535 lines
74 KiB
XML
535 lines
74 KiB
XML
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<mods:title id="BEC79C29C21A88C9C822EE15F6E18FE0">3378</mods:title>
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<mods:date id="ADEC5831721F8D9AA1020C628C7C707D">2012</mods:date>
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<emphasis id="B93E0F56C522FFFCF39BFF3AFE44FF2B" bold="true" box="[151,449,151,177]" italics="true" pageId="29" pageNumber="30">Hemidactylus hajarensis</emphasis>
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(
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<figureCitation id="1371CFC1C522FFFCF393FF17FF6AFF48" box="[159,239,186,210]" captionStart="FIGURE 2" captionStartId="13.[151,244,1580,1601]" captionTargetBox="[160,1436,197,1542]" captionTargetId="figure-120@13.[151,1436,191,1544]" captionTargetPageId="13" captionText="FIGURE 2. Distribution map of Hemidactylus luqueorum sp. nov., H. hajarensis sp. nov., H. alkiyumii sp. nov. and H. festivus sp. nov. Color dots indicate specimens included in the molecular analyses and listed in Table 1. Stars indicate Museum specimens included in the morphological analyses only and listed in Appendix I: 1) BMNH1975.916; 2) BMNH1975.41, BMNH1980.558; 3) BMNH1976.1404; 4) BMNH1977.35; 5) BMNH1983.706; 6) BMNH1977.975; 7) BMNH1977.976–981; 8) BMNH1977.963–966; 9) BMNH1977.956–959; 10) BMNH1977.972–973 and BMNH1976.1409. Dashed lines delimit the different genetic lineages found within H. hajarensis (B1 and B2) and H. alkiyumii (C1, C2 and C3) and shown in Fig. 5 and Appendix III." pageId="29" pageNumber="30">Figs. 2</figureCitation>
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,
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<figureCitation id="1371CFC1C522FFFCF3F6FF17FE98FF48" box="[250,285,186,210]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza & Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="29" pageNumber="30">5B</figureCitation>
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,
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<figureCitation id="1371CFC1C522FFFCF225FF17FEBCFF48" box="[297,313,186,210]" captionStart="FIGURE 6" captionStartId="20.[151,244,1945,1966]" captionTargetBox="[192,1390,199,1918]" captionTargetId="figure-83@20.[191,1396,191,1928]" captionTargetPageId="20" captionText="FIGURE 6. Statistical parsimony nuclear allele networks of c-mos, mc1r and rag2 loci with colors corresponding to species H. persicus, H. luqueorum sp. nov. and H. hajarensis sp. nov.. Circle sizes are proportional to the number of alleles. White circles represent mutational steps. B1 and B2 refer to the different genetic lineages found within H. hajarensis (see Fig. 5 and Appendix III)." pageId="29" pageNumber="30">6</figureCitation>
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,
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<figureCitation id="1371CFC1C522FFFCF249FF17FE0FFF48" box="[325,394,186,210]" captionStart="FIGURE 9" captionStartId="25.[151,244,1730,1751]" captionTargetBox="[151,1436,191,1708]" captionTargetId="figure-78@25.[151,1436,191,1708]" captionTargetPageId="25" captionText="FIGURE 9. A) male, Holotype of H. luqueorum sp. nov. from Sayq, Jebel Akhdar (BMNH2005.1660); B) left: male of H. luqueorum Wadi Sayq, Jebel Akhdar (BMNH1980.558); right: female of H. hajarensis sp. nov. from Wadi Sabt, Jebel Akhdar (BMNH1977.35); C) above, H. hajarensis (BMNH1977.35); below H. luqueorum (BMNH1980.558); D) underside of right hind feet; left: H. luqueorum (BMNH1980.558), right: H. hajarensis (BMNH1977.35)." pageId="29" pageNumber="30">9B–D</figureCitation>
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,
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<figureCitation id="1371CFC1C522FFFCF29AFF17FE5AFF48" box="[406,479,186,210]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStartId-0="30.[151,244,1936,1957]" captionStartId-1="32.[151,244,1884,1905]" captionTargetBox-0="[211,1376,191,1915]" captionTargetBox-1="[345,1247,191,1868]" captionTargetId-0="figure-62@30.[211,1376,191,1915]" captionTargetId-1="figure-114@32.[345,1247,191,1868]" captionTargetPageId-0="30" captionTargetPageId-1="32" captionText-0="FIGURE 12. Live specimens of H. hajarensis sp. nov. A) male from Wadi Tanuf (IBES7151); B) male from Wadi Bani Khalid (IBES7335); C) female from Wadi Tanuf (IBES8064); D) male, Holotype, from Wadi Bani Khalid (BMNH2008.714); E) same specimen as in B." captionText-1="FIGURE 13. Different localities where H. hajarensis sp. nov. has been found A) Wadi Tanuf, Jebel Akhdar. In this locality H. hajarensis has been found in several occasion during the night on the rocky sides of the wadi; B) Wadi Bani Khalid, Eastern Hajar Mountains, where H. hajarensis was found at night on the ground and on boulders; D) 9 km N of Al Chayan, in the Eastern Hajar Mountains, where H. hajarensis was relatively abundant and several specimens were found on the rocky substrate and on boulders and rocky sides of the small wadis." pageId="29" pageNumber="30">12–13</figureCitation>
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; Table 1; Appendix I; Appendix IIIB)
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MorphoBank
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<emphasis id="B93E0F56C522FFFCF39BFEE7FEFBFEFA" box="[151,382,330,352]" italics="true" pageId="29" pageNumber="30">Hemidactylus persicus</emphasis>
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(part.);
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(part.);
