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<document id="1061322C329E558C3FEA7421C7F925B6" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFFD500EFF23FED5FC3841B2" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus ivelonanus Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="15" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="14" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="EC032AB586AAF11C673803A5AA3EB48F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2EC63A73B5D0FC2A7E54846AACC2B083">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="CC9207806A2F5FF5136187CEE5A53986">2024</mods:date>
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<mods:classification id="459E74F3D5F144721403473D5286FCF0">journal article</mods:classification>
<mods:identifier id="FB150579634DA5AFD2B67C6A429379A5" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
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<treatment id="03E7FF50FFFD500EFF23FED5FC3841B2" ID-DOI="http://doi.org/10.5281/zenodo.13746003" ID-Zenodo-Dep="13746003" LSID="urn:lsid:plazi:treatment:03E7FF50FFFD500EFF23FED5FC3841B2" httpUri="http://treatment.plazi.org/id/03E7FF50FFFD500EFF23FED5FC3841B2" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C3541DCDFFFD500DFF23FED5FD234540" box="[151,742,281,308]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BF14E46FFFD500DFF23FED5FD234540" blockId="13.[151,742,281,344]" box="[151,742,281,308]" pageId="13" pageNumber="14">
<heading id="D0B9F92AFFFD500DFF23FED5FD234540" bold="true" box="[151,742,281,308]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="B93A9254FFFD500DFF23FED5FD234540" bold="true" box="[151,742,281,308]" pageId="13" pageNumber="14">
<taxonomicName id="4C4E35C5FFFD500DFF23FED5FDAB4540" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,622,281,308]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ivelonanus" status="comb. nov.">
<emphasis id="B93A9254FFFD500DFF23FED5FDC24547" bold="true" box="[151,519,281,308]" italics="true" pageId="13" pageNumber="14">Anhoraeomorphus ivelonanus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFFD500DFDCFFED5FD234540" box="[635,742,281,307]" pageId="13" pageNumber="14" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFD500DFF23FEF1FEE2452B" box="[151,295,317,344]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BF14E46FFFD500DFF23FEF1FEE2452B" blockId="13.[151,742,281,344]" box="[151,295,317,344]" pageId="13" pageNumber="14">
(
<figureCitation id="137552C3FFFD500DFF2BFEF1FEDA452B" box="[159,287,317,344]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="13" pageNumber="14">Figs 28–35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFD500DFF23FE4EFDBD45E9" box="[151,632,386,410]" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BF14E46FFFD500DFF23FE4EFDBD45E9" blockId="13.[151,632,386,410]" box="[151,632,386,410]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB5E6968FFFD500DFF23FE4EFDBD45E9" box="[151,632,386,410]" pageId="13" pageNumber="14">
<treatmentCitation id="0AEF6857FFFD500DFF23FE4EFDB145E9" author="Franz, H." box="[151,628,386,410]" page="171" pageId="13" pageNumber="14" year="1986">
<taxonomicName id="4C4E35C5FFFD500DFF23FE4EFDB145E9" ID-CoL="3MNKB" authority="Franz, 1986 b: 171" authorityName="Franz" authorityPageNumber="171" authorityYear="1986" box="[151,628,386,410]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFD500DFF23FE4EFE7745E9" box="[151,434,386,410]" italics="true" pageId="13" pageNumber="14">Horaeomorphus ivelonanus</emphasis>
<bibRefCitation id="EFDF33B7FFFD500DFE0CFE4EFDB145E9" author="Franz, H." box="[440,628,386,410]" pageId="13" pageNumber="14" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 171</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFD500EFF23FE04FB524516" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BF14E46FFFD500DFF23FE04FEC8463D" blockId="13.[151,1436,456,590]" pageId="13" pageNumber="14">
<emphasis id="B93A9254FFFD500DFF23FE04FE774591" bold="true" box="[151,434,456,482]" pageId="13" pageNumber="14">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFFD500DFE74FE04FEC8463D" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="13" pageNumber="14" specimenCount="58" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFFD500DFE74FE04FD2E4591" bold="true" box="[448,747,456,482]" pageId="13" pageNumber="14">
<typeStatus id="54F5F0E4FFFD500DFE74FE04FDE84591" box="[448,557,456,482]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(Madagascar):
</emphasis>
<specimenCount id="9D4885CFFFFD500DFD4EFE05FCD44590" box="[762,785,457,483]" count="1" pageId="13" pageNumber="14" type="male">♂</specimenCount>
, four labels: “Réserenat. III / Andranomalaza / Valeé d’Ivelona / X.
