FE1A6F34FE23" author="Oguma, K." box="[1247,1403,403,429]" pageId="9" pageNumber="578" pagination="96 - 112" refId="ref6962" refString="Oguma, K. (1922) The Japanese dragonfly-fauna of the family Libellulidae. Deutsche Entomologische Zeitschrift, 1, 96 - 112. https: // doi. org / 10.1002 / mmnd. 192219220110" type="journal article" year="1922">Oguma, 1922</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFA7CFE1A6B52FE5E" authorityName="Tang, Yeh &amp; Chen" authorityYear="2013" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="nantouensis">
<emphasis id="B92DEF7FFFBC503EFA7CFE1A6B52FE5E" italics="true" pageId="9" pageNumber="578">S. nantouensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFED0FE3E6BE3FE5F" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[299,428,439,465]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="orientale">
<emphasis id="B92DEF7FFFBC503EFED0FE3E6BE3FE5F" box="[299,428,439,465]" italics="true" pageId="9" pageNumber="578">S. orientale</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5948EEFFBC503EFE41FE3E6BB4FE5E" authorityName="Bartenef" authorityYear="1914" box="[442,507,439,465]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="risi">
<emphasis id="B92DEF7FFFBC503EFE41FE3E6BB4FE5E" box="[442,507,439,465]" italics="true" pageId="9" pageNumber="578">S. risi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5948EEFFBC503EFDC9FE3E68C9FE5E" authorityName="Han &amp; Zhu" authorityYear="1997" box="[562,646,439,465]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="xiaoi">
<emphasis id="B92DEF7FFFBC503EFDC9FE3E68C9FE5E" box="[562,646,439,465]" italics="true" pageId="9" pageNumber="578">S. xiaoi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC84E9CFFBC503EFD6FFE3E699BFE5F" author="Pilgrim, E. M. &amp; von Dohlen, C. D." box="[660,980,439,465]" pageId="9" pageNumber="578" pagination="281 - 295" refId="ref7118" refString="Pilgrim, E. M. &amp; von Dohlen, C. D. (2012) Phylogeny of the dragonfly genus Sympetrum (Odonata: Libellulidae). Organisms, Diversity and Evolution, 12 (3), 281 - 295. https: // doi. org / 10.1007 / s 13127 - 012 - 0081 - 7" type="journal article" year="2012">Pilgrim &amp; von Dohlen 2012</bibRefCitation>
;
<bibRefCitation id="EFC84E9CFFBC503EFC24FE3E6ED6FE5F" author="Tang, H. - C. &amp; Yeh, W. - C. &amp; Chen, S. - L." box="[991,1177,439,465]" pageId="9" pageNumber="578" pagination="351 - 357" refId="ref7367" refString="Tang, H. - C., Yeh, W. - C. &amp; Chen, S. - L. (2013) Description of an endemic and endangered new Sympetrum species (Odonata: Libellulidae) from the subtropical area of Taiwan. Zootaxa, 3693 (3), 351 - 357. https: // doi. org / 10.11646 / zootaxa. 3693.3.5" type="journal article" year="2013">
Tang
<emphasis id="B92DEF7FFFBC503EFBE6FE3E6E19FE5F" box="[1053,1110,439,465]" italics="true" pageId="9" pageNumber="578">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BE6336DFFBC503EFF3CFE52684FFD27" blockId="9.[151,1436,151,825]" pageId="9" pageNumber="578">
<taxonomicName id="4C5948EEFFBC503EFF3CFE526BAEFE7B" authorityName="Makbun" authorityYear="2023" box="[199,481,475,501]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B92DEF7FFFBC503EFF3CFE526BAEFE7B" box="[199,481,475,501]" italics="true" pageId="9" pageNumber="578">Sympetrum thailandensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C5948EEFFBC503EFD55FE526910FE7B" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[686,863,475,501]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="darwinianum">
<emphasis id="B92DEF7FFFBC503EFD55FE526910FE7B" box="[686,863,475,501]" italics="true" pageId="9" pageNumber="578">S. darwinianum</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C5948EEFFBC503EFC50FE526E69FE7B" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[939,1062,475,501]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="infuscatum">
<emphasis id="B92DEF7FFFBC503EFC50FE526E69FE7B" box="[939,1062,475,501]" italics="true" pageId="9" pageNumber="578">infuscatum</emphasis>
</taxonomicName>
-group. They share synthorax and abdomen yellow to red; S89 red with middorsal spots; wing tip hyaline and cornua with exposed apices curved downward. However, the new species can be separated from the congener by a combination of following characters (character of
<taxonomicName id="4C5948EEFFBC503EFED1FDCE6B95FDEF" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[298,474,583,609]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="darwinianum">
<emphasis id="B92DEF7FFFBC503EFED1FDCE6B95FDEF" box="[298,474,583,609]" italics="true" pageId="9" pageNumber="578">S. darwinianum</emphasis>
</taxonomicName>
for comparison is in paratheses): in males, cerci curved with pointed tip in lateral view (vs. slightly curved and parallel-sided); mesepisternum reddish brown (vs. red) and cerci are black with red basally (vs. entirely pale red or yellow).