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<bibRefCitation id="EFDBAEB5C522FFFCF21FFEC4FDD8FEE5" author="Carranza, S. & Arnold, E. N." box="[275,605,361,383]" pageId="29" pageNumber="30" pagination="531 - 545" refId="ref42847" refString="Carranza, S. & Arnold, E. N. (2006) Systematics, biogeography, and evolution of Hemidactylus geckos (Reptilia: Gekkonidae) elucidated using mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 38, 531 - 545." type="journal article" year="2006">Carranza and Arnold, 2006: 536</bibRefCitation>
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;
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|
||
<bibRefCitation id="EFDBAEB5C522FFFCF16BFEC4FC51FEE5" author="Sindaco, R. & Jeremcenko, V. K." box="[615,980,361,383]" pageId="29" pageNumber="30" refId="ref45930" refString="Sindaco, R. & Jeremcenko, V. K. (2008) The reptiles of the Western Palearctic. Annotated checklist and distributional atlas of the turtles, crocodiles, amphisbaenians and lizards of Europe, North Africa, Middle East and Central Asia. Monografie della Societas Herpetologica Italica - I. 579 pp." type="book" year="2008">Sindaco and Jeremcenko, 2008: 115</bibRefCitation>
|
||
</treatmentCitation>
|
||
(part.).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35080CFC522FFC0F39BFE01FA89FCDA" lastPageId="33" lastPageNumber="34" pageId="29" pageNumber="30" type="materials_examined">
|
||
<paragraph id="8BF5D344C522FFFCF39BFE01FF7BFE5E" blockId="29.[151,1436,428,631]" box="[151,254,428,452]" pageId="29" pageNumber="30">
|
||
<emphasis id="B93E0F56C522FFFCF39BFE01FF7BFE5E" box="[151,254,428,452]" italics="true" pageId="29" pageNumber="30">
|
||
<typeStatus id="54F16DE6C522FFFCF39BFE01FF7BFE5E" box="[151,254,428,452]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF3CBFE62FC69FDED" blockId="29.[151,1436,428,631]" pageId="29" pageNumber="30">
|
||
<materialsCitation id="3B22D919C522FFFCF3CBFE62FC69FDED" accessionNumber="M99903, M99917, M99885, M99893, M99894, M99902, M99969, M99976" collectingDate="2011-05" collectionCode="BMNH, ONHM, IBES" collectorName="S. Carranza & E. Gomez-Diaz & F. Amat & MorphoBank & Paratypes" country="Oman" county="North" elevation="647" latitude="59.09371" location="Wadi Bani Khalid" municipality="Eastern Hajar" pageId="29" pageNumber="30" specimenCode="BMNH2008.714, ONHM3706, IBES7335, IBES7336" specimenCount="3" typeStatus="paratype">
|
||
<specimenCode id="DBEC7B3FC522FFFCF3CBFE62FE02FE7D" box="[199,391,463,487]" collectionCode="BMNH" pageId="29" pageNumber="30">BMNH2008.714</specimenCode>
|
||
, male from
|
||
<location id="8E95859FC522FFFCF102FE62FD5EFE7D" LSID="urn:lsid:plazi:treatment:03E36252C522FFC0F39BFF3AFA89FCDA:8E95859FC522FFFCF102FE62FD5EFE7D" box="[526,731,463,487]" country="Oman" county="North" latitude="59.09371" municipality="Eastern Hajar" name="Wadi Bani Khalid" pageId="29" pageNumber="30">Wadi Bani Khalid</location>
|
||
,
|
||
<quantity id="4CB27EA1C522FFFCF1EAFE62FCB5FE7D" box="[742,816,463,487]" metricMagnitude="2" metricUnit="m" metricValue="6.47" pageId="29" pageNumber="30" unit="m" value="647.0">
|
||
<elevation id="00673477C522FFFCF1EAFE62FCB5FE7D" box="[742,816,463,487]" metricMagnitude="2" metricUnit="m" metricValue="6.47" pageId="29" pageNumber="30" unit="m" value="647.0">647 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMunicipality id="6B91493EC522FFFCF030FE7DFC50FE7D" box="[828,981,463,487]" pageId="29" pageNumber="30">Eastern Hajar</collectingMunicipality>
|
||
(
|
||
<collectingCounty id="6294ABC8C522FFFCF0E8FE62FBA2FE7D" box="[996,1063,463,487]" pageId="29" pageNumber="30">North</collectingCounty>
|
||
<collectingCountry id="F35D93D4C522FFFCF722FE62FBF2FE7D" box="[1070,1143,463,487]" name="Oman" pageId="29" pageNumber="30">Oman</collectingCountry>
|
||
), 22.61609’
|
||
<geoCoordinate id="EE7EB583C522FFFCF7F0FE62FA03FE7D" box="[1276,1414,463,487]" degrees="59.09371" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="59.09371">N 59.09371</geoCoordinate>
|
||
’E WGS84, collected in
|
||
<date id="FFF4F584C522FFFCF28AFE59FE72FD96" box="[390,503,500,524]" pageId="29" pageNumber="30" value="2011-05">
|
||
<collectingDate id="EFB00C6CC522FFFCF28AFE59FE72FD96" box="[390,503,500,524]" pageId="29" pageNumber="30" value="2011-05">May 2011</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26BFB692C522FFFCF113FE59FD21FD96" box="[543,676,500,524]" pageId="29" pageNumber="30">S. Carranza</collectorName>
|
||
,
|
||
<collectorName id="26BFB692C522FFFCF1A3FE59FCD9FD96" box="[687,860,500,524]" pageId="29" pageNumber="30">E. Gómez-Díaz</collectorName>
|
||
and
|
||
<collectorName id="26BFB692C522FFFCF09EFE59FC6EFD96" box="[914,1003,500,524]" pageId="29" pageNumber="30">F. Amat</collectorName>
|
||
(
|
||
<collectorName id="26BFB692C522FFFCF0F5FE59FB15FD96" box="[1017,1168,500,524]" pageId="29" pageNumber="30">MorphoBank</collectorName>
|
||
<accessionNumber id="94194EA7C522FFFCF79AFE59FB73FD96" box="[1174,1270,500,524]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99903" pageId="29" pageNumber="30" type="EnaNcbi">M99903</accessionNumber>
|
||
–
|
||
<accessionNumber id="94194EA7C522FFFCF609FE59FAE0FD96" box="[1285,1381,500,524]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99917" pageId="29" pageNumber="30" type="EnaNcbi">M99917</accessionNumber>
|
||
;
|
||
<figureCitation id="1371CFC1C522FFFCF67CFE59FF4EFDB5" captionStart="FIGURE 12" captionStartId="30.[151,244,1936,1957]" captionTargetBox="[211,1376,191,1915]" captionTargetId="figure-62@30.[211,1376,191,1915]" captionTargetPageId="30" captionText="FIGURE 12. Live specimens of H. hajarensis sp. nov. A) male from Wadi Tanuf (IBES7151); B) male from Wadi Bani Khalid (IBES7335); C) female from Wadi Tanuf (IBES8064); D) male, Holotype, from Wadi Bani Khalid (BMNH2008.714); E) same specimen as in B." pageId="29" pageNumber="30">Fig. 12D</figureCitation>
|
||
).
