treatments-xml/data/03/E7/DD/03E7DD69FFF78677241C786AFD8C3FF0.xml
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<mods:title id="57182CCE298A0CBBE45FFC2700F0B18D">A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: Pinnipeds and Cetaceans</mods:title>
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<mods:namePart id="0EE9CE190EFB567DB6EFFCE05ED8FE7C">Boessenecker, Robert W.</mods:namePart>
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<treatment id="03E7DD69FFF78677241C786AFD8C3FF0" ID-DOI="http://doi.org/10.5281/zenodo.4818614" ID-GBIF-Taxon="182400954" ID-Zenodo-Dep="4818614" LSID="urn:lsid:plazi:treatment:03E7DD69FFF78677241C786AFD8C3FF0" httpUri="http://treatment.plazi.org/id/03E7DD69FFF78677241C786AFD8C3FF0" lastPageId="42" lastPageNumber="856" pageId="39" pageNumber="854">
<subSubSection id="C3543FF4FFF7867A241C786AFB983F11" box="[859,1027,1039,1065]" pageId="39" pageNumber="854" type="nomenclature">
<paragraph id="8BF16C7FFFF7867A241C786AFB983F11" blockId="39.[859,1027,1039,1065]" box="[859,1027,1039,1065]" pageId="39" pageNumber="854">
<heading id="D0B9DB13FFF7867A241C786AFB983F11" box="[859,1027,1039,1065]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="854" reason="2">
cf.
<taxonomicName id="4C4E17FCFFF7867A243B786AFB983F11" ID-CoL="37H3" box="[892,1027,1039,1065]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="854" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF7867A243B786AFB983F11" box="[892,1027,1039,1065]" italics="true" pageId="39" pageNumber="854">Balaenoptera</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3543FF4FFF7867A25E2782CFB3F3FC2" pageId="39" pageNumber="854" type="materials_examined">
<paragraph id="8BF16C7FFFF7867A25E2782CFCD03FAD" blockId="39.[677,1209,1096,1173]" pageId="39" pageNumber="854">
REFERRED MATERIAL. — UCMP 219136, a right petrosal, collected by R.W. Boessenecker from UCMP locality
<accessionNumber id="941DF19CFFF7867A25B47818FCDC3FAD" box="[755,839,1149,1173]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V99851" pageId="39" pageNumber="854" type="EnaNcbi">V99851</accessionNumber>
.
</paragraph>
<paragraph id="8BF16C7FFFF7867A25E278CBFB3F3FC2" blockId="39.[675,1210,1197,1274]" pageId="39" pageNumber="854">
STRATIGRAPHIC OCCURRENCE. — Lowermost part of the San Gregorio section of the Purisima Formation, latest Miocene (6.4-5.6 Ma; Messinian equivalent;
<figureCitation id="137570FAFFF7867A23197887FB033FC2" box="[1118,1176,1250,1274]" captionStart="FIG" captionStartId="6.[113,124,1221,1239]" captionTargetBox="[761,1205,215,1066]" captionTargetId="figure-149@6.[759,1205,592,972]" captionTargetPageId="6" captionText="FIG. 2. — Stratigraphy of the Purisima Formation: A, Stratigraphic column of the San Gregorio section of the Purisima Formation (in meters), showing stratigraphic position of UCMP vertebrate localities, dated ash beds, diatom floras, and species of Carcharodon; thickness measurements start over above missing section in uppermost part due lack of overlapping strata; epoch and stage boundaries marked as &quot;?&quot; are estimated; modified from Boessenecker (2011a); Pliocene-Pleistocene boundary after Hilgen et al. (2012); B, photograph of 3-4 m thick ash within Purisima Formation at mouth of San Gregorio Creek correlated with 2.5 Ma Ishi Tuff (Powell et al.2007); C, trace fossils in pervasively bioturbated sandstone,Purisima Formation,mouth of San Gregorio Creek; D, swaley cross-stratified sandstone, mouth of San Gregorio Creek. Abbreviations: Qt, Pleistocene terrace deposits; Tp, Purisima Formation; C, claystone; St, siltstone; Ss, sandstone. Scale bars: A, 2 m; B, C, 10 cm." figureDoi="http://doi.org/10.5281/zenodo.4538204" httpUri="https://zenodo.org/record/4538204/files/figure.png" pageId="39" pageNumber="854">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3543FF4FFF7867525E27948FBB43D9C" lastPageId="40" lastPageNumber="855" pageId="39" pageNumber="854" type="description">
