treatments-xml/data/03/B7/87/03B787EBFFD70514D4F80AC1FE09F844.xml

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<mods:title id="CC0E0EBA627EF619AB9082C2313595DC">A new species of the genus Raphidopus Stimpson, 1858 (Crustacea: Decapoda: Anomura: Porcellanidae) from Peninsular Malaysia, with additional records of R. johnsoni Ng &amp; Nakasone, 1994 from Southeast Asia, and a key to species in the genus</mods:title>
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<mods:namePart id="77B1BCE40AAB040C6D976FD0E6721611">Ng, Peter K. L.</mods:namePart>
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<paragraph id="8BA136FDFFD70511D4F80AC1FE36FF2A" blockId="8.[151,476,151,211]" box="[151,476,151,178]" pageId="8" pageNumber="119">
<heading id="D0E98191FFD70511D4F80AC1FE36FF2A" bold="true" box="[151,476,151,178]" fontSize="11" level="1" pageId="8" pageNumber="119" reason="1">
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<taxonomicName id="4C1E4D7EFFD70511D4F80AC1FE78FF29" ID-CoL="bffae44c-be3c-4af9-af99-3d6e0aa78da9" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[151,402,151,177]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD70511D4F80AC1FE78FF29" bold="true" box="[151,402,151,177]" italics="true" pageId="8" pageNumber="119">Raphidopus brevipes</emphasis>
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<paragraph id="8BA136FDFFD70511D4F80AECFE8AFF4B" blockId="8.[151,476,151,211]" box="[151,352,186,211]" pageId="8" pageNumber="119">
(
<figureCitation id="13252A78FFD70511D4F00AECFEE1FF4B" box="[159,267,186,211]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="8.[151,250,1655,1677]" captionStartId-1="10.[151,250,1684,1706]" captionStartId-2="11.[151,250,1500,1522]" captionTargetBox-0="[171,1421,664,1604]" captionTargetBox-1="[165,1420,199,1640]" captionTargetBox-2="[175,1409,208,1434]" captionTargetId-0="figure@8.[151,1436,654,1633]" captionTargetId-1="figure@10.[150,1436,193,1662]" captionTargetId-2="figure@11.[150,1436,193,1478]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." captionText-1="FIGURE 7. Raphidopus brevipes n. sp., paratypes, chelipeds detached from bodies, ZRC 1992.10522–10526, Penang. A, largest larger cheliped, dorsal vieW; B, largest larger cheliped, merUs and carpUs, ventral vieW (sUrface strUctUre not draWn); C, largest larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, dorsal vieW (sUrface strUctUre partially draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, second largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." captionText-2="FIGURE 8. Raphidopus brevipes, n. sp. Paratypes, ambUlatory legs detached from bodies, ZRC 1992.10522–10526, Penang (A–G); male (cl 4.1 mm), ZRC 2016.528, Selangor (H–K). A, left sUpposed second pereopod, lateral vieW; B, left sUpposed second pereopod, merUs, mesial vieW (setae not draWn); C, left sUpposed second pereopod, dactylUs, lateral vieW (setae not draWn); D, left sUpposed third pereopod, lateral vieW (setae not draWn); E, left sUpposed third pereopod, merUs, lateral perpendicUlar vieW; F, left sUpposed foUrth pereopod, lateral vieW (setae not draWn); G, left sUpposed foUrth pereopod, merUs, lateral perpendicUlar vieW; H–J, meri of left second to foUrth pereopods, lateral perpendicUlar vieW (setae not draWn); K, dactylUs of left foUrth pereopod, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri-0="https://zenodo.org/record/1287147/files/figure.png" httpUri-1="https://zenodo.org/record/1287149/files/figure.png" httpUri-2="https://zenodo.org/record/1287151/files/figure.png" pageId="8" pageNumber="119">Figs. 6–8</figureCitation>
, 
<figureCitation id="13252A78FFD70511D5790AEDFEBDFF4B" box="[278,343,187,211]" captionStart="FIGURE 9" captionStartId="12.[151,250,1662,1684]" captionTargetBox="[151,1436,193,1641]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURE 9. Preserved specimens of three Raphidopus species. Entire animal, dorsal vieW (A–D); carapace, dorsal vieW (E); larger and smaller chelipeds, dorsal vieW (F). A, Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246; B, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669; C, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, female (cl 5.9 mm), CBM-ZC 12485; D, Raphidopus brevipes n. sp., female (cl 5.1 mm), ZRC 2016.528; E, Raphidopus brevipes n. sp., male, holotype, male (cl 5.8 mm), ZRC 1992.10521; F, Raphidopus brevipes n. sp., paratypes, ZRC 1992.10522–10526." httpUri="https://zenodo.org/record/1287153/files/figure.png" pageId="8" pageNumber="119">9D–F</figureCitation>
)
</paragraph>
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<paragraph id="8BA136FDFFD70511D4F80B54FBFBFE80" blockId="8.[151,1041,258,280]" box="[151,1041,258,280]" pageId="8" pageNumber="119">
<treatmentCitationGroup id="AB0E11D3FFD70511D4F80B54FBFBFE80" box="[151,1041,258,280]" pageId="8" pageNumber="119">
<emphasis id="B96AEAEFFFD70511D4F80B54FE96FE80" box="[151,380,258,280]" italics="true" pageId="8" pageNumber="119">
<taxonomicName id="4C1E4D7EFFD70511D4F80B54FEB4FE80" ID-CoL="445049fb-994e-4886-ba7b-e2521935c222" authorityName="Henderson" authorityYear="1893" box="[151,350,258,280]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="indicus">Raphidopus indicus</taxonomicName>
.—
</emphasis>
Ng &amp; 
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<bibRefCitation id="EF8F4B0CFFD70511D5D40B54FDB1FE80" box="[443,603,258,280]" pageId="8" pageNumber="119" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone 1994</bibRefCitation>
: 7
</treatmentCitation>
, fig. 3 (not 
<taxonomicName id="4C1E4D7EFFD70511D6850B55FBEFFE80" ID-CoL="445049fb-994e-4886-ba7b-e2521935c222" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[746,1029,258,280]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD70511D6850B55FCA5FE80" box="[746,847,258,280]" italics="true" pageId="8" pageNumber="119">R. indicus</emphasis>
<bibRefCitation id="EF8F4B0CFFD70511D73A0B54FBEFFE80" box="[853,1029,258,280]" pageId="8" pageNumber="119" refId="ref6699" refString="Henderson, J. R. (1893) A contribUtion to Indian carcinology. Transaction of the Linnean Society, London, Zoology, Series 2, 5, 325 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article">Henderson, 1893</bibRefCitation>
</taxonomicName>
).
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</paragraph>
</subSubSection>
<subSubSection id="C3046576FFD70511D4F80B13FE31FDC1" pageId="8" pageNumber="119" type="materials_examined">
<paragraph id="8BA136FDFFD70511D4F80B13FCCCFE1A" blockId="8.[151,1437,325,602]" pageId="8" pageNumber="119">
<materialsCitation id="3B763CA0FFD70511D4F80B13FCCCFE1A" ID-GBIF-Occurrence="1848744100" collectingDate="1966-03-10" collectionCode="ZRC" collectorName="J. A. Bullock." country="Malaysia" location="Tanjong Tokong" pageId="8" pageNumber="119" specimenCode="ZRC 1992.10521" specimenCount="2" stateProvince="Penang" typeStatus="holotype">
<emphasis id="B96AEAEFFFD70511D4F80B13FEA1FEC6" bold="true" box="[151,331,325,350]" pageId="8" pageNumber="119">
<typeStatus id="54A5885FFFD70511D4F80B13FF38FEC6" box="[151,210,325,350]" pageId="8" pageNumber="119">Type</typeStatus>
material.
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<typeStatus id="54A5885FFFD70511D5370B13FE2FFEC6" box="[344,453,325,350]" pageId="8" pageNumber="119" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="DBB89E86FFD70511D5B90B10FD48FEC6" box="[470,674,325,350]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="119">ZRC 1992.10521</specimenCode>
, male (cl 
<quantity id="4CE69B18FFD70511D74D0B13FC68FEC6" box="[802,898,325,350]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="8" pageNumber="119" unit="mm" value="5.8">5.8 mm</quantity>
), 
<location id="8EC16026FFD70511D7F30B13FBB6FEC6" LSID="urn:lsid:plazi:treatment:03B787EBFFD70514D4F80AC1FE09F844:8EC16026FFD70511D7F30B13FBB6FEC6" box="[924,1116,325,350]" country="Malaysia" name="Tanjong Tokong" pageId="8" pageNumber="119" stateProvince="Penang">Tanjong Tokong</location>
, 
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, 
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, among terebellid worm tubes, 
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<date id="FFA0103DFFD70511D5F60B3FFDACFE1A" box="[409,582,361,386]" pageId="8" pageNumber="119" value="1966-03-10">10 March 1966</date>
</collectingDate>
, coll. 
