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<document id="926A2C7420EF849B8A79CAFADDE97BE3" ID-CLB-Dataset="45355" ID-DOI="10.11646/zootaxa.3826.1.6" ID-GBIF-Dataset="c83f335b-ffc1-4cc7-af45-083caed1da51" ID-ISSN="1175-5326" ID-Zenodo-Dep="251844" ID-ZooBank="4C7900FD-656C-4180-80DA-449C310CD2B8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460445643758" checkinUser="plazi" docAuthor="Lawrence, John F., Escalona, Hermes E., Leschen, Richard A. B. &amp; Ślipiński, Adam" docDate="2014" docId="03B787B1FFF7FFD1ACC5FABEFA10BFE6" docLanguage="en" docName="zt03826p229.pdf" docOrigin="Zootaxa 3826 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mycetophagidae" docType="key" docVersion="9" lastPageNumber="211" masterDocId="FF8EFFC9FFF9FFC1AC52FFDEFF8CB91C" masterDocTitle="Review of the genera of Mycetophagidae (Coleoptera: Tenebrionoidea) with descriptions of new genera and a world generic key" masterLastPageNumber="229" masterPageNumber="195" pageNumber="209" updateTime="1698333336532" updateUser="ExternalLinkService">
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<mods:title id="AE911CB41B20C6623731A8F3D3859462">Review of the genera of Mycetophagidae (Coleoptera: Tenebrionoidea) with descriptions of new genera and a world generic key</mods:title>
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<mods:namePart id="A2F69D33112E0CF9764FB15C841A749E">Lawrence, John F.</mods:namePart>
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<mods:namePart id="D5A0BBDCE176E56EE041D984753CBD32">Escalona, Hermes E.</mods:namePart>
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<mods:namePart id="483C182F208B2718C24FCB02CE8F044A">Leschen, Richard A. B.</mods:namePart>
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<mods:namePart id="579B2080D206319D974E8ADC66C02EF1">Ślipiński, Adam</mods:namePart>
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<mods:title id="6B129743D86DCF8D1299C1F808139BD9">Zootaxa</mods:title>
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<mods:date id="5E2B1BABC8B55BFA57E1A34CF435455C">2014</mods:date>
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<mods:identifier id="E13A3C52D20739F9AB915E0A40D83925" type="CLB-Dataset">45355</mods:identifier>
<mods:identifier id="9348E4B1AC841ECB2E41F3545A7AC17B" type="DOI">10.11646/zootaxa.3826.1.6</mods:identifier>
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<treatment id="03B787B1FFF7FFD1ACC5FABEFA10BFE6" ID-DOI="http://doi.org/10.5281/zenodo.6141930" ID-GBIF-Taxon="119509367" ID-Zenodo-Dep="6141930" LSID="urn:lsid:plazi:treatment:03B787B1FFF7FFD1ACC5FABEFA10BFE6" httpUri="http://treatment.plazi.org/id/03B787B1FFF7FFD1ACC5FABEFA10BFE6" lastPageId="16" lastPageNumber="211" pageId="14" pageNumber="209">
<subSubSection id="C304652CFFF7FFCFACC5FABEFBA0BCFF" pageId="14" pageNumber="209" type="nomenclature">
<paragraph id="8BA136A7FFF7FFCFACC5FABEFD35BC66" blockId="14.[151,697,1376,1402]" box="[151,697,1376,1402]" pageId="14" pageNumber="209">
<heading id="D0E981CBFFF7FFCFACC5FABEFD35BC66" bold="true" box="[151,697,1376,1402]" fontSize="11" level="1" pageId="14" pageNumber="209" reason="1">
<emphasis id="B96AEAB5FFF7FFCFACC5FABEFD35BC66" bold="true" box="[151,697,1376,1402]" pageId="14" pageNumber="209">
Key to the world genera of
<taxonomicName id="4C1E4D24FFF7FFCFADB9FABEFD35BC66" ID-CoL="D25" ID-ENA="219427" box="[491,697,1376,1402]" class="Insecta" family="Mycetophagidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="family">Mycetophagidae</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA136A7FFF7FFCFACC5FA78FBA0BCFF" blockId="14.[151,1436,1446,1507]" pageId="14" pageNumber="209">
This key is based primarily on dissected specimens but characteristics of the genera
<taxonomicName id="4C1E4D24FFF7FFCFA803FA78FF5CBCFF" authority="Nikitsky" authorityName="Nikitsky" authorityYear="1989" class="Insecta" family="Mycetophagidae" genus="Triphyllopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF7FFCFA803FA78FB6EBCA3" box="[1105,1250,1446,1471]" italics="true" pageId="14" pageNumber="209">Triphyllopsis</emphasis>
<bibRefCitation id="EF8F4B56FFF7FFCFA8B9FA78FA14BCA3" author="Nikitsky" box="[1259,1432,1446,1471]" pageId="14" pageNumber="209" refString="Nikitsky, N. B. (1989) New taxa of beetles from the families Mycetophagidae and Tetratomidae of the USSR. fauna. Zoologicheskii Zhurnal, 68 (11), 56 - 64." type="journal article" year="1989">Nikitsky (1989</bibRefCitation>
,
<bibRefCitation id="EF8F4B56FFF7FFCFACC5FA15FF5CBCFF" author="Nikitsky" box="[151,208,1483,1507]" pageId="14" pageNumber="209" refString="Nikitsky, N. B. (1993) Mushroom beetles (Coleoptera, Mycetophagidae) of the Fauna of Russia and Contiguous Land. Izdatel'stvo Moskovskogo Universiteta, Moscow 184 pp. [in Russian]" type="book" year="1993">1993</bibRefCitation>
</taxonomicName>
) and
<taxonomicName id="4C1E4D24FFF7FFCFAD5CFA14FD2ABCFF" authority="Nikitsky (1996)" authorityName="Nikitsky" authorityYear="1996" box="[270,678,1482,1507]" class="Insecta" family="Mycetophagidae" genus="Pseudotriphyllopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF7FFCFAD5CFA14FE60BCFF" box="[270,492,1482,1507]" italics="true" pageId="14" pageNumber="209">Pseudotriphyllopsis</emphasis>
<bibRefCitation id="EF8F4B56FFF7FFCFADA6FA14FD2ABCFF" author="Nikitsky" box="[500,678,1482,1507]" pageId="14" pageNumber="209" refString="Nikitsky, N. B. (1996) New Coleoptera from China. Zoologicheskii Zhurnal, 75, 1366 - 1373. [in Russian; translated in Entomological Review, 76, 1231 - 1237]" type="journal article" year="1996">Nikitsky (1996)</bibRefCitation>
</taxonomicName>
were extracted from the literature.
</paragraph>
</subSubSection>
<subSubSection id="C304652CFFF7FFD1ACC5F9CCFA10BFE6" lastPageId="16" lastPageNumber="211" pageId="14" pageNumber="209" type="key">
<keyStep id="30EA2E02FFF7FFCFACC5F9CCFA17BE0E" pageId="14" pageNumber="209">
<paragraph id="8BA136A7FFF7FFCFACC5F9CCFA10BF80" blockId="14.[151,1437,1554,2015]" pageId="14" pageNumber="209">
<keyLead id="30EF9592FFF7FFCFACC5F9CCFA10BF80" pageId="14" pageNumber="209">
1. Head (
<figureCitation id="13252A22FFF7FFCFAD72F9CCFE1BBF3B" box="[288,407,1554,1575]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1C, 1K</figureCitation>
) only slightly narrower than greatest pronotal width, more or less abruptly constricted behind eyes to form broad neck and short to moderately long temples (
<figureCitation id="13252A22FFF7FFCFAE89F9F1FCC7BF58" box="[731,843,1583,1604]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">Figs 7A–B</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFAF05F9F1FC1FBF58" box="[855,915,1583,1604]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">8A–B</figureCitation>
); pronotum more or less quadrate with very weakly defined lateral carinae, distinctly narrower than combined elytral bases (
<figureCitation id="13252A22FFF7FFCFAFC3F993FC6CBF7E" box="[913,992,1613,1634]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">Figs 7D</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFAFB8F993FB85BF7E" box="[1002,1033,1613,1634]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">8D</figureCitation>
); male with pair of slender, oblique setose impressions between procoxae and anterior pronotal angles (
<figureCitation id="13252A22FFF7FFCFAF74F9B4FCF8BF63" box="[806,884,1642,1663]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">Figs 7E</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFAF2DF9B4FC11BF63" box="[895,925,1642,1663]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">8E</figureCitation>
); antennal club 2-segmented (
<figureCitation id="13252A22FFF7FFCFA890F9B4FA9EBF63" box="[1218,1298,1642,1663]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1K</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFA94CF9B4FAB7BF63" box="[1310,1339,1642,1663]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">7C</figureCitation>
) OR tarsi 3-3-3 (
<figureCitation id="13252A22FFF7FFCFAD70F959FEE2BF80" box="[290,366,1671,1692]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">Fig. 8C</figureCitation>
); length less than
<quantity id="4CE69B42FFF7FFCFAE48F959FDD8BF80" box="[538,596,1671,1692]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="209" unit="mm" value="2.0">2 mm</quantity>
......................................................... ...