<paragraph id="8BF16C7FFFF7867A25E27948FCAE3E7D" blockId="39.[674,1212,1325,1700]" box="[677,821,1325,1351]" pageId="39" pageNumber="854">DESCRIPTION</paragraph>
<paragraph id="8BF16C7FFFF7867525E27929FBB43D9C" blockId="39.[674,1212,1325,1700]" lastBlockId="40.[674,1212,1324,1700]" lastPageId="40" lastPageNumber="855" pageId="39" pageNumber="854">
Ŋis large partial petrosal exhibits a low, and ventrally flattened pars cochlearis that is roughly oval-shaped in ventral aspect and somewhat elongated dorsally (
<figureCitation id="137570FAFFF7867A25EB79CEFD623EFD" box="[684,761,1451,1477]" captionStart="FIG" captionStartId="41.[113,123,1071,1089]" captionTargetBox="[116,1198,216,1032]" captionTargetId="figure-275@41.[161,734,214,751]" captionTargetPageId="41" captionText="FIG. 20.— Right petrosal of cf.Balaenoptera (UCMP 219146);A, medial view;B, lateral view;C, ventral view;D, dorsal view.Scale bar:3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538251" httpUri="https://zenodo.org/record/4538251/files/figure.png" pageId="39" pageNumber="854">Fig. 20</figureCitation>
;
<tableCitation id="C6CC59C4FFF7867A244479CEFCC93EFE" box="[771,850,1451,1478]" captionStart="TABLE" captionStartId="38.[113,124,219,237]" captionTargetPageId="38" captionText="TABLE 7. — Measurements (in cm) of mysticete petrosals. Measurements given to nearest tenth of a millimeter. Total length refers to length of the petrosal body, and does not include the posterior process (preserved only in UCMP 219114 among these specimens)." httpUri="http://table.plazi.org/id/DF313CF7FFF6867B27367CBEFB2D3A3C" pageId="39" pageNumber="854" tableUuid="DF313CF7FFF6867B27367CBEFB2D3A3C">Table 7</tableCitation>
). A small, thin, and bladelike posterior cochlear crest occurs posteroventrally on the pars cochlearis. Ŋe fenestra cochleae is small and circular and occurs within a small fossa. Ŋe anterior process is acutely triangular, transversely thick, and a longitudinal keel occurs on the ventral side at its apex. Ŋe anterior pedicle sits atop a rectangular plateau on the medial surface of the anterior process.A well- developed and knob-like lateral tuberosity is present on the lateral margin of the anterior process, anterior to the position of the fenestra vestibuli.Ŋe oval-shaped fenestra vestibuli is positioned at the posteroventral end of the pars cochlearis, and a posteriorly oriented facial sulcus is present dorsolaterally to the fenestra vestibuli. A small, rectangular, and concave fossa for the stapedial muscle is positioned lateral to the posterior cochlear rest and posterior to the fenestra vestibuli. A relatively small, oval shaped, and deep suprameatal fossa occurs on the anterodorsal side of the pars cochlearis. Ŋe vestibulocochlear canal and facial canal are located on the posterodorsal portion of the pars cochlearis. Both canals are circular and small, closely appressed, with a thin crista transversa. Ŋe endolymphatic and perilymphatic foramina are positioned posterior to the vestibulocochlear nerve canal, and occur in a deep fossa which is deeper at the dorsal end near the endolymphatic foramen (
<figureCitation id="137570FAFFF8867525E97A6FFC8F3D1D" box="[686,788,1546,1573]" captionStart="FIG" captionStartId="41.[113,123,1071,1089]" captionTargetBox="[116,1198,216,1032]" captionTargetId="figure-275@41.[161,734,214,751]" captionTargetPageId="41" captionText="FIG. 20.— Right petrosal of cf.Balaenoptera (UCMP 219146);A, medial view;B, lateral view;C, ventral view;D, dorsal view.Scale bar:3 cm." figureDoi="http://doi.org/10.5281/zenodo.4538251" httpUri="https://zenodo.org/record/4538251/files/figure.png" pageId="40" pageNumber="855">Fig. 20D</figureCitation>
). Both foramina are slit-like and open dorsally. Ŋe shallowly concave stylomastoid fossa is small, rectangular, and positioned dorsal to the fenestra cochleae. Ŋe dorsal side of the petrosal and anterior process is convex and rugose.
