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<document id="97D62F65FA7830977F305F55D243ECD3" ID-DOI="10.5281/zenodo.175235" ID-GBIF-Dataset="823fc002-72b6-4b9f-b133-1ec97a85a0b1" ID-ISSN="1175­5326" ID-Zenodo-Dep="175235" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459802972764" checkinUser="plazi" docAuthor="Ying, Wang &amp; Dong, Ren" docDate="2007" docId="03D8522EFF97FFCE41DF0313FB9AF80F" docLanguage="en" docName="zt01390p049.pdf" docOrigin="Zootaxa 1390" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Quadraticossus longicaulis Ying &amp; Dong, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="46" masterDocId="FFE12A56FF95FFCB4148041AFF8AFFE6" masterDocTitle="Two new genera of fossil palaeontinids from the Middle Jurassic in Daohugou, Inner Mongolia, China (Hemiptera, Palaeontinidae)" masterLastPageNumber="49" masterPageNumber="41" pageNumber="43" updateTime="1698220193496" updateUser="plazi">
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<mods:title id="48E84C15C4D16AD77B5D895330AFBE37">Two new genera of fossil palaeontinids from the Middle Jurassic in Daohugou, Inner Mongolia, China (Hemiptera, Palaeontinidae)</mods:title>
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<mods:name id="4FFD1DD0079E781D034B012670EECF13" type="personal">
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<mods:namePart id="3567D15E11DB175050072A10806C4752">Ying, Wang</mods:namePart>
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<mods:name id="6A786BA1E6D413F487DF11D457217F5E" type="personal">
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<mods:namePart id="5682DDE5868744361475FFB10BC97D72">Dong, Ren</mods:namePart>
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<mods:typeOfResource id="85F7A420D0B7F4BC068B03EDE9223AC6">text</mods:typeOfResource>
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<mods:title id="C627E0B30C8C056628CC89592DA1C9D2">Zootaxa</mods:title>
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<mods:part id="B866013FF25AF767FAACF780B190FA3F">
<mods:date id="7F062841DFDCE0ED42CE09096884C93C">2007</mods:date>
<mods:detail id="2A1E12767BADE216FA3BE60CCFEB3597" type="volume">
<mods:number id="B02280B5FEE4B90567C7EABBB8E92D16">1390</mods:number>
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<mods:classification id="7FF3FE8D7C335A9D1829590DC7B5628E">journal article</mods:classification>
<mods:identifier id="7FB90205088F372028DBD90414A340A8" type="DOI">10.5281/zenodo.175235</mods:identifier>
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<mods:identifier id="2DDBB5C1E200E14A4BF901B94A707A60" type="ISSN">1175­5326</mods:identifier>
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<treatment id="03D8522EFF97FFCE41DF0313FB9AF80F" ID-DOI="http://doi.org/10.5281/zenodo.6243395" ID-GBIF-Taxon="119352198" ID-Zenodo-Dep="6243395" LSID="urn:lsid:plazi:treatment:03D8522EFF97FFCE41DF0313FB9AF80F" httpUri="http://treatment.plazi.org/id/03D8522EFF97FFCE41DF0313FB9AF80F" lastPageId="5" lastPageNumber="46" pageId="2" pageNumber="43">
<subSubSection id="C36BB0B3FF97FFC941DF0313FEACF8A3" pageId="2" pageNumber="43" type="nomenclature">
<paragraph id="8BCEE338FF97FFC941DF0313FDCEF8C4" blockId="2.[151,580,1800,1861]" box="[151,580,1800,1827]" pageId="2" pageNumber="43">
<heading id="D0865454FF97FFC941DF0313FDCEF8C4" bold="true" box="[151,580,1800,1827]" fontSize="11" level="1" pageId="2" pageNumber="43" reason="1">
<emphasis id="B9053F2AFF97FFC941DF0313FDCEF8C4" bold="true" box="[151,580,1800,1827]" pageId="2" pageNumber="43">
<taxonomicName id="4C7198BBFF97FFC941DF0313FE6AF8C5" ID-CoL="4QZ3L" box="[151,480,1801,1827]" class="Insecta" family="Palaeontinidae" genus="Quadraticossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="longicaulis" status="sp. nov.">
<emphasis id="B9053F2AFF97FFC941DF0313FE6AF8C5" bold="true" box="[151,480,1801,1827]" italics="true" pageId="2" pageNumber="43">Quadraticossus longicaulis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2368251FF97FFC940AE0312FDCEF8C4" box="[486,580,1800,1826]" pageId="2" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCEE338FF97FFC941DF0331FEACF8A3" blockId="2.[151,580,1800,1861]" box="[151,294,1835,1861]" pageId="2" pageNumber="43">
(
<figureCitation id="134AFFBDFF97FFC941D70331FE94F8A3" box="[159,286,1835,1861]" captionStart="FIGURE 2. A, B" captionStartId="5.[151,255,1406,1430]" captionTargetBox="[270,1329,184,1405]" captionTargetId="figure@5.[268,1327,785,1410]" captionTargetPageId="5" captionText="FIGURE 2. A, B, Quadraticossus longicaulis gen. &amp; sp. nov., holotype: A, photograph of part CNU­H­NN 2006094; B, line drawing." httpUri="https://zenodo.org/record/175238/files/figure.png" pageId="2" pageNumber="43">Fig. 2AB</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36BB0B3FF97FFC941DF0361FEDEF80D" pageId="2" pageNumber="43" type="diagnosis">
<paragraph id="8BCEE338FF97FFC941DF0361FEDEF80D" blockId="2.[151,1435,1915,2031]" pageId="2" pageNumber="43">
<emphasis id="B9053F2AFF97FFC941DF0361FE93F873" bold="true" box="[151,281,1915,1941]" pageId="2" pageNumber="43">Diagnosis:</emphasis>
Sc with weak branches, terminating on R1 near the middle of forewing. The stem of M is long and M bifurcated into M1+2 and M3+4 at the same level of Sc fusing with R1.Crossvein m­cua near point of Sc arising from Sc+R1.
</paragraph>
</subSubSection>
<caption id="DF0EB3B0FF96FFC841DF02FFFBE1F8F9" httpUri="https://zenodo.org/record/175236/files/figure.png" pageId="3" pageNumber="44" targetBox="[262,1323,182,1750]" targetPageId="3">
<paragraph id="8BCEE338FF96FFC841DF02FFFBE1F8F9" blockId="3.[151,1439,1765,1823]" pageId="3" pageNumber="44">
<emphasis id="B9053F2AFF96FFC841DF02FFFED9F91B" bold="true" box="[151,339,1765,1789]" pageId="3" pageNumber="44">FIGURE 1 AC.</emphasis>
<taxonomicName id="4C7198BBFF96FFC8401102FCFDB4F91B" box="[345,574,1766,1789]" class="Insecta" family="Palaeontinidae" genus="Quadraticossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="fangi">
<emphasis id="B9053F2AFF96FFC8401102FCFDB4F91B" box="[345,574,1766,1789]" italics="true" pageId="3" pageNumber="44">Quadraticossus fangi</emphasis>
</taxonomicName>
gen. &amp; sp. nov., holotype: A, photograph of part CNU­H­NN2006093­1; B, photograph of counterpart CNU­H­NN2006093­2; C, photograph of subcosta (Sc) on forewing.
</paragraph>
</caption>
<subSubSection id="C36BB0B3FF96FFCE418F0348FCB2F9E7" lastPageId="5" lastPageNumber="46" pageId="3" pageNumber="44" type="description">
<paragraph id="8BCEE338FF96FFCF418F0348FD62FEC9" blockId="3.[151,1438,1874,2031]" lastBlockId="4.[151,1437,152,303]" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
<emphasis id="B9053F2AFF96FFC8418F0348FEE8F88A" bold="true" box="[199,354,1874,1900]" pageId="3" pageNumber="44">Description:</emphasis>
Forewing triangular with distinct CP and nodal line. Sc with weak branches, separated from R at base, fused with R at the level of M bifurcation into M1+2 and M3+4, extending at indentation and terminating on R1. M branched into M1+2 and M3+4 little after point of R bifurcation into R1 and Rs. M1+2 and M3+4 separated at same level. Crossvein r­m near point of M1+2 branching into M1 and M2. CuA curved joined with M by a crossvein m­cua and also connected with M4 by long horizontal crossvein. Nodal line traceable as indentation across point of Sc arising from Sc+R1, along with R1 for a short distance, then across Rs to M1+2, along with latter to separating point of M, separating discal cells. Forewing with prominent color pattern consisting of hyaline patches on a dark ground. (
<figureCitation id="134AFFBDFF91FFCF431B050FFD50FEC9" box="[595,730,277,303]" captionStart="FIGURE 2. A, B" captionStartId="5.[151,255,1406,1430]" captionTargetBox="[270,1329,184,1405]" captionTargetId="figure@5.[268,1327,785,1410]" captionTargetPageId="5" captionText="FIGURE 2. A, B, Quadraticossus longicaulis gen. &amp; sp. nov., holotype: A, photograph of part CNU­H­NN 2006094; B, line drawing." httpUri="https://zenodo.org/record/175238/files/figure.png" pageId="4" pageNumber="45">Fig.2. A, B</figureCitation>
).
</paragraph>
<caption id="DF0EB3B0FF91FFCF41DF03D3FADFF806" box="[151,1365,1993,2017]" httpUri="https://zenodo.org/record/175237/files/figure.png" pageId="4" pageNumber="45" targetBox="[276,1301,360,1966]" targetPageId="4">
<paragraph id="8BCEE338FF91FFCF41DF03D3FADFF806" blockId="4.[151,1365,1993,2017]" box="[151,1365,1993,2017]" pageId="4" pageNumber="45">
<emphasis id="B9053F2AFF91FFCF41DF03D3FEC5F807" bold="true" box="[151,335,1993,2017]" pageId="4" pageNumber="45">FIGURE 1 DF.</emphasis>
<taxonomicName id="4C7198BBFF91FFCF401D03D3FDB0F806" box="[341,570,1993,2016]" class="Insecta" family="Palaeontinidae" genus="Quadraticossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="fangi">
<emphasis id="B9053F2AFF91FFCF401D03D3FDB0F806" box="[341,570,1993,2016]" italics="true" pageId="4" pageNumber="45">Quadraticossus fangi</emphasis>
</taxonomicName>
gen. &amp; sp. nov., holotype: D, body with wings; E, forewing; F, hind wing.
</paragraph>
</caption>
<caption id="DF0EB3B0FF90FFCE41DF0164FEAEFA5E" httpUri="https://zenodo.org/record/175238/files/figure.png" pageId="5" pageNumber="46" targetBox="[270,1329,184,1405]" targetPageId="5">
<paragraph id="8BCEE338FF90FFCE41DF0164FEAEFA5E" blockId="5.[151,1436,1406,1464]" pageId="5" pageNumber="46">
<emphasis id="B9053F2AFF90FFCE41DF0164FE93FA70" bold="true" box="[151,281,1406,1430]" pageId="5" pageNumber="46">FIGURE 2.</emphasis>
A, B,
<taxonomicName id="4C7198BBFF90FFCE40280165FD0EFA70" box="[352,644,1407,1430]" class="Insecta" family="Palaeontinidae" genus="Quadraticossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="longicaulis">
<emphasis id="B9053F2AFF90FFCE40280165FD0EFA70" box="[352,644,1407,1430]" italics="true" pageId="5" pageNumber="46">Quadraticossus longicaulis</emphasis>
</taxonomicName>
gen. &amp; sp. nov., holotype: A, photograph of part CNU­H­NN2006094; B, line drawing.
</paragraph>
</caption>
<paragraph id="8BCEE338FF90FFCE418F01FDFCB2F9E7" blockId="5.[151,1438,1511,2025]" box="[199,824,1511,1537]" pageId="5" pageNumber="46">
Forewing (preserved part)
<quantity id="4C894EDDFF90FFCE434801FDFDD9F9E7" box="[512,595,1511,1537]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="46" unit="mm" value="40.0">40 mm</quantity>
long,
<quantity id="4C894EDDFF90FFCE43D501FDFD7AF9E7" box="[669,752,1511,1537]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="46" unit="mm" value="19.0">19 mm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="C36BB0B3FF90FFCE418F0214FDA3F9B7" pageId="5" pageNumber="46" type="etymology">
<paragraph id="8BCEE338FF90FFCE418F0214FDA3F9B7" blockId="5.[151,1438,1511,2025]" pageId="5" pageNumber="46">
<emphasis id="B9053F2AFF90FFCE418F0214FED2F9CE" bold="true" box="[199,344,1550,1576]" pageId="5" pageNumber="46">Etymology:</emphasis>
Specific epithet is a combination of Latin prefix longi­ (long) and ­caulis (stem), referring to the long stem of forewing vein M.
</paragraph>
</subSubSection>
<subSubSection id="C36BB0B3FF90FFCE418F0244FD1CF917" pageId="5" pageNumber="46" type="materials_examined">
<paragraph id="8BCEE338FF90FFCE418F0244FDB6F947" blockId="5.[151,1438,1511,2025]" pageId="5" pageNumber="46">
<emphasis id="B9053F2AFF90FFCE418F0244FEB7F99E" bold="true" box="[199,317,1630,1656]" pageId="5" pageNumber="46">Material:</emphasis>
The new species is known from a single forewing.
<typeStatus id="54CA5D9AFF90FFCE42D20245FB86F99F" box="[922,1036,1631,1657]" pageId="5" pageNumber="46" type="holotype">Holotype</typeStatus>
: A single forewing, with anal area missing, No.: CNU­H­NN2006094.
</paragraph>
<paragraph id="8BCEE338FF90FFCE418F02B4FD1CF917" blockId="5.[151,1438,1511,2025]" pageId="5" pageNumber="46">
<emphasis id="B9053F2AFF90FFCE418F02B4FE5CF92E" bold="true" box="[199,470,1710,1736]" pageId="5" pageNumber="46">Horizon and locality:</emphasis>
Jiulongshan Formation, Middle Jurassic, Daohugou Village, Shantou Township, Ningcheng County, Inner
<collectingCountry id="F366A3A8FF90FFCE408F02CDFDB6F917" box="[455,572,1751,1777]" name="Mongolia" pageId="5" pageNumber="46">Mongolia</collectingCountry>
,
<collectingCountry id="F366A3A8FF90FFCE430002CDFD18F917" box="[584,658,1751,1777]" name="China" pageId="5" pageNumber="46">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36BB0B3FF90FFCE418F02E4FB9AF80F" pageId="5" pageNumber="46" type="discussion">
<paragraph id="8BCEE338FF90FFCE418F02E4FD46F870" blockId="5.[151,1438,1511,2025]" pageId="5" pageNumber="46">
<emphasis id="B9053F2AFF90FFCE418F02E4FEC9F8FE" bold="true" box="[199,323,1790,1816]" pageId="5" pageNumber="46">Remarks:</emphasis>
This species is similar to
<taxonomicName id="4C7198BBFF90FFCE433302E5FD56F8FE" box="[635,732,1791,1816]" class="Insecta" family="Palaeontinidae" genus="Quadraticossus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="fangi" status="sp. nov.">
<emphasis id="B9053F2AFF90FFCE433302E5FD56F8FE" box="[635,732,1791,1816]" italics="true" pageId="5" pageNumber="46">Q. fangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2368251FF90FFCE43AD02E5FCB6F8FF" box="[741,828,1791,1817]" pageId="5" pageNumber="46" rank="species">sp. nov</taxonomicNameLabel>
, but can be differentiated from the latter in the following features on the forewing: Sc with weak branches, terminating on R1 near the middle of wing; M bifurcated into M1+2 and M3+4 at the same level of Sc fusing with R1; crossvein r­m near the point of M1+2 branching into M1 and M2; m4­cua is shorter than the latter.
</paragraph>
<paragraph id="8BCEE338FF90FFCE418F03BDFB9AF80F" blockId="5.[151,1438,1511,2025]" pageId="5" pageNumber="46">This new species is erected based on a single incomplete forewing, but we anticipate discovery of more well­preserved specimens that will allow us to restudy this species in detail.</paragraph>
</subSubSection>
</treatment>
</document>