<taxonomicName id="4C1E4D24FFF7FFCFA940F959FA0DBF80" box="[1298,1409,1671,1692]" class="Insecta" family="Mycetophagidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="subFamily" subFamily="Bergininae">Bergininae</taxonomicName>
...2
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFF7FFCFACC5F97BFA17BE0E" blockId="14.[151,1437,1554,2015]" pageId="14" pageNumber="209">
<keyLead id="30EF9592FFF7FFCFACC5F97BFA17BE0E" pageId="14" pageNumber="209">
- Head (
<figureCitation id="13252A22FFF7FFCFAD73F97BFE41BFA6" box="[289,461,1701,1722]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1A–B, 1D–J</figureCitation>
,
<date id="FFA01067FFF7FFCFAD8AF97BFE60BFA6" box="[472,492,1701,1722]" pageId="14" pageNumber="209">
<figureCitation id="13252A22FFF7FFCFAD8AF97BFE6FBFA6" box="[472,483,1701,1722]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="14" pageNumber="209">5</figureCitation>
I
</date>
–J, 5M–O) distinctly narrower than greatest pronotal width, not abruptly constricted behind eyes; pronotum (
<figureCitation id="13252A22FFF7FFCFAD1EF91CFE74BFCB" box="[332,504,1730,1751]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1A–B, 1D–J</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFAE51F91CFD82BFCB" box="[515,526,1730,1751]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="14" pageNumber="209">5</figureCitation>
M–N) with more distinctly rounded sides and distinct lateral carinae, not or only slightly narrower than combined elytral bases; male without paired ventral prothoracic impressions; antennal club with at least 3 segments (
<figureCitation id="13252A22FFF7FFCFACB5F923FE18BE0E" box="[231,404,1789,1810]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1A–B, 1E–F</figureCitation>
); tarsi 4-4-
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female, 3-4-
<quantity id="4CE69B42FFF7FFCFAEC9F923FD4EBE0E" box="[667,706,1789,1810]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="14" pageNumber="209" unit="in" value="4.0">4 in</quantity>
male....................................................... 3
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<paragraph id="8BA136A7FFF7FFCFACC5F8C4FA17BE76" blockId="14.[151,1437,1554,2015]" pageId="14" pageNumber="209">
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2(1) Temples shorter than eye or neck (
<figureCitation id="13252A22FFF7FFCFAE7AF8C4FDFFBE33" box="[552,627,1818,1839]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">Fig. 7B</figureCitation>
); antennal club 2-segmented (
<figureCitation id="13252A22FFF7FFCFAFC3F8C4FC6CBE33" box="[913,992,1818,1839]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1K</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFAFB8F8C4FB85BE33" box="[1002,1033,1818,1839]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">7C</figureCitation>
); tarsi 4-4-
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female, 3-4-
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male; dorsal surfaces clothed with distinct, thickened setae, usually more or less decumbent (
<figureCitation id="13252A22FFF7FFCFA85DF8E9FBECBE50" box="[1039,1120,1847,1868]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1K</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFA839F8E9FB00BE50" box="[1131,1164,1847,1868]" captionStart="FIGURES 7 A – H" captionStartId="23.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@23.[151,1436,193,1781]" captionTargetPageId="23" captionText="FIGURES 7 A – H, Berginus sp. (Panama): A, head, dorsal; B, head, ventral; C, antenna; D, prothorax, dorsal; E, prothorax, ventral; F, mesotibia and mesotarsus; G, elytron; H, portion of mesothorax and metathorax, showing lateral closure of mesocoxal cavity. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251851/files/figure.png" pageId="14" pageNumber="209">7G</figureCitation>
); length more than
<quantity id="4CE69B42FFF7FFCFA918F8E9FA14BE50" box="[1354,1432,1847,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="209" unit="mm" value="1.5">1.5 mm</quantity>
; North and Central
<collectingCountry id="F3097637FFF7FFCFADC6F88BFE65BE76" box="[404,489,1877,1898]" name="United States of America" pageId="14" pageNumber="209">America</collectingCountry>
, West Indies, southern Europe, Caucasus,
<collectingCountry id="F3097637FFF7FFCFAF2CF88BFC3EBE76" box="[894,946,1877,1898]" name="India" pageId="14" pageNumber="209">India</collectingCountry>
,
<collectingCountry id="F3097637FFF7FFCFAFEEF88BFB97BE76" box="[956,1051,1877,1898]" name="Sri Lanka" pageId="14" pageNumber="209">Sri Lanka</collectingCountry>
, East Indies.......
<taxonomicName id="4C1E4D24FFF7FFCFA8B9F88BFA17BE76" ID-CoL="39NR" authority="Erichson" authorityName="Erichson" box="[1259,1435,1877,1898]" class="Insecta" family="Mycetophagidae" genus="Berginus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF7FFCFA8B9F88BFACCBE76" box="[1259,1344,1877,1898]" italics="true" pageId="14" pageNumber="209">Berginus</emphasis>
Erichson
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</paragraph>
<paragraph id="8BA136A7FFF7FFCFACC5F8ACFA01BEDE" blockId="14.[151,1437,1554,2015]" pageId="14" pageNumber="209">
<keyLead id="30EF9592FFF7FFCFACC5F8ACFA01BEDE" pageId="14" pageNumber="209">
- Temples much longer than eye and about as long as neck (
<figureCitation id="13252A22FFF7FFCFAF4FF8ACFCE0BE9B" box="[797,876,1906,1927]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="14" pageNumber="209">Figs 1C</figureCitation>
,
<figureCitation id="13252A22FFF7FFCFAF2AF8ACFC1ABE9B" box="[888,918,1906,1927]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">8B</figureCitation>
); antennal club 3-segmented (
<figureCitation id="13252A22FFF7FFCFA8EEF8ACFA85BE9B" box="[1212,1289,1906,1927]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">Fig. 8H</figureCitation>
); tarsi 3-3-
<quantity id="4CE69B42FFF7FFCFA927F8ACFA10BE9B" box="[1397,1436,1906,1927]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="209" unit="in" value="3.0">3 in</quantity>
both sexes; dorsal surfaces clothed with minute, fine setae barely visible at lower magnifications (
<figureCitation id="13252A22FFF7FFCFA8F1F851FB7EBEB8" box="[1187,1266,1935,1956]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">Fig. 8D</figureCitation>
); lateral edges of each elytron with row of deep pores (
<figureCitation id="13252A22FFF7FFCFAE15F873FD1FBEDE" box="[583,659,1965,1986]" captionStart="FIGURES 8 A – H" captionStartId="24.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@24.[151,1436,193,1781]" captionTargetPageId="24" captionText="FIGURES 8 A – H, Nesolathrus sp. (McIlwraith Range, Queensland, Australia): A, head, dorsal; B, head, ventral; C, mesotibia and mesotarsus; D, prothorax, dorsal; E, prothorax, ventral; F, pterothorax, ventral with mesocoxae removed; G, left elytron, lateral; H, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251852/files/figure.png" pageId="14" pageNumber="209">Fig. 8G</figureCitation>
); length less than
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;
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, Christmas I.,
<collectingCountry id="F3097637FFF7FFCFA8D8F873FB6EBEDE" box="[1162,1250,1965,1986]" name="Australia" pageId="14" pageNumber="209">Australia</collectingCountry>
<taxonomicName id="4C1E4D24FFF7FFCFA8B0F873FA01BEDE" authority="Scott" authorityName="Scott" box="[1250,1421,1965,1986]" class="Insecta" family="Mycetophagidae" genus="Nesolathrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF7FFCFA8B0F873FADABEDE" box="[1250,1366,1965,1986]" italics="true" pageId="14" pageNumber="209">Nesolathrus</emphasis>
Scott
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</paragraph>
</keyStep>
<keyStep id="30EA2E02FFF7FFCEACC5F814FA10B866" lastPageId="15" lastPageNumber="210" pageId="14" pageNumber="209">
<paragraph id="8BA136A7FFF7FFCFACC5F814FA11BEC3" blockId="14.[151,1437,1554,2015]" box="[151,1437,1994,2015]" pageId="14" pageNumber="209">
<keyLead id="30EF9592FFF7FFCFACC5F814FA11BEC3" box="[151,1437,1994,2015]" pageId="14" pageNumber="209">
3(1). Metacoxal length (parallel to long axis of body) more than half its width (perpendicular to long axis of body) (
<figureCitation id="13252A22FFF7FFCFA95EF814FAD4BEC3" box="[1292,1368,1994,2015]" captionStart="FIGURES 6 A – G" captionStartId="22.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@22.[151,1436,193,1781]" captionTargetPageId="22" captionText="FIGURES 6 A – G, Esarcus leprieuri Reiche: A, head, ventral; B, anterior portion of head; C, pterothorax with right mesocoxa and left metacoxa removed; D, prothorax, ventral, with right coxa removed; E, scutellar shield; F, abdominal ventrites; G, right elytron. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251850/files/figure.png" pageId="14" pageNumber="209">Fig. 6C</figureCitation>
), well-
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFF6FFCEACB3FF49FA10B866" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
separated from elytral epipleura; metanepisternum relatively short and broad, its length less than 2.5 times its width; metaventrite distinctly shorter than abdominal ventrite 1; sides of pronotum usually distinctly serrate (
<figureCitation id="13252A22FFF6FFCEA808FF6BFB2BB9D6" box="[1114,1191,181,202]" captionStart="FIGURES 6 A – G" captionStartId="22.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@22.[151,1436,193,1781]" captionTargetPageId="22" captionText="FIGURES 6 A – G, Esarcus leprieuri Reiche: A, head, ventral; B, anterior portion of head; C, pterothorax with right mesocoxa and left metacoxa removed; D, prothorax, ventral, with right coxa removed; E, scutellar shield; F, abdominal ventrites; G, right elytron. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251850/files/figure.png" pageId="15" pageNumber="210">Fig. 6D</figureCitation>
); scutellar shield strongly transverse (
<figureCitation id="13252A22FFF6FFCEAD02FF0CFE16B9FB" box="[336,410,210,231]" captionStart="FIGURES 6 A – G" captionStartId="22.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@22.[151,1436,193,1781]" captionTargetPageId="22" captionText="FIGURES 6 A – G, Esarcus leprieuri Reiche: A, head, ventral; B, anterior portion of head; C, pterothorax with right mesocoxa and left metacoxa removed; D, prothorax, ventral, with right coxa removed; E, scutellar shield; F, abdominal ventrites; G, right elytron. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251850/files/figure.png" pageId="15" pageNumber="210">Fig. 6E</figureCitation>
) or absent; elytral punctures large, nude and distinctly seriate (
<figureCitation id="13252A22FFF6FFCEA853FF0CFBDCB9FB" box="[1025,1104,210,231]" captionStart="FIGURES 6 A – G" captionStartId="22.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@22.[151,1436,193,1781]" captionTargetPageId="22" captionText="FIGURES 6 A – G, Esarcus leprieuri Reiche: A, head, ventral; B, anterior portion of head; C, pterothorax with right mesocoxa and left metacoxa removed; D, prothorax, ventral, with right coxa removed; E, scutellar shield; F, abdominal ventrites; G, right elytron. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251850/files/figure.png" pageId="15" pageNumber="210">Fig. 6G</figureCitation>
); pterothorax and abdomen sometimes with large setose cavities (
<figureCitation id="13252A22FFF6FFCEAE45FF31FD00B818" box="[535,652,239,260]" captionStart="FIGURES 6 A – G" captionStartId="22.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@22.[151,1436,193,1781]" captionTargetPageId="22" captionText="FIGURES 6 A – G, Esarcus leprieuri Reiche: A, head, ventral; B, anterior portion of head; C, pterothorax with right mesocoxa and left metacoxa removed; D, prothorax, ventral, with right coxa removed; E, scutellar shield; F, abdominal ventrites; G, right elytron. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251850/files/figure.png" pageId="15" pageNumber="210">Figs 6C, 6F</figureCitation>
); hind wings usually absent; southern Europe, North Africa
<taxonomicName id="4C1E4D24FFF6FFCEA89FFF31FAA7B818" box="[1229,1323,239,260]" class="Insecta" family="Mycetophagidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="subFamily" subFamily="Esarcinae">Esarcinae</taxonomicName>
.................................................................................................