</paragraph>
</subSubSection>
<caption id="DF313CF7FFF88675273678DBFEE03FDE" ID-DOI="http://doi.org/10.5281/zenodo.4538249" ID-Zenodo-Dep="4538249" httpUri="https://zenodo.org/record/4538249/files/figure.png" pageId="40" pageNumber="855" startId="40.[113,124,1214,1232]" targetBox="[120,1203,215,1176]" targetPageId="40">
<paragraph id="8BF16C7FFFF88675273678DBFEE03FDE" blockId="40.[113,1211,1213,1254]" pageId="40" pageNumber="855">
FIG. 19. — Right petrosal of “
<taxonomicName id="4C4E17FCFFF88675262B78DBFE403FE8" box="[364,475,1214,1232]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="855" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF88675262B78DBFE403FE8" box="[364,475,1214,1232]" italics="true" pageId="40" pageNumber="855">Balaenoptera</emphasis>
</taxonomicName>
<emphasis id="B93AB06DFFF8867526AD78DBFDB93FE8" box="[490,546,1214,1232]" italics="true" pageId="40" pageNumber="855">cortesi</emphasis>
“var.”
<emphasis id="B93AB06DFFF88675251B78DBFD143FE8" box="[604,655,1214,1232]" italics="true" pageId="40" pageNumber="855">portisi</emphasis>
(UCMP 219135);
<emphasis id="B93AB06DFFF88675246E78DBFCAD3FE8" bold="true" box="[809,822,1214,1232]" pageId="40" pageNumber="855">A</emphasis>
, medial view;
<emphasis id="B93AB06DFFF8867524F578DBFC243FE8" bold="true" box="[946,959,1214,1232]" pageId="40" pageNumber="855">B</emphasis>
, lateral view;
<emphasis id="B93AB06DFFF88675237178D8FBDF3FF7" bold="true" box="[1078,1092,1213,1231]" pageId="40" pageNumber="855">C</emphasis>
, ventral view;
<emphasis id="B93AB06DFFF88675273678B1FFE43FDE" bold="true" box="[113,127,1236,1254]" pageId="40" pageNumber="855">D</emphasis>
, dorsal view. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="DF313CF7FFF986742736784AFB203F79" ID-DOI="http://doi.org/10.5281/zenodo.4538251" ID-Zenodo-Dep="4538251" box="[113,1211,1071,1089]" httpUri="https://zenodo.org/record/4538251/files/figure.png" pageId="41" pageNumber="856" startId="41.[113,123,1071,1089]" targetBox="[116,1198,216,1032]" targetPageId="41">
<paragraph id="8BF16C7FFFF986742736784AFB203F79" blockId="41.[113,1211,1071,1089]" box="[113,1211,1071,1089]" pageId="41" pageNumber="856">
FIG. 20.— Right petrosal of cf.
<taxonomicName id="4C4E17FCFFF986742626784AFE573F79" box="[353,460,1071,1089]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF986742626784AFE573F79" box="[353,460,1071,1089]" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
(UCMP 219146);
<emphasis id="B93AB06DFFF986742511784AFDF93F79" bold="true" box="[598,610,1071,1089]" pageId="41" pageNumber="856">A</emphasis>
, medial view;
<emphasis id="B93AB06DFFF986742596784AFD453F79" bold="true" box="[721,734,1071,1089]" pageId="41" pageNumber="856">B</emphasis>
, lateral view;
<emphasis id="B93AB06DFFF986742400784AFCCF3F79" bold="true" box="[839,852,1071,1089]" pageId="41" pageNumber="856">C</emphasis>
, ventral view;
<emphasis id="B93AB06DFFF986742484784AFC4B3F79" bold="true" box="[963,976,1071,1089]" pageId="41" pageNumber="856">D</emphasis>
, dorsal view.Scale bar:3 cm.