<specimenCount id="9D4885CFFFFD500DFE82FE20FEB44674" box="[310,369,492,519]" count="57" pageId="13" pageNumber="14" type="generic">57 P.</specimenCount>
Soga” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFFD500DFBB6FE20FB0C4675" box="[1026,1225,492,518]" name="Madagascar" pageId="13" pageNumber="14">MADAGASCAR</collectingCountry>
” [bluish, printed], “
<taxonomicName id="4C4E35C5FFFD500DFF15FDDCFE9D4659" box="[161,344,528,554]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ ivelonanus m. / det.
<determinerName id="7E2DA282FFFD500DFDE4FDDCFD684659" box="[592,685,528,554]" pageId="13" pageNumber="14">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFFD500DFF2BFDF8FEC7463D" box="[159,258,564,590]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF311ECEFFFD500DFF23F8B9FEF543A5" ID-DOI="http://doi.org/10.5281/zenodo.13745931" ID-Zenodo-Dep="13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,264,1909,1933]" targetBox="[151,1434,614,1874]" targetPageId="13" targetType="figure">
<paragraph id="8BF14E46FFFD500DFF23F8B9FEF543A5" blockId="13.[151,1436,1909,2006]" pageId="13" pageNumber="14">
<emphasis id="B93A9254FFFD500DFF23F8B9FE9643FD" bold="true" box="[151,339,1909,1934]" pageId="13" pageNumber="14">FIGURES 28–35.</emphasis>
<taxonomicName id="4C4E35C5FFFD500DFEECF8BAFD1C43FD" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[344,729,1910,1934]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFD500DFEECF8BAFD4D43FD" box="[344,648,1910,1934]" italics="true" pageId="13" pageNumber="14">Anhoraeomorphus ivelonanus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFFD500DFB82F8BAFB8A43FD" bold="true" box="[1078,1103,1910,1934]" pageId="13" pageNumber="14">28</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFD500DFAC7F8BAFA4943FD" bold="true" box="[1395,1420,1910,1934]" pageId="13" pageNumber="14">29</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFFD500DFCE3F856FCB543C1" bold="true" box="[855,880,1946,1970]" pageId="13" pageNumber="14">30</emphasis>
); antenna (
<emphasis id="B93A9254FFFD500DFC57F856FC3943C1" bold="true" box="[995,1020,1946,1970]" pageId="13" pageNumber="14">31</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFD500DFB50F856FAE643C1" bold="true" box="[1252,1315,1946,1970]" pageId="13" pageNumber="14">32, 34</emphasis>
) and lateral (
<emphasis id="B93A9254FFFD500DFF14F872FF1A43A5" bold="true" box="[160,223,1982,2006]" pageId="13" pageNumber="14">33, 35</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFFE500EFF73FF5AFB524516" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FF5AFE6344C3" bold="true" box="[199,422,150,176]" pageId="14" pageNumber="15">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFFE500EFDB0FF5BFD9F44C2" box="[516,602,151,177]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) slender; head (
<figureCitation id="137552C3FFFE500EFCA9FF5BFCB644C2" box="[797,883,151,177]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 29</figureCitation>
) slightly elongate, with vertex semicircular, in anterodorsal view its posterior margin evenly rounded; pronotum (
<figureCitation id="137552C3FFFE500EFC3AFF77FC2444A6" box="[910,993,187,213]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) subtrapezoidal, broadest at base, with two pairs of deep antebasal pits; antennomeres 1–7 distinctly elongate, 8–10 each about as long as broad; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFFE500EFCEEFECFFC19456E" box="[858,988,258,285]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Figs 32–35</figureCitation>
) in ventral view broadest near middle, with apical region of dorsal wall weakly curved distoventrally, distolateral endophallic sclerites curved mesad and overlapping at middle, each with rounded apex, each paramere with three long apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF73FEA2FE5B409A" pageId="14" pageNumber="15" type="description">
<paragraph id="8BF14E46FFFE500EFF73FEA2FD5845DE" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FEA2FEB045FB" bold="true" box="[199,373,366,392]" pageId="14" pageNumber="15">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFFE500EFDABFEA3FDAA45FA" box="[543,623,367,393]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) light brown, tarsi and palps distinctly lighter, antennae slightly infuscate in distal halves, setae yellowish; BL