<taxonomicName id="4C1E4D24FFF6FFCEA954FED3FA10B83E" ID-CoL="62RSY" authority="Reiche" authorityName="Reiche" box="[1286,1436,269,290]" class="Insecta" family="Mycetophagidae" genus="Esarcus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF6FFCEA954FED3FADFB83E" box="[1286,1363,269,290]" italics="true" pageId="15" pageNumber="210">Esarcus</emphasis>
Reiche
</taxonomicName>
<keyLead id="30EF9592FFF6FFCEACC5FEF4FA10B866" pageId="15" pageNumber="210">
- Metacoxal length (parallel to long axis of body) much less than half its width (perpendicular to long axis of body) (
<figureCitation id="13252A22FFF6FFCEA91BFEF4FA14B823" box="[1353,1432,298,319]" captionStart="FIGURES 9 A – G" captionStartId="25.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@25.[151,1436,193,1781]" captionTargetPageId="25" captionText="FIGURES 9 A – G, Mycetophagus punctatus Say: A, head, dorsal; B, prothorax, ventral; C, mesotibial apex; D, antenna; E, prothorax, dorsal; F, portion of meso- and metathorax, showing laterally open mesocoxal cavity; G, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251853/files/figure.png" pageId="15" pageNumber="210">Figs 9G</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEACB3FE99FE84B840" box="[225,264,327,348]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">11E</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAD41FE99FEB3B840" box="[275,319,327,348]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="15" pageNumber="210">14H</figureCitation>
); large, nude, seriate elytral punctures and setose pterothoracic and abdominal cavities always absent.................................................................................................
<taxonomicName id="4C1E4D24FFF6FFCEA88FFEBBFA0CB866" box="[1245,1408,357,378]" class="Insecta" family="Mycetophagidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="subFamily" subFamily="Mycetophaginae">Mycetophaginae</taxonomicName>
...4
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</paragraph>
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<keyStep id="30EA2E02FFF6FFCEACC5FE5CFA17B8CE" pageId="15" pageNumber="210">
<paragraph id="8BA136A7FFF6FFCEACC5FE5CFA17B8A8" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FE5CFA17B8A8" pageId="15" pageNumber="210">
4(3). Mesepimera not reaching mesocoxal cavities, which are closed laterally by the meeting of mesoventrite and metaventrite (
<figureCitation id="13252A22FFF6FFCEA923FE5CFEB2B8A8" captionStart="FIGURES 13 A – L" captionStartId="29.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@29.[151,1436,193,1781]" captionTargetPageId="29" captionText="FIGURES 13 A – L, Afrotyphaeola natalensis sp. n., ♀: A, head, dorsal; B, head, ventral; C, prothorax, dorsal; D, pterothorax, ventral with right mesocoxa removed; E, antenna; F, prothorax, ventral; G, left mesocoxa, showing lateral closure of cavity by mesoventrite and metaventrite; H, abdomen, ventral; I, right procoxal cavity; J, mesotibial apex and mesotarsus; K, protibial apex and protarsus, L, scutellar shield. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251857/files/figure.png" pageId="15" pageNumber="210">Figs 13D, 13G</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAD1AFE41FE2AB8A8" box="[328,422,415,436]" captionStart="FIGURES 15 A – C" captionStartId="31.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@31.[151,1436,193,1781]" captionTargetPageId="31" captionText="FIGURES 15 A – C, Typhaea stercorea (Linnaeus): A, pterothorax, ventral; B, head, dorsal; C, mesocoxa, showing lateral closure of cavity by meso- and metaventrites. Figures 15 D – G, Thrimolus minutus Casey: D, prothorax, ventral, with right coxa removed; E, pterothorax and base of abdomen, ventral; F, head, dorsal; G, left metacoxa and left portion of abdominal ventrite 1, with pore field vaguely indicated. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251859/files/figure.png" pageId="15" pageNumber="210">15A, 15C</figureCitation>
).................................................................................. 5
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFF6FFCEACC5FE63FA17B8CE" blockId="15.[151,1436,151,2020]" box="[151,1435,445,466]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FE63FA17B8CE" box="[151,1435,445,466]" pageId="15" pageNumber="210">
- Mesepimera reaching mesocoxal cavities (
<figureCitation id="13252A22FFF6FFCEAE2AFE63FD53B8CE" box="[632,735,445,466]" captionStart="FIGURES 9 A – G" captionStartId="25.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@25.[151,1436,193,1781]" captionTargetPageId="25" captionText="FIGURES 9 A – G, Mycetophagus punctatus Say: A, head, dorsal; B, prothorax, ventral; C, mesotibial apex; D, antenna; E, prothorax, dorsal; F, portion of meso- and metathorax, showing laterally open mesocoxal cavity; G, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251853/files/figure.png" pageId="15" pageNumber="210">Figs 9F–G</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAEBAFE63FC9CB8CE" box="[744,784,445,466]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">11E</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAF48FE63FCC9B8CE" box="[794,837,445,466]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="15" pageNumber="210">12D</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAF1DFE63FCF6B8CE" box="[847,890,445,466]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="15" pageNumber="210">14H</figureCitation>
)............................................ 8
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</paragraph>
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<keyStep id="30EA2E02FFF6FFCEACC5FE04FA17BB9E" pageId="15" pageNumber="210">
<paragraph id="8BA136A7FFF6FFCEACC5FE04FA10BB36" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FE04FA10BB36" pageId="15" pageNumber="210">
5(4). Base of pronotum distinctly narrower than combined elytral bases (
<figureCitation id="13252A22FFF6FFCEAF36FE04FC11B8F3" box="[868,925,474,495]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="15" pageNumber="210">Fig. 5</figureCitation>
O); antennomeres 9 and 10 about 1.5 times as long as wide, antennomere 11 shorter and narrower than 9 or 10 (
<figureCitation id="13252A22FFF6FFCEAF5AFE29FCC9BB10" box="[776,837,503,524]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">Figs 2</figureCitation>
H, 5O); mesoventrite carinate for most of its length (
<figureCitation id="13252A22FFF6FFCEA916FE29FAF1BB10" box="[1348,1405,503,524]" captionStart="FIGURES 3 A – B" captionStartId="19.[151,264,1802,1825]" captionTargetBox="[151,1431,193,1778]" captionTargetId="figure@19.[151,1436,193,1781]" captionTargetPageId="19" captionText="FIGURES 3 A – B, portion of elytral surface: A, Typhaea stercorea (Linnaeus); B, Nototriphyllus araucania sp. n. Figure 3 C, Zeclaviger explanatus sp. n., pterothorax, ventral, with right metanepisternum and right metacoxa and hind leg removed. Figure 3 D, Nototriphyllus araucania sp. n., prothorax, dorsal, with both coxae removed. Figure 3 E, Zeclaviger explanatus sp. n., left halves of abdominal ventrites 1 and 2, showing pore field in anterolateral corner of ventrite 1. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251847/files/figure.png" pageId="15" pageNumber="210">Fig. 3</figureCitation>
C); vestiture of moderately long erect and inclined setae (
<figureCitation id="13252A22FFF6FFCEAEB1FDCBFC90BB36" box="[739,796,533,554]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="15" pageNumber="210">Fig. 5</figureCitation>
O);
<collectingCountry id="F3097637FFF6FFCEAF10FDCBFC4EBB36" box="[834,962,533,554]" name="New Zealand" pageId="15" pageNumber="210">New Zealand</collectingCountry>
......................
<taxonomicName id="4C1E4D24FFF6FFCEA887FDCBFAB0BB36" box="[1237,1340,533,554]" class="Insecta" family="Mycetophagidae" genus="Zeclaviger" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96AEAB5FFF6FFCEA887FDCBFAB0BB36" box="[1237,1340,533,554]" italics="true" pageId="15" pageNumber="210">Zeclaviger</emphasis>
</taxonomicName>
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<emphasis id="B96AEAB5FFF6FFCEA910FDCAFA10BB36" bold="true" box="[1346,1436,532,554]" pageId="15" pageNumber="210">gen. nov.</emphasis>
</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BA136A7FFF6FFCEACC5FDECFA17BB9E" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FDECFA17BB9E" pageId="15" pageNumber="210">- Base of pronotum not or barely narrower than combined elytral bases; antennomeres 9 and 10 transverse or only slightly longer than wide; antennomere 11 at least slightly longer than 9 or10; mesoventrite not carinate, sometimes with short anterior linear prosternal rest; vestiture without long, erect setae......................................................... 6</keyLead>
</paragraph>
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<keyStep id="30EA2E02FFF6FFCEACC5FD54FA17BA70" pageId="15" pageNumber="210">
<paragraph id="8BA136A7FFF6FFCEACC5FD54FA17BBEB" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FD54FA17BBEB" pageId="15" pageNumber="210">
6(5). Elytral vestiture dual, consisting of longer and shorter, inclined or decumbent setae, the longer setae forming about 12 longitudinal rows separated by at least two rows of shorter setae (
<figureCitation id="13252A22FFF6FFCEAF59FD79FCD6BBA0" box="[779,858,679,700]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1H</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAF36FD79FC08BBA0" box="[868,900,679,700]" captionStart="FIGURES 3 A – B" captionStartId="19.[151,264,1802,1825]" captionTargetBox="[151,1431,193,1778]" captionTargetId="figure@19.[151,1436,193,1781]" captionTargetPageId="19" captionText="FIGURES 3 A – B, portion of elytral surface: A, Typhaea stercorea (Linnaeus); B, Nototriphyllus araucania sp. n. Figure 3 C, Zeclaviger explanatus sp. n., pterothorax, ventral, with right metanepisternum and right metacoxa and hind leg removed. Figure 3 D, Nototriphyllus araucania sp. n., prothorax, dorsal, with both coxae removed. Figure 3 E, Zeclaviger explanatus sp. n., left halves of abdominal ventrites 1 and 2, showing pore field in anterolateral corner of ventrite 1. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251847/files/figure.png" pageId="15" pageNumber="210">3A</figureCitation>
); apical maxillary palpomere more or less parallel-sided and not wider than subapical one (
<figureCitation id="13252A22FFF6FFCEAE67FD1BFD02BBC6" box="[565,654,709,730]" captionStart="FIGURES 15 A – C" captionStartId="31.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@31.[151,1436,193,1781]" captionTargetPageId="31" captionText="FIGURES 15 A – C, Typhaea stercorea (Linnaeus): A, pterothorax, ventral; B, head, dorsal; C, mesocoxa, showing lateral closure of cavity by meso- and metaventrites. Figures 15 D – G, Thrimolus minutus Casey: D, prothorax, ventral, with right coxa removed; E, pterothorax and base of abdomen, ventral; F, head, dorsal; G, left metacoxa and left portion of abdominal ventrite 1, with pore field vaguely indicated. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251859/files/figure.png" pageId="15" pageNumber="210">Fig. 15B</figureCitation>
); shortest distance between mesocoxal and metacoxal cavities distinctly greater than length of abdominal ventrite 1 at same point; hind wings well-developed; cosmopolitan (
<figureCitation id="13252A22FFF6FFCEA831FD3CFB21BBEB" box="[1123,1197,738,759]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Fig. 1H</figureCitation>
)....