<collectorName id="26EB532BFFD70511D6E50B3CFCCCFE1A" box="[650,806,361,386]" pageId="8" pageNumber="119">J. A. Bullock.</collectorName>
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</paragraph>
<paragraph id="8BA136FDFFD70511D4A80BD8FDA9FE52" blockId="8.[151,1437,325,602]" pageId="8" pageNumber="119">
Paratypes: ZRC 1992.10522–10526, 1 male (cl 
<quantity id="4CE69B18FFD70511D6890BDBFCABFE3E" box="[742,833,397,422]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="119" unit="mm" value="5.7">5.7 mm</quantity>
), 
<materialsCitation id="3B763CA0FFD70511D73A0BD8FDAAFE52" ID-GBIF-Occurrence="1848744177" collectingDate="1966-03-10" collectionCode="ZRC" collectorName="J. A. Bullock." country="Malaysia" location="Tanjong Tokong" pageId="8" pageNumber="119" specimenCount="2" specimenCount-female="2" stateProvince="Penang">
<specimenCount id="9D18FD74FFD70511D73A0BD8FC28FE3E" box="[853,962,397,422]" pageId="8" pageNumber="119" type="female">2 females</specimenCount>
(cl 6.1, 
<quantity id="4CE69B18FFD70511D04C0BDBFB94FE3E" box="[1059,1150,397,422]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="119" unit="mm" value="6.2">6.2 mm</quantity>
), 2 oVigerous females (cl 6.4, 
<quantity id="4CE69B18FFD70511D4A70BE7FEC8FE51" box="[200,290,433,458]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="119" unit="mm" value="6.8">6.8 mm</quantity>
), collected with holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BA136FDFFD70511D4A80B83FE31FDC1" blockId="8.[151,1437,325,602]" pageId="8" pageNumber="119">
<materialsCitation id="3B763CA0FFD70511D4A80B83FE31FDC1" ID-GBIF-Occurrence="1848744202" collectingDate="2016-07" collectingDateMax="2016-10-31" collectingDateMin="2016-07" collectionCode="ZRC" collectorName="J. J. Eeo." country="Malaysia" county="Other" latitude="3.2241666" location="Jeram Polychaete Reef" longLatPrecision="21" longitude="101.30361" municipality="Jeram" pageId="8" pageNumber="119" specimenCode="ZRC 2016.528" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Selangor">
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<collectingCounty id="62C04E71FFD70511D4A80B83FEE5FE76" box="[199,271,469,494]" pageId="8" pageNumber="119">Other</collectingCounty>
material examined.
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<collectingCountry id="F309766DFFD70511D66A0B83FD9AFE76" box="[517,624,469,494]" name="Malaysia" pageId="8" pageNumber="119">Malaysia</collectingCountry>
: 
<specimenCode id="DBB89E86FFD70511D6140B80FCCFFE76" box="[635,805,469,494]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="119">ZRC 2016.528</specimenCode>
, 
<specimenCount id="9D18FD74FFD70511D75E0B80FC62FE76" box="[817,904,469,494]" pageId="8" pageNumber="119" type="male">7 males</specimenCount>
(cl 4.1–5.0 mm), 
<specimenCount id="9D18FD74FFD70511D03A0B80FB2BFE76" box="[1109,1217,469,494]" pageId="8" pageNumber="119" type="female">3 females</specimenCount>
(cl 
<quantity id="4CE69B18FFD70511D0800B83FA90FE76" box="[1263,1402,469,494]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="8" pageNumber="119" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.1–5.1 mm</quantity>
), 2 oVigerous females (cl 4.4, 
<quantity id="4CE69B18FFD70511D58D0BAFFDAFFD89" box="[482,581,505,530]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="119" unit="mm" value="4.5">4.5 mm</quantity>
), 
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, Kuala 
<collectingRegion id="49DAF81FFFD70511D7630BAFFC9BFD8A" box="[780,881,505,530]" country="Malaysia" name="Selangor" pageId="8" pageNumber="119">Selangor</collectingRegion>
, 
<collectingRegion id="49DAF81FFFD70511D7EA0BAFFC00FD8A" box="[901,1002,505,530]" country="Malaysia" name="Selangor" pageId="8" pageNumber="119">Selangor</collectingRegion>
, 
<location id="8EC16026FFD70511D7910BACFAF9FD8A" LSID="urn:lsid:plazi:treatment:03B787EBFFD70514D4F80AC1FE09F844:8EC16026FFD70511D7910BACFAF9FD8A" box="[1022,1299,505,530]" country="Malaysia" county="Other" latitude="3.2241666" longLatPrecision="21" longitude="101.30361" municipality="Jeram" name="Jeram Polychaete Reef" pageId="8" pageNumber="119" stateProvince="Selangor">Jeram Polychaete Reef</location>
, station A, 
<geoCoordinate id="EE2A503AFFD70511D4F80848FEF2FDAE" box="[151,280,542,566]" direction="north" orientation="latitude" pageId="8" pageNumber="119" precision="15" value="3.2241666">3°13’27’’N</geoCoordinate>
, 
<geoCoordinate id="EE2A503AFFD70511D54A0848FE55FDAE" box="[293,447,542,566]" direction="east" orientation="longitude" pageId="8" pageNumber="119" precision="15" value="101.30361">101°18’13’’E</geoCoordinate>
, lower intertidal, mud, associated with tubes of 
<taxonomicName id="4C1E4D7EFFD70511D7B20848FAB9FDAE" authority="Caullery, 1944" authorityName="Caullery" authorityYear="1944" box="[989,1363,541,566]" class="Polychaeta" family="Terebellidae" genus="Loimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="8" pageNumber="119" phylum="Annelida" rank="species" species="verrucosa">
<emphasis id="B96AEAEFFFD70511D7B20848FB4EFDAD" box="[989,1188,542,565]" italics="true" pageId="8" pageNumber="119">Loimia verrucosa</emphasis>
<bibRefCitation id="EF8F4B0CFFD70511D0C4084BFAB9FDAE" box="[1195,1363,541,566]" pageId="8" pageNumber="119" refId="ref6502" refString="CaUllery, M. (1944) Polychetes Sedentaires de l'Expedition dU Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, OWeniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24, 2 bis, 1 - 204." type="journal article">Caullery, 1944</bibRefCitation>
</taxonomicName>
, 
<collectingDate id="EFE4E9D5FFD70511D130084BFED9FDC2" pageId="8" pageNumber="119" value="2016-07" valueMax="2016-10-31" valueMin="2016-07-00">
<date id="FFA0103DFFD70511D130084BFED9FDC2" pageId="8" pageNumber="119" value="2016-07" valueMax="2016-10-31" valueMin="2016-07-00">July– October 2016</date>
</collectingDate>
, coll. 