|
||
<emphasis id="B93E0F56C522FFFCF3EBFDB5FEDAFDAA" bold="true" box="[231,351,536,560]" pageId="29" pageNumber="30">
|
||
<typeStatus id="54F16DE6C522FFFCF3EBFDB5FEDAFDAA" box="[231,351,536,560]" pageId="29" pageNumber="30" type="paratype">
|
||
<collectorName id="26BFB692C522FFFCF3EBFDB5FEDAFDAA" box="[231,351,536,560]" pageId="29" pageNumber="30">Paratypes</collectorName>
|
||
</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCode id="DBEC7B3FC522FFFCF27BFDBAFD82FDB5" box="[375,519,535,559]" collectionCode="ONHM" pageId="29" pageNumber="30">ONHM3706</specimenCode>
|
||
, male, same collecting data as Holotype (
|
||
<collectorName id="26BFB692C522FFFCF714FDBAFB2AFDB5" box="[1048,1199,535,559]" pageId="29" pageNumber="30">MorphoBank</collectorName>
|
||
<accessionNumber id="94194EA7C522FFFCF7B3FDBAFAA5FDB5" box="[1215,1312,535,559]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99885" pageId="29" pageNumber="30" type="EnaNcbi">M99885</accessionNumber>
|
||
–
|
||
<accessionNumber id="94194EA7C522FFFCF623FDBAFA15FDB5" box="[1327,1424,535,559]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99893" pageId="29" pageNumber="30" type="EnaNcbi">M99893</accessionNumber>
|
||
);
|
||
<specimenCode id="DBEC7B3FC522FFFCF39BFD91FE8BFDCE" box="[151,270,572,596]" collectionCode="IBES" pageId="29" pageNumber="30">IBES7335</specimenCode>
|
||
, male, same collecting data as
|
||
<typeStatus id="54F16DE6C522FFFCF19FFD91FD7EFDCE" box="[659,763,572,596]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectorName id="26BFB692C522FFFCF01DFD91FC2DFDCE" box="[785,936,572,596]" pageId="29" pageNumber="30">MorphoBank</collectorName>
|
||
<accessionNumber id="94194EA7C522FFFCF0BAFD91FB93FDCE" box="[950,1046,572,596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99894" pageId="29" pageNumber="30" type="EnaNcbi">M99894</accessionNumber>
|
||
–
|
||
<accessionNumber id="94194EA7C522FFFCF729FD91FB00FDCE" box="[1061,1157,572,596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99902" pageId="29" pageNumber="30" type="EnaNcbi">M99902</accessionNumber>
|
||
;
|
||
<figureCitation id="1371CFC1C522FFFCF794FD91FA80FDCE" box="[1176,1285,572,596]" captionStart="FIGURE 11" captionStartId="28.[151,244,1906,1927]" captionTargetBox="[228,1358,191,1884]" captionTargetId="figure-118@28.[228,1358,191,1884]" captionTargetPageId="28" captionText="FIGURE 11. Different localities where H. luqueorum sp. nov. has been found A) 1 km E of Hat, Jebel Akhdar. In this locality H. luqueorum was found out during the day close to crevices and in small caves on the large boulders at the sides of the narrow Wadi; B) Wadi Bani Habib, Jebel Akhdar. In this locality H. luqueorum was found at night in man made constructions and on boulders; C) Wadi al Khahafa, Jebel Akhdar, where two specimens of H. luqueorum have been found at just 492 m altitude on the rocky sides of the wadi." pageId="29" pageNumber="30">Fig. 11D</figureCitation>
|
||
);
|
||
<specimenCode id="DBEC7B3FC522FFFCF62DFD91FA1DFDCE" box="[1313,1432,572,596]" collectionCode="IBES" pageId="29" pageNumber="30">IBES7336</specimenCode>
|
||
, female, same collecting data as
|
||
<typeStatus id="54F16DE6C522FFFCF2F7FDF2FDE6FDED" box="[507,611,607,631]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectorName id="26BFB692C522FFFCF17EFDF2FC8CFDED" box="[626,777,607,631]" pageId="29" pageNumber="30">MorphoBank</collectorName>
|
||
<accessionNumber id="94194EA7C522FFFCF01CFDF2FCF4FDED" box="[784,881,607,631]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99969" pageId="29" pageNumber="30" type="EnaNcbi">M99969</accessionNumber>
|
||
–
|
||
<accessionNumber id="94194EA7C522FFFCF08CFDF2FC64FDED" box="[896,993,607,631]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99976" pageId="29" pageNumber="30" type="EnaNcbi">M99976</accessionNumber>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF39BFD05FE35FD5A" blockId="29.[151,1436,680,812]" box="[151,432,680,704]" pageId="29" pageNumber="30">
|
||
<emphasis id="B93E0F56C522FFFCF39BFD05FE35FD5A" box="[151,432,680,704]" italics="true" pageId="29" pageNumber="30">Other material examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF3CBFD61FD43FCB6" blockId="29.[151,1436,680,812]" pageId="29" pageNumber="30">
|
||
Eighteen vouchers listed in Appendix I under
|
||
<taxonomicName id="4C4AA8C7C522FFFCF1E8FD60FCF8FD7E" authority="Carranza & Arnold, 2012" authorityName="Carranza & Arnold" authorityYear="2012" box="[740,893,716,740]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis" status="sp. nov.">
|
||
<emphasis id="B93E0F56C522FFFCF1E8FD60FCF8FD7E" box="[740,893,716,740]" italics="true" pageId="29" pageNumber="30">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20DB22DC522FFFCF08BFD61FC67FD79" box="[903,994,716,739]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and not mentioned above. Specimens
|
||
<collectionCode id="ED5B4B81C522FFFCF39BFD42FF48FC9D" box="[151,205,751,775]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
|
||
227612,
|
||
<collectionCode id="ED5B4B81C522FFFCF222FD42FEE1FC9D" box="[302,356,751,775]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="30" type="Museum">CAS</collectionCode>
|
||
227614,
|
||
<collectionCode id="ED5B4B81C522FFFCF2CAFD42FD9FFC9D" box="[454,538,751,775]" pageId="29" pageNumber="30">BMNH</collectionCode>
|
||
2008.705 (juvenile) and samples JS65, JS81, JS98 JS99, S7321, and S6061 were included in the molecular analyses only (Table 1).
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF39BFCF0FE8CFCEE" blockId="29.[151,1436,861,1435]" box="[151,265,861,884]" pageId="29" pageNumber="30">
|
||
<emphasis id="B93E0F56C522FFFCF39BFCF0FE8CFCEE" box="[151,265,861,884]" italics="true" pageId="29" pageNumber="30">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF3CBFCD2FB65FB24" blockId="29.[151,1436,861,1435]" pageId="29" pageNumber="30">
|
||
A medium-sized
|
||
<taxonomicName id="4C4AA8C7C522FFFCF283FC2DFDACFC02" authorityName="Oken" authorityYear="1817" box="[399,553,896,920]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93E0F56C522FFFCF283FC2DFDACFC02" box="[399,553,896,920]" italics="true" pageId="29" pageNumber="30">Hemidactylus</emphasis>
|
||
</taxonomicName>
|
||
with a maximum recorded SVL of
|
||
<quantity id="4CB27EA1C522FFFCF0DDFCD2FBB9FC0D" box="[977,1084,895,919]" metricMagnitude="-2" metricUnit="m" metricValue="6.69" pageId="29" pageNumber="30" unit="mm" value="66.9">66.9 mm</quantity>
|
||
; with a mean of 14.2 (13–15) longitudinal rows of enlarged dorsal tubercles at mid-body; adhesive pads fairly broad, in adults maximum width of pad on the fourth toe about 0.4–0.5 of its length; lamellae under the 1
|
||
<superScript id="7C3F7E0CC522FFFCF0B6FC6AFC41FC4F" attach="left" box="[954,964,967,981]" fontSize="6" pageId="29" pageNumber="30">st</superScript>
|
||
toe of pes mean 8.0 (7–9); lamellae under the 4
|
||
<superScript id="7C3F7E0CC522FFFCF3D9FC43FF64FC66" attach="left" box="[213,225,1006,1020]" fontSize="6" pageId="29" pageNumber="30">th</superScript>
|
||
toe of pes mean 12.1 (11–14); preanal pores mean 5.5 (4–6); expanded subcaudal scales extending proximally as far as the second or third whorl after the vent and starting just after the hemipenial bulge in males; dorsum grey-buff with irregular small spots; a dark stripe from the nostril, through the eye, on to cheek above ear and often on to neck; tail with small irregular dark blotches basally and numerous transverse dark bands more distally, the total number being around 17. Underside of tail pale but large subcaudals may be suffused with grey formed by dark chromatophores that increase in intensity distally; underside of toe pads also grey.