<taxonomicName id="4C1E4D24FFF6FFCEA8BDFD3CFA17BBEB" ID-CoL="8472" authority="Stephens" authorityName="Stephens" box="[1263,1435,738,759]" class="Insecta" family="Mycetophagidae" genus="Typhaea" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF6FFCEA8BDFD3CFACCBBEB" box="[1263,1344,738,759]" italics="true" pageId="15" pageNumber="210">Typhaea</emphasis>
Stephens
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- Elytral vestiture single, consisting of inclined or decumbent setae of more or less equal length (
<figureCitation id="13252A22FFF6FFCEA827FD21FB4FBA08" box="[1141,1219,767,788]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">Fig. 2A</figureCitation>
); apical maxillary palpomere distinctly wider than subapical one (
<figureCitation id="13252A22FFF6FFCEAEC0FCC3FC81BA2E" box="[658,781,797,818]" captionStart="FIGURES 13 A – L" captionStartId="29.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@29.[151,1436,193,1781]" captionTargetPageId="29" captionText="FIGURES 13 A – L, Afrotyphaeola natalensis sp. n., ♀: A, head, dorsal; B, head, ventral; C, prothorax, dorsal; D, pterothorax, ventral with right mesocoxa removed; E, antenna; F, prothorax, ventral; G, left mesocoxa, showing lateral closure of cavity by mesoventrite and metaventrite; H, abdomen, ventral; I, right procoxal cavity; J, mesotibial apex and mesotarsus; K, protibial apex and protarsus, L, scutellar shield. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251857/files/figure.png" pageId="15" pageNumber="210">Figs 13A–B</figureCitation>
); shortest distance between mesocoxal and metacoxal cavities subequal to or less than length of abdominal ventrite 1 at the same point (
<figureCitation id="13252A22FFF6FFCEAFD2FCE4FC55BA53" box="[896,985,826,847]" captionStart="FIGURES 13 A – L" captionStartId="29.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@29.[151,1436,193,1781]" captionTargetPageId="29" captionText="FIGURES 13 A – L, Afrotyphaeola natalensis sp. n., ♀: A, head, dorsal; B, head, ventral; C, prothorax, dorsal; D, pterothorax, ventral with right mesocoxa removed; E, antenna; F, prothorax, ventral; G, left mesocoxa, showing lateral closure of cavity by mesoventrite and metaventrite; H, abdomen, ventral; I, right procoxal cavity; J, mesotibial apex and mesotarsus; K, protibial apex and protarsus, L, scutellar shield. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251857/files/figure.png" pageId="15" pageNumber="210">Fig. 13D</figureCitation>
); hind wings more or less reduced (apical field shortened or absent, medial field reduced, radial cell sometimes absent).......................................... 7
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7(6). Pronotum widest near middle; lateral pronotal carinae distinctly serrate; posterior pronotal angles acute; lateral edges of elytra visible from above for most of their lengths; elytral apex obliquely truncate; protarsomere
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male distinctly expanded; abdominal ventrite
<quantity id="4CE69B42FFF6FFCEADCEFC71FE4EBAD8" box="[412,450,943,964]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="210" unit="in" value="1.0">1 in</quantity>
male with two large patches of setigerous spicules accompanied by glandular openings (
<figureCitation id="13252A22FFF6FFCEA948FC71FA0CBAD8" box="[1306,1408,943,964]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="15" pageNumber="210">Figs 4C, 4</figureCitation>
F); southern Europe, North Africa..........................................................
<taxonomicName id="4C1E4D24FFF6FFCEA893FC13FA10BAFE" authority="Ganglbauer" authorityName="Ganglbauer" box="[1217,1436,973,994]" class="Insecta" family="Mycetophagidae" genus="Typhaeola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
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Ganglbauer
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- Pronotum widest well behind middle (
<figureCitation id="13252A22FFF6FFCEAE0FFC34FD21BAE3" box="[605,685,1002,1023]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">Fig. 2A</figureCitation>
); lateral pronotal carinae simple; posterior pronotal angles not acute; lateral edges of elytra not visible from above; elytral apex broadly rounded; protarsomere
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male not expanded; abdominal ventrite
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male with two slender transverse pore fields (
<figureCitation id="13252A22FFF6FFCEAEEDFBFBFCAABD26" box="[703,806,1061,1082]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="15" pageNumber="210">Figs 4A, 4</figureCitation>
D); southern Africa.................
<taxonomicName id="4C1E4D24FFF6FFCEA8E1FBFBFAB7BD26" box="[1203,1339,1061,1082]" class="Insecta" family="Mycetophagidae" genus="Afrotyphaeola" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus" status="gen. nov.">
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8(4). Antennae gradually expanded apically or with weak club composed of 4 or more antennomeres (
<figureCitation id="13252A22FFF6FFCEA8C6FB9CFB71BD4B" box="[1172,1277,1090,1111]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1D–F</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEA95AFB9CFA98BD4B" box="[1288,1300,1090,1111]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="15" pageNumber="210">5</figureCitation>
J, 9D); elytral punctation seriate, but puncture rows sometimes not clearly visible due to dense interstrial punctation and recumbent setae (
<figureCitation id="13252A22FFF6FFCEACB5FBA3FEDCBD8E" box="[231,336,1149,1170]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1D–F</figureCitation>
).......................................................................................... 9
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- Antennae with distinct 3-segmented club (
<figureCitation id="13252A22FFF6FFCEAE24FB44FC9BBDB3" box="[630,791,1178,1199]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">
Figs 1A–B,
<date id="FFA01067FFF6FFCEAEBEFB44FD71BDB3" box="[748,765,1178,1199]" pageId="15" pageNumber="210">1I</date>
–J
</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAF73FB44FCA0BDB3" box="[801,812,1178,1199]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">2</figureCitation>
F–G,
<date id="FFA01067FFF6FFCEAF33FB44FCFABDB3" box="[865,886,1178,1199]" pageId="15" pageNumber="210">5I</date>
, 5M–N, 14B); punctation seriate or not............ 11
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9(8). Frontoclypeal suture somewhat angulate and not deeply impressed; paired basal pronotal impressions absent; posterior pronotal angles more or less embracing elytral bases (
<figureCitation id="13252A22FFF6FFCEAEFAFB0BFD6EBDF6" box="[680,738,1237,1258]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="15" pageNumber="210">Fig. 5</figureCitation>
J); shortest distance between mesocoxal and metacoxal cavities distinctly less than length of abdominal ventrite 1 at same point; tibial spurs biserrate; hind wings reduced and strap-like; Juan Fernandez Is.........................................................................
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Leschen &amp; Lawrence
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- Frontoclypeal suture deeply impressed and straight or curved (
<figureCitation id="13252A22FFF6FFCEAF11FAF3FC0DBC5E" box="[835,897,1325,1346]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">Figs 2</figureCitation>
E, 9A); paired basal pronotal impressions present (
<figureCitation id="13252A22FFF6FFCEA923FAF3FE95BC43" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1E–F</figureCitation>
); shortest distance between mesocoxal and metacoxal cavities distinctly greater than length of abdominal ventrite 1 at same point (
<figureCitation id="13252A22FFF6FFCEAD04FAB9FE28BC60" box="[342,420,1383,1404]" captionStart="FIGURES 9 A – G" captionStartId="25.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@25.[151,1436,193,1781]" captionTargetPageId="25" captionText="FIGURES 9 A – G, Mycetophagus punctatus Say: A, head, dorsal; B, prothorax, ventral; C, mesotibial apex; D, antenna; E, prothorax, dorsal; F, portion of meso- and metathorax, showing laterally open mesocoxal cavity; G, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251853/files/figure.png" pageId="15" pageNumber="210">Fig. 9G</figureCitation>
); tibial spurs simple (
<figureCitation id="13252A22FFF6FFCEAE20FAB9FD31BC60" box="[626,701,1383,1404]" captionStart="FIGURES 9 A – G" captionStartId="25.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@25.[151,1436,193,1781]" captionTargetPageId="25" captionText="FIGURES 9 A – G, Mycetophagus punctatus Say: A, head, dorsal; B, prothorax, ventral; C, mesotibial apex; D, antenna; E, prothorax, dorsal; F, portion of meso- and metathorax, showing laterally open mesocoxal cavity; G, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251853/files/figure.png" pageId="15" pageNumber="210">Fig. 9C</figureCitation>
); hind wings well-developed................................... 10
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</paragraph>
</keyStep>
<keyStep id="30EA2E02FFF6FFCEACC5FA5BFA17BF30" pageId="15" pageNumber="210">
<paragraph id="8BA136A7FFF6FFCEACC5FA5BFA10BCC8" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FA5BFA10BCC8" pageId="15" pageNumber="210">
10(9). Basal pronotal impressions very small and sometimes indistinct; elytron with sutural stria; antennal club indistinct; eyes rounded; elytra usually unicolorous red or brown (
<figureCitation id="13252A22FFF6FFCEAE96FA7CFC9CBCAB" box="[708,784,1442,1463]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Fig. 1D</figureCitation>
); eastern Palaearctic, Caucasus, North Africa,
<collectingCountry id="F3097637FFF6FFCEA89DFA7CFA8ABCAB" box="[1231,1286,1442,1463]" name="Japan" pageId="15" pageNumber="210">Japan</collectingCountry>
...............................................................................................