</paragraph>
<paragraph id="8BE6336DFFBC503EFF3CFD3A6A9DFCB7" blockId="9.[151,1436,151,825]" pageId="9" pageNumber="578">
In females,
<taxonomicName id="4C5948EEFFBC503EFEB7FD3A6BB1FD43" authorityName="Makbun" authorityYear="2023" box="[332,510,691,717]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B92DEF7FFFBC503EFEB7FD3A6BB1FD43" box="[332,510,691,717]" italics="true" pageId="9" pageNumber="578">S. thailandensis</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C5948EEFFBC503EFCC8FD3A69ABFD43" baseAuthorityName="Selys" baseAuthorityYear="1883" box="[819,996,691,717]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="darwinianum">
<emphasis id="B92DEF7FFFBC503EFCC8FD3A69ABFD43" box="[819,996,691,717]" italics="true" pageId="9" pageNumber="578">S. darwinianum</emphasis>
</taxonomicName>
by a set of characters as follows: cerci entirely black (vs. entirely yellowish or reddish); mesepisternum unicoloured and unmarked (vs. dark brown or black with pale yellow stripe) and valvulae vulva short with bluntly triangular-tip bilobe (vs. very small or vestigial).
</paragraph>
<caption id="DF2663E5FFBC503EFF6CF9496ED7F8AF" ID-DOI="http://doi.org/10.5281/zenodo.7984442" ID-Zenodo-Dep="7984442" httpUri="https://zenodo.org/record/7984442/files/figure.png" pageId="9" pageNumber="578" startId="9.[151,250,1728,1752]" targetBox="[151,1436,849,1705]" targetPageId="9" targetType="figure">
<paragraph id="8BE6336DFFBC503EFF6CF9496ED7F8AF" blockId="9.[151,1436,1728,1825]" pageId="9" pageNumber="578">
<emphasis id="B92DEF7FFFBC503EFF6CF9496B5CF956" bold="true" box="[151,275,1728,1752]" pageId="9" pageNumber="578">FIGURE 8.</emphasis>
<taxonomicName id="4C5948EEFFBC503EFEE1F9486855F957" authority="Makbun, 2023" authorityName="Makbun" authorityYear="2023" box="[282,538,1729,1753]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B92DEF7FFFBC503EFEE1F9486855F957" box="[282,538,1729,1753]" italics="true" pageId="9" pageNumber="578">Sympetrum thailandensis</emphasis>
</taxonomicName>
<emphasis id="B92DEF7FFFBC503EFDDAF948683EF957" bold="true" box="[545,625,1729,1753]" pageId="9" pageNumber="578">
<taxonomicNameLabel id="A21E5204FFBC503EFDDAF948683EF957" box="[545,625,1729,1753]" pageId="9" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life, males (ab), females (cd); (a) Hin Tung, Muang, Nakhon Nayok province, Thailand, 3-xi-2018 (photo by Noppadon Makbun); (b) same locality, 24-x-2018 (photo by Dennis Farrell); (c) same locality, 24-x-2018 (photo by Dennis Farrell); (d) same locality, 3-xi-2018 (photo by Noppadon Makbun).
</paragraph>
</caption>
<paragraph id="8BE6336DFFBC503DFF3CF8C66F11FF5B" blockId="9.[151,1437,1871,2041]" lastBlockId="10.[151,1436,151,213]" lastPageId="10" lastPageNumber="579" pageId="9" pageNumber="578">
<emphasis id="B92DEF7FFFBC503EFF3CF8C66BF5F8E7" bold="true" box="[199,442,1871,1897]" pageId="9" pageNumber="578">Habitat and ecology.</emphasis>
The habitat of
<taxonomicName id="4C5948EEFFBC503EFD9AF8C6695EF8E7" authorityName="Makbun" authorityYear="2023" box="[609,785,1871,1897]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B92DEF7FFFBC503EFD9AF8C6695EF8E7" box="[609,785,1871,1897]" italics="true" pageId="9" pageNumber="578">S. thailandensis</emphasis>
</taxonomicName>
is a small seasonal pond surrounded trees and grasses near the parking area of the tourist attraction. It coexists with
<taxonomicName id="4C5948EEFFBC503EFCE1F8FA69E1F803" baseAuthorityName="Selys" baseAuthorityYear="1884" box="[794,942,1907,1933]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="hypomelas">
<emphasis id="B92DEF7FFFBC503EFCE1F8FA69E1F803" box="[794,942,1907,1933]" italics="true" pageId="9" pageNumber="578">S. hypomelas</emphasis>
</taxonomicName>
at restricted altitude (
<quantity id="4CA19E88FFBC503EFB5EF8FA6F1FF803" box="[1189,1360,1907,1933]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="9" pageNumber="578" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">
<geoCoordinate id="EE6D55AAFFBC503EFB5EF8FA6EABF803" box="[1189,1252,1907,1933]" degrees="1,200" direction="north" orientation="latitude" pageId="9" pageNumber="578" precision="55" value="1.2">1,200</geoCoordinate>
<geoCoordinate id="EE6D55AAFFBC503EFB1FF8FA6F7DF803" box="[1252,1330,1907,1933]" degrees="1,300" direction="west" orientation="longitude" pageId="9" pageNumber="578" precision="55" value="-1.3">1,300</geoCoordinate>
m
</quantity>
asl) at the
<typeStatus id="54E28DCFFFBC503EFF44F81E6AA0F83F" box="[191,239,1943,1969]" pageId="9" pageNumber="578">type</typeStatus>
locality. It is found perching on leaf in sunny day but disappears when it is overcast. So far, this new species is distributed restrictedly, compared to the congener species,
<taxonomicName id="4C5948EEFFBC503EFCC0F8326981F85B" baseAuthorityName="Selys" baseAuthorityYear="1884" box="[827,974,1979,2005]" class="Insecta" family="Libellulidae" genus="Sympetrum" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="hypomelas">
<emphasis id="B92DEF7FFFBC503EFCC0F8326981F85B" box="[827,974,1979,2005]" italics="true" pageId="9" pageNumber="578">S. hypomelas</emphasis>
</taxonomicName>
, in
<collectingCountry id="F34E73FDFFBC503EFC0CF8326E14F85B" box="[1015,1115,1979,2005]" name="Thailand" pageId="9" pageNumber="578">Thailand</collectingCountry>
(
<figureCitation id="13622FE8FFBC503EFB92F8326EE3F85B" box="[1129,1196,1979,2005]" captionStart="FIGURE 9" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[151,1436,237,1201]" captionTargetId="figure-58@10.[151,1436,237,1201]" captionTargetPageId="10" captionText="FIGURE 9. Distribution map of Sympetrum spp. in Thailand; S. hypomelas (yellow triangle); S. thailandensis sp. nov. (purple circle)." figureDoi="http://doi.org/10.5281/zenodo.7984444" httpUri="https://zenodo.org/record/7984444/files/figure.png" pageId="9" pageNumber="578">Fig. 9</figureCitation>
). The emergence was observed in late July. Mating and oviposition was recorded from late October to early November. An ovipositing female, with guarding male, was observed hovering about a meter above the water for a while. Then, when ready, the female flicked a light green egg into the water at a time. The flight season ranges from July to November.
</paragraph>
</subSubSection>
</treatment>
</document>