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF3CBFB64FC7FFA01" blockId="29.[151,1436,861,1435]" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C4AA8C7C522FFFCF3CBFB64FE65FB7B" authorityName="Carranza & Arnold" authorityYear="2012" box="[199,480,1225,1249]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF3CBFB64FE65FB7B" box="[199,480,1225,1249]" italics="true" pageId="29" pageNumber="30">Hemidactylus hajarensis</emphasis>
|
||
</taxonomicName>
|
||
is generally similar to
|
||
<taxonomicName id="4C4AA8C7C522FFFCF1F0FB67FC04FB7B" authorityName="Arnold" authorityYear="1986" box="[764,897,1226,1249]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B93E0F56C522FFFCF1F0FB67FC04FB7B" box="[764,897,1226,1249]" italics="true" pageId="29" pageNumber="30">H. persicus</emphasis>
|
||
</taxonomicName>
|
||
in the number of its moderately-sized dorsal tubercles across mid-body, and large adhesive pads on toes but differs from it in its reduced number of preanal pores in males (mean 5.5, 4–6, compared with mean 9.2, 8–11).
|
||
<taxonomicName id="4C4AA8C7C522FFFCF0B6FABCFB53FAB3" authorityName="Carranza & Arnold" authorityYear="2012" box="[954,1238,1297,1321]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF0B6FABCFB53FAB3" box="[954,1238,1297,1321]" italics="true" pageId="29" pageNumber="30">Hemidactylus hajarensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4AA8C7C522FFFCF68DFABFFE95FAD4" authority="Carranza & Arnold, 2012" authorityName="Carranza & Arnold" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="luqueorum" status="sp. nov.">
|
||
<emphasis id="B93E0F56C522FFFCF68DFABFFE95FAD4" italics="true" pageId="29" pageNumber="30">H. luqueorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20DB22DC522FFFCF21BFA9BFEEBFAD7" box="[279,366,1334,1357]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in its smaller size (SVL mean
|
||
<quantity id="4CB27EA1C522FFFCF1C5FA98FC9FFAD7" box="[713,794,1333,1357]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="29" pageNumber="30" unit="mm" value="54.0">54 mm</quantity>
|
||
, max.
|
||
<quantity id="4CB27EA1C522FFFCF068FA98FC4FFAD7" box="[868,970,1333,1357]" metricMagnitude="-2" metricUnit="m" metricValue="6.69" pageId="29" pageNumber="30" unit="mm" value="66.9">66.9 mm</quantity>
|
||
, compared with mean.
|
||
<quantity id="4CB27EA1C522FFFCF7DCFA98FAB3FAD7" box="[1232,1334,1333,1357]" metricMagnitude="-2" metricUnit="m" metricValue="7.68" pageId="29" pageNumber="30" unit="mm" value="76.8">76.8 mm</quantity>
|
||
, max.
|
||
<quantity id="4CB27EA1C522FFFCF68CFA98FF42FAEE" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="29" pageNumber="30" unit="mm" value="88.0">88 mm</quantity>
|
||
), and in having fewer lamellae under the 1
|
||
<superScript id="7C3F7E0CC522FFFCF1ABFAF4FD34FAFD" attach="left" box="[679,689,1369,1383]" fontSize="6" pageId="29" pageNumber="30">st</superScript>
|
||
toe of pes (mean 8.0, 7–9, compared with mean 10.3, 10–11), and under the 4
|
||
<superScript id="7C3F7E0CC522FFFCF219FA2DFEA4FA14" attach="left" box="[277,289,1408,1422]" fontSize="6" pageId="29" pageNumber="30">th</superScript>
|
||
toe of pes (mean 12.1, 11–14, compared with mean 13.6, 13–14)
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF39BFA66FE8AFA79" blockId="29.[151,1435,1483,1579]" box="[151,271,1483,1507]" pageId="29" pageNumber="30">
|
||
<emphasis id="B93E0F56C522FFFCF39BFA66FE8AFA79" box="[151,271,1483,1507]" italics="true" pageId="29" pageNumber="30">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF3CBFA42FE01F9B1" blockId="29.[151,1435,1483,1579]" pageId="29" pageNumber="30">
|
||
The species epithet “
|
||
<taxonomicName id="4C4AA8C7C522FFFCF2B9FA42FDACF99D" authorityName="Carranza & Arnold" authorityYear="2012" box="[437,553,1519,1543]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF2B9FA42FDACF99D" box="[437,553,1519,1543]" italics="true" pageId="29" pageNumber="30">hajarensis</emphasis>
|
||
</taxonomicName>
|
||
” is an adjective that refers to the mountain range where the species is found, the Hajar Mountains.
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFCF39BF9F6FDC0F9E9" blockId="29.[151,1436,1627,2012]" box="[151,581,1627,1651]" pageId="29" pageNumber="30">
|
||
<emphasis id="B93E0F56C522FFFCF39BF9F6FDC0F9E9" box="[151,581,1627,1651]" italics="true" pageId="29" pageNumber="30">Genetic and phylogeographic remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C522FFFEF3CBF9D2FE3FFDC5" blockId="29.[151,1436,1627,2012]" lastBlockId="31.[151,1436,151,607]" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C4AA8C7C522FFFCF3CBF9D2FE5BF90D" authorityName="Carranza & Arnold" authorityYear="2012" box="[199,478,1663,1687]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF3CBF9D2FE5BF90D" box="[199,478,1663,1687]" italics="true" pageId="29" pageNumber="30">Hemidactylus hajarensis</emphasis>
|
||
</taxonomicName>
|
||
is monophyletic in the phylogenetic analyses of Dataset 1 (
|
||
<figureCitation id="1371CFC1C522FFFCF798F9D2FB69F90D" box="[1172,1260,1663,1687]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza & Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="29" pageNumber="30">Fig. 5B</figureCitation>
|
||
) and Dataset 3 (Appendix
|
||
<collectionCode id="ED5B4B81C522FFFCF21AF90EFECCF921" box="[278,329,1699,1723]" pageId="29" pageNumber="30">IIIB</collectionCode>
|
||
). In both phylogenetic trees it forms a clade together with
|
||
<taxonomicName id="4C4AA8C7C522FFFCF0EDF909FBE4F921" authorityName="Arnold" authorityYear="1986" box="[993,1121,1700,1723]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B93E0F56C522FFFCF0EDF909FBE4F921" box="[993,1121,1700,1723]" italics="true" pageId="29" pageNumber="30">H. persicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4AA8C7C522FFFCF796F909FAB0F921" authority="Carranza & Arnold, 2012" authorityName="Carranza & Arnold" authorityYear="2012" box="[1178,1333,1699,1723]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="luqueorum" status="sp. nov.">
|
||
<emphasis id="B93E0F56C522FFFCF796F909FAB0F921" box="[1178,1333,1699,1723]" italics="true" pageId="29" pageNumber="30">H. luqueorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20DB22DC522FFFCF632F90EFA1DF920" box="[1342,1432,1699,1722]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, although bootstrap support and pp values are very low (see
|
||
<figureCitation id="1371CFC1C522FFFCF021F96AFCEBF945" box="[813,878,1735,1759]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza & Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="29" pageNumber="30">Fig. 5</figureCitation>
|
||
and Appendix
|
||
<collectionCode id="ED5B4B81C522FFFCF714F96AFBBDF945" box="[1048,1080,1735,1759]" pageId="29" pageNumber="30">III</collectionCode>
|
||