<taxonomicName id="4C1E4D24FFF6FFCEA896FA61FA10BCC8" authority="Motschulsky" authorityName="Motschulsky" box="[1220,1436,1471,1492]" class="Insecta" family="Mycetophagidae" genus="Eulagius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF6FFCEA896FA61FA95BCC8" box="[1220,1305,1471,1492]" italics="true" pageId="15" pageNumber="210">Eulagius</emphasis>
Motschulsky
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFF6FFCEACC5FA03FA17BF30" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5FA03FA17BF30" pageId="15" pageNumber="210">
- Basal pronotal impressions large and distinct (
<figureCitation id="13252A22FFF6FFCEAECCFA03FC8ABCEE" box="[670,774,1501,1522]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1E–F</figureCitation>
); elytra without sutural stria; antennal club usually distinctly 4- to 6- segmented (
<figureCitation id="13252A22FFF6FFCEAD05FA24FE24BF13" box="[343,424,1530,1551]" captionStart="FIGURES 9 A – G" captionStartId="25.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@25.[151,1436,193,1781]" captionTargetPageId="25" captionText="FIGURES 9 A – G, Mycetophagus punctatus Say: A, head, dorsal; B, prothorax, ventral; C, mesotibial apex; D, antenna; E, prothorax, dorsal; F, portion of meso- and metathorax, showing laterally open mesocoxal cavity; G, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251853/files/figure.png" pageId="15" pageNumber="210">Fig. 9D</figureCitation>
); eyes elongate-oval, usually slightly emarginate (
<figureCitation id="13252A22FFF6FFCEAFCBFA24FC66BF13" box="[921,1002,1530,1551]" captionStart="FIGURES 9 A – G" captionStartId="25.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@25.[151,1436,193,1781]" captionTargetPageId="25" captionText="FIGURES 9 A – G, Mycetophagus punctatus Say: A, head, dorsal; B, prothorax, ventral; C, mesotibial apex; D, antenna; E, prothorax, dorsal; F, portion of meso- and metathorax, showing laterally open mesocoxal cavity; G, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251853/files/figure.png" pageId="15" pageNumber="210">Fig. 9A</figureCitation>
); elytra usually with dark and light pattern, occasionally black; widely distributed except
<collectingCountry id="F3097637FFF6FFCEAEC5F9C9FD63BF30" box="[663,751,1559,1580]" name="Australia" pageId="15" pageNumber="210">Australia</collectingCountry>
and
<collectingCountry id="F3097637FFF6FFCEAF4CF9C9FC12BF30" box="[798,926,1559,1580]" name="New Zealand" pageId="15" pageNumber="210">New Zealand</collectingCountry>
.......................
<taxonomicName id="4C1E4D24FFF6FFCEA8EEF9C9FA17BF30" ID-CoL="5W2X" authority="Hellwig" authorityName="Hellwig" box="[1212,1435,1559,1580]" class="Insecta" family="Mycetophagidae" genus="Mycetophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFF6FFCEA8EEF9C9FAC4BF30" box="[1212,1352,1559,1580]" italics="true" pageId="15" pageNumber="210">Mycetophagus</emphasis>
Hellwig
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</paragraph>
</keyStep>
<keyStep id="30EA2E02FFF6FFCEACC5F9EBFA17BEB6" pageId="15" pageNumber="210">
<paragraph id="8BA136A7FFF6FFCEACC5F9EBFA17BE0B" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5F9EBFA17BE0B" pageId="15" pageNumber="210">
11(8). Pronotum widest at base, which is as wide as or wider than combined elytral bases (
<figureCitation id="13252A22FFF6FFCEA855F9EBFBEFBF56" box="[1031,1123,1589,1610]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">
Figs
<date id="FFA01067FFF6FFCEA865F9EBFBC5BF56" box="[1079,1097,1589,1610]" pageId="15" pageNumber="210">1I</date>
–J
</figureCitation>
); posterior pronotal angles more or less embracing elytral humeri (
<figureCitation id="13252A22FFF6FFCEAE7CF98CFD24BF7B" box="[558,680,1618,1639]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">Figs 11C–D</figureCitation>
); pronotal hypomera (
<figureCitation id="13252A22FFF6FFCEAFD0F98CFB9CBF7B" box="[898,1040,1618,1639]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="15" pageNumber="210">Figs 12F, 12H</figureCitation>
) and elytral epipleura (
<figureCitation id="13252A22FFF6FFCEA8A5F98CFACBBF7B" box="[1271,1351,1618,1639]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="15" pageNumber="210">
Fig.
<date id="FFA01067FFF6FFCEA974F98CFACBBF7B" box="[1318,1351,1618,1639]" pageId="15" pageNumber="210" value="2012-01">12I</date>
</figureCitation>
) usually strongly oblique or almost vertical, not visible in lateral view; pronotal disc with pair of elongate, slit-like, basal impressions, usually darkly pigmented (
<figureCitation id="13252A22FFF6FFCEADB0F953FD91BFBE" box="[482,541,1677,1698]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">Fig. 2</figureCitation>
C); metaventrite not shorter than first abdominal ventrite (
<figureCitation id="13252A22FFF6FFCEA819F953FB28BFBE" box="[1099,1188,1677,1698]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="15" pageNumber="210">Fig. 12D</figureCitation>
); frontoclypeal suture not deeply impressed, forming a fine curved or angulate line (
<figureCitation id="13252A22FFF6FFCEAF4CF974FCF4BFA3" box="[798,888,1706,1727]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">Fig. 11A</figureCitation>
); scutellar shield widest at base and broadly rounded at apex (
<figureCitation id="13252A22FFF6FFCEAD4EF919FEFABFC0" box="[284,374,1735,1756]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="15" pageNumber="210">Fig. 12K</figureCitation>
); antennal club usually slender, often with antennomeres 9–11 at least slightly elongate (
<figureCitation id="13252A22FFF6FFCEA89DF919FA92BFC0" box="[1231,1310,1735,1756]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">
Fig.
<date id="FFA01067FFF6FFCEA8AEF919FA92BFC0" box="[1276,1310,1735,1756]" pageId="15" pageNumber="210" value="2011-01">11I</date>
</figureCitation>
); tibial spurs usually bipectinate and sometimes highly complex (
<figureCitation id="13252A22FFF6FFCEAE8FF93BFCE6BFE6" box="[733,874,1765,1786]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">Figs 11B, 11G</figureCitation>
–HN, 12B); body more or less flattened, often with complex, dual vestiture (
<figureCitation id="13252A22FFF6FFCEADF6F8DCFDA6BE0B" box="[420,554,1794,1815]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="15" pageNumber="210">
Figs 11A,
<date id="FFA01067FFF6FFCEAE58F8DCFDA6BE0B" box="[522,554,1794,1815]" pageId="15" pageNumber="210" value="2011-01">11I</date>
</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAE66F8DCFDD3BE0B" box="[564,607,1794,1815]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="15" pageNumber="210">12A</figureCitation>
).................................................................. 12
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</paragraph>
<paragraph id="8BA136A7FFF6FFCEACC5F8C1FA17BEB6" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
<keyLead id="30EF9592FFF6FFCEACC5F8C1FA17BEB6" pageId="15" pageNumber="210">
- Pronotum usually widest anterior to base (
<figureCitation id="13252A22FFF6FFCEAED5F8C1FCACBE28" box="[647,800,1823,1844]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1G–H, 1L</figureCitation>
,
<date id="FFA01067FFF6FFCEAF7CF8C1FCCFBE28" box="[814,835,1823,1844]" pageId="15" pageNumber="210">
<figureCitation id="13252A22FFF6FFCEAF7CF8C1FCB6BE28" box="[814,826,1823,1844]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="15" pageNumber="210">5</figureCitation>
I
</date>
, 5N); IF pronotum widest at base, THEN posterior pronotal angles not embracing elytral humeri (
<figureCitation id="13252A22FFF6FFCEAE02F8E3FD2DBE4E" box="[592,673,1853,1874]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="15" pageNumber="210">Figs 1A</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAEFFF8E3FD34BE4E" box="[685,696,1853,1874]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="15" pageNumber="210">5</figureCitation>
M) OR metaventrite distinctly shorter than first abdominal ventrite; base of pronotal disc without or with subcircular paired impressions; frontoclypeal suture usually more deeply impressed; elytral epipleura more or less flattened, often partly visible in lateral view; scutellar shield usually widest behind base and apically truncate (
<figureCitation id="13252A22FFF6FFCEAD46F84BFEE1BEB6" box="[276,365,1941,1962]" captionStart="FIGURES 10 A – H" captionStartId="26.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@26.[151,1436,193,1781]" captionTargetPageId="26" captionText="FIGURES 10 A – H, Pseudochrodes suturalis Reitter: A, head, dorsal; B, prothorax, dorsal; C, mesotibial apex; D, antenna; E, prothorax, ventral; F, left elytron, dorsal; G, scutellar shield; H, male abdomen showing setose sex patches. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251854/files/figure.png" pageId="15" pageNumber="210">Figs 10G</figureCitation>
,
<figureCitation id="13252A22FFF6FFCEAD25F84BFE2FBEB6" box="[375,419,1941,1962]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="15" pageNumber="210">14D</figureCitation>
), body usually more convex with simple vestiture........................................... 13
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<paragraph id="8BA136A7FFF6FFCEACC5F86CFA17BEF8" blockId="15.[151,1436,151,2020]" pageId="15" pageNumber="210">
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12(11). Outer edge of protibia lined with closely-spaced, spine-like setae; protibial apex with one long bipectinate spur and another much shorter spur (
<figureCitation id="13252A22FFF6FFCEADCBF811FE5ABEF8" box="[409,470,1999,2020]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="15" pageNumber="210">
Figs
<date id="FFA01067FFF6FFCEAD9BF811FE5ABEF8" box="[457,470,1999,2020]" pageId="15" pageNumber="210">2</date>
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I, 12B); apical maxillary palpomere slightly longer than preapical one and strongly obliquely truncate
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<paragraph id="8BA136A7FFE9FFD1ACB3FF49FA17B823" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
so that outer edge is acute (
<figureCitation id="13252A22FFE9FFD1ADB6FF49FDB0B9B0" box="[484,572,151,172]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="16" pageNumber="211">Fig. 12C</figureCitation>
); apical labial palpomere as wide as or slightly wider than preapical one and attached at its apex (
<figureCitation id="13252A22FFE9FFD1AD49FF6BFEFFB9D6" box="[283,371,181,202]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="16" pageNumber="211">Fig. 12C</figureCitation>
); elytra with distinct rows of punctures (
<figureCitation id="13252A22FFE9FFD1AEA4FF6BFCB0B9D6" box="[758,828,181,202]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="16" pageNumber="211">
Figs
<date id="FFA01067FFE9FFD1AF75FF6BFCB0B9D6" box="[807,828,181,202]" pageId="16" pageNumber="211">1I</date>
</figureCitation>
,
<figureCitation id="13252A22FFE9FFD1AF15FF6BFCE0B9D6" box="[839,876,181,202]" captionStart="FIGURES 12 A – L" captionStartId="28.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@28.[151,1436,193,1781]" captionTargetPageId="28" captionText="FIGURES 12 A – L, Litargops sp. (Brindabella Range, ACT, Australia): A, head, dorsal; B, apex of protibia; C, head, ventral; D, pterothorax, ventral with right mesocoxa removed; E, apex of mesotibia; F, prothorax, ventral, with right coxa removed; G, mesotibial apex and mesotarsus; H, prothorax, anterior; I, left elytron, ventral; J, right elytron, dorsal; K, scutellar shield; L, antenna. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251856/files/figure.png" pageId="16" pageNumber="211">12J</figureCitation>
); Madeira,
<collectingCountry id="F3097637FFE9FFD1AF8BFF6BFB9EB9D6" box="[985,1042,181,202]" name="Japan" pageId="16" pageNumber="211">Japan</collectingCountry>
,
<collectingCountry id="F3097637FFE9FFD1A849FF6BFBFFB9D6" box="[1051,1139,181,202]" name="Australia" pageId="16" pageNumber="211">Australia</collectingCountry>
..........