</paragraph>
</caption>
<subSubSection id="C3543FF4FFF98677273678EAFD8C3FF0" lastPageId="42" lastPageNumber="857" pageId="41" pageNumber="856" type="discussion">
<paragraph id="8BF16C7FFFF98674273678EAFE3C3F9F" blockId="41.[111,647,1167,1700]" box="[113,423,1167,1193]" pageId="41" pageNumber="856">REMARKS AND COMPARISONS</paragraph>
<paragraph id="8BF16C7FFFF98677273678CBFD8C3FF0" blockId="41.[111,647,1167,1700]" lastBlockId="42.[112,646,913,1224]" lastPageId="42" lastPageNumber="857" pageId="41" pageNumber="856">
Ŋis petrosal is identified as a balaenopterid because of the relatively wide and triangular anterior process and the medially elongated pars cochlearis. It differs from “
<taxonomicName id="4C4E17FCFFF9867427FF7968FEA13E1F" box="[184,314,1293,1319]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF9867427FF7968FEA13E1F" box="[184,314,1293,1319]" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
<emphasis id="B93AB06DFFF986742609796BFE103E1F" box="[334,395,1294,1319]" italics="true" pageId="41" pageNumber="856">cortesi</emphasis>
“var.”
<emphasis id="B93AB06DFFF986742694796BFD8A3E1F" box="[467,529,1294,1319]" italics="true" pageId="41" pageNumber="856">portisi</emphasis>
(including UCMP 219135, described above) and
<taxonomicName id="4C4E17FCFFF98674255D7948FF6B3E5E" authorityName="Brisson" authorityYear="1762" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="novaeangliae">
<emphasis id="B93AB06DFFF98674255D7948FF6B3E5E" italics="true" pageId="41" pageNumber="856">Megaptera novaeangliae</emphasis>
</taxonomicName>
in lacking extreme medial elongation of the pars cochlearis, and in having a transversely thick anterior process. Ŋe cranial nerve canals of UCMP 219136 are located on the posterodorsal portion of the pars cochlearis, like extant
<taxonomicName id="4C4E17FCFFF9867426EE79AEFDB13EDD" box="[425,554,1483,1509]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF9867426EE79AEFDB13EDD" box="[425,554,1483,1509]" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
spp. and
<taxonomicName id="4C4E17FCFFF986742737798EFE463D3D" authority="(Ekdale et al. 2011)" baseAuthorityName="Ekdale" baseAuthorityYear="2011" box="[112,477,1515,1541]" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="novaeangliae">
<emphasis id="B93AB06DFFF986742737798EFE883D3D" box="[112,275,1515,1541]" italics="true" pageId="41" pageNumber="856">M. novaeangliae</emphasis>
(
<bibRefCitation id="EFDF118EFFF986742659798EFE4E3D3D" author="EKDALE E. G. &amp; BERTA A. &amp; DEMERE T. A." box="[286,469,1515,1541]" pageId="41" pageNumber="856" pagination="1 - 42" refId="ref68110" refString="EKDALE E. G., BERTA A. &amp; DEMERE T. A. 2011. - Ne comparative osteology of the petrotympanic complex (ear region) of extant baleen whales (Cetacea: Mysticeti). Plos One 6 (6): 1 - 42." type="journal article" year="2011">
Ekdale
<emphasis id="B93AB06DFFF98674262D798EFE023D3D" box="[362,409,1515,1541]" italics="true" pageId="41" pageNumber="856">et al.</emphasis>
2011
</bibRefCitation>
)
</taxonomicName>
; this differs from many extinct balaenopterids including “
<emphasis id="B93AB06DFFF98674255B7A6EFDA83D1C" box="[540,563,1547,1572]" italics="true" pageId="41" pageNumber="856">B.</emphasis>
<emphasis id="B93AB06DFFF98674250E7A6EFD1C3D1C" box="[585,647,1547,1572]" italics="true" pageId="41" pageNumber="856">cortesi</emphasis>
“var.”
<emphasis id="B93AB06DFFF9867427EC7A4EFF7D3D7C" box="[171,230,1579,1604]" italics="true" pageId="41" pageNumber="856">portisi</emphasis>
,
<taxonomicName id="4C4E17FCFFF9867427B67A4EFE7B3D7C" baseAuthorityName="Bosselaers &amp; Post" baseAuthorityYear="2010" box="[241,480,1578,1604]" class="Mammalia" family="Balaenopteridae" genus="Diunatans" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="luctoretmurgo">
<emphasis id="B93AB06DFFF9867427B67A4EFE7B3D7C" box="[241,480,1578,1604]" italics="true" pageId="41" pageNumber="856">Diunatans luctoretmurgo</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4E17FCFFF9867425527A4FFEA83D5C" authorityName="Bisconti" authorityYear="2010" class="Mammalia" family="Balaenopteridae" genus="Plesiobalaenoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="quarantellii">
<emphasis id="B93AB06DFFF9867425527A4FFEA83D5C" italics="true" pageId="41" pageNumber="856">Plesiobalaenoptera quarantellii</emphasis>
</taxonomicName>
, in which the cranial nerve canals are centrally positioned on the dorsal face of the pars cochlearis (
<bibRefCitation id="EFDF118EFFF9867427A67AECFE4F3D9B" author="BOSSELAERS M. &amp; POST K." box="[225,468,1673,1700]" pageId="41" pageNumber="856" pagination="331 - 363" refId="ref66734" refString="BOSSELAERS M. &amp; POST K. 2010. - A new fossil rorqual (Mammalia, Cetacea, Balaenopteridae) from the Early Pliocene of the North Sea, with a review of the rorqual species described by Owen and Van Beneden. Geodiversitas 32: 331 - 363. http: // dx. doi. org / 10.5252 / g 2010 n 2 a 6" type="journal article" year="2010">Bosselaers &amp; Post 2010</bibRefCitation>
;
<bibRefCitation id="EFDF118EFFF98674269A7AECFDE73D9B" author="BISCONTI M." box="[477,636,1673,1699]" pageId="41" pageNumber="856" pagination="943 - 958" refId="ref66164" refString="BISCONTI M. 2010 a. - A new balaenopterid whale from the Late Miocene of the Stirone River, Northern Italy (Mammalia, Cetacea, Mysticeti). Journal of Vertebrate Paleontology 30: 943 - 958." type="journal article" year="2010">Bisconti 2010a</bibRefCitation>
). Ŋis specimen is tentatively referred to
<taxonomicName id="4C4E17FCFFF98674231B78EBFD423FF0" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF98674231B78EBFD423FF0" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
owing to several features. Ŋe pars cochlearis of UCMP 219136 is more similar to extant
<taxonomicName id="4C4E17FCFFF98674233278A8FD693E3F" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF98674233278A8FD693E3F" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C4E17FCFFF98674240D788BFC2C3E3F" box="[842,951,1262,1287]" class="Mammalia" family="Balaenopteridae" genus="Megaptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF98674240D788BFC2C3E3F" box="[842,951,1262,1287]" italics="true" pageId="41" pageNumber="856">Megaptera</emphasis>
</taxonomicName>
or any described extinct balaenopterids; however, the anterior process differs from every modern species within
<taxonomicName id="4C4E17FCFFF98674236A7948FB233E7F" box="[1069,1208,1325,1351]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF98674236A7948FB233E7F" box="[1069,1208,1325,1351]" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
in being dorsally rather than anteriorly directed. Among extant
<taxonomicName id="4C4E17FCFFF9867424107909FC723EBE" box="[855,1001,1388,1414]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF9867424107909FC723EBE" box="[855,1001,1388,1414]" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
, the morphology of the pars cochlearis is closest to
<taxonomicName id="4C4E17FCFFF98674236A79E9FCB63EFD" authorityName="Lacepede" authorityYear="1804" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="acutorostrata">
<emphasis id="B93AB06DFFF98674236A79E9FCB63EFD" italics="true" pageId="41" pageNumber="856">Balaenoptera acutorostrata</emphasis>
</taxonomicName>
; although the anterior process is of similarly short length between
<taxonomicName id="4C4E17FCFFF9867424AB79A9FB173EDD" authorityName="Lacepede" authorityYear="1804" box="[1004,1164,1484,1509]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="acutorostrata">
<emphasis id="B93AB06DFFF9867424AB79A9FB173EDD" box="[1004,1164,1484,1509]" italics="true" pageId="41" pageNumber="856">B. acutorostrata</emphasis>
</taxonomicName>
and UCMP 219136, it is anteriorly directed in extant
<taxonomicName id="4C4E17FCFFF9867425E47A6EFCCB3D1C" authorityName="Lacepede" authorityYear="1804" box="[675,848,1547,1572]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="species" species="acutorostrata">
<emphasis id="B93AB06DFFF9867425E47A6EFCCB3D1C" box="[675,848,1547,1572]" italics="true" pageId="41" pageNumber="856">B. acutorostrata</emphasis>
</taxonomicName>
and anterodorsally deflected in UCMP219136. UCMP 219136 also differs from all extant species of
<taxonomicName id="4C4E17FCFFF9867424C37A2FFB963D5C" box="[900,1037,1610,1636]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="41" pageNumber="856" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFF9867424C37A2FFB963D5C" box="[900,1037,1610,1636]" italics="true" pageId="41" pageNumber="856">Balaenoptera</emphasis>
</taxonomicName>
in this feature. Ŋis indicates that at least two balaenopterids are represented in the San Gregorio assemblage based on petrosal morphology. UCMP 219136 is nearly identical in morphology to a fossil balaenopterid (SDNHM 80102) formerly identified as
<taxonomicName id="4C4E17FCFFFA8677256E7FB5FD9F3F33" authority="(Demere et al. 2005)" baseAuthorityName="Demere" baseAuthorityYear="2005" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="42" pageNumber="857" phylum="Chordata" rank="species" species="davidsonii">
<emphasis id="B93AB06DFFFA8677256E7FB5FE813F32" italics="true" pageId="42" pageNumber="857">Balaenoptera davidsonii</emphasis>
(Deméré
<emphasis id="B93AB06DFFFA867726C17F94FE213F32" box="[390,442,1008,1034]" italics="true" pageId="42" pageNumber="857">et al.</emphasis>
2005)
</taxonomicName>
, which now appears to represent a separate balaenopterid taxon (T. A. Deméré pers. comm. 2011). Ŋe skull of SDNHM 80102 lacks squamosal creases and has elongate zygomatic processes, unlike
<taxonomicName id="4C4E17FCFFFA867726B9780BFF543F91" authority="bertae n." authorityName="BERTAE N." class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="42" pageNumber="850" phylum="Chordata" rank="genus">
<emphasis id="B93AB06DFFFA867726B9780BFF2A3F90" italics="true" pageId="42" pageNumber="857">Balaenoptera bertae</emphasis>
<taxonomicNameLabel id="A2090D16FFFA867727FD78EAFF543F91" box="[186,207,1167,1193]" pageId="42" pageNumber="857" rank="species">n.</taxonomicNameLabel>
</taxonomicName>
sp., and it is thus unlikely that UCMP 219136 is a petrosal of
<taxonomicName id="4C4E17FCFFFA8677263378CAFE483FF0" authority="Boessenecker, 2013" authorityName="Boessenecker" authorityYear="2013" box="[372,467,1198,1224]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="42" pageNumber="857" phylum="Chordata" rank="species" species="bertae" status="sp. nov.">
<emphasis id="B93AB06DFFFA8677263378CAFE483FF0" box="[372,467,1198,1224]" italics="true" pageId="42" pageNumber="857">B. bertae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2090D16FFFA8677269C78CBFD8C3FF0" box="[475,535,1198,1224]" pageId="42" pageNumber="857" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>