). According to the results of the dating analysis inferred with Dataset 2, these three species split about 12.6 mya (95% HPD: 7.7–17.9). According to
|
||
<figureCitation id="1371CFC1C522FFFCF3B8F8A2FF72F8BD" box="[180,247,1807,1831]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza & Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="29" pageNumber="30">Fig. 5</figureCitation>
|
||
and Appendix
|
||
<collectionCode id="ED5B4B81C522FFFCF2AAF8BDFE41F8BD" box="[422,452,1808,1831]" pageId="29" pageNumber="30">III</collectionCode>
|
||
,
|
||
<taxonomicName id="4C4AA8C7C522FFFCF2DDF8BDFDE2F8BD" authorityName="Carranza & Arnold" authorityYear="2012" box="[465,615,1807,1831]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF2DDF8BDFDE2F8BD" box="[465,615,1807,1831]" italics="true" pageId="29" pageNumber="30">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
is more closely related to
|
||
<taxonomicName id="4C4AA8C7C522FFFCF09AF8BDFBB7F8BD" authorityName="Carranza & Arnold" authorityYear="2012" box="[918,1074,1807,1831]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="luqueorum">
|
||
<emphasis id="B93E0F56C522FFFCF09AF8BDFBB7F8BD" box="[918,1074,1807,1831]" italics="true" pageId="29" pageNumber="30">H. luqueorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1371CFC1C522FFFCF74FF8A2FB1FF8BD" box="[1091,1178,1807,1831]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza & Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="29" pageNumber="30">Fig. 5B</figureCitation>
|
||
, Appendix
|
||
<collectionCode id="ED5B4B81C522FFFCF612F8A2FAD5F8BD" box="[1310,1360,1807,1831]" pageId="29" pageNumber="30">IIIB</collectionCode>
|
||
), from which it split approximately 9.6 mya (95% HPD: 5.7–13.8). The level of genetic variability within
|
||
<taxonomicName id="4C4AA8C7C522FFFCF7E0F899FA06F8D1" authorityName="Carranza & Arnold" authorityYear="2012" box="[1260,1411,1843,1867]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF7E0F899FA06F8D1" box="[1260,1411,1843,1867]" italics="true" pageId="29" pageNumber="30">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
is very high, 6.1% in the
|
||
<emphasis id="B93E0F56C522FFFCF29AF8FAFE40F8F5" box="[406,453,1879,1903]" italics="true" pageId="29" pageNumber="30">cytb</emphasis>
|
||
and 2.6% in the
|
||
<emphasis id="B93E0F56C522FFFCF18EF8FAFD29F8F5" box="[642,684,1879,1903]" italics="true" pageId="29" pageNumber="30">12S</emphasis>
|
||
. As shown in
|
||
<figureCitation id="1371CFC1C522FFFCF040F8FAFC25F8F5" box="[844,928,1879,1903]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza & Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="29" pageNumber="30">Fig. 5B</figureCitation>
|
||
and Appendix
|
||
<collectionCode id="ED5B4B81C522FFFCF740F8F5FBF8F8F5" box="[1100,1149,1880,1903]" pageId="29" pageNumber="30">IIIB</collectionCode>
|
||
,
|
||
<taxonomicName id="4C4AA8C7C522FFFCF784F8F5FA9BF8F5" authorityName="Carranza & Arnold" authorityYear="2012" box="[1160,1310,1879,1903]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C522FFFCF784F8F5FA9BF8F5" box="[1160,1310,1879,1903]" italics="true" pageId="29" pageNumber="30">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
consists of two very well differentiated and well-supported clades, B1 and B2. The uncorrected genetic distances between these two clades are 10.5% in the
|
||
<emphasis id="B93E0F56C522FFFCF128F832FDD6F82D" box="[548,595,1951,1975]" italics="true" pageId="29" pageNumber="30">cytb</emphasis>
|
||
and 5% in the
|
||
<emphasis id="B93E0F56C522FFFCF00BF832FCB7F82D" box="[775,818,1951,1975]" italics="true" pageId="29" pageNumber="30">12S</emphasis>
|
||
. As shown in
|
||
<figureCitation id="1371CFC1C522FFFCF0D0F832FBA7F82D" box="[988,1058,1951,1975]" captionStart="FIGURE 2" captionStartId="13.[151,244,1580,1601]" captionTargetBox="[160,1436,197,1542]" captionTargetId="figure-120@13.[151,1436,191,1544]" captionTargetPageId="13" captionText="FIGURE 2. Distribution map of Hemidactylus luqueorum sp. nov., H. hajarensis sp. nov., H. alkiyumii sp. nov. and H. festivus sp. nov. Color dots indicate specimens included in the molecular analyses and listed in Table 1. Stars indicate Museum specimens included in the morphological analyses only and listed in Appendix I: 1) BMNH1975.916; 2) BMNH1975.41, BMNH1980.558; 3) BMNH1976.1404; 4) BMNH1977.35; 5) BMNH1983.706; 6) BMNH1977.975; 7) BMNH1977.976–981; 8) BMNH1977.963–966; 9) BMNH1977.956–959; 10) BMNH1977.972–973 and BMNH1976.1409. Dashed lines delimit the different genetic lineages found within H. hajarensis (B1 and B2) and H. alkiyumii (C1, C2 and C3) and shown in Fig. 5 and Appendix III." pageId="29" pageNumber="30">Fig. 2</figureCitation>
|
||
, the geographical limits between clades B1 and B2 are not very clear, being clade B1 present in the Jebel Akhdar and in the coastal areas close to
|
||
<collectingCountry id="F35D93D4C520FFFEF39BFF35FF6CFF35" box="[151,233,152,175]" name="Oman" pageId="31" pageNumber="32">Muscat</collectingCountry>
|
||
(Wadi Mayh, Jebel Abu Daud and a wadi North of Qurayyat), while clade B2 seems restricted to the Eastern Hajars. According to the calibrations, clades B1 and B2 split approximately 4.7 mya (95% HPD: 2.6–7.0). The uncorrected genetic distances between
|
||
<taxonomicName id="4C4AA8C7C520FFFEF18DFF4CFC92FF62" authorityName="Carranza & Arnold" authorityYear="2012" box="[641,791,224,248]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF18DFF4CFC92FF62" box="[641,791,224,248]" italics="true" pageId="31" pageNumber="32">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4AA8C7C520FFFEF043FF4CFC6EFF62" authorityName="Carranza & Arnold" authorityYear="2012" box="[847,1003,224,248]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="luqueorum">
|
||
<emphasis id="B93E0F56C520FFFEF043FF4CFC6EFF62" box="[847,1003,224,248]" italics="true" pageId="31" pageNumber="32">H. luqueorum</emphasis>
|
||
</taxonomicName>
|
||
are 15.6% in the
|
||
<emphasis id="B93E0F56C520FFFEF7B8FF4DFB67FF62" box="[1204,1250,224,248]" italics="true" pageId="31" pageNumber="32">cytb</emphasis>
|
||
and 6.9% in the
|
||
<emphasis id="B93E0F56C520FFFEF39BFEA9FF44FE86" box="[151,193,260,284]" italics="true" pageId="31" pageNumber="32">12S</emphasis>
|
||
; and between
|
||
<taxonomicName id="4C4AA8C7C520FFFEF268FEA8FE7CFE86" authorityName="Carranza & Arnold" authorityYear="2012" box="[356,505,260,284]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF268FEA8FE7CFE86" box="[356,505,260,284]" italics="true" pageId="31" pageNumber="32">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4AA8C7C520FFFEF13CFEA8FD2AFE86" authorityName="Arnold" authorityYear="1986" box="[560,687,261,284]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="persicus">
|
||