<taxonomicName id="4C1E4D24FFE9FFD1A8AAFF6BFA10B9D6" authority="Reitter" authorityName="Reitter" box="[1272,1436,181,202]" class="Insecta" family="Mycetophagidae" genus="Litargops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A8AAFF6BFAD9B9D6" box="[1272,1365,181,202]" italics="true" pageId="16" pageNumber="211">Litargops</emphasis>
Reitter
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- Outer edge of protibia lined with finer, less closely spaced setae (
<figureCitation id="13252A22FFE9FFD1AF37FF0CFC33B9FB" box="[869,959,210,231]" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="16" pageNumber="211">Fig. 11G</figureCitation>
); protibial apex with two bipectinate spurs (
<figureCitation id="13252A22FFE9FFD1A923FF0CFEA6B818" captionStart="FIGURES 11 A – I" captionStartId="27.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@27.[151,1436,193,1781]" captionTargetPageId="27" captionText="FIGURES 11 A – I, Litargus irregularis (Sharp): A, head, dorsal; B, mesotibia and mesotarsus; C, pronotum, dorsal; D, prothrax, ventral; E, pterothorax, ventral, with right meso- and metacoxae removed; F, antenna; G, protibia and protarsus; H, apex of protibia; I, area of dorsal head surface. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251855/files/figure.png" pageId="16" pageNumber="211">Figs 11G–H</figureCitation>
); apical maxillary palpomere distinctly longer than preapical one and not strongly, obliquely truncate (
<figureCitation id="13252A22FFE9FFD1A951FF31FAB1B818" box="[1283,1341,239,260]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Fig. 2</figureCitation>
K); apical labial palpomere distinctly narrower than preapical one and attached subapically (
<figureCitation id="13252A22FFE9FFD1A852FED3FBB7B83E" box="[1024,1083,269,290]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Fig. 2</figureCitation>
J); elytra punctures not forming distinct rows (
<figureCitation id="13252A22FFE9FFD1AD1FFEF4FE1CB823" box="[333,400,298,319]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="16" pageNumber="211">Fig. 1J</figureCitation>
); widely distributed........................................................
<taxonomicName id="4C1E4D24FFE9FFD1A8BEFEF4FA17B823" ID-CoL="5FK6" authority="Erichson" authorityName="Erichson" box="[1260,1435,298,319]" class="Insecta" family="Mycetophagidae" genus="Litargus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A8BEFEF4FACCB823" box="[1260,1344,298,319]" italics="true" pageId="16" pageNumber="211">Litargus</emphasis>
Erichson
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13(11). Body about 2.5 times as long as wide (
<figureCitation id="13252A22FFE9FFD1AE0DFE99FD21B840" box="[607,685,327,348]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="16" pageNumber="211">Fig. 1G</figureCitation>
); lateral pronotal carinae dentate (
<figureCitation id="13252A22FFE9FFD1AFA9FE99FB01B840" box="[1019,1165,327,348]" captionStart="FIGURES 10 A – H" captionStartId="26.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@26.[151,1436,193,1781]" captionTargetPageId="26" captionText="FIGURES 10 A – H, Pseudochrodes suturalis Reitter: A, head, dorsal; B, prothorax, dorsal; C, mesotibial apex; D, antenna; E, prothorax, ventral; F, left elytron, dorsal; G, scutellar shield; H, male abdomen showing setose sex patches. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251854/files/figure.png" pageId="16" pageNumber="211">Figs 10B, 10E</figureCitation>
); sides of elytra parallel for most of their lengths; elytral punctation distinctly seriate (
<figureCitation id="13252A22FFE9FFD1AF58FEBBFCD3B866" box="[778,863,357,378]" captionStart="FIGURES 10 A – H" captionStartId="26.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@26.[151,1436,193,1781]" captionTargetPageId="26" captionText="FIGURES 10 A – H, Pseudochrodes suturalis Reitter: A, head, dorsal; B, prothorax, dorsal; C, mesotibial apex; D, antenna; E, prothorax, ventral; F, left elytron, dorsal; G, scutellar shield; H, male abdomen showing setose sex patches. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251854/files/figure.png" pageId="16" pageNumber="211">Fig. 10F</figureCitation>
); abdominal ventrites
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male each with a pair of small, circular setose patches near midline (
<figureCitation id="13252A22FFE9FFD1AE16FE5CFD12B88B" box="[580,670,386,407]" captionStart="FIGURES 10 A – H" captionStartId="26.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@26.[151,1436,193,1781]" captionTargetPageId="26" captionText="FIGURES 10 A – H, Pseudochrodes suturalis Reitter: A, head, dorsal; B, prothorax, dorsal; C, mesotibial apex; D, antenna; E, prothorax, ventral; F, left elytron, dorsal; G, scutellar shield; H, male abdomen showing setose sex patches. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251854/files/figure.png" pageId="16" pageNumber="211">Fig. 10H</figureCitation>
);
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and
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............................
<taxonomicName id="4C1E4D24FFE9FFD1A890FE5CFA10B88B" authority="Reitter" authorityName="Reitter" box="[1218,1436,386,407]" class="Insecta" family="Silvanidae" genus="Pseudochrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A890FE5CFAD8B88B" box="[1218,1364,386,407]" italics="true" pageId="16" pageNumber="211">Pseudochrodes</emphasis>
Reitter
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFE9FFD1ACC5FE41FA17B8F3" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FE41FA17B8F3" pageId="16" pageNumber="211">
- Body shorter and broader (
<figureCitation id="13252A22FFE9FFD1ADB1FE41FDFAB8A8" box="[483,630,415,436]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="16" pageNumber="211">Figs 1A–B, 1L</figureCitation>
,
<date id="FFA01067FFE9FFD1AED3FE41FD1AB8A8" box="[641,662,415,436]" pageId="16" pageNumber="211">
<figureCitation id="13252A22FFE9FFD1AED3FE41FD02B8A8" box="[641,654,415,436]" captionStart="FIGURES 5 A – H" captionStartId="21.[151,264,1784,1807]" captionTargetBox="[160,1421,196,1771]" captionTargetId="figure@21.[151,1436,193,1781]" captionTargetPageId="21" captionText="FIGURES 5 A – H, mycetophagid male genitalia: A – B, Nototriphyllus araucania sp. n.: A, basale (phallobase) and apicale (partly fused parameres); B, penis; C – E, Neotriphyllus confusus (Horn): C, basale and apicale; D, penis; E, segments IX and X with long spiculum gastrale; F – H, Zeclaviger explanatus sp. n.: F, basale and apicale; G, penis; H, segments IX and X with short spiculum gastrale. FIGURES 5 I – J, habitus, dorsal: I, Nototriphyllus hispidellus (Broun), length 1.9 mm, © Landcare New Zealand; J, Filicivora chilensis (Philippi &amp; Philippi), length 5.3 mm, © CSIRO Australia. FIGURES 5 K – L, Afrotyphaeola natalensis sp. n.: K, basale and apicale; L, penis. FIGURES 5 M – O, habitus: M, Neotriphyllus confusus (Horn), length 3.4 mm, © Landcare New Zealand; N, Nototriphyllus araucania sp. n., length 2.8 mm, © CSIRO Australia; O, Zeclaviger explanatus sp. n., length 1.6 mm, © Landcare New Zealand." httpUri="https://zenodo.org/record/251849/files/figure.png" pageId="16" pageNumber="211">5</figureCitation>
I
</date>
, 5M–N); lateral pronotal carinae simple or minutely denticulate (
<figureCitation id="13252A22FFE9FFD1A959FE41FAC4B8A8" box="[1291,1352,415,436]" captionStart="FIGURES 3 A – B" captionStartId="19.[151,264,1802,1825]" captionTargetBox="[151,1431,193,1778]" captionTargetId="figure@19.[151,1436,193,1781]" captionTargetPageId="19" captionText="FIGURES 3 A – B, portion of elytral surface: A, Typhaea stercorea (Linnaeus); B, Nototriphyllus araucania sp. n. Figure 3 C, Zeclaviger explanatus sp. n., pterothorax, ventral, with right metanepisternum and right metacoxa and hind leg removed. Figure 3 D, Nototriphyllus araucania sp. n., prothorax, dorsal, with both coxae removed. Figure 3 E, Zeclaviger explanatus sp. n., left halves of abdominal ventrites 1 and 2, showing pore field in anterolateral corner of ventrite 1. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251847/files/figure.png" pageId="16" pageNumber="211">Figs 3</figureCitation>
D, 14E); sides of elytra more or less curved for most of their lengths; elytral punctation seriate or not; abdominal ventrites in male without paired, subcircular setose patches..................................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EA2E02FFE9FFD1ACC5FE29FA17BB36" pageId="16" pageNumber="211">
<paragraph id="8BA136A7FFE9FFD1ACC5FE29FA17BB10" blockId="16.[151,1437,151,1786]" box="[151,1435,503,524]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FE29FA17BB10" box="[151,1435,503,524]" pageId="16" pageNumber="211">14(13). Elytra with distinct puncture rows....................................................................... 15</keyLead>
</paragraph>
<paragraph id="8BA136A7FFE9FFD1ACC5FDCBFA17BB36" blockId="16.[151,1437,151,1786]" box="[151,1435,533,554]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FDCBFA17BB36" box="[151,1435,533,554]" pageId="16" pageNumber="211">- Elytra without distinct puncture rows (occasionally with indistinct or incomplete rows)............................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EA2E02FFE9FFD1ACC5FDECFA10BBEB" pageId="16" pageNumber="211">
<paragraph id="8BA136A7FFE9FFD1ACC5FDECFA10BB9E" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FDECFA10BB9E" pageId="16" pageNumber="211">
15(14). Shortest distance between mesocoxal and metacoxal cavities distinctly less than length of abdominal ventrite 1 at same point; sides of elytra distinctly converging beyond middle; elytra with short scutellary striole; pronotum at least 0.65 times as long as wide; hind wings absent; abdominal ventrites in male without pore fields; central Asia..............
<taxonomicName id="4C1E4D24FFE9FFD1A894FDB3FA10BB9E" authority="Nikitsky" authorityName="Nikitsky" box="[1222,1436,621,642]" class="Insecta" family="Mycetophagidae" genus="Triphyllopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A894FDB3FACFBB9E" box="[1222,1347,621,642]" italics="true" pageId="16" pageNumber="211">Triphyllopsis</emphasis>
Nikitsky
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFE9FFD1ACC5FD54FA10BBEB" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FD54FA10BBEB" pageId="16" pageNumber="211">
- Shortest distance between mesocoxal and metacoxal cavities distinctly greater than length of abdominal ventrite 1 at same point; sides of elytra more or less parallel, converging only at posterior fourth; elytra without scutellary striole; pronotum less than half as long as wide; hind wings well-developed; abdominal ventrites
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male each with a pair of transversely oriented V- or U-shaped pore fields (as in
<figureCitation id="13252A22FFE9FFD1AE08FD3CFD29BBEB" box="[602,677,738,759]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="16" pageNumber="211">Fig. 4B</figureCitation>
);
<collectingCountry id="F3097637FFE9FFD1AEE4FD3CFD61BBEB" box="[694,749,738,759]" name="Japan" pageId="16" pageNumber="211">Japan</collectingCountry>
......................................
<taxonomicName id="4C1E4D24FFE9FFD1A8E7FD3CFA10BBEB" authority="Miyatake" authorityName="Miyatake" box="[1205,1436,738,759]" class="Insecta" family="Mycetophagidae" genus="Triphyllioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A8E7FD3CFAB6BBEB" box="[1205,1338,738,759]" italics="true" pageId="16" pageNumber="211">Triphyllioides</emphasis>
Miyatake
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EA2E02FFE9FFD1ACC5FD21FA17BABB" pageId="16" pageNumber="211">
<paragraph id="8BA136A7FFE9FFD1ACC5FD21FA17BA53" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FD21FA17BA53" pageId="16" pageNumber="211">16(14). Posterior pronotal angles at least slightly acute and sometimes embracing elytral bases; basal pronotal foveae absent; shortest distance between mesocoxal and metacoxal cavities distinctly less than length of abdominal ventrite 1; scutellar shield strongly transverse; hind wings absent........................................................................... 17</keyLead>
</paragraph>
<paragraph id="8BA136A7FFE9FFD1ACC5FC89FA17BABB" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FC89FA17BABB" pageId="16" pageNumber="211">- Posterior pronotal angles right or obtuse, not embracing elytral bases; basal pronotal foveae usually present; shortest distance between mesocoxal and metacoxal cavities distinctly greater than length of abdominal ventrite 1; scutellar shield slightly transverse; hind wings present.............................................................................. 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EA2E02FFE9FFD1ACC5FC71FA10BD4B" pageId="16" pageNumber="211">
<paragraph id="8BA136A7FFE9FFD1ACC5FC71FA10BAE3" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FC71FA10BAE3" pageId="16" pageNumber="211">
17(16). Mesoventrite carinate; posterior pronotal angles not or only slightly embracing elytral bases (
<figureCitation id="13252A22FFE9FFD1A8D7FC71FB58BAD8" box="[1157,1236,943,964]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="16" pageNumber="211">Fig. 1A</figureCitation>
); prosternal process extending well beyond procoxae, narrowly rounded at apex; prosternum in male with pubescent fovea (as in
<figureCitation id="13252A22FFE9FFD1A8BEFC13FACDBAFE" box="[1260,1345,973,994]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Fig. 14E</figureCitation>
); abdominal ventrite
<quantity id="4CE69B42FFE9FFD1AD08FC34FEF3BAE3" box="[346,383,1002,1023]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="211" unit="in" value="1.0">1 in</quantity>
male with pair of U-shaped pore fields (as in
<figureCitation id="13252A22FFE9FFD1AF65FC34FC17BAE3" box="[823,923,1002,1023]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="16" pageNumber="211">Figs 4B, 4</figureCitation>
E); Caucasus.................
<taxonomicName id="4C1E4D24FFE9FFD1A8B8FC34FA10BAE3" authority="Reitter" authorityName="Reitter" box="[1258,1436,1002,1023]" class="Insecta" family="Mycetophagidae" genus="Triphyllina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A8B8FC34FAD9BAE3" box="[1258,1365,1002,1023]" italics="true" pageId="16" pageNumber="211">Triphyllina</emphasis>
Reitter
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA136A7FFE9FFD1ACC5FBD9FA10BD4B" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FBD9FA10BD4B" pageId="16" pageNumber="211">
- Mesoventrite not carinae, with small subtriangular prosternal rest; posterior pronotal angles distinctly embracing elytra; prosternal process not extending beyond procoxae, truncate at apex; prosternum and abdominal ventrite
<quantity id="4CE69B42FFE9FFD1A8CDFBFBFB48BD26" box="[1183,1220,1061,1082]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="211" unit="in" value="1.0">1 in</quantity>
male simple;
<collectingCountry id="F3097637FFE9FFD1A919FBFBFA08BD26" box="[1355,1412,1061,1082]" name="China" pageId="16" pageNumber="211">China</collectingCountry>
................................................................................
<taxonomicName id="4C1E4D24FFE9FFD1A8D3FB9CFA10BD4B" authority="Nikitsky" authorityName="Nikitsky" box="[1153,1436,1090,1111]" class="Insecta" family="Mycetophagidae" genus="Pseudotriphyllopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A8D3FB9CFACFBD4B" box="[1153,1347,1090,1111]" italics="true" pageId="16" pageNumber="211">Pseudotriphyllopsis</emphasis>
Nikitsky
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EA2E02FFE9FFD1ACC5FB81FA17BC1B" pageId="16" pageNumber="211">
<paragraph id="8BA136A7FFE9FFD1ACC5FB81FA11BDD0" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5FB81FA11BDD0" pageId="16" pageNumber="211">
18(16). Antennomeres 9 and 10 slightly longer than wide and slightly expanded apically with sensory field at apex, 11 distinctly longer than wide and distinctly longer than 9 or 10 (
<figureCitation id="13252A22FFE9FFD1AEF6FBA3FD6CBD8E" box="[676,736,1149,1170]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Fig. 2</figureCitation>
F); mesoventrite more or less concave, with small anterior subquadrate prosternal rest; vertex in male with pubescent fovea (
<figureCitation id="13252A22FFE9FFD1AEB1FB44FCADBDB3" box="[739,801,1178,1199]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Figs 2</figureCitation>
E, 5M); prosternum and abdomen in male simple; southern North
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and Central
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..........................................................
<taxonomicName id="4C1E4D24FFE9FFD1A8E9FB69FAB3BDD0" box="[1211,1343,1207,1228]" class="Insecta" family="Mycetophagidae" genus="Neotriphyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96AEAB5FFE9FFD1A8E9FB69FAB3BDD0" box="[1211,1343,1207,1228]" italics="true" pageId="16" pageNumber="211">Neotriphyllus</emphasis>
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- Antennomeres 9 and 10 transverse (
<figureCitation id="13252A22FFE9FFD1AE6FFB0BFDFBBDF6" box="[573,631,1237,1258]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Fig. 2</figureCitation>
G); mesoventrite with linear or triangular prosternal rest (
<figureCitation id="13252A22FFE9FFD1A8F1FB0BFB73BDF6" box="[1187,1279,1237,1258]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Figs 14H</figureCitation>
,
<figureCitation id="13252A22FFE9FFD1A958FB0BFAAEBDF6" box="[1290,1314,1237,1258]" captionStart="FIGURES 15 A – C" captionStartId="31.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@31.[151,1436,193,1781]" captionTargetPageId="31" captionText="FIGURES 15 A – C, Typhaea stercorea (Linnaeus): A, pterothorax, ventral; B, head, dorsal; C, mesocoxa, showing lateral closure of cavity by meso- and metaventrites. Figures 15 D – G, Thrimolus minutus Casey: D, prothorax, ventral, with right coxa removed; E, pterothorax and base of abdomen, ventral; F, head, dorsal; G, left metacoxa and left portion of abdominal ventrite 1, with pore field vaguely indicated. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251859/files/figure.png" pageId="16" pageNumber="211">15</figureCitation>
E); vertex in male without pubescent fovea; abdomen in male with pore fields (
<figureCitation id="13252A22FFE9FFD1AF08FB2CFC4CBC1B" box="[858,960,1266,1287]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="16" pageNumber="211">Figs 4B, 4</figureCitation>
E).................................... 19
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19(18). Posterior pronotal angles broadly rounded (
<figureCitation id="13252A22FFE9FFD1AE2DFAD1FD41BC38" box="[639,717,1295,1316]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Figs 2B</figureCitation>
,
<figureCitation id="13252A22FFE9FFD1AE84FAD1FD62BC38" box="[726,750,1295,1316]" captionStart="FIGURES 15 A – C" captionStartId="31.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@31.[151,1436,193,1781]" captionTargetPageId="31" captionText="FIGURES 15 A – C, Typhaea stercorea (Linnaeus): A, pterothorax, ventral; B, head, dorsal; C, mesocoxa, showing lateral closure of cavity by meso- and metaventrites. Figures 15 D – G, Thrimolus minutus Casey: D, prothorax, ventral, with right coxa removed; E, pterothorax and base of abdomen, ventral; F, head, dorsal; G, left metacoxa and left portion of abdominal ventrite 1, with pore field vaguely indicated. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251859/files/figure.png" pageId="16" pageNumber="211">15</figureCitation>
D); mesoventrite with large triangular prosternal rest occupying most of ventrite (
<figureCitation id="13252A22FFE9FFD1AD68FAF3FE0EBC5E" box="[314,386,1325,1346]" captionStart="FIGURES 15 A – C" captionStartId="31.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@31.[151,1436,193,1781]" captionTargetPageId="31" captionText="FIGURES 15 A – C, Typhaea stercorea (Linnaeus): A, pterothorax, ventral; B, head, dorsal; C, mesocoxa, showing lateral closure of cavity by meso- and metaventrites. Figures 15 D – G, Thrimolus minutus Casey: D, prothorax, ventral, with right coxa removed; E, pterothorax and base of abdomen, ventral; F, head, dorsal; G, left metacoxa and left portion of abdominal ventrite 1, with pore field vaguely indicated. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251859/files/figure.png" pageId="16" pageNumber="211">Fig. 15</figureCitation>
E); abdominal ventrite
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male with very slender, curved linear pore fields extending from anterolateral angles to mesal fourth (
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G); length less than
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; North and Central
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.................