<collectorName id="26EB532BFFD70511D5190814FE31FDC1" box="[374,475,578,601]" pageId="8" pageNumber="119">J. J. Eeo.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF616675FFD70511D4F80C21FCBCF890" httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="8" pageNumber="119" startId="8.[151,250,1655,1677]" targetBox="[171,1421,664,1604]" targetPageId="8">
<paragraph id="8BA136FDFFD70511D4F80C21FCBCF890" blockId="8.[151,1436,1655,1800]" pageId="8" pageNumber="119">
<emphasis id="B96AEAEFFFD70511D4F80C21FEFEF914" bold="true" box="[151,276,1655,1677]" pageId="8" pageNumber="119">FIGURE 6.</emphasis>
<taxonomicName id="4C1E4D7EFFD70511D5730C21FE18F915" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[284,498,1655,1677]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD70511D5730C21FE18F915" box="[284,498,1655,1677]" italics="true" pageId="8" pageNumber="119">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD70511D5950C21FDDEF914" box="[506,564,1655,1676]" pageId="8" pageNumber="119" rank="species">
<emphasis id="B96AEAEFFFD70511D5950C21FDDEF914" bold="true" box="[506,564,1655,1676]" pageId="8" pageNumber="119">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C3046576FFD70514D4A80D0EFDADFDF8" lastPageId="13" lastPageNumber="124" pageId="8" pageNumber="119" type="description">
<paragraph id="8BA136FDFFD70510D4A80D0EFCE9FE84" blockId="8.[151,1436,1880,2013]" lastBlockId="9.[151,1437,151,1869]" lastPageId="9" lastPageNumber="120" pageId="8" pageNumber="119">
<emphasis id="B96AEAEFFFD70511D4A80D0EFEB3F8E9" bold="true" box="[199,345,1880,1905]" pageId="8" pageNumber="119">Description.</emphasis>
Carapace (
<figureCitation id="13252A78FFD70511D5B60D0EFDD8F8E9" box="[473,562,1880,1905]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="8" pageNumber="119">Fig. 6A</figureCitation>
) 1.3 times as broad as long; dorsal surface slightly conVex from side to side, with scattered, short or Very short striae; pair of protogastric ridges low, obtuse; cerVical grooVes moderately deep; anterior branchial region with small tubercles and short rows of beaded tubercles; posterior branchial and intestinal regions with long, tuberculate ridges; hepatic margin broadly conVex, granular; branchial margins strongly conVex, narrowly laminar, each with row of small, rounded tubercles; epigastric angle blunt, shallowly notched; lateral surfaces below branchial margins with dense, soft plumose setae. Rostrum (
<figureCitation id="13252A78FFD60510D7800AEAFBACFF4D" box="[1007,1094,188,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6B</figureCitation>
) bent Ventrally, trilobate, with all lobes rounded on anterior margins; median lobe reaching or slightly oVerreaching leVel of apices of lateral lobes; anterior margin with row of short plumose setae.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A80B71FD02FEFD" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Pterygostomial flap entire, with longitudinal ridges of small tubercles and dense, soft plumose setae; anterior margin terminating in small subacute apex (
<figureCitation id="13252A78FFD60510D6ED0B1AFD31FEFD" box="[642,731,332,357]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="8BA136FDFFD60510D4A80B39FB7AFE6D" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Third thoracic sternite (
<figureCitation id="13252A78FFD60510D58E0B39FDD5FE10" box="[481,575,367,392]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6D</figureCitation>
) 3.9–4.9 times broader than long measured along midline, with surface somewhat depressed medially; anterior margin trilobate, median lobe broad, low, lateral lobes strongly produced anteriorly and each with rounded apex. Fourth thoracic sternite with anterior margin concaVe, anteromedian part transVerse; surface depressed medially, with submarginal row of small tubercles anteriorly.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A80BA9FCE1FDA5" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Telson (
<figureCitation id="13252A78FFD60510D5490BA9FE95FD80" box="[294,383,511,536]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6E</figureCitation>
) distinctly broader than long, composed of 7 plates; with scattered, Very short striae on extensor surface; each posterior plate broader than long.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A8081EFE77FD1D" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Eyes (
<figureCitation id="13252A78FFD60510D57F0811FE65FDF8" box="[272,399,583,608]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6A, G</figureCitation>
) small, at most slightly exceeding lateral limits of orbits; dorsodistal projection of ocular peduncle low, rounded.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A808D9FE5FFC8D" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Basal article of antennular peduncle (
<figureCitation id="13252A78FFD60510D60208D9FD2BFD30" box="[621,705,655,680]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6F</figureCitation>
) approximately as long as broad; Ventral surface with transVerse row of small tubercles on lateromedian part, anterior part slightly concaVe and delimited by transVerse ridges of small tubercles; anterior margin obtuse, with row of small tubercles, but without distinct lobes or teeth; anterior mesiodistal margin blunt.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A80949FB9BFC3D" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Antennal peduncle (
<figureCitation id="13252A78FFD60510D5DD0949FDE5FCA0" box="[434,527,799,824]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6G</figureCitation>
) slender, weakly striate on surface. First article largest, as long as combined second to fourth articles, produced forward in lateral View, broadly in contact with lower orbital margin; surface slightly concaVe, with row of small tubercles along Ventral margin. Second article short, subrectangular. Third article oblong, about 1.5 times longer than second article. Fourth article short, suboVate.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A809F9FC58FB38" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Third maxilliped (
<figureCitation id="13252A78FFD60510D5FD09F9FE00FC50" box="[402,490,943,968]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="9" pageNumber="120">Fig. 6H</figureCitation>
) moderately stout, with short ridges on lateral surfaces of ischium to carpus. Ischium broad; distolateral margin strongly produced, narrowly rounded. Merus with rounded lobe proximal to median part of Ventral margin. Carpus with small subtriangular projection medially on inflated Ventral margin; dorsal margin slightly undulate. Propodus moderately stout, slightly tapering; lateral surface with row of short ridges along dorsal margin. Dactylus short, subtriangular. Merus to dactylus with long plumose setae on Ventral margin. Exopod broad, 0.7 width of ischium, reaching nearly to distal margin of merus, with distal flagellum; lateral surface with small, low tubercles and striae; dorsally margin strongly conVex on distal 0.8.
</paragraph>
<paragraph id="8BA136FDFFD60510D4A80EFAFD89F8D5" blockId="9.[151,1437,151,1869]" pageId="9" pageNumber="120">
Chelipeds (
<figureCitation id="13252A78FFD60510D5280EFAFE29FB5C" box="[327,451,1196,1221]" captionStart="FIGURE 7" captionStartId="10.[151,250,1684,1706]" captionTargetBox="[165,1420,199,1640]" captionTargetId="figure@10.[150,1436,193,1662]" captionTargetPageId="10" captionText="FIGURE 7. Raphidopus brevipes n. sp., paratypes, chelipeds detached from bodies, ZRC 1992.10522–10526, Penang. A, largest larger cheliped, dorsal vieW; B, largest larger cheliped, merUs and carpUs, ventral vieW (sUrface strUctUre not draWn); C, largest larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, dorsal vieW (sUrface strUctUre partially draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, second largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287149/files/figure.png" pageId="9" pageNumber="120">Fig. 7A–H</figureCitation>
) unequal in size; Ventral surface of merus to chela with scattered, numerous, short and Very short rows of granules; dense, soft plumose setae on anterior and posterior margins of merus to chela, dorsoanterior and Ventroposterior surfaces of chela, and cutting regions of fixed finger and dactylus. Merus with moderately narrow, rounded lobe on dorsal disto-anterior margin; dorsal surface with short and long, transVerse rows off small tubercles and with transVerse crest medially; Ventro-anterior margin conVex, tuberculate, with strong but short spine medially. Carpus 1.5–1.9 (larger cheliped) and 1.5–1.8 (smaller cheliped) times as long as broad; dorsal surface somewhat conVex, with short and moderately long rows of small tubercles entirely, and with low (sometimes indistinct), blunt ridge on longitudinal midline, shallow sulcus sometimes present posterior along median longitudinal ridge; dorso-anterior margin slightly concaVe medially, with row of small tubercles; dorsoposterior distal lobe broadly rounded or roundly truncate; posterior margin unarmed, rounded entirely (larger) and somewhat bluntly angled on distal part (smaller). Chela 2.3–2.4 (larger) and 2.5–2.6 (smaller) times longer than broad, with curVed claw; dorsal surface conVex, with numerous, coarse, small tubercles and with slightly eleVated ridge of small tubercles along posterior margin; anterior margin nearly straight on posterior half and fixed finger; cutting edge of fixed finger with row of moderately large, blunt teeth (larger, proximal and distal teeth usually largest) or with row of small, low teeth (smaller), proximal tooth prominent. Dactylus 0.4–0.5 (larger) and 0.5–0.6 (smaller) length of chela, opening at oblique angle (more acute angle in smaller cheliped), with curVed claw; dorsal surface with scattered, flattened, small tubercles and with slightly eleVated ridge of small tubercles along posterior margin; cutting edge with 2 distinct, blunt teeth on proximal part (larger) and with row of small, low teeth (smaller); claw distally curVed.
</paragraph>
<caption id="DF616675FFD50513D4F80CC2FC42F8FA" httpUri="https://zenodo.org/record/1287149/files/figure.png" pageId="10" pageNumber="121" startId="10.[151,250,1684,1706]" targetBox="[165,1420,199,1640]" targetPageId="10">
<paragraph id="8BA136FDFFD50513D4F80CC2FC42F8FA" blockId="10.[151,1436,1684,1890]" pageId="10" pageNumber="121">
<emphasis id="B96AEAEFFFD50513D4F80CC2FEFCF931" bold="true" box="[151,278,1684,1706]" pageId="10" pageNumber="121">FIGURE 7.</emphasis>
<taxonomicName id="4C1E4D7EFFD50513D54F0CC2FE12F932" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[288,504,1684,1706]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="121" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD50513D54F0CC2FE12F932" box="[288,504,1684,1706]" italics="true" pageId="10" pageNumber="121">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD50513D66E0CC2FDD4F931" box="[513,574,1684,1705]" pageId="10" pageNumber="121" rank="species">
<emphasis id="B96AEAEFFFD50513D66E0CC2FDD4F931" bold="true" box="[513,574,1684,1705]" pageId="10" pageNumber="121">n. sp.</emphasis>
</taxonomicNameLabel>
, paratypes, chelipeds detached from bodies, ZRC 1992.10522–10526, Penang. A, largest larger cheliped, dorsal vieW; B, largest larger cheliped, merUs and carpUs, ventral vieW (sUrface strUctUre not draWn); C, largest larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, dorsal vieW (sUrface strUctUre partially draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, second largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF616675FFD40512D4F80F8AFC1EF932" httpUri="https://zenodo.org/record/1287151/files/figure.png" pageId="11" pageNumber="122" startId="11.[151,250,1500,1522]" targetBox="[175,1409,208,1434]" targetPageId="11">
<paragraph id="8BA136FDFFD40512D4F80F8AFC1EF932" blockId="11.[151,1436,1500,1706]" pageId="11" pageNumber="122">
<emphasis id="B96AEAEFFFD40512D4F80F8AFEFEFA69" bold="true" box="[151,276,1500,1522]" pageId="11" pageNumber="122">FIGURE 8.</emphasis>
<taxonomicName id="4C1E4D7EFFD40512D5740F8AFE1BFA6A" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[283,497,1500,1522]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="122" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD40512D5740F8AFE1BFA6A" box="[283,497,1500,1522]" italics="true" pageId="11" pageNumber="122">Raphidopus brevipes</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2595794FFD40512D5900F8AFDD3FA69" box="[511,569,1500,1521]" pageId="11" pageNumber="122" rank="species">
<emphasis id="B96AEAEFFFD40512D5900F8AFDD3FA69" bold="true" box="[511,569,1500,1521]" pageId="11" pageNumber="122">n. sp.</emphasis>
</taxonomicNameLabel>
Paratypes, ambUlatory legs detached from bodies, ZRC 1992.10522–10526, Penang (A–G); male (cl 4.1 mm), ZRC 2016.528, Selangor (H–K). A, left sUpposed second pereopod, lateral vieW; B, left sUpposed second pereopod, merUs, mesial vieW (setae not draWn); C, left sUpposed second pereopod, dactylUs, lateral vieW (setae not draWn); D, left sUpposed third pereopod, lateral vieW (setae not draWn); E, left sUpposed third pereopod, merUs, lateral perpendicUlar vieW; F, left sUpposed foUrth pereopod, lateral vieW (setae not draWn); G, left sUpposed foUrth pereopod, merUs, lateral perpendicUlar vieW; H–J, meri of left second to foUrth pereopods, lateral perpendicUlar vieW (setae not draWn); K, dactylUs of left foUrth pereopod, lateral vieW (setae not draWn). Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF616675FFD30515D4F80C28FDEDF8A4" httpUri="https://zenodo.org/record/1287153/files/figure.png" pageId="12" pageNumber="123" startId="12.[151,250,1662,1684]" targetBox="[151,1436,193,1641]" targetPageId="12">
<paragraph id="8BA136FDFFD30515D4F80C28FDEDF8A4" blockId="12.[151,1436,1662,1852]" pageId="12" pageNumber="123">
<emphasis id="B96AEAEFFFD30515D4F80C28FEFEF90B" bold="true" box="[151,276,1662,1684]" pageId="12" pageNumber="123">FIGURE 9.</emphasis>
Preserved specimens of three 
<taxonomicName id="4C1E4D7EFFD30515D63F0C28FD23F90C" authorityName="Johnsoni Ng &amp; Nakasone" authorityYear="1994" box="[592,713,1662,1684]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFD30515D63F0C28FD23F90C" box="[592,713,1662,1684]" italics="true" pageId="12" pageNumber="123">Raphidopus</emphasis>
</taxonomicName>
species. Entire animal, dorsal vieW (A–D); carapace, dorsal vieW (E); larger and smaller chelipeds, dorsal vieW (F). A, 
<taxonomicName id="4C1E4D7EFFD30515D6F20CC9FBCFF92D" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[669,1061,1695,1717]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD30515D6F20CC9FCFCF92D" box="[669,790,1695,1717]" italics="true" pageId="12" pageNumber="123">Raphidopus</emphasis>
<emphasis id="B96AEAEFFFD30515D7700CC9FC82F92D" box="[799,872,1695,1717]" italics="true" pageId="12" pageNumber="123">indicus</emphasis>
Henderson, 1893
</taxonomicName>
, holotype, male (cl 6.5 mm), NHM 1892.7.15.246; B, 
<taxonomicName id="4C1E4D7EFFD30515D53B0C97FCF2F94F" authority="Johnsoni Ng &amp; Nakasone, 1994" authorityName="Johnsoni Ng &amp; Nakasone" authorityYear="1994" box="[340,792,1729,1751]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFD30515D53B0C97FDC1F94F" box="[340,555,1729,1751]" italics="true" pageId="12" pageNumber="123">Raphidopus Johnsoni</emphasis>
Ng &amp; Nakasone, 1994
</taxonomicName>
, holotype, male (cl 3.8 mm), ZRC 1992.10669; C, 
<taxonomicName id="4C1E4D7EFFD30515D14C0C97FE3DF960" authority="Johnsoni Ng &amp; Nakasone, 1994" authorityName="Johnsoni Ng &amp; Nakasone" authorityYear="1994" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFD30515D14C0C97FF04F960" italics="true" pageId="12" pageNumber="123">Raphidopus Johnsoni</emphasis>
Ng &amp; Nakasone, 1994
</taxonomicName>
, female (cl 5.9 mm), CBM-ZC 12485; D, 
<taxonomicName id="4C1E4D7EFFD30515D7100CB4FBB9F960" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[895,1107,1762,1784]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD30515D7100CB4FBB9F960" box="[895,1107,1762,1784]" italics="true" pageId="12" pageNumber="123">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD30515D0360CB4FB78F96F" box="[1113,1170,1762,1783]" pageId="12" pageNumber="123" rank="species">
<emphasis id="B96AEAEFFFD30515D0360CB4FB78F96F" bold="true" box="[1113,1170,1762,1783]" pageId="12" pageNumber="123">n. sp.</emphasis>
</taxonomicNameLabel>
, female (cl 5.1 mm), ZRC 2016.528; E, 
<taxonomicName id="4C1E4D7EFFD30515D54E0D55FE1CF881" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[289,502,1795,1817]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD30515D54E0D55FE1CF881" box="[289,502,1795,1817]" italics="true" pageId="12" pageNumber="123">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD30515D5920D52FDDDF881" box="[509,567,1796,1817]" pageId="12" pageNumber="123" rank="species">
<emphasis id="B96AEAEFFFD30515D5920D52FDDDF881" bold="true" box="[509,567,1796,1817]" pageId="12" pageNumber="123">n. sp.</emphasis>
</taxonomicNameLabel>
, male, holotype, male (cl 5.8 mm), ZRC 1992.10521; F, 
<taxonomicName id="4C1E4D7EFFD30515D0EF0D55FABFF881" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[1152,1365,1795,1817]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="123" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD30515D0EF0D55FABFF881" box="[1152,1365,1795,1817]" italics="true" pageId="12" pageNumber="123">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD30515D1330D52FA7CF881" box="[1372,1430,1796,1817]" pageId="12" pageNumber="123" rank="species">
<emphasis id="B96AEAEFFFD30515D1330D52FA7CF881" bold="true" box="[1372,1430,1796,1817]" pageId="12" pageNumber="123">n. sp.</emphasis>
</taxonomicNameLabel>
, paratypes, ZRC 1992.10522–10526.
</paragraph>
</caption>
<paragraph id="8BA136FDFFD30514D4A80DC7FAD6FE1F" blockId="12.[151,1436,1937,2034]" lastBlockId="13.[151,1437,151,2013]" lastPageId="13" lastPageNumber="124" pageId="12" pageNumber="123">
Ambulatory legs (
<figureCitation id="13252A78FFD30515D5FC0DC7FDEEF831" box="[403,516,1937,1962]" captionStart="FIGURE 8" captionStartId="11.[151,250,1500,1522]" captionTargetBox="[175,1409,208,1434]" captionTargetId="figure@11.[150,1436,193,1478]" captionTargetPageId="11" captionText="FIGURE 8. Raphidopus brevipes, n. sp. Paratypes, ambUlatory legs detached from bodies, ZRC 1992.10522–10526, Penang (A–G); male (cl 4.1 mm), ZRC 2016.528, Selangor (H–K). A, left sUpposed second pereopod, lateral vieW; B, left sUpposed second pereopod, merUs, mesial vieW (setae not draWn); C, left sUpposed second pereopod, dactylUs, lateral vieW (setae not draWn); D, left sUpposed third pereopod, lateral vieW (setae not draWn); E, left sUpposed third pereopod, merUs, lateral perpendicUlar vieW; F, left sUpposed foUrth pereopod, lateral vieW (setae not draWn); G, left sUpposed foUrth pereopod, merUs, lateral perpendicUlar vieW; H–J, meri of left second to foUrth pereopods, lateral perpendicUlar vieW (setae not draWn); K, dactylUs of left foUrth pereopod, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287151/files/figure.png" pageId="12" pageNumber="123">Fig. 8A–J</figureCitation>
) comparatiVely stout, somewhat flattened laterally, decreasing in size posteriorly, second pereopod longest; lateral and mesial surfaces with scattered, Very short, weak striae, small pits, and sparse, soft plumose setae, on fourth pereopod, setae most numerous; margins without spines or teeth; dense, soft plumose setae on anterior and posterior margins of ischium to dactylus. Meri 3.1–3.3, 2.8–3.0, 2.4–2.6 times longer than high on respectiVe second to fourth pereopods, somewhat narrowed distally in lateral View; lateral and mesial surfaces with sparse, short plumose setae; anterior and posterior margins somewhat conVex; posterolateral and posteromesial margins rounded; posterodistal margin rounded. Carpi relatiVely slender. Propodi 2.6–2.8, 2.4–2.6, 2.2–2.4 times longer than high on respectiVe second to fourth pereopods, somewhat narrowed distally in lateral View. Dactyli 0.6–0.8 length of propodi, each with short, distal claw; anterior and posterior margins nearly straight; lateral and mesial surfaces each with blunt, median, longitudinal ridge flanked by rows of short stiff setae.
</paragraph>
<paragraph id="8BA136FDFFD20514D4A80BC2FA7CFE34" blockId="13.[151,1437,151,2013]" box="[199,1430,404,429]" pageId="13" pageNumber="124">Fifth pereopod slender, chelate; propodus with numerous short simple setae and 1 or 2 subdistal hooked setae.</paragraph>
<paragraph id="8BA136FDFFD20514D4A80BE1FE75FE6D" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">Male with pair of gonopods on second abdominal segment; endopod elongate oVate, rounded on distal margin; exopod small, rounded.</paragraph>
<paragraph id="8BA136FDFFD20514D4A80BA9FBFDFD80" blockId="13.[151,1437,151,2013]" box="[199,1047,511,536]" pageId="13" pageNumber="124">Female with well deVeloped pleopods on third to fifth abdominal segments.</paragraph>
<paragraph id="8BA136FDFFD20514D4A80872FDADFDF8" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<emphasis id="B96AEAEFFFD20514D4A80872FEA2FDA5" bold="true" box="[199,328,548,573]" pageId="13" pageNumber="124">Coloration</emphasis>
. In freshly preserVed specimens (
<figureCitation id="13252A78FFD20514D6A90872FCCAFDA4" box="[710,800,548,573]" captionStart="FIGURE 9" captionStartId="12.[151,250,1662,1684]" captionTargetBox="[151,1436,193,1641]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURE 9. Preserved specimens of three Raphidopus species. Entire animal, dorsal vieW (A–D); carapace, dorsal vieW (E); larger and smaller chelipeds, dorsal vieW (F). A, Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246; B, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669; C, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, female (cl 5.9 mm), CBM-ZC 12485; D, Raphidopus brevipes n. sp., female (cl 5.1 mm), ZRC 2016.528; E, Raphidopus brevipes n. sp., male, holotype, male (cl 5.8 mm), ZRC 1992.10521; F, Raphidopus brevipes n. sp., paratypes, ZRC 1992.10522–10526." httpUri="https://zenodo.org/record/1287153/files/figure.png" pageId="13" pageNumber="124">Fig. 9D</figureCitation>
), the body and pereopods are generally pale brown, and the soft plumose setae are dark brown.
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFD20514D4A8083AFBA5FC18" pageId="13" pageNumber="124" type="distribution">
<paragraph id="8BA136FDFFD20514D4A8083AFE68FD30" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<emphasis id="B96AEAEFFFD20514D4A8083AFE8AFD1D" bold="true" box="[199,352,620,645]" pageId="13" pageNumber="124">Distribution.</emphasis>
At present, known only from the island of 
<collectingRegion id="49DAF81FFFD20514D712083BFC3AFD1D" box="[893,976,621,645]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
and Kuala 
<collectingRegion id="49DAF81FFFD20514D00C083AFB22FD1D" box="[1123,1224,620,645]" country="Malaysia" name="Selangor" pageId="13" pageNumber="124">Selangor</collectingRegion>
, western coast of Peninsular 
<collectingCountry id="F309766DFFD20514D57B08D9FE94FD30" box="[276,382,655,680]" name="Malaysia" pageId="13" pageNumber="124">Malaysia</collectingCountry>
.
</paragraph>
<paragraph id="8BA136FDFFD20514D4A808E2FBA5FC18" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<emphasis id="B96AEAEFFFD20514D4A808E2FEC0FD55" bold="true" box="[199,298,692,717]" pageId="13" pageNumber="124">Habitat.</emphasis>
The specimens from 
<collectingRegion id="49DAF81FFFD20514D64C08E3FD9CFD55" box="[547,630,693,717]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
were collected together with unidentified terebellids, but the nature of the association with the polychaetes was not specified on the label (cf. Ng &amp; 
<bibRefCitation id="EF8F4B0CFFD20514D0650881FB56FD68" box="[1034,1212,727,752]" pageId="13" pageNumber="124" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone 1994</bibRefCitation>
). The Jeram (Kuala 
<collectingRegion id="49DAF81FFFD20514D4F808AAFF16FC8D" box="[151,252,764,789]" country="Malaysia" name="Selangor" pageId="13" pageNumber="124">Selangor</collectingRegion>
) specimens were collected from the lower intertidal region of muddy shore and associated with tubes of the terebellid 
<taxonomicName id="4C1E4D7EFFD20514D55C0976FD47FCA0" authority="Caullery, 1944" authorityName="Caullery" authorityYear="1944" box="[307,685,799,824]" class="Polychaeta" family="Terebellidae" genus="Loimia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="13" pageNumber="124" phylum="Annelida" rank="species" species="verrucosa">
<emphasis id="B96AEAEFFFD20514D55C0976FE11FCA0" box="[307,507,800,824]" italics="true" pageId="13" pageNumber="124">Loimia verrucosa</emphasis>
<bibRefCitation id="EF8F4B0CFFD20514D66C0949FD47FCA0" box="[515,685,799,824]" pageId="13" pageNumber="124" refId="ref6502" refString="CaUllery, M. (1944) Polychetes Sedentaires de l'Expedition dU Siboga: Ariciidae, Spionidae, Chaetopteridae, Chlorhaemidae, Opheliidae, OWeniidae, Sabellariidae, Sternaspidae, Amphictenidae, Ampharetidae, Terebellidae. Siboga-Expeditie Uitkomsten op Zoologisch, Bonatisch, Oceanographisch en Geologisch gebied verzameld in Nederlandsch Oost-Indie 1899 - 1900, 24, 2 bis, 1 - 204." type="journal article">Caullery, 1944</bibRefCitation>
</taxonomicName>
. Polgar 
<emphasis id="B96AEAEFFFD20514D7660977FCA9FCA0" box="[777,835,800,824]" italics="true" pageId="13" pageNumber="124">et al.</emphasis>
(2015) described and illustrated the enVironments of the Jeram beach from 
<date id="FFA0103DFFD20514D5CD0912FDB3FCC4" box="[418,601,836,861]" pageId="13" pageNumber="124" value="2010-12">December 2010</date>
to 
<date id="FFA0103DFFD20514D6E90912FCEDFCC4" box="[646,775,836,861]" pageId="13" pageNumber="124" value="2013-04">April 2013</date>
, and reported that the bottom condition and community composition of polychaetes in the area underwent substantial changes oVer the years.
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFD20514D4A809DAFAE6FC50" pageId="13" pageNumber="124" type="etymology">
<paragraph id="8BA136FDFFD20514D4A809DAFAE6FC50" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<emphasis id="B96AEAEFFFD20514D4A809DAFEA4FC3D" bold="true" box="[199,334,908,933]" pageId="13" pageNumber="124">Etymology.</emphasis>
The specific name is deriVed from a combination of the Latin, 
<emphasis id="B96AEAEFFFD20514D05409DAFB94FC3C" box="[1083,1150,908,932]" italics="true" pageId="13" pageNumber="124">brevis</emphasis>
(short) and 
<emphasis id="B96AEAEFFFD20514D17F09DBFADFFC3C" box="[1296,1333,909,932]" italics="true" pageId="13" pageNumber="124">pes</emphasis>
(leg), in reference to the proportionally stout ambulatory legs of the new species; used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFD20514D4A80982FA65F89D" pageId="13" pageNumber="124" type="discussion">
<paragraph id="8BA136FDFFD20514D4A80982FC45FA95" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<emphasis id="B96AEAEFFFD20514D4A80982FED1FC75" bold="true" box="[199,315,980,1005]" pageId="13" pageNumber="124">Remarks.</emphasis>
As preViously mentioned, the 
<collectingRegion id="49DAF81FFFD20514D6F70983FD01FC75" box="[664,747,981,1005]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
specimens reported as 
<taxonomicName id="4C1E4D7EFFD20514D79A0983FB8CFC74" authorityName="Henderson" authorityYear="1893" box="[1013,1126,980,1004]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD20514D79A0983FB8CFC74" box="[1013,1126,980,1004]" italics="true" pageId="13" pageNumber="124">R. indicus</emphasis>
</taxonomicName>
by Ng &amp; 
<bibRefCitation id="EF8F4B0CFFD20514D0B40982FA76FC74" box="[1243,1436,980,1005]" pageId="13" pageNumber="124" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone (1994)</bibRefCitation>
are referred to the present new species. The largest male from 
<collectingRegion id="49DAF81FFFD20514D73B09AEFC4DFB88" box="[852,935,1016,1040]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
(cl 
<quantity id="4CE69B18FFD20514D7BC09A1FBC1FB97" box="[979,1067,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="124" unit="mm" value="5.8">5.8 mm</quantity>
, ZRC 1992.10521) is designated as the 
<typeStatus id="54A5885FFFD20514D48D0E4AFEAEFBAD" box="[226,324,1052,1077]" pageId="13" pageNumber="124" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C1E4D7EFFD20514D5030E4BFE01FBAC" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[364,491,1052,1076]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D5030E4BFE01FBAC" box="[364,491,1052,1076]" italics="true" pageId="13" pageNumber="124">R. brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D59B0E4AFDDEFBAD" box="[500,564,1052,1077]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D59B0E4AFDDEFBAD" bold="true" box="[500,564,1052,1077]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
, and preserVes only the body, cephalic appendages, and third maxillipeds as noted and illustrated by Ng &amp; 
<bibRefCitation id="EF8F4B0CFFD20514D5810E69FD40FBC0" box="[494,682,1087,1112]" pageId="13" pageNumber="124" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone (1994</bibRefCitation>
: 8, fig. 3A–D, J, K). The other 
<collectingRegion id="49DAF81FFFD20514D0630E16FBB4FBC0" box="[1036,1118,1088,1112]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
specimens including bodies of one male and four females with detached pereopods (ZRC 1992.10522–10526) are all designated as 
<typeStatus id="54A5885FFFD20514D17C0E33FA94FBE5" box="[1299,1406,1125,1149]" pageId="13" pageNumber="124" type="paratype">paratypes</typeStatus>
of the new species. The specimens recently collected from Jeram include intact indiViduals but are excluded from the 
<typeStatus id="54A5885FFFD20514D4F80EFBFF2DFB5D" box="[151,199,1197,1221]" pageId="13" pageNumber="124">type</typeStatus>
series because they were collected from the different locality and are much smaller than the 
<typeStatus id="54A5885FFFD20514D0A40EFBFAECFB5D" box="[1227,1286,1197,1221]" pageId="13" pageNumber="124">types</typeStatus>
from 
<collectingRegion id="49DAF81FFFD20514D1250EFBFA76FB5D" box="[1354,1436,1197,1221]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
(
<quantity id="4CE69B18FFD20514D4F00E99FEC3FB70" box="[159,297,1231,1256]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.1" metricValueMin="4.1" pageId="13" pageNumber="124" unit="mm" value="4.6" valueMax="5.1" valueMin="4.1">4.1–5.1 mm</quantity>
Versus 
<quantity id="4CE69B18FFD20514D5ED0E99FDE0FB70" box="[386,522,1231,1256]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.8" metricValueMin="5.7" pageId="13" pageNumber="124" unit="mm" value="6.25" valueMax="6.8" valueMin="5.7">5.7–6.8 mm</quantity>
in the carapace length). The Jeram and 
<collectingRegion id="49DAF81FFFD20514D7B80E87FBC0FB71" box="[983,1066,1233,1257]" country="Malaysia" name="Pulau Pinang" pageId="13" pageNumber="124">Penang</collectingRegion>
specimens agree well with each other in all diagnostic characters, and are here regarded as conspecific.
</paragraph>
<paragraph id="8BA136FDFFD20514D4A80F4EFA65F89D" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<taxonomicName id="4C1E4D7EFFD20514D4A80F4EFE5FFAA8" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[199,437,1304,1328]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D4A80F4EFE5FFAA8" box="[199,437,1304,1328]" italics="true" pageId="13" pageNumber="124">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D5AF0F41FDE8FAA8" box="[448,514,1303,1328]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D5AF0F41FDE8FAA8" bold="true" box="[448,514,1303,1328]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
is similar to 
<taxonomicName id="4C1E4D7EFFD20514D6CC0F4FFCFDFAA8" authorityName="Henderson" authorityYear="1893" box="[675,791,1304,1328]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD20514D6CC0F4FFCFDFAA8" box="[675,791,1304,1328]" italics="true" pageId="13" pageNumber="124">R. indicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1E4D7EFFD20514D73B0F4FFC3DFAA8" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[852,983,1304,1328]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD20514D73B0F4FFC3DFAA8" box="[852,983,1304,1328]" italics="true" pageId="13" pageNumber="124">R. johnsoni</emphasis>
</taxonomicName>
in the carapace branchial margins and posterior margins of the carpi of both chelipeds lacking distinct spines, but can be distinguished by the detailed morphology of the carapace, chelipeds, and ambulatory legs. The carapace is proportionally narrower in 
<taxonomicName id="4C1E4D7EFFD20514D1EB0F37FF1FFA04" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D1EB0F37FF1FFA04" italics="true" pageId="13" pageNumber="124">R. brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D4930FD2FED1FA05" box="[252,315,1412,1437]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D4930FD2FED1FA05" bold="true" box="[252,315,1412,1437]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
than the two known species; i.e., 1.3 Versus 1.4 times breadth of the length (
<figureCitation id="13252A78FFD20514D0F60FD2FB17FA04" box="[1177,1277,1412,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1860]" captionTargetBox="[165,1421,435,1783]" captionTargetId="figure@2.[150,1436,409,1817]" captionTargetPageId="2" captionText="FIGURE 1. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges on right side not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, telson, external vieW; E, basal article of left antennUlar pedUncle, ventral vieW; F, left ocUlar and antennal pedUncles, dorsolateral vieW; G, left third maxilliped, lateral vieW; H–J, right second to foUrth pereopods, lateral vieW. Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287137/files/figure.png" pageId="13" pageNumber="124">Figs. 1A</figureCitation>
, 
<figureCitation id="13252A78FFD20514D1660FD2FAC7FA04" box="[1289,1325,1412,1436]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="13" pageNumber="124">3A</figureCitation>
, 
<figureCitation id="13252A78FFD20514D1560FD2FAB4FA04" box="[1337,1374,1412,1436]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="13" pageNumber="124">4A</figureCitation>
, 
<figureCitation id="13252A78FFD20514D1050FD2FA65FA05" box="[1386,1423,1412,1437]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="13" pageNumber="124">6A</figureCitation>
). On each cheliped, the carpus usually has a low, blunt ridge on the dorsal longitudinal midline and the palm possesses small, closely-set tubercles on the dorsal surface in 
<taxonomicName id="4C1E4D7EFFD20514D7230F9BFC23FA7C" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[844,969,1484,1508]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D7230F9BFC23FA7C" box="[844,969,1484,1508]" italics="true" pageId="13" pageNumber="124">R. brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D7BF0F9AFBE5FA7D" box="[976,1039,1484,1509]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D7BF0F9AFBE5FA7D" bold="true" box="[976,1039,1484,1509]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13252A78FFD20514D0700F9AFB72FA7C" box="[1055,1176,1484,1509]" captionStart="FIGURE 7" captionStartId="10.[151,250,1684,1706]" captionTargetBox="[165,1420,199,1640]" captionTargetId="figure@10.[150,1436,193,1662]" captionTargetPageId="10" captionText="FIGURE 7. Raphidopus brevipes n. sp., paratypes, chelipeds detached from bodies, ZRC 1992.10522–10526, Penang. A, largest larger cheliped, dorsal vieW; B, largest larger cheliped, merUs and carpUs, ventral vieW (sUrface strUctUre not draWn); C, largest larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, dorsal vieW (sUrface strUctUre partially draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, second largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287149/files/figure.png" pageId="13" pageNumber="124">Fig. 7A, E</figureCitation>
), whereas in 
<taxonomicName id="4C1E4D7EFFD20514D1430F9BFA76FA7C" authorityName="Henderson" authorityYear="1893" box="[1324,1436,1484,1508]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD20514D1430F9BFA76FA7C" box="[1324,1436,1484,1508]" italics="true" pageId="13" pageNumber="124">R. indicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1E4D7EFFD20514D4A90FA7FEAFF990" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[198,325,1520,1544]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD20514D4A90FA7FEAFF990" box="[198,325,1520,1544]" italics="true" pageId="13" pageNumber="124">R. johnsoni</emphasis>
</taxonomicName>
, the dorsomedian ridge on the carpus is absent, and more coarsely distributed tubercles are present on the palm (
<figureCitation id="13252A78FFD20514D5440C42FE65F9B4" box="[299,399,1556,1581]" captionStart="FIGURE 2" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[166,1420,215,1289]" captionTargetId="figure@3.[150,1436,193,1295]" captionTargetPageId="3" captionText="FIGURE 2. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287139/files/figure.png" pageId="13" pageNumber="124">Figs. 2A</figureCitation>
, 
<figureCitation id="13252A78FFD20514D5F40C42FE2BF9B5" box="[411,449,1556,1581]" captionStart="FIGURE 5" captionStartId="7.[151,250,1377,1399]" captionTargetBox="[173,1420,218,1300]" captionTargetId="figure@7.[150,1436,193,1356]" captionTargetPageId="7" captionText="FIGURE 5. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 4.7 mm), CBM-ZC 12485, PhUket. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre on palm and dactylUs not draWn); D, smaller cheliped, dorsal vieW (sUrface strUctUre not draWn); E, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287145/files/figure.png" pageId="13" pageNumber="124">5A</figureCitation>
). The cutting edge of the dactylus of the larger cheliped possesses two large, blunt teeth on the proximal part in 
<taxonomicName id="4C1E4D7EFFD20514D5F30C6FFDF3F9C8" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[412,537,1592,1616]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D5F30C6FFDF3F9C8" box="[412,537,1592,1616]" italics="true" pageId="13" pageNumber="124">R. brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D6700C61FDB7F9C8" box="[543,605,1591,1616]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D6700C61FDB7F9C8" bold="true" box="[543,605,1591,1616]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13252A78FFD20514D6030C61FD0DF9C8" box="[620,743,1591,1616]" captionStart="FIGURE 7" captionStartId="10.[151,250,1684,1706]" captionTargetBox="[165,1420,199,1640]" captionTargetId="figure@10.[150,1436,193,1662]" captionTargetPageId="10" captionText="FIGURE 7. Raphidopus brevipes n. sp., paratypes, chelipeds detached from bodies, ZRC 1992.10522–10526, Penang. A, largest larger cheliped, dorsal vieW; B, largest larger cheliped, merUs and carpUs, ventral vieW (sUrface strUctUre not draWn); C, largest larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, dorsal vieW (sUrface strUctUre partially draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, second largest larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287149/files/figure.png" pageId="13" pageNumber="124">Fig. 7D, H</figureCitation>
), while it is armed with only one distinct, proximal tooth in 
<taxonomicName id="4C1E4D7EFFD20514D1EB0C6FFF02F9EC" authorityName="Henderson" authorityYear="1893" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD20514D1EB0C6FFF02F9EC" italics="true" pageId="13" pageNumber="124">R. indicus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1E4D7EFFD20514D54C0C0BFE4FF9EC" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[291,421,1628,1652]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD20514D54C0C0BFE4FF9EC" box="[291,421,1628,1652]" italics="true" pageId="13" pageNumber="124">R. johnsoni</emphasis>
</taxonomicName>
(
<figureCitation id="13252A78FFD20514D5DA0C0AFDF6F9EC" box="[437,540,1628,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[166,1420,215,1289]" captionTargetId="figure@3.[150,1436,193,1295]" captionTargetPageId="3" captionText="FIGURE 2. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287139/files/figure.png" pageId="13" pageNumber="124">Figs. 2D</figureCitation>
, 
<figureCitation id="13252A78FFD20514D6460C0AFDA4F9EC" box="[553,590,1628,1652]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="13" pageNumber="124">3H</figureCitation>
, 
<figureCitation id="13252A78FFD20514D6330C0AFD6AF9ED" box="[604,640,1628,1653]" captionStart="FIGURE 5" captionStartId="7.[151,250,1377,1399]" captionTargetBox="[173,1420,218,1300]" captionTargetId="figure@7.[150,1436,193,1356]" captionTargetPageId="7" captionText="FIGURE 5. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 4.7 mm), CBM-ZC 12485, PhUket. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre on palm and dactylUs not draWn); D, smaller cheliped, dorsal vieW (sUrface strUctUre not draWn); E, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287145/files/figure.png" pageId="13" pageNumber="124">5C</figureCitation>
). The ambulatory legs are proportionally stouter in 
<taxonomicName id="4C1E4D7EFFD20514D0BC0C0BFAB8F9EC" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[1235,1362,1628,1652]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D0BC0C0BFAB8F9EC" box="[1235,1362,1628,1652]" italics="true" pageId="13" pageNumber="124">R. brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D1340C0AFA76F9ED" box="[1371,1436,1628,1653]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D1340C0AFA76F9ED" bold="true" box="[1371,1436,1628,1653]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
than in the other two species; particular in the fourth pereopod: 2.4–2.6 Versus 2.8–3.1 times and 2.2–2.4 Versus 2.7–3.0 times longer than high on the merus and propodus, respectiVely (
<figureCitation id="13252A78FFD20514D7800CF2FBA6F924" box="[1007,1100,1700,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1860]" captionTargetBox="[165,1421,435,1783]" captionTargetId="figure@2.[150,1436,409,1817]" captionTargetPageId="2" captionText="FIGURE 1. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges on right side not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, telson, external vieW; E, basal article of left antennUlar pedUncle, ventral vieW; F, left ocUlar and antennal pedUncles, dorsolateral vieW; G, left third maxilliped, lateral vieW; H–J, right second to foUrth pereopods, lateral vieW. Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287137/files/figure.png" pageId="13" pageNumber="124">Figs. 1J</figureCitation>
, 
<figureCitation id="13252A78FFD20514D0330CF2FB49F924" box="[1116,1187,1700,1724]" captionStart="FIGURE 2" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[166,1420,215,1289]" captionTargetId="figure@3.[150,1436,193,1295]" captionTargetPageId="3" captionText="FIGURE 2. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287139/files/figure.png" pageId="13" pageNumber="124">2K, L</figureCitation>
, 
<figureCitation id="13252A78FFD20514D0DD0CF2FB16F924" box="[1202,1276,1700,1724]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="13" pageNumber="124">4N, O</figureCitation>
, 
<figureCitation id="13252A78FFD20514D1630CF2FA7AF924" box="[1292,1424,1700,1724]" captionStart="FIGURE 8" captionStartId="11.[151,250,1500,1522]" captionTargetBox="[175,1409,208,1434]" captionTargetId="figure@11.[150,1436,193,1478]" captionTargetPageId="11" captionText="FIGURE 8. Raphidopus brevipes, n. sp. Paratypes, ambUlatory legs detached from bodies, ZRC 1992.10522–10526, Penang (A–G); male (cl 4.1 mm), ZRC 2016.528, Selangor (H–K). A, left sUpposed second pereopod, lateral vieW; B, left sUpposed second pereopod, merUs, mesial vieW (setae not draWn); C, left sUpposed second pereopod, dactylUs, lateral vieW (setae not draWn); D, left sUpposed third pereopod, lateral vieW (setae not draWn); E, left sUpposed third pereopod, merUs, lateral perpendicUlar vieW; F, left sUpposed foUrth pereopod, lateral vieW (setae not draWn); G, left sUpposed foUrth pereopod, merUs, lateral perpendicUlar vieW; H–J, meri of left second to foUrth pereopods, lateral perpendicUlar vieW (setae not draWn); K, dactylUs of left foUrth pereopod, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287151/files/figure.png" pageId="13" pageNumber="124">8F, G, J, K</figureCitation>
). 
<taxonomicName id="4C1E4D7EFFD20514D4F80C9EFE68F978" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[151,386,1736,1760]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD20514D4F80C9EFE68F978" box="[151,386,1736,1760]" italics="true" pageId="13" pageNumber="124">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD20514D5E50C91FE22F978" box="[394,456,1735,1760]" pageId="13" pageNumber="124" rank="species">
<emphasis id="B96AEAEFFFD20514D5E50C91FE22F978" bold="true" box="[394,456,1735,1760]" pageId="13" pageNumber="124">n. sp.</emphasis>
</taxonomicNameLabel>
can be further separated from 
<taxonomicName id="4C1E4D7EFFD20514D74B0C9FFC4EF978" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[804,932,1736,1760]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD20514D74B0C9FFC4EF978" box="[804,932,1736,1760]" italics="true" pageId="13" pageNumber="124">R. johnsoni</emphasis>
</taxonomicName>
by its comparatiVely narrower third thoracic sternite, 3.9–4.9 Versus 5.3–7.0 times broader than the length measured along the midline (
<figureCitation id="13252A78FFD20514D0E00CBAFB1AF89C" box="[1167,1264,1772,1797]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1860]" captionTargetBox="[165,1421,435,1783]" captionTargetId="figure@2.[150,1436,409,1817]" captionTargetPageId="2" captionText="FIGURE 1. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges on right side not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, telson, external vieW; E, basal article of left antennUlar pedUncle, ventral vieW; F, left ocUlar and antennal pedUncles, dorsolateral vieW; G, left third maxilliped, lateral vieW; H–J, right second to foUrth pereopods, lateral vieW. Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287137/files/figure.png" pageId="13" pageNumber="124">Figs. 1C</figureCitation>
, 
<figureCitation id="13252A78FFD20514D0920CBAFACAF89C" box="[1277,1312,1772,1796]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="13" pageNumber="124">3C</figureCitation>
, 
<figureCitation id="13252A78FFD20514D1430CBAFABBF89C" box="[1324,1361,1772,1796]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="13" pageNumber="124">4D</figureCitation>
, 
<figureCitation id="13252A78FFD20514D1330CBAFA68F89D" box="[1372,1410,1772,1797]" captionStart="FIGURE 6" captionStartId="8.[151,250,1655,1677]" captionTargetBox="[171,1421,664,1604]" captionTargetId="figure@8.[151,1436,654,1633]" captionTargetPageId="8" captionText="FIGURE 6. Raphidopus brevipes n. sp., holotype, male (cl 5.8 mm), ZRC 1992.10521, Penang. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (setae not draWn); B, rostrUm, dorso-anterior vieW; C, anterior part of left pterygostomial flap, ventrolateral vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW (setae not draWn); H, left third maxilliped, lateral vieW (setae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287147/files/figure.png" pageId="13" pageNumber="124">6D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFD20514D4A80D46FE09F844" pageId="13" pageNumber="124" type="description">
<paragraph id="8BA136FDFFD20514D4A80D46FE09F844" blockId="13.[151,1437,151,2013]" pageId="13" pageNumber="124">
<taxonomicName id="4C1E4D7EFFD20514D4A80D46FE5EF8B0" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[199,436,1808,1832]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD20514D4A80D46FE5EF8B0" box="[199,436,1808,1832]" italics="true" pageId="13" pageNumber="124">Raphidopus johnsoni</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C1E4D7EFFD20514D6250D47FD51F8B0" authorityName="Henderson" authorityYear="1893" box="[586,699,1808,1832]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD20514D6250D47FD51F8B0" box="[586,699,1808,1832]" italics="true" pageId="13" pageNumber="124">R. indicus</emphasis>
</taxonomicName>
in haVing the relatiVely larger eyes reaching at least to the lateral limits of the orbits (
<figureCitation id="13252A78FFD20514D51C0D62FE3DF8D4" box="[371,471,1844,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="13" pageNumber="124">Figs. 3A</figureCitation>
, 
<figureCitation id="13252A78FFD20514D58C0D62FDE3F8D4" box="[483,521,1844,1868]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="13" pageNumber="124">4A</figureCitation>
), the third thoracic sternite being proportionally broader (5.3–7.0 Versus 4.6 times broader than the length measured along the midline; 
<figureCitation id="13252A78FFD20514D6910D01FC8BF8E8" box="[766,865,1879,1904]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1860]" captionTargetBox="[165,1421,435,1783]" captionTargetId="figure@2.[150,1436,409,1817]" captionTargetPageId="2" captionText="FIGURE 1. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges on right side not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, telson, external vieW; E, basal article of left antennUlar pedUncle, ventral vieW; F, left ocUlar and antennal pedUncles, dorsolateral vieW; G, left third maxilliped, lateral vieW; H–J, right second to foUrth pereopods, lateral vieW. Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287137/files/figure.png" pageId="13" pageNumber="124">Figs. 1C</figureCitation>
, 
<figureCitation id="13252A78FFD20514D71E0D0EFC79F8E8" box="[881,915,1880,1904]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="13" pageNumber="124">3C</figureCitation>
, 
<figureCitation id="13252A78FFD20514D7CD0D0EFC22F8E8" box="[930,968,1880,1904]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="13" pageNumber="124">4D</figureCitation>
), and the dactylus of the larger cheliped possessing a distinct blunt tooth on the median part of the cutting edge (
<figureCitation id="13252A78FFD20514D7DA0D2AFBF2F80C" box="[949,1048,1916,1941]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="13" pageNumber="124">Figs. 3H</figureCitation>
, 
<figureCitation id="13252A78FFD20514D04C0D2AFBA2F80D" box="[1059,1096,1916,1941]" captionStart="FIGURE 5" captionStartId="7.[151,250,1377,1399]" captionTargetBox="[173,1420,218,1300]" captionTargetId="figure@7.[150,1436,193,1356]" captionTargetPageId="7" captionText="FIGURE 5. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 4.7 mm), CBM-ZC 12485, PhUket. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre on palm and dactylUs not draWn); D, smaller cheliped, dorsal vieW (sUrface strUctUre not draWn); E, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287145/files/figure.png" pageId="13" pageNumber="124">5C</figureCitation>
). In 
<taxonomicName id="4C1E4D7EFFD20514D0170D2BFB02F80C" authorityName="Henderson" authorityYear="1893" box="[1144,1256,1916,1940]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="124" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD20514D0170D2BFB02F80C" box="[1144,1256,1916,1940]" italics="true" pageId="13" pageNumber="124">R. indicus</emphasis>
</taxonomicName>
, the eyes do not reach the lateral limits of the orbits (
<figureCitation id="13252A78FFD20514D6430DF6FD6BF820" box="[556,641,1952,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1860]" captionTargetBox="[165,1421,435,1783]" captionTargetId="figure@2.[150,1436,409,1817]" captionTargetPageId="2" captionText="FIGURE 1. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges on right side not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, telson, external vieW; E, basal article of left antennUlar pedUncle, ventral vieW; F, left ocUlar and antennal pedUncles, dorsolateral vieW; G, left third maxilliped, lateral vieW; H–J, right second to foUrth pereopods, lateral vieW. Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287137/files/figure.png" pageId="13" pageNumber="124">Fig. 1A</figureCitation>
) and the dactylus of the larger cheliped has no prominent median tooth on the cutting edge (
<figureCitation id="13252A78FFD20514D5130D92FE3DF844" box="[380,471,1988,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1316,1338]" captionTargetBox="[166,1420,215,1289]" captionTargetId="figure@3.[150,1436,193,1295]" captionTargetPageId="3" captionText="FIGURE 2. Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246, Madras. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); D, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); E, smaller cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); F, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); G, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287139/files/figure.png" pageId="13" pageNumber="124">Fig. 2D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>