<emphasis id="B93E0F56C520FFFEF13CFEA8FD2AFE86" box="[560,687,261,284]" italics="true" pageId="31" pageNumber="32">H. persicus</emphasis>
|
||
</taxonomicName>
|
||
15.1% in the
|
||
<emphasis id="B93E0F56C520FFFEF040FEA9FCFEFE86" box="[844,891,260,284]" italics="true" pageId="31" pageNumber="32">cytb</emphasis>
|
||
and 10.7% in the
|
||
<emphasis id="B93E0F56C520FFFEF74BFEA9FBF4FE86" box="[1095,1137,260,284]" italics="true" pageId="31" pageNumber="32">12S</emphasis>
|
||
. The results of the nuclear networks presented in
|
||
<figureCitation id="1371CFC1C520FFFEF2ACFE8AFE5AFEA5" box="[416,479,295,319]" captionStart="FIGURE 6" captionStartId="20.[151,244,1945,1966]" captionTargetBox="[192,1390,199,1918]" captionTargetId="figure-83@20.[191,1396,191,1928]" captionTargetPageId="20" captionText="FIGURE 6. Statistical parsimony nuclear allele networks of c-mos, mc1r and rag2 loci with colors corresponding to species H. persicus, H. luqueorum sp. nov. and H. hajarensis sp. nov.. Circle sizes are proportional to the number of alleles. White circles represent mutational steps. B1 and B2 refer to the different genetic lineages found within H. hajarensis (see Fig. 5 and Appendix III)." pageId="31" pageNumber="32">Fig 6</figureCitation>
|
||
and a network analysis including all specimens from Dataset 1 (data not shown) indicate that all alleles of
|
||
<taxonomicName id="4C4AA8C7C520FFFEF2B7FEE0FDD5FEFE" authorityName="Carranza & Arnold" authorityYear="2012" box="[443,592,332,356]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF2B7FEE0FDD5FEFE" box="[443,592,332,356]" italics="true" pageId="31" pageNumber="32">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
for all three independent loci analyzed (c
|
||
<emphasis id="B93E0F56C520FFFEF724FEE0FBDBFEFE" box="[1064,1118,333,356]" italics="true" pageId="31" pageNumber="32">-mos</emphasis>
|
||
,
|
||
<emphasis id="B93E0F56C520FFFEF761FEE1FB23FEFE" box="[1133,1190,332,356]" italics="true" pageId="31" pageNumber="32">mc1r</emphasis>
|
||
and
|
||
<emphasis id="B93E0F56C520FFFEF7D2FEE1FA96FEFE" box="[1246,1299,332,356]" italics="true" pageId="31" pageNumber="32">rag2</emphasis>
|
||
) are private (not shared with any other species included in the present analyses). It is also interesting to notice that, although clades B1 and B2 share alleles of the c
|
||
<emphasis id="B93E0F56C520FFFEF147FE38FD04FE36" box="[587,641,405,428]" italics="true" pageId="31" pageNumber="32">-mos</emphasis>
|
||
gene, specimens of these two clades do not share a single allele of the
|
||
<emphasis id="B93E0F56C520FFFEF39BFE15FF55FE4A" box="[151,208,440,464]" italics="true" pageId="31" pageNumber="32">mc1r</emphasis>
|
||
and
|
||
<emphasis id="B93E0F56C520FFFEF20AFE15FEBEFE4A" box="[262,315,440,464]" italics="true" pageId="31" pageNumber="32">rag2</emphasis>
|
||
nuclear genes. The high genetic differentiation between clades B1 and B2 of
|
||
<taxonomicName id="4C4AA8C7C520FFFEF7ACFE14FAB3FE4A" authorityName="Carranza & Arnold" authorityYear="2012" box="[1184,1334,440,464]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF7ACFE14FAB3FE4A" box="[1184,1334,440,464]" italics="true" pageId="31" pageNumber="32">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
suggests long separation of the two units. However, the absence of clear morphological differences between these two clades and the relatively low number of available vouchers to carry out a thorough morphological analysis prevents us from taking any taxonomic decisions at present. Future studies should clarify the taxonomic status of these two clades (work in progress).
|
||
</paragraph>
|
||
<caption id="DF3583CCC521FFFFF39BF83DFEC5F872" pageId="30" pageNumber="31" startId="30.[151,244,1936,1957]" targetBox="[211,1376,191,1915]" targetPageId="30">
|
||
<paragraph id="8BF5D344C521FFFFF39BF83DFEC5F872" blockId="30.[151,1436,1936,2024]" pageId="30" pageNumber="31">
|
||
<emphasis id="B93E0F56C521FFFFF39BF83DFE99F83F" bold="true" box="[151,284,1936,1957]" pageId="30" pageNumber="31">FIGURE 12.</emphasis>
|
||
Live specimens of
|
||
<taxonomicName id="4C4AA8C7C521FFFFF2E8F83CFDE2F83C" authority="Carranza & Arnold, 2012" authorityName="Carranza & Arnold" authorityYear="2012" box="[484,615,1937,1958]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="hajarensis" status="sp. nov.">
|
||
<emphasis id="B93E0F56C521FFFFF2E8F83CFDE2F83C" box="[484,615,1937,1958]" italics="true" pageId="30" pageNumber="31">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20DB22DC521FFFFF17CF83DFD3BF83F" box="[624,702,1936,1957]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A) male from Wadi Tanuf (IBES7151); B) male from Wadi Bani Khalid (IBES7335); C) female from Wadi Tanuf (IBES8064); D) male, Holotype, from Wadi Bani Khalid (BMNH2008.714); E) same specimen as in B.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF5D344C520FFFEF39BFD3DFEA5FD32" blockId="31.[151,1436,656,895]" box="[151,288,656,680]" pageId="31" pageNumber="32">
|
||
<emphasis id="B93E0F56C520FFFEF39BFD3DFEA5FD32" box="[151,288,656,680]" italics="true" pageId="31" pageNumber="32">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C520FFFEF3CBFD19FEDAFCE5" blockId="31.[151,1436,656,895]" pageId="31" pageNumber="32">
|
||
The species is widespread in the mountains of North
|
||
<collectingCountry id="F35D93D4C520FFFEF046FD19FC0BFD56" box="[842,910,692,716]" name="Oman" pageId="31" pageNumber="32">Oman</collectingCountry>
|
||
from the Jebel Akhdar to the East. Despite intensive surveys across the Hajar Mountain range, it has never been found to the West of the Jebel Akhdar, in the Western Hajars or in the
|
||
<collectingRegion id="498E1DA6C520FFFEF2BAFD51FDB7FC8E" box="[438,562,764,788]" country="Oman" name="Musandam" pageId="31" pageNumber="32">Musandam</collectingRegion>
|
||
Peninsula. Several localities exist for the coastal wadis near
|
||
<collectingCountry id="F35D93D4C520FFFEF7E2FD51FAC5FC89" box="[1262,1344,764,787]" name="Oman" pageId="31" pageNumber="32">Muscat</collectingCountry>
|
||
and the Eastern Hajars, from Jebel Al Abyad in the West to Jebel Qahwan in the extreme East. Across its distribution range it has been recorded from almost sea level (
|
||
<quantity id="4CB27EA1C520FFFEF198FCE9FD57FCC1" box="[660,722,836,860]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="31" pageNumber="32" unit="m" value="22.0">22 m</quantity>
|
||
in Wadi Mayh) up to
|
||
<quantity id="4CB27EA1C520FFFEF0D5FCE9FBB4FCC1" box="[985,1073,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.683" pageId="31" pageNumber="32" unit="m" value="1683.0">1683 m</quantity>
|
||
in a locality
|
||
<quantity id="4CB27EA1C520FFFEF7C2FCE9FA8CFCC6" box="[1230,1289,836,860]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="km" value="9.0">9 km</quantity>
|
||
North of Al Chayan (Table 1).