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<emphasis id="B96AEAB5FFE9FFD1A8AAFA94FAD7BC43" box="[1272,1371,1354,1375]" italics="true" pageId="16" pageNumber="211">Thrimolus</emphasis>
Casey
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- Posterior pronotal angles not broadly rounded (
<figureCitation id="13252A22FFE9FFD1AEF1FAB9FD52BC60" box="[675,734,1383,1404]" captionStart="FIGURES 3 A – B" captionStartId="19.[151,264,1802,1825]" captionTargetBox="[151,1431,193,1778]" captionTargetId="figure@19.[151,1436,193,1781]" captionTargetPageId="19" captionText="FIGURES 3 A – B, portion of elytral surface: A, Typhaea stercorea (Linnaeus); B, Nototriphyllus araucania sp. n. Figure 3 C, Zeclaviger explanatus sp. n., pterothorax, ventral, with right metanepisternum and right metacoxa and hind leg removed. Figure 3 D, Nototriphyllus araucania sp. n., prothorax, dorsal, with both coxae removed. Figure 3 E, Zeclaviger explanatus sp. n., left halves of abdominal ventrites 1 and 2, showing pore field in anterolateral corner of ventrite 1. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251847/files/figure.png" pageId="16" pageNumber="211">Figs 3</figureCitation>
D, 14E); mesoventrite with linear or small, triangular prosternal rest (
<figureCitation id="13252A22FFE9FFD1A927FAB9FE82BC86" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Fig. 14H</figureCitation>
); abdominal ventrite
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male with shorter and broader linear or C-shaped pore fields (
<figureCitation id="13252A22FFE9FFD1A836FA5BFB58BC86" box="[1124,1236,1413,1434]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="16" pageNumber="211">Figs 4A–B</figureCitation>
); length greater than
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; Holarctic and Notogean........................................................................ 20
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20(19). Antennomeres 9 and 10 slightly transverse, slightly expanded apically, with sensory fields at apex; anterior edge of mesoventrite with 2 small carinae; prosternum in male without setose fovea; abdominal ventrites
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male each with pair of Vshaped or linear pore fields; habitus (
<figureCitation id="13252A22FFE9FFD1AE16FA24FD02BF13" box="[580,654,1530,1551]" captionStart="FIGURES 1 A – L" captionStartId="17.[151,264,1788,1811]" captionTargetBox="[151,1426,206,1773]" captionTargetId="figure@17.[151,1436,195,1783]" captionTargetPageId="17" captionText="FIGURES 1 A – L, habitus, dorsal: A, Triphyllina lederi Reitter, length 2.0 mm, © K. V. Makarov; B, Pseudotriphyllus colchicus (Reitter), length 2.3 mm, © K. V. Makarov; C, Nesolathrus sp. (Australia: QLD: McIlwraith Range), length 1.2 mm, © CSIRO Australia; D, Eulagius ussuriensis Nikitsky, length 4 mm, © K. V. Makarov; E, Mycetophagus decempunctatus Fabricius, length 4.2 mm, © K. V. Makarov; F, Mycetophagus irroratus Reitter, length 3.8 mm, © K. V. Makarov; G, Pseudochrodes suturalis Reitter, length 2.5 mm, © CSIRO Australia; H, Typhaea stercorea (Linnaeus), length 3.1 mm, © CSIRO Australia; I, Litargops maculosus Reitter, length 3.4 mm, © K. V. Makarov; J, Litargus connexus Geoffroy, length 3.5 mm, © K. V. Makarov; K, Berginus sp. (Panama: Paraiso), length 1.4 mm, © CSIRO Australia; L, Triphyllus bicolor Fabricius, length 3.9 mm, © K. V. Makarov." httpUri="https://zenodo.org/record/251845/files/figure.png" pageId="16" pageNumber="211">Fig. 1L</figureCitation>
); Eurasia...........................................
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Dejean
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- Antennomeres 9 and 10 distinctly transverse and widest at middle, with sensory fields often occupying half of antennomere (
<figureCitation id="13252A22FFE9FFD1ACB5F9EBFEAFBF56" box="[231,291,1589,1610]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Figs 2</figureCitation>
G, 14B); anterior edge of mesoventrite with short linear or narrowly triangular elevation (
<figureCitation id="13252A22FFE9FFD1A82AF9EBFB5DBF56" box="[1144,1233,1589,1610]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Fig. 14H</figureCitation>
); prosternum in male with setose fovea (
<figureCitation id="13252A22FFE9FFD1ADC6F98CFE67BF7B" box="[404,491,1618,1639]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Fig. 14E</figureCitation>
); abdomen with U-shaped or linear pore fields on ventrite 1 only (
<figureCitation id="13252A22FFE9FFD1A837F98CFB46BF7B" box="[1125,1226,1618,1639]" captionStart="FIGURES 4 A – C" captionStartId="20.[151,264,1744,1767]" captionTargetBox="[152,1431,193,1718]" captionTargetId="figure@20.[151,1436,193,1723]" captionTargetPageId="20" captionText="FIGURES 4 A – C, right half of abdominal ventrite 1: A, Afrotyphaeola natalensis sp. n. with linear pore field; B, Nototriphyllus araucania sp. n., with C-shaped pore field; C, Typhaeola maculata Perris, with pores at setal bases. FIGURES 4 D – F, all five abdominal ventrites of same three species. Figure 4 G, Nototriphyllus araucania sp. n., hind wing. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251848/files/figure.png" pageId="16" pageNumber="211">Figs 4B, 4</figureCitation>
E)............. 21
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<keyLead id="30EF9592FFE9FFD1ACC5F9B1FA10BFBE" pageId="16" pageNumber="211">
21(20). Antennomere 11 distinctly narrower than 9 or 10 (
<figureCitation id="13252A22FFE9FFD1AE91F9B1FD71BF98" box="[707,765,1647,1668]" captionStart="FIGURES 2 A – B" captionStartId="18.[151,264,1802,1825]" captionTargetBox="[151,1421,207,1780]" captionTargetId="figure@18.[151,1436,193,1781]" captionTargetPageId="18" captionText="FIGURES 2 A – B, habitus, dorsal: A, Afrotyphaeola natalensis sp. n., length 1.6 mm; B, Thrimolus minutus Casey, length 1.2 mm. FIGURES 2 C – D, Litargus balteatus LeConte, prothoraces: C, dorsal, showing paired slit-like impressions; D, posterior, showing notal attachments of posterior apodemes and cryptopleura. Figure 2 E, Neotriphyllus confusus (Horn), head, dorsal. FIGURES 2 F – H, antennal apex: F, Neotriphyllus confusus (Horn); G, Pseudotriphyllus suturalis Reitter; H, Zeclaviger explanatus sp. n. Figure 2 I, Litargops sp. (Australia: ACT: Picadilly Circus, Brindabella Range), protibial apex. FIGURES 2 J – K, Litargus balteatus LeConte: J, labium; K, maxilla. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251846/files/figure.png" pageId="16" pageNumber="211">Fig. 2</figureCitation>
G); prosternal process parallel-sided, slightly emarginate at apex; procoxal cavity without slit-like extension; Eurasia and eastern North
<collectingCountry id="F3097637FFE9FFD1AF32F953FC3EBFBE" box="[864,946,1677,1698]" name="United States of America" pageId="16" pageNumber="211">America</collectingCountry>
....................
<taxonomicName id="4C1E4D24FFE9FFD1A8E0F953FA10BFBE" ID-CoL="73XD" authority="Reitter" authorityName="Reitter" box="[1202,1436,1677,1698]" class="Insecta" family="Mycetophagidae" genus="Pseudotriphyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB5FFE9FFD1A8E0F953FAD9BFBE" box="[1202,1365,1677,1698]" italics="true" pageId="16" pageNumber="211">Pseudotriphyllus</emphasis>
Reitter
</taxonomicName>
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</paragraph>
<paragraph id="8BA136A7FFE9FFD1ACC5F974FA10BFE6" blockId="16.[151,1437,151,1786]" pageId="16" pageNumber="211">
<keyLead id="30EF9592FFE9FFD1ACC5F974FA10BFE6" pageId="16" pageNumber="211">
- Antennomeres 9–11 of equal width; prosternal process slightly expanded almost to apex, then narrowed and apically truncate (
<figureCitation id="13252A22FFE9FFD1ACBAF919FEB3BFC0" box="[232,319,1735,1756]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Fig. 14E</figureCitation>
); procoxal cavity with short, curved, slit-like extension (
<figureCitation id="13252A22FFE9FFD1AF0EF919FC3FBFC0" box="[860,947,1735,1756]" captionStart="FIGURES 14 A – H" captionStartId="30.[151,264,1802,1825]" captionTargetBox="[151,1436,193,1779]" captionTargetId="figure@30.[151,1436,193,1781]" captionTargetPageId="30" captionText="FIGURES 14 A – H, Nototriphyllus araucania sp. n.: A, head, dorsal; B, antenna; C, head, ventral; D, scutellar shield; E, male prothorax, ventral with left coxa removed, showing prosternal sex patch; F, right elytron, dorsal; G, mesotibial apex and mesotarsus; H, pterothorax, ventral. All figures © CSIRO Australia." httpUri="https://zenodo.org/record/251858/files/figure.png" pageId="16" pageNumber="211">Fig. 14E</figureCitation>
);
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,
<collectingCountry id="F3097637FFE9FFD1A803F919FB48BFC0" box="[1105,1220,1735,1756]" name="Madagascar" pageId="16" pageNumber="211">Madagascar</collectingCountry>
and
<collectingCountry id="F3097637FFE9FFD1A8A1F919FAABBFC0" box="[1267,1319,1735,1756]" name="Chile" pageId="16" pageNumber="211">Chile</collectingCountry>
.................................................................................
<taxonomicName id="4C1E4D24FFE9FFD1A8E3F93BFAB7BFE6" box="[1201,1339,1765,1786]" class="Insecta" family="Mycetophagidae" genus="Nototriphyllus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="211" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96AEAB5FFE9FFD1A8E3F93BFAB7BFE6" box="[1201,1339,1765,1786]" italics="true" pageId="16" pageNumber="211">Nototriphyllus</emphasis>
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<emphasis id="B96AEAB5FFE9FFD1A910F93AFA10BFE6" bold="true" box="[1346,1436,1764,1786]" pageId="16" pageNumber="211">gen. nov.</emphasis>
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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