<quantity id="4CB6E3A3FFFE500EFD86FE5FFD5C45DE" box="[562,665,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="mm" value="1.6">1.60 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FE7BFBB846DA" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Head (
<figureCitation id="137552C3FFFE500EFEA6FE7BFE5645A2" box="[274,403,439,465]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Figs 29–30</figureCitation>
) in anterodorsal view subtrapezoidal, broadest at eyes, HL
<quantity id="4CB6E3A3FFFE500EFB87FE7BFB5F45A2" box="[1075,1178,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="4CB6E3A3FFFE500EFB68FE7BFA8145A2" box="[1244,1348,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="15" unit="mm" value="0.28">0.28 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFFE500EFEFDFE17FE5E4586" box="[329,411,475,501]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and slightly elongate; posterior margin of vertex weakly and evenly rounded; each eye large and strongly convex, with small but distinct posteromedian emargination. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; frons and vertex densely covered with thick bristles directed posteriorly.Antenna (
<figureCitation id="137552C3FFFE500EFB91FD8BFBB04612" box="[1061,1141,583,609]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 31</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFFE500EFE77FDA7FDEF46F6" box="[451,554,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="15" unit="mm" value="0.78">0.78 mm</quantity>
, antennomeres 1–7 each elongate, 8–10 each about as long as broad; 11 much shorter than 9 and 10 combined, slightly narrower than 10, nearly twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FD7FFA57472E" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Pronotum (
<figureCitation id="137552C3FFFE500EFEF3FD7FFE5F46BE" box="[327,410,691,717]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) subtrapezoidal, equally broad from base to middle; PL
<quantity id="4CB6E3A3FFFE500EFB96FD7FFB4F46BE" box="[1058,1162,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="mm" value="0.4">0.40 mm</quantity>
, PW
<quantity id="4CB6E3A3FFFE500EFB7EFD7FFAF746BE" box="[1226,1330,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="14" pageNumber="15" unit="mm" value="0.33">0.33 mm</quantity>
. Anterior margin weakly arcuate, lateral margins slightly rounded in anterior half and strongly diverging posteriorly, in posterior half parallel; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FCABFF3947BA" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Elytra (
<figureCitation id="137552C3FFFE500EFEAFFCABFEA947F2" box="[283,364,871,897]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
) together oval, broadest near middle; EL
<quantity id="4CB6E3A3FFFE500EFC89FCABFC6647F2" box="[829,931,871,897]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="15" unit="mm" value="0.9">0.90 mm</quantity>
, EW
<quantity id="4CB6E3A3FFFE500EFC54FCAAFB8347F2" box="[992,1094,870,897]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="15" unit="mm" value="0.65">0.65 mm</quantity>
, EI 1.38; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FC1FFBCC479E" blockId="14.[151,1437,150,1473]" box="[199,1033,979,1005]" pageId="14" pageNumber="15">Legs moderately long, slender, unmodified except slightly recurved tibiae.</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FC3BFA8F40B6" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
Aedeagus (
<figureCitation id="137552C3FFFE500EFEFDFC3BFE084062" box="[329,461,1014,1041]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="14" pageNumber="15">Figs 32–35</figureCitation>
) stout, AeL
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; in ventral view broadest near middle and weakly narrowing towards step-wise delimited apical region, which is subrectangular, transverse and shallowly emarginate; in lateral view apical region of dorsal plate weakly curved distoventrally; endophallus with slightly asymmetrical and elongate distolateral sclerites, symmetrical lateral groups of long and dense basal sclerites and asymmetrical weakly sclerotized median structures, distolateral sclerites strongly curved mesad and overlapping at middle, each with rounded apex; parameres thick, much shorter than median lobe, each paramere with three long apical setae.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FB03FE5B409A" blockId="14.[151,1437,150,1473]" box="[199,414,1231,1257]" pageId="14" pageNumber="15">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF73FB3EFC8A417E" box="[199,847,1266,1293]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8BF14E46FFFE500EFF73FB3EFC8A417E" blockId="14.[151,1437,150,1473]" box="[199,847,1266,1293]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FB3EFEA4417F" bold="true" box="[199,353,1266,1292]" pageId="14" pageNumber="15">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFFE500EFE75FB3FFD8D417E" box="[449,584,1267,1293]" name="Madagascar" pageId="14" pageNumber="15">Madagascar</collectingCountry>
(Analamanga Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF73FADBFC3841B2" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BF14E46FFFE500EFF73FADBFA444142" blockId="14.[151,1437,150,1473]" box="[199,1409,1303,1329]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73FADBFEFE4142" bold="true" box="[199,315,1303,1329]" pageId="14" pageNumber="15">Remarks.</emphasis>
According to
<bibRefCitation id="EFDF33B7FFFE500EFE6EFADBFDA04142" author="Franz, H." box="[474,613,1303,1329]" pageId="14" pageNumber="15" pagination="1 - 93" refId="ref52970" refString="Franz, H. (1963) Scydmaeniden (Coleoptera) aus Ostafrika, dem Kongo und der Tschadrepublik (vorwiegend aus den im Sommer 1962 vom Verfasser selbst gesammelten Ausbeuten). Stuttgarter Beitrage zur Naturkunde, 120, 1 - 93." type="journal article" year="1963">Franz (1963</bibRefCitation>
b), the word “Réserenat.” on the label is a misspelling of ‘Réserve nat’.
</paragraph>
<paragraph id="8BF14E46FFFE500EFF73FAF7FC3841B2" blockId="14.[151,1437,150,1473]" pageId="14" pageNumber="15">
<taxonomicName id="4C4E35C5FFFE500EFF73FAF7FDDF4126" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,538,1339,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFE500EFF73FAF7FDDF4126" box="[199,538,1339,1365]" italics="true" pageId="14" pageNumber="15">Anhoraeomorphus ivelonanus</emphasis>
</taxonomicName>
is externally slightly similar to
<taxonomicName id="4C4E35C5FFFE500EFC39FAF7FBCE4126" box="[909,1035,1339,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="grucheti">
<emphasis id="B93A9254FFFE500EFC39FAF7FBCE4126" box="[909,1035,1339,1365]" italics="true" pageId="14" pageNumber="15">A. grucheti</emphasis>
</taxonomicName>
, but it can be distinguished by the stronger elongate pronotum and distinctly elongate antennomeres 4–6 (about as long as broad in
<taxonomicName id="4C4E35C5FFFE500EFABFFA93FA49410A" box="[1291,1420,1375,1401]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="grucheti">
<emphasis id="B93A9254FFFE500EFABFFA93FA49410A" box="[1291,1420,1375,1401]" italics="true" pageId="14" pageNumber="15">A. grucheti</emphasis>
</taxonomicName>
). Another externally similar species is
<taxonomicName id="4C4E35C5FFFE500EFD86FA4FFD7741EF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[562,690,1411,1437]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFE500EFD86FA4FFD7741EF" box="[562,690,1411,1437]" italics="true" pageId="14" pageNumber="15">A. procerus</emphasis>
</taxonomicName>
. However, in
<taxonomicName id="4C4E35C5FFFE500EFCF9FA4FFC0841EF" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[845,973,1411,1437]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFE500EFCF9FA4FFC0841EF" box="[845,973,1411,1437]" italics="true" pageId="14" pageNumber="15">A. procerus</emphasis>
</taxonomicName>
, the posterior margin of vertex is truncate and straight, whereas that in
<taxonomicName id="4C4E35C5FFFE500EFE6DFA6BFDB441B2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[473,625,1447,1473]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFFE500EFE6DFA6BFDB441B2" box="[473,625,1447,1473]" italics="true" pageId="14" pageNumber="15">A. ivelonanus</emphasis>
</taxonomicName>
is rounded and posteriorly convex.
</paragraph>
</subSubSection>
</treatment>
</document>
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