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C520FFFEF39BFC04FF67FC5B" blockId="31.[151,1436,937,1249]" box="[151,226,937,961]" pageId="31" pageNumber="32">
|
||
<emphasis id="B93E0F56C520FFFEF39BFC04FF67FC5B" box="[151,226,937,961]" italics="true" pageId="31" pageNumber="32">Habits</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C520FFFEF3CBFC60FD13FB7B" blockId="31.[151,1436,937,1249]" pageId="31" pageNumber="32">
|
||
<taxonomicName id="4C4AA8C7C520FFFEF3CBFC60FE59FC7F" authorityName="Carranza & Arnold" authorityYear="2012" box="[199,476,973,997]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF3CBFC60FE59FC7F" box="[199,476,973,997]" italics="true" pageId="31" pageNumber="32">Hemidactylus hajarensis</emphasis>
|
||
</taxonomicName>
|
||
has been found at sides of wadis low on rocks that were interspersed and sometimes partly overhung with vegetation. The species was also sometimes seen on gravel floors of wadis (
|
||
<figureCitation id="1371CFC1C520FFFEF7EDFC5CFAB0FB93" box="[1249,1333,1009,1033]" captionStart="FIGURE 13" captionStartId="32.[151,244,1884,1905]" captionTargetBox="[345,1247,191,1868]" captionTargetId="figure-114@32.[345,1247,191,1868]" captionTargetPageId="32" captionText="FIGURE 13. Different localities where H. hajarensis sp. nov. has been found A) Wadi Tanuf, Jebel Akhdar. In this locality H. hajarensis has been found in several occasion during the night on the rocky sides of the wadi; B) Wadi Bani Khalid, Eastern Hajar Mountains, where H. hajarensis was found at night on the ground and on boulders; D) 9 km N of Al Chayan, in the Eastern Hajar Mountains, where H. hajarensis was relatively abundant and several specimens were found on the rocky substrate and on boulders and rocky sides of the small wadis." pageId="31" pageNumber="32">Fig. 13</figureCitation>
|
||
). Strictly nocturnal, it has never been recorded during the day.
|
||
<taxonomicName id="4C4AA8C7C520FFFEF1F0FBBBFC16FBB7" authorityName="Carranza & Arnold" authorityYear="2012" box="[764,915,1045,1069]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF1F0FBBBFC16FBB7" box="[764,915,1045,1069]" italics="true" pageId="31" pageNumber="32">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
moves quickly and is very agile, fleeing to a nearby refuge seconds after being spotted. In Wadi Tiwi and Wadi Hebaheba,
|
||
<taxonomicName id="4C4AA8C7C520FFFEF72CFB97FB3CFBCB" authorityName="Carranza & Arnold" authorityYear="2012" box="[1056,1209,1081,1105]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C520FFFEF72CFB97FB3CFBCB" box="[1056,1209,1081,1105]" italics="true" pageId="31" pageNumber="32">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
occurs also low on rocks. As
|
||
<taxonomicName id="4C4AA8C7C520FFFEF204FBF3FE21FBEF" authorityName="Carranza & Arnold" authorityYear="2012" box="[264,420,1117,1141]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="luqueorum">
|
||
<emphasis id="B93E0F56C520FFFEF204FBF3FE21FBEF" box="[264,420,1117,1141]" italics="true" pageId="31" pageNumber="32">H. luqueorum</emphasis>
|
||
</taxonomicName>
|
||
, it losses the skin very easily when being handled and sometimes specimens have scars of regenerated skin on the back, probably as a results of fights with conspecifics or attacks from predators (
|
||
<figureCitation id="1371CFC1C520FFFEF62BFB2CFA15FB03" box="[1319,1424,1153,1177]" captionStart="FIGURE 12" captionStartId="30.[151,244,1936,1957]" captionTargetBox="[211,1376,191,1915]" captionTargetId="figure-62@30.[211,1376,191,1915]" captionTargetPageId="30" captionText="FIGURE 12. Live specimens of H. hajarensis sp. nov. A) male from Wadi Tanuf (IBES7151); B) male from Wadi Bani Khalid (IBES7335); C) female from Wadi Tanuf (IBES8064); D) male, Holotype, from Wadi Bani Khalid (BMNH2008.714); E) same specimen as in B." pageId="31" pageNumber="32">Fig. 12D</figureCitation>
|
||
). This species has never been found in sympatry with
|
||
<taxonomicName id="4C4AA8C7C520FFFEF01FFB0BFC31FB27" authorityName="Carranza & Arnold" authorityYear="2012" box="[787,948,1189,1213]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="luqueorum">
|
||
<emphasis id="B93E0F56C520FFFEF01FFB0BFC31FB27" box="[787,948,1189,1213]" italics="true" pageId="31" pageNumber="32">H. luqueorum</emphasis>
|
||
</taxonomicName>
|
||
, however, it shares habitat with
|
||
<taxonomicName id="4C4AA8C7C520FFFEF64CFB08FEB0FB7B" authorityName="Arnold and Gardner" authorityYear="1994" class="Reptilia" family="Phyllodactylidae" genus="Asaccus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="platyrhynchus">
|
||
<emphasis id="B93E0F56C520FFFEF64CFB08FEB0FB7B" italics="true" pageId="31" pageNumber="32">Asaccus platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4AA8C7C520FFFEF267FB64FD0AFB7B" baseAuthorityName="Donndorf" baseAuthorityYear="1798" box="[363,655,1225,1249]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="hasselquistii">
|
||
<emphasis id="B93E0F56C520FFFEF267FB64FD0AFB7B" box="[363,655,1225,1249]" italics="true" pageId="31" pageNumber="32">Ptyodactylus hasselquistii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C520FFFEF39BFAA6FE9EFAB8" blockId="31.[151,1437,1291,2034]" box="[151,283,1291,1314]" pageId="31" pageNumber="32">
|
||
<emphasis id="B93E0F56C520FFFEF39BFAA6FE9EFAB8" box="[151,283,1291,1314]" italics="true" pageId="31" pageNumber="32">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C520FFFEF3CBFA83FB8DFA28" blockId="31.[151,1437,1291,2034]" pageId="31" pageNumber="32">Head and body markedly depressed. Head breadth variable and neck well defined. Head length about 24–31% of SVL (mean males 28%, mean females 27%), head width 65–85% of head length (mean males 71%, mean females 72%), and head height 36–55% of head length (mean males and females 42%). Adhesive pads fairly broad; in adults maximum width of pad on fourth hind toe about 0.4–0.5 of its length.</paragraph>
|
||
<paragraph id="8BF5D344C520FFFEF3CBFA13FC81F96C" blockId="31.[151,1437,1291,2034]" pageId="31" pageNumber="32">Nostril between rostral, supranasal and two superposed postnasals, with the first supralabial scale usually also entering narrowly into its border. Usually one scale separating supranasals on midline. About 14–18 scales in a straight line from postnasal to edge of orbit. Small conical tubercles scattered in orbital area, crown of head and temporal area above the level of ear opening and immediately in front of the upper part of this. Ear opening with its longest axis running upwards and backwards, smooth-edged, usually about half or more of eye diameter. Supralabial scales mean 10.7 (9–12), infralabials mean 8.8 (7–10). Mental scale broadly triangular posteriorly bordered by two large postmentals making contact behind it, a second pair of more lateral postmentals also present, the postmentals contacting the first and second supralabials; second and more posterior lower labials bordered by more irregular and smaller enlarged scales. Gulars fine.</paragraph>
|
||
<paragraph id="8BF5D344C520FFC0F3CBF8AFFB14FF60" blockId="31.[151,1437,1291,2034]" lastBlockId="33.[151,1436,151,685]" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
|
||
Enlarged tubercles present on back, arranged in 14.2 (mean) (13–15) longitudinal rows at mid-body, which also form backwardly directed oblique rows from near midline to flank, and
|
||
<quantity id="4CB27EA1C520FFFEF0FDF88BFBD1F8A4" box="[1009,1108,1830,1854]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="31" pageNumber="32" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
|
||
a paravertebral row from the level of the axilla to that of the groin, where they are separated by spaces of about their own length. Enlarged tubercles strongly keeled and trihedral but becoming smaller and more conical on flanks. Ventrals small, but larger than dorsals and more imbricate, about
|
||
<quantity id="4CB27EA1C520FFFEF150F83FFD44F830" box="[604,705,1938,1962]" metricMagnitude="-1" metricUnit="m" metricValue="9.779" metricValueMax="10.922" metricValueMin="8.636" pageId="31" pageNumber="32" unit="in" value="38.5" valueMax="43.0" valueMin="34.0">34–43 in</quantity>
|
||
a transverse row at mid body between lateral folds. Males with 4–6 preanal pores (mean 5.5) separated by one or two scales giving a formula of 2+2 or 3+3. Scales on upper forelimb small and imbricate, interspersed with enlarged tubercles on distal section. Scales on front of thigh and beneath hind leg about same size as belly scales and imbricate, enlarged tubercles present on upper surface of both femur and tibia. Lamellae under the toes of pes: 1
|
||
<superScript id="7C3F7E0CC51EFFC0F176FF72FD01FF77" attach="left" box="[634,644,223,237]" fontSize="6" pageId="33" pageNumber="34">st</superScript>
|
||
toe mean 8.0 (7–9), 4
|
||
<superScript id="7C3F7E0CC51EFFC0F076FF72FC03FF77" attach="left" box="[890,902,223,237]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
toe mean 12.1 (11–14).
|
||
</paragraph>
|
||
<caption id="DF3583CCC51FFFC1F39BF8F1FE45F86E" pageId="32" pageNumber="33" startId="32.[151,244,1884,1905]" targetBox="[345,1247,191,1868]" targetPageId="32">
|
||
<paragraph id="8BF5D344C51FFFC1F39BF8F1FE45F86E" blockId="32.[151,1437,1884,2036]" pageId="32" pageNumber="33">
|
||
<emphasis id="B93E0F56C51FFFC1F39BF8F1FE99F8EB" bold="true" box="[151,284,1884,1905]" pageId="32" pageNumber="33">FIGURE 13.</emphasis>
|
||
Different localities where
|
||
<taxonomicName id="4C4AA8C7C51FFFC1F127F8F1FD2AF8EB" authority="Carranza & Arnold, 2012" authorityName="Carranza & Arnold" authorityYear="2012" box="[555,687,1884,1905]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="hajarensis" status="sp. nov.">
|
||
<emphasis id="B93E0F56C51FFFC1F127F8F1FD2AF8EB" box="[555,687,1884,1905]" italics="true" pageId="32" pageNumber="33">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20DB22DC51FFFC1F1B4F8F1FC82F8EB" box="[696,775,1884,1905]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been found A) Wadi Tanuf, Jebel Akhdar. In this locality
|
||
<taxonomicName id="4C4AA8C7C51FFFC1F688F8F1FF7FF80B" authorityName="Carranza & Arnold" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C51FFFC1F688F8F1FF7FF80B" italics="true" pageId="32" pageNumber="33">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
has been found in several occasion during the night on the rocky sides of the wadi; B) Wadi Bani Khalid, Eastern Hajar Mountains, where
|
||
<taxonomicName id="4C4AA8C7C51FFFC1F241F831FE4AF82B" authorityName="Carranza & Arnold" authorityYear="2012" box="[333,463,1948,1969]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C51FFFC1F241F831FE4AF82B" box="[333,463,1948,1969]" italics="true" pageId="32" pageNumber="33">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
was found at night on the ground and on boulders; D) 9 km N of Al Chayan, in the Eastern Hajar Mountains, where
|
||
<taxonomicName id="4C4AA8C7C51FFFC1F245F811FE4CF84B" authorityName="Carranza & Arnold" authorityYear="2012" box="[329,457,1980,2001]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="hajarensis">
|
||
<emphasis id="B93E0F56C51FFFC1F245F811FE4CF84B" box="[329,457,1980,2001]" italics="true" pageId="32" pageNumber="33">H. hajarensis</emphasis>
|
||
</taxonomicName>
|
||
was relatively abundant and several specimens were found on the rocky substrate and on boulders and rocky sides of the small wadis.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF5D344C51EFFC0F3CBFEAAFAACFE17" blockId="33.[151,1436,151,685]" pageId="33" pageNumber="34">
|
||
Tail relatively slender; between eight and six enlarged, keeled and pointed tubercles on each whorl on tail base, dropping to four from about 4–10
|
||
<superScript id="7C3F7E0CC51EFFC0F101FE87FD9CFEA2" attach="left" box="[525,537,298,312]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
whorl after vent. Tubercles about one third the length of basal whorls, becoming smaller and placed more posteriorly on whorls distally. Subcaudal scales enlarged and broad, extending proximally as far as the second or third whorl after the vent and the starting just after the hemipenial bulge in males.
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C51EFFC0F3C3FE37FD1BFD37" blockId="33.[151,1436,151,685]" pageId="33" pageNumber="34">In spirit pale grey-buff; a dark stripe from the nostril, through the eye, on to cheek above ear and often on to neck; body with irregular small spots; some tubercles on forebody and vertebral area have opaque white coloring on one side and dark coloring on the other. Belly pale but there may be a very fine slight stipple at the sides, the dark punctuate spots being much smaller than the scales. Tail with small irregular dark blotches basally and numerous irregular dark bands more distally, the total number being around 17. Underside of tail pale but large subcaudals may be suffused in some places with grey formed by dark chromatophores that increase in intensity distally. Underside of toe pads also grey. In life, animals from many localities have dark blotches or bars on the upper surface that are suffused yellow-orange.</paragraph>
|
||
<paragraph id="8BF5D344C51EFFC0F39BFD73FE7EFD6C" blockId="33.[151,1436,734,832]" box="[151,507,734,758]" pageId="33" pageNumber="34">
|
||
<emphasis id="B93E0F56C51EFFC0F39BFD73FE7EFD6C" box="[151,507,734,758]" italics="true" pageId="33" pageNumber="34">
|
||
Distinctive features of
|
||
<typeStatus id="54F16DE6C51EFFC0F298FD73FE7EFD6C" box="[404,507,734,758]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5D344C51EFFC0F3CBFCACFA89FCDA" blockId="33.[151,1436,734,832]" pageId="33" pageNumber="34">
|
||
Male,
|
||
<quantity id="4CB27EA1C51EFFC0F202FCACFEF4FC83" box="[270,369,769,793]" metricMagnitude="-2" metricUnit="m" metricValue="5.9799999999999995" pageId="33" pageNumber="34" unit="mm" value="59.8">59.8 mm</quantity>
|
||
SVL; tail missing from the base. Supralabial scales 10/10; infralabials 9/9; 14 rows of enlarged tubercles at mid-back; 6 (3+3) preanal pores; lamellae under the 1
|
||
<superScript id="7C3F7E0CC51EFFC0F076FC88FC01FCA9" attach="left" box="[890,900,805,819]" fontSize="6" pageId="33" pageNumber="34">st</superScript>
|
||
toe of pes 8/8, 4
|
||
<superScript id="7C3F7E0CC51EFFC0F732FC88FBCFFCA9" attach="left" box="[1086,1098,805,819]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
toe of pes 12/12.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |