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<document id="261E76E61C3135D3C114058CFA28AC3A" ID-CLB-Dataset="26897" ID-DOI="10.11646/zootaxa.4603.1.7" ID-GBIF-Dataset="100f9b88-b979-4a43-ad71-5ebaaf477b2d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2673381" ID-ZooBank="49EB4D9F-89AD-462B-8879-EA258C810B15" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557294626015" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03B787B0FFB48C4178C6FCD5FBCF4ABA" docLanguage="en" docName="zootaxa.4603.1.7.pdf" docOrigin="Zootaxa 4603 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Catalinus Casey 1897" docType="treatment" docVersion="7" lastPageNumber="151" masterDocId="FF8EFFC8FFB58C477851FFA8FF9A4C59" masterDocTitle="Notes on Nearctic genera Catalinus Casey and Parascydmus Casey (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="158" masterPageNumber="145" pageNumber="146" updateTime="1698726535999" updateUser="ExternalLinkService">
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<mods:title id="4E2E95A49787856BB01BE2BA72666D71">Notes on Nearctic genera Catalinus Casey and Parascydmus Casey (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="F05ECA60C5C629832DABFBEC8EA76FE4">Jałoszyński, Paweł</mods:namePart>
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<treatment id="03B787B0FFB48C4178C6FCD5FBCF4ABA" ID-GBIF-Taxon="157063823" LSID="urn:lsid:plazi:treatment:03B787B0FFB48C4178C6FCD5FBCF4ABA" httpUri="http://treatment.plazi.org/id/03B787B0FFB48C4178C6FCD5FBCF4ABA" lastPageId="6" lastPageNumber="151" pageId="1" pageNumber="146">
<subSubSection id="C304652DFFB48C4678C6FCD5FE2C4FCE" box="[151,438,893,920]" pageId="1" pageNumber="146" type="nomenclature">
<paragraph id="8BA136A6FFB48C4678C6FCD5FE2C4FCE" blockId="1.[151,438,893,920]" box="[151,438,893,920]" pageId="1" pageNumber="146">
<heading id="D0E981CAFFB48C4678C6FCD5FE2C4FCE" bold="true" box="[151,438,893,920]" fontSize="11" level="1" pageId="1" pageNumber="146" reason="1">
<emphasis id="B96AEAB4FFB48C4678C6FCD5FE2C4FCE" bold="true" box="[151,438,893,920]" pageId="1" pageNumber="146">
Genus
<taxonomicName id="4C1E4D25FFB48C4678BFFCD5FE2C4FCE" ID-CoL="8XK9B" authority="Casey" authorityName="Casey" authorityYear="1897" box="[238,438,893,920]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB48C4678BFFCD5FEF94FC1" bold="true" box="[238,355,893,920]" italics="true" pageId="1" pageNumber="146">Catalinus</emphasis>
Casey
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304652DFFB48C4678C6FC6AFBC44F83" box="[151,1118,961,986]" pageId="1" pageNumber="146" type="reference_group">
<paragraph id="8BA136A6FFB48C4678C6FC6AFBC44F83" blockId="1.[151,1118,961,986]" box="[151,1118,961,986]" pageId="1" pageNumber="146">
<treatmentCitationGroup id="AB0E1188FFB48C4678C6FC6AFBC44F83" box="[151,1118,961,986]" pageId="1" pageNumber="146">
<taxonomicName id="4C1E4D25FFB48C4678C6FC6AFE284F83" ID-CoL="8XK9B" authority="Casey, 1897: 492" authorityName="Casey" authorityPageNumber="492" authorityYear="1897" box="[151,434,962,986]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB48C4678C6FC6AFF604F83" box="[151,250,962,986]" italics="true" pageId="1" pageNumber="146">Catalinus</emphasis>
<treatmentCitation id="0ABF10B7FFB48C467951FC6AFE284F83" author="Casey, T. L." box="[256,434,962,986]" page="492" pageId="1" pageNumber="146" year="1897">
<bibRefCitation id="EF8F4B57FFB48C467951FC6AFEE54F83" author="Casey, T. L." box="[256,383,962,986]" pageId="1" pageNumber="146" pagination="285 - 684" refId="ref7607" refString="Casey, T. L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Sciences, 9, 285 - 684. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" type="journal article" year="1897">Casey, 1897</bibRefCitation>
: 492
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54A58804FFB48C4679EAFC6AFE774F83" box="[443,493,962,986]" pageId="1" pageNumber="146">Type</typeStatus>
species:
<taxonomicName id="4C1E4D25FFB48C467A1BFC6AFC434F83" ID-CoL="6XZT7" authority="LeConte, 1852" authorityName="LeConte" authorityYear="1852" box="[586,985,961,986]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="angustulus">
<emphasis id="B96AEAB4FFB48C467A1BFC6AFCA64F83" box="[586,828,962,986]" italics="true" pageId="1" pageNumber="146">Scydmaenus angustulus</emphasis>
<bibRefCitation id="EF8F4B57FFB48C467B13FC6AFC434F83" author="LeConte, J. L." box="[834,985,961,986]" pageId="1" pageNumber="146" pagination="149 - 157" refId="ref8464" refString="LeConte, J. L. (1852) Synopsis of the Scydmaenidae of the United States. Proceedings of the Academy of Natural Scences, Philiadelphia, 6, 149 - 157." type="journal article" year="1852">LeConte, 1852</bibRefCitation>
</taxonomicName>
(monotypy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C304652DFFB48C4678C6FBAFFE094A42" pageId="1" pageNumber="146" type="diagnosis">
<paragraph id="8BA136A6FFB48C4678C6FBAFFE094A42" blockId="1.[151,1437,1031,2031]" pageId="1" pageNumber="146">
<emphasis id="B96AEAB4FFB48C4678C6FBAFFEE9487B" bold="true" box="[151,371,1031,1058]" pageId="1" pageNumber="146">Revised diagnosis.</emphasis>
Body slender, with distinct constrictions between head and pronotum and between pronotum and elytra; head and prothorax lacking thick bristles; head subtriangular, not bulging posterodorsad; eyes located in posterior portion of head; mandible subtriangular; antennae gradually thickened; lateral sutures of submentum present, their posterior ends broadly separated; hypomeral ridges complete; posterior tentorial pits hidden in transverse groove demarcating ventrally ‘neck’ region; occipital constriction only slightly narrower than vertex; pronotum broadest distinctly in front of middle, lacking lateral and sublateral carinae, antebasal pits and grooves; procoxal cavities closed; prosternal process narrowly carinate, diffuse, weakly elevated, hidden between procoxae; basisternal part of prosternum much shorter than coxal part; hypomeral ridges marked only near anterior margins of mesocoxae, posteriorly obliterated; notosternal sutures complete; mesoventrite with asetose procoxal rests, deep setose impressions behind the asetose area and carinate mesoventral intercoxal process extending posteriorly to the middle of mesocoxae; anterior metaventral process absent; metacoxae subcontiguous, i.e., narrowly separated by short metaventral intercoxal process with deep median notch dividing it into a pair of short, blunt subtriangular projections; each elytron with one large, deep and asetose basal fovea near suture; mesoscutellum not exposed; aedeagus with symmetrical median lobe, asymmetrical endophallic sclerites and presumably with parameres (broken off in the studied specimen).
</paragraph>
</subSubSection>
<subSubSection id="C304652DFFB48C437896F98BFE4149AA" lastPageId="4" lastPageNumber="149" pageId="1" pageNumber="146" type="description">
<paragraph id="8BA136A6FFB48C467896F98BFD954ADE" blockId="1.[151,1437,1031,2031]" pageId="1" pageNumber="146">
<emphasis id="B96AEAB4FFB48C467896F98BFEEF4A67" bold="true" box="[199,373,1571,1598]" pageId="1" pageNumber="146">Redescription.</emphasis>
Body (
<figureCitation id="13252A23FFB48C467996F98CFD9D4A67" box="[455,519,1572,1598]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1983,2008]" captionTargetBox="[169,1422,735,1935]" captionTargetId="figure@2.[164,1423,735,1935]" captionTargetPageId="2" captionText="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="1" pageNumber="146">Fig. 1</figureCitation>
) elongate and slender, flattened, with deep constrictions between head and pronotum and between pronotum and elytra, brown and covered with setae, without thick bristles, adult body length of the only known species
<quantity id="4CE69B43FFB48C4679F5F9C4FD914ADE" box="[420,523,1644,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="146" unit="mm" value="1.1">1.10 mm</quantity>
.
</paragraph>
<paragraph id="8BA136A6FFB48C467896F938FE814B07" blockId="1.[151,1437,1031,2031]" pageId="1" pageNumber="146">
Head capsule (
<figureCitation id="13252A23FFB48C46793EF938FE4F4AF3" box="[367,469,1680,1706]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStartId-0="2.[151,264,1983,2008]" captionStartId-1="3.[151,264,1341,1366]" captionTargetBox-0="[169,1422,735,1935]" captionTargetBox-1="[153,1435,192,1300]" captionTargetId-0="figure@2.[164,1423,735,1935]" captionTargetId-1="figure@3.[151,1436,192,1300]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." captionText-1="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri-0="https://zenodo.org/record/2673383/files/figure.png" httpUri-1="https://zenodo.org/record/2673385/files/figure.png" pageId="1" pageNumber="146">Figs 2–3</figureCitation>
) with anterior part (in front of occiput) subtriangular in shape. Occipital constriction slightly narrower than vertex; tempora subequal to eyes, rounded; vertex transverse and convex, lacking posterior ridge, not bulging posterodorsad, anteriorly confluent with frons, which is subtrapezoidal and anteriorly narrowing; frontoclypeal groove present, short; anterior margin of clypeus rounded; supraantennal tubercles distinct but weakly elevated; antennal insertions broadly separated. Eyes large, coarsely faceted, weakly projecting from the silhouette of the head.
</paragraph>
<paragraph id="8BA136A6FFB48C467896F8C0FC7D4BFF" blockId="1.[151,1437,1031,2031]" pageId="1" pageNumber="146">
Gular plate (
<figureCitation id="13252A23FFB48C467904F8C0FE014BDB" box="[341,411,1896,1922]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="1" pageNumber="146">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB48C4679F9F8C1FE5E4BDB" box="[424,452,1897,1922]" italics="true" pageId="1" pageNumber="146">gp</emphasis>
) large and subtrapezoidal; gular sutures (
<figureCitation id="13252A23FFB48C467BC8F8C0FC444BDB" box="[921,990,1896,1922]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="1" pageNumber="146">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB48C467BBAF8C1FB9E4BDB" box="[1003,1028,1897,1922]" italics="true" pageId="1" pageNumber="146">gs</emphasis>
) distinct; posterior tentorial pits not exposed, hidden in a transverse groove demarcating ‘neck’ region ventrally.
</paragraph>
<paragraph id="8BA136A6FFB48C457896F818FC1A4D8A" blockId="1.[151,1437,1031,2031]" lastBlockId="2.[151,1436,153,683]" lastPageId="2" lastPageNumber="147" pageId="1" pageNumber="146">
Labrum transverse with anterior margin arcuate. Mandibles (
<figureCitation id="13252A23FFB48C467B24F818FC224B93" box="[885,952,1968,1994]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="1" pageNumber="146">Fig. 3</figureCitation>
) poorly visible in the only available specimen, appear symmetrical, each subtriangular, mesal margin not exposed. Maxilla (
<figureCitation id="13252A23FFB48C467C7FF87CFBEA4BB7" box="[1070,1136,2004,2030]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="1" pageNumber="146">Fig. 3</figureCitation>
) generalized, as in all gen- era of
<taxonomicName id="4C1E4D25FFB78C45788FFF31FEEC4CEA" box="[222,374,153,179]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
, with moderately long maxillary palp composed of minute and distinctly elongate palpomere I, strongly elongate, pedunculate palpomere II, broad palpomere III broadest near distal third, and strongly elongate and slender subconical and pointed palpomere IV longer than half length of palpomere III. Labium (
<figureCitation id="13252A23FFB78C457D45FF49FAC24CA2" box="[1300,1368,225,251]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="2" pageNumber="147">Fig. 3</figureCitation>
) with trapezoidal submentum (
<figureCitation id="13252A23FFB78C4579FFFEADFE694D46" box="[430,499,261,287]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="2" pageNumber="147">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB78C457A51FEAEFDB74D46" box="[512,557,262,287]" italics="true" pageId="2" pageNumber="147">smn</emphasis>
) demarcated laterally from postcardinal regions by lateral sutures (
<figureCitation id="13252A23FFB78C457D72FEADFAF24D46" box="[1315,1384,261,287]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="2" pageNumber="147">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB78C457D24FEADFA094D46" box="[1397,1427,261,287]" italics="true" pageId="2" pageNumber="147">lss</emphasis>
) which are strongly convergent posteriorly but their ends are broadly separated; mentum (
<figureCitation id="13252A23FFB78C457C20FE81FB2E4D1A" box="[1137,1204,297,323]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="2" pageNumber="147">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB78C457C91FE82FB784D1A" box="[1216,1250,298,323]" italics="true" pageId="2" pageNumber="147">mn</emphasis>
) subtrapezoidal, transverse; prementum short, bearing labial palps with all palpomeres distinctly elongate (palpomere II the longest); anteromedian region of prementum not exposed in the studied specimen. Hypostomal ridges (
<figureCitation id="13252A23FFB78C457CF5FED9FB724DD2" box="[1188,1256,369,395]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="2" pageNumber="147">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB78C457CA2FED9FA964DD2" box="[1267,1292,369,395]" italics="true" pageId="2" pageNumber="147">hr</emphasis>
) distinct and complete, extending posteromesad up to posterior ends of lateral sutures of submentum and posteriorly confluent with posteroventral margin of the anterior portion of head capsule.
</paragraph>
<paragraph id="8BA136A6FFB78C457896FE75FC394DAE" blockId="2.[151,1436,153,683]" box="[199,931,477,503]" pageId="2" pageNumber="147">
Antennae (
<figureCitation id="13252A23FFB78C457913FE75FE184DAE" box="[322,386,477,503]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1983,2008]" captionTargetBox="[169,1422,735,1935]" captionTargetId="figure@2.[164,1423,735,1935]" captionTargetPageId="2" captionText="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="2" pageNumber="147">Fig. 1</figureCitation>
) slender and weakly gradually thickened distad.
</paragraph>
<paragraph id="8BA136A6FFB78C457896FDA9FE284EF2" blockId="2.[151,1436,153,683]" pageId="2" pageNumber="147">
Pronotum (
<figureCitation id="13252A23FFB78C457916FDA9FE374E42" box="[327,429,513,539]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1983,2008]" captionTargetBox="[169,1422,735,1935]" captionTargetId="figure@2.[164,1423,735,1935]" captionTargetPageId="2" captionText="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="2" pageNumber="147">Figs 1–2</figureCitation>
) in dorsal view approximately bell-shaped, strongly elongate and broadest distinctly in front of middle; anterior margin arcuate and laterally confluent with sides of pronotum without any indication of anterior corners; lateral margins rounded in anterior half and indistinctly sinuate posteriorly; posterior corners indistinct, broadly rounded, obtuse-angled; posterior margin arcuate. Pronotal base lacking pits and grooves; lateral and sublateral carinae absent.
</paragraph>
<caption id="DF61662EFFB78C4578C6F817FD5D4BA5" httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="2" pageNumber="147" startId="2.[151,264,1983,2008]" targetBox="[169,1422,735,1935]" targetPageId="2">
<paragraph id="8BA136A6FFB78C4578C6F817FD5D4BA5" blockId="2.[151,1436,1983,2044]" pageId="2" pageNumber="147">
<emphasis id="B96AEAB4FFB78C4578C6F817FEA04B81" bold="true" box="[151,314,1983,2008]" pageId="2" pageNumber="147">FIGURES 1–2.</emphasis>
Morphological structures of
<taxonomicName id="4C1E4D25FFB78C457A0EF868FCBB4B81" box="[607,801,1984,2008]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B96AEAB4FFB78C457A0EF868FCBB4B81" box="[607,801,1984,2008]" italics="true" pageId="2" pageNumber="147">Catalinus angustus</emphasis>
</taxonomicName>
. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea.
</paragraph>
</caption>
<caption id="DF61662EFFB68C4478C6FA95FB77499B" httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148" startId="3.[151,264,1341,1366]" targetBox="[153,1435,192,1300]" targetPageId="3">
<paragraph id="8BA136A6FFB68C4478C6FA95FB77499B" blockId="3.[151,1436,1341,1474]" pageId="3" pageNumber="148">
<emphasis id="B96AEAB4FFB68C4478C6FA95FEA0490F" bold="true" box="[151,314,1341,1366]" pageId="3" pageNumber="148">FIGURES 3–4.</emphasis>
Morphological structures of
<taxonomicName id="4C1E4D25FFB68C447A31FA96FCB8490F" box="[608,802,1342,1366]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B96AEAB4FFB68C447A31FA96FCB8490F" box="[608,802,1342,1366]" italics="true" pageId="3" pageNumber="148">Catalinus angustus</emphasis>
</taxonomicName>
. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum.
</paragraph>
</caption>
<paragraph id="8BA136A6FFB68C447896FA59FAEF4AC3" blockId="3.[151,1437,1520,2015]" pageId="3" pageNumber="148">
Prosternum (
<figureCitation id="13252A23FFB68C447909FA58FE064A52" box="[344,412,1520,1547]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
) with basisternal part (
<figureCitation id="13252A23FFB68C447AF3FA58FD7D4A52" box="[674,743,1520,1547]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447AA5FA59FC8F4A52" box="[756,789,1521,1547]" italics="true" pageId="3" pageNumber="148">bst</emphasis>
) much shorter than coxal part (but not vestigial) and distinctly demarcated from procoxal cavities; median portion of sternum with weakly elevated and diffuse longitudinal carina, in intact specimens hidden between procoxae (
<figureCitation id="13252A23FFB68C447B5FF990FCCF4A0A" box="[782,853,1592,1619]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
); procoxal sockets presumably closed; notosternal sutures (
<figureCitation id="13252A23FFB68C4478A4F9F4FEA04A2E" box="[245,314,1628,1655]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447917F9F5FEF04A2F" box="[326,362,1629,1654]" italics="true" pageId="3" pageNumber="148">nss</emphasis>
) complete, each posteriorly overlapped by inner portion of prothoracic hypomeron which forms a lobe projecting mesad; hypomeral ridges (
<figureCitation id="13252A23FFB68C447AD2F928FD524AC2" box="[643,712,1664,1691]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447A85F929FD634AC2" box="[724,761,1665,1691]" italics="true" pageId="3" pageNumber="148">hyr</emphasis>
) incomplete, marked only anteriorly, largely obliterated.
</paragraph>
<paragraph id="8BA136A6FFB68C447896F90CFC4F4AE7" blockId="3.[151,1437,1520,2015]" box="[199,981,1700,1727]" pageId="3" pageNumber="148">Mesoscutellar shield not visible between elytral bases in intact beetle.</paragraph>
<paragraph id="8BA136A6FFB68C447896F960FC284B17" blockId="3.[151,1437,1520,2015]" pageId="3" pageNumber="148">
Mesoventrite (
<figureCitation id="13252A23FFB68C447938F960FE314ABA" box="[361,427,1736,1763]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 4</figureCitation>
) with narrow anterior ridge (
<figureCitation id="13252A23FFB68C447ABAF960FCB54ABA" box="[747,815,1736,1763]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 4</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447B6BF961FCC94ABB" box="[826,851,1737,1762]" italics="true" pageId="3" pageNumber="148">ar</emphasis>
); large, asetose transverse area not divided at middle and functioning as procoxal rest (
<figureCitation id="13252A23FFB68C447A4FF944FDFC4B5E" box="[542,614,1772,1799]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447A27F945FD014B5F" box="[630,667,1773,1798]" italics="true" pageId="3" pageNumber="148">pcr</emphasis>
), and behind it with a pair of large, transverse setose impressions separated by a carinate mesoventral intercoxal process (
<figureCitation id="13252A23FFB68C447B54F8B8FCD34B72" box="[773,841,1808,1835]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447B04F8B9FC154B73" box="[853,911,1809,1834]" italics="true" pageId="3" pageNumber="148">msvp</emphasis>
), the latter moderately elevated (not keel-like), posteriorly reaching middle of mesocoxae, with a distinct posterior tip.
</paragraph>
<paragraph id="8BA136A6FFB68C447896F8F1FE494B2B" blockId="3.[151,1437,1520,2015]" box="[199,467,1880,1907]" pageId="3" pageNumber="148">Metanotum not studied.</paragraph>
<paragraph id="8BA136A6FFB68C447896F8D4FB574B86" blockId="3.[151,1437,1520,2015]" pageId="3" pageNumber="148">
Metaventrite (
<figureCitation id="13252A23FFB68C447939F8D4FE374BCE" box="[360,429,1916,1943]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
) subtrapezoidal, distinctly broadening posteriorly, lacking anterior metaventral process; metaventral intercoxal process (
<figureCitation id="13252A23FFB68C447A53F808FDD24BE2" box="[514,584,1952,1979]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="3" pageNumber="148">Fig. 3</figureCitation>
;
<emphasis id="B96AEAB4FFB68C447A07F809FD164BE3" box="[598,652,1953,1978]" italics="true" pageId="3" pageNumber="148">mtvp</emphasis>
) narrow, subtrapezoidal, with a deep median notch dividing it into a pair of short lateral subtriangular and blunt projections. Metanepisterna and metepimera narrow.
</paragraph>
<paragraph id="8BA136A6FFB18C437896FF31FD324CEA" blockId="4.[151,1436,153,539]" box="[199,680,153,180]" pageId="4" pageNumber="149">Metendosternite (= metafurca) not studied.</paragraph>
<paragraph id="8BA136A6FFB18C437896FF15FD434CA2" blockId="4.[151,1436,153,539]" pageId="4" pageNumber="149">
Elytra (
<figureCitation id="13252A23FFB18C43794CFF15FE1B4C8E" box="[285,385,189,215]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1983,2008]" captionTargetBox="[169,1422,735,1935]" captionTargetId="figure@2.[164,1423,735,1935]" captionTargetPageId="2" captionText="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="4" pageNumber="149">Figs 1–2</figureCitation>
) oval, lacking humeral calli and basal impressions, each with one distinct and deep asetose basal fovea (
<figureCitation id="13252A23FFB18C437974FF49FEF14CA2" box="[293,363,225,251]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1983,2008]" captionTargetBox="[169,1422,735,1935]" captionTargetId="figure@2.[164,1423,735,1935]" captionTargetPageId="2" captionText="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="4" pageNumber="149">Fig. 2</figureCitation>
;
<emphasis id="B96AEAB4FFB18C437927FF49FE024CA2" box="[374,408,225,251]" italics="true" pageId="4" pageNumber="149">bef</emphasis>
), apices separately rounded.
</paragraph>
<paragraph id="8BA136A6FFB18C437896FEADFE494D46" blockId="4.[151,1436,153,539]" box="[199,467,261,287]" pageId="4" pageNumber="149">Hind wings not studied.</paragraph>
<paragraph id="8BA136A6FFB18C437896FE81FC464DD2" blockId="4.[151,1436,153,539]" pageId="4" pageNumber="149">
Legs (
<figureCitation id="13252A23FFB18C43795CFE81FECE4D1A" box="[269,340,297,323]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1983,2008]" captionTargetBox="[169,1422,735,1935]" captionTargetId="figure@2.[164,1423,735,1935]" captionTargetPageId="2" captionText="FIGURES 1–2. Morphological structures of Catalinus angustus. Dorsal habitus of male (1); head, prothorax and elytral base in dorsal view (2). Abbreviations: bef, basal elytral fovea." httpUri="https://zenodo.org/record/2673383/files/figure.png" pageId="4" pageNumber="149">Figs 1</figureCitation>
,
<figureCitation id="13252A23FFB18C437932FE81FE144D1A" box="[355,398,297,323]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1341,1366]" captionTargetBox="[153,1435,192,1300]" captionTargetId="figure@3.[151,1436,192,1300]" captionTargetPageId="3" captionText="FIGURES 3–4. Morphological structures of Catalinus angustus. Head, prothorax and anterior portion of pterothorax in ventral view (3); pterothorax in ventral view (4). Abbreviations: ar, anterior ridge; bst, basisternal portion of prosternum; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lss, lateral suture of labium; mn, mentum; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; smn, submentum." httpUri="https://zenodo.org/record/2673385/files/figure.png" pageId="4" pageNumber="149">3–4</figureCitation>
) long and slender; pro- and mesocoxae in ventral view oval, metacoxae strongly transverse, laterally reaching lateral margins of metaventrite; all trochanters short, subtriangular; all femora moderately strongly and gradually clavate; tibiae straight or nearly so; tarsi long and slender.
</paragraph>
<paragraph id="8BA136A6FFB18C437896FE3DFDA34DF6" blockId="4.[151,1436,153,539]" box="[199,569,405,431]" pageId="4" pageNumber="149">Abdominal sternites unmodified.</paragraph>
<paragraph id="8BA136A6FFB18C437896FE11FCE64E42" blockId="4.[151,1436,153,539]" pageId="4" pageNumber="149">
Aedeagus (
<figureCitation id="13252A23FFB18C437916FE11FE374D8A" box="[327,429,440,467]" captionStart="FIGURES 5–7" captionStartId="4.[151,264,1390,1415]" captionTargetBox="[169,1400,613,1315]" captionTargetId="figure@4.[151,1436,598,1327]" captionTargetPageId="4" captionText="FIGURES 5–7. Aedeagus of Catalinus angustus in ventral (5) and lateral (6) views and original set of labels of the presumable male syntype (7)." httpUri="https://zenodo.org/record/2673387/files/figure.png" pageId="4" pageNumber="149">Figs 5–6</figureCitation>
) with symmetrical median lobe, its apical region undivided, endophallus asymmetrical, with a group of elongate sclerites; parameres presumably present, but in the only available specimen broken off during a previous preparation, dorsal foramen in sub-basal region.
</paragraph>
<caption id="DF61662EFFB18C4378C6FAC6FED249F2" httpUri="https://zenodo.org/record/2673387/files/figure.png" pageId="4" pageNumber="149" startId="4.[151,264,1390,1415]" targetBox="[169,1400,613,1315]" targetPageId="4">
<paragraph id="8BA136A6FFB18C4378C6FAC6FED249F2" blockId="4.[151,1436,1390,1451]" pageId="4" pageNumber="149">
<emphasis id="B96AEAB4FFB18C4378C6FAC6FEA049DE" bold="true" box="[151,314,1390,1415]" pageId="4" pageNumber="149">FIGURES 5–7.</emphasis>
Aedeagus of
<taxonomicName id="4C1E4D25FFB18C437994FAC7FD1D49DE" box="[453,647,1391,1415]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B96AEAB4FFB18C437994FAC7FD1D49DE" box="[453,647,1391,1415]" italics="true" pageId="4" pageNumber="149">Catalinus angustus</emphasis>
</taxonomicName>
in ventral (5) and lateral (6) views and original set of labels of the presumable male syntype (7).
</paragraph>
</caption>
<paragraph id="8BA136A6FFB18C437896FA70FE4149AA" blockId="4.[151,1437,1496,2028]" box="[199,475,1496,1523]" pageId="4" pageNumber="149">
<emphasis id="B96AEAB4FFB18C437896FA70FEFD49AA" bold="true" box="[199,359,1496,1523]" pageId="4" pageNumber="149">Composition.</emphasis>
1 species.
</paragraph>
</subSubSection>
<subSubSection id="C304652DFFB18C437896FA54FD6F4A41" box="[199,757,1532,1560]" pageId="4" pageNumber="149" type="distribution">
<paragraph id="8BA136A6FFB18C437896FA54FD6F4A41" blockId="4.[151,1437,1496,2028]" box="[199,757,1532,1560]" pageId="4" pageNumber="149">
<emphasis id="B96AEAB4FFB18C437896FA54FD814A4E" bold="true" box="[199,539,1532,1559]" pageId="4" pageNumber="149">Distribution. North America</emphasis>
(
<collectingCountry id="F3097636FFB18C437A7BFA55FDFD4A4E" box="[554,615,1533,1559]" name="United States of America" pageId="4" pageNumber="149">USA</collectingCountry>
:
<collectingRegion id="49DAF844FFB18C437A23FA55FD734A41" box="[626,745,1533,1560]" country="United States of America" name="California" pageId="4" pageNumber="149">California</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C304652DFFB18C437896F989FEAE4BFA" pageId="4" pageNumber="149" type="discussion">
<paragraph id="8BA136A6FFB18C437896F989FF7A4B61" blockId="4.[151,1437,1496,2028]" pageId="4" pageNumber="149">
<emphasis id="B96AEAB4FFB18C437896F989FEA14A62" bold="true" box="[199,315,1569,1595]" pageId="4" pageNumber="149">Remarks.</emphasis>
The status of the type specimen preserved at MCZ (with labels shown in
<figureCitation id="13252A23FFB18C437C23F989FB2E4A65" box="[1138,1204,1569,1596]" captionStart="FIGURES 5–7" captionStartId="4.[151,264,1390,1415]" captionTargetBox="[169,1400,613,1315]" captionTargetId="figure@4.[151,1436,598,1327]" captionTargetPageId="4" captionText="FIGURES 5–7. Aedeagus of Catalinus angustus in ventral (5) and lateral (6) views and original set of labels of the presumable male syntype (7)." httpUri="https://zenodo.org/record/2673387/files/figure.png" pageId="4" pageNumber="149">Fig. 7</figureCitation>
) is unclear.
<bibRefCitation id="EF8F4B57FFB18C437D6DF989FF784A06" author="LeConte, J. L." pageId="4" pageNumber="149" pagination="149 - 157" refId="ref8464" refString="LeConte, J. L. (1852) Synopsis of the Scydmaenidae of the United States. Proceedings of the Academy of Natural Scences, Philiadelphia, 6, 149 - 157." type="journal article" year="1852">LeConte (1852)</bibRefCitation>
did not specify the number of specimens. In descriptions of other species in the same paper, LeConte either specified the number of specimens (most frequently as one, only in a few cases as two and four), or gave only the locality (most frequently as the name of state only). Based on this information, the type specimen should be treated as a
<typeStatus id="54A58804FFB18C43789AF91AFEBC4A95" box="[203,294,1714,1740]" pageId="4" pageNumber="149" type="syntype">syntype</typeStatus>
. However,
<bibRefCitation id="EF8F4B57FFB18C4379F4F919FDA44A92" author="Casey, T. L." box="[421,574,1713,1739]" pageId="4" pageNumber="149" pagination="285 - 684" refId="ref7607" refString="Casey, T. L. (1897) Coleopterological notices, VII. Annals of the New York Academy of Sciences, 9, 285 - 684. https: // doi. org / 10.1111 / j. 1749 - 6632.1896. tb 55435. x" type="journal article" year="1897">Casey (1897)</bibRefCitation>
, who clearly studied the LeConte collection, stated that “the sex of the individual described is not determinable”, suggesting that there was only one specimen used for the species description, and therefore it would be a
<typeStatus id="54A58804FFB18C437999F951FDB44B4A" box="[456,558,1785,1811]" pageId="4" pageNumber="149" type="holotype">holotype</typeStatus>
. This question is here left open, as the purpose of this paper is to redescribe the genus.
</paragraph>
<paragraph id="8BA136A6FFB18C437896F8E8FEAE4BFA" blockId="4.[151,1437,1496,2028]" pageId="4" pageNumber="149">
<emphasis id="B96AEAB4FFB18C437896F8E8FEC54B02" bold="true" box="[199,351,1856,1883]" pageId="4" pageNumber="149">Conclusions.</emphasis>
<taxonomicName id="4C1E4D25FFB18C437939F8E9FE4F4B02" authorityName="Casey" authorityYear="1897" box="[360,469,1857,1883]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB18C437939F8E9FE4F4B02" box="[360,469,1857,1883]" italics="true" pageId="4" pageNumber="149">Catalinus</emphasis>
</taxonomicName>
is here maintained as a separate genus. Although defined only by a combination of synapomorphies known to occur in other glandulariines, this set of features is unique and allows for unambiguous identification.
</paragraph>
</subSubSection>
<subSubSection id="C304652DFFB18C417896F804FBCF4ABA" lastPageId="6" lastPageNumber="151" pageId="4" pageNumber="149" type="description">
<paragraph id="8BA136A6FFB18C437896F804FAD44BB2" blockId="4.[151,1437,1496,2028]" pageId="4" pageNumber="149">
<emphasis id="B96AEAB4FFB18C437896F804FE2B4B9E" bold="true" box="[199,433,1964,1991]" pageId="4" pageNumber="149">Comparative notes.</emphasis>
<taxonomicName id="4C1E4D25FFB18C4379E8F805FDBC4B9E" authorityName="Casey" authorityYear="1897" box="[441,550,1965,1991]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB18C4379E8F805FDBC4B9E" box="[441,550,1965,1991]" italics="true" pageId="4" pageNumber="149">Catalinus</emphasis>
</taxonomicName>
belongs to a group of glandulariine genera with lateral sutures of submentum. This group includes 19 other genera;
<taxonomicName id="4C1E4D25FFB18C437A6BF879FD3D4BB2" authorityName="Casey" authorityYear="1897" box="[570,679,2001,2027]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB18C437A6BF879FD3D4BB2" box="[570,679,2001,2027]" italics="true" pageId="4" pageNumber="149">Catalinus</emphasis>
</taxonomicName>
differs from each of them in the following major characters:
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FF31FAEF4D3E" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FF31FE9A4CEA" bold="true" box="[151,256,152,179]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427956FF30FDBA4CEA" authority="Franz, 1980 a" authorityName="Franz" authorityYear="1980" box="[263,544,152,179]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427956FF30FE1F4CEA" bold="true" box="[263,389,152,179]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
Franz, 1980a
</taxonomicName>
in the pronotum lacking antebasal pits and groove (present in
<taxonomicName id="4C1E4D25FFB08C427CB7FF30FAFB4CEB" authorityName="Franz" authorityYear="1980" box="[1254,1377,152,178]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CB7FF30FAFB4CEB" box="[1254,1377,152,178]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
</taxonomicName>
); the hypomeral ridges marked only anteriorly, posteriorly largely obliterated (complete in
<taxonomicName id="4C1E4D25FFB08C427CF3FF14FA874C8F" authorityName="Franz" authorityYear="1980" box="[1186,1309,188,214]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CF3FF14FA874C8F" box="[1186,1309,188,214]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
</taxonomicName>
); the large setose impressions behind asetose procoxal rests on the mesoventrite (in
<taxonomicName id="4C1E4D25FFB08C427BBCFF48FBF24CA3" authorityName="Franz" authorityYear="1980" box="[1005,1128,224,250]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BBCFF48FBF24CA3" box="[1005,1128,224,250]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
</taxonomicName>
with large asetose procoxal rests and very short, additional impressions behind them, with only their posterior margin setose); each elytron with one large basal fovea (in
<taxonomicName id="4C1E4D25FFB08C427A45FE80FD154D1B" authorityName="Franz" authorityYear="1980" box="[532,655,296,322]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A45FE80FD154D1B" box="[532,655,296,322]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
</taxonomicName>
two vestigial foveae); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB08C42797FFEE4FE334D3F" authorityName="Franz" authorityYear="1980" box="[302,425,332,358]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C42797FFEE4FE334D3F" box="[302,425,332,358]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
</taxonomicName>
exposed); the aedeagus lacking the basal ‘pumping apparatus’ (present in
<taxonomicName id="4C1E4D25FFB08C427CB8FEE4FAFE4D3F" authorityName="Franz" authorityYear="1980" box="[1257,1380,332,358]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CB8FEE4FAFE4D3F" box="[1257,1380,332,358]" italics="true" pageId="5" pageNumber="150">Alloraphes</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FED9FDFF4DAE" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FED9FE9A4DD2" bold="true" box="[151,256,368,395]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427958FED8FDFD4DD2" authority="Franz, 1971" authorityName="Franz" authorityYear="1971" box="[265,615,368,395]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427958FED8FE494DD2" bold="true" box="[265,467,368,395]" italics="true" pageId="5" pageNumber="150">Austrostenichnus</emphasis>
<bibRefCitation id="EF8F4B57FFB08C42798FFED9FDFD4DD2" author="Franz, H." box="[478,615,369,395]" pageId="5" pageNumber="150" pagination="43 - 157" refId="ref7730" refString="Franz, H. (1971) Untersuchungen uber die Scydmaenidenfauna Neukaledoniens (Col., Scydm.). Koleopterologische Rundschau, 49, 43 - 157." type="journal article" year="1971">
Franz,
<quantity id="4CE69B43FFB08C427A7EFED9FDFD4DD2" box="[559,615,369,395]" metricMagnitude="1" metricUnit="m" metricValue="5.00634" pageId="5" pageNumber="150" unit="in" value="1971.0">1971</quantity>
</bibRefCitation>
</taxonomicName>
in the tempora subequal to eyes, and eyes near middle of head length (tempora very short, vestigial, and eyes in posterior half of head in
<taxonomicName id="4C1E4D25FFB08C427B9FFE3CFB144DF7" authorityName="Franz" authorityYear="1971" box="[974,1166,404,430]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427B9FFE3CFB144DF7" box="[974,1166,404,430]" italics="true" pageId="5" pageNumber="150">Austrostenichnus</emphasis>
</taxonomicName>
); the pronotum lacking antebasal pits and groove (present in
<taxonomicName id="4C1E4D25FFB08C427A39FE10FCB24D8B" authorityName="Franz" authorityYear="1971" box="[616,808,440,466]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A39FE10FCB24D8B" box="[616,808,440,466]" italics="true" pageId="5" pageNumber="150">Austrostenichnus</emphasis>
</taxonomicName>
); the mesoventrite with carinate mesoventral intercoxal process (absent in
<taxonomicName id="4C1E4D25FFB08C4279C5FE74FDCE4DAF" authorityName="Franz" authorityYear="1971" box="[404,596,476,502]" class="Insecta" family="Staphylinidae" genus="Austrostenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C4279C5FE74FDCE4DAF" box="[404,596,476,502]" italics="true" pageId="5" pageNumber="150">Austrostenichnus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FDA9FA9C4EDE" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FDA9FECA4E42" bold="true" box="[151,336,512,539]" pageId="5" pageNumber="150">
- from
<taxonomicName id="4C1E4D25FFB08C427954FDA8FECA4E42" authorityName="Casey" authorityYear="1897" box="[261,336,512,539]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427954FDA8FECA4E42" bold="true" box="[261,336,512,539]" italics="true" pageId="5" pageNumber="150">Delius</emphasis>
</taxonomicName>
</emphasis>
in the head and prothorax lacking thick bristles (present in
<taxonomicName id="4C1E4D25FFB08C427BB1FDA8FBB34E43" authorityName="Casey" authorityYear="1897" box="[992,1065,512,538]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BB1FDA8FBB34E43" box="[992,1065,512,538]" italics="true" pageId="5" pageNumber="150">Delius</emphasis>
</taxonomicName>
); the pronotum lacking antebasal pits (present in
<taxonomicName id="4C1E4D25FFB08C427923FD8CFE214E67" authorityName="Casey" authorityYear="1897" box="[370,443,548,574]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427923FD8CFE214E67" box="[370,443,548,574]" italics="true" pageId="5" pageNumber="150">Delius</emphasis>
</taxonomicName>
); the basisternal part of prosternum much shorter than the coxal part (subequal in length in
<taxonomicName id="4C1E4D25FFB08C4278B3FDE0FEB14E3B" authorityName="Casey" authorityYear="1897" box="[226,299,584,610]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C4278B3FDE0FEB14E3B" box="[226,299,584,610]" italics="true" pageId="5" pageNumber="150">Delius</emphasis>
</taxonomicName>
); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB08C427B81FDE0FB834E3B" authorityName="Casey" authorityYear="1897" box="[976,1049,584,610]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427B81FDE0FB834E3B" box="[976,1049,584,610]" italics="true" pageId="5" pageNumber="150">Delius</emphasis>
</taxonomicName>
exposed); the metacoxae laterally reaching lateral margins of metaventrite (broadly separated from sides of metaventrite in
<taxonomicName id="4C1E4D25FFB08C427CFDFDC4FB6F4EDF" authorityName="Casey" authorityYear="1897" box="[1196,1269,620,646]" class="Insecta" family="Staphylinidae" genus="Delius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CFDFDC4FB6F4EDF" box="[1196,1269,620,646]" italics="true" pageId="5" pageNumber="150">Delius</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FD39FAA94F62" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FD39FE9A4EF2" bold="true" box="[151,256,656,683]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427954FD38FD134EF2" authority="Croissandeau, 1898" authorityName="Croissandeau" authorityYear="1898" box="[261,649,656,683]" class="Insecta" family="Staphylinidae" genus="Leptoderoides" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427954FD38FE3D4EF2" bold="true" box="[261,423,656,683]" italics="true" pageId="5" pageNumber="150">Leptoderoides</emphasis>
<bibRefCitation id="EF8F4B57FFB08C4279FCFD38FD134EF2" author="Croissandeau, J." box="[429,649,656,683]" pageId="5" pageNumber="150" pagination="402 - 430" refId="ref7655" refString="Croissandeau, J. (1898) Monographie des Scydmaenidae. Annales de la Societe Entomologique de France, 66, 402 - 430, pls. 12 - 16." type="journal article" year="1898">
Croissandeau,
<quantity id="4CE69B43FFB08C427A00FD39FD134EF2" box="[593,649,657,683]" metricMagnitude="1" metricUnit="m" metricValue="4.820919999999999" pageId="5" pageNumber="150" unit="in" value="1898.0">1898</quantity>
</bibRefCitation>
</taxonomicName>
in the head and prothorax lacking thick bristles (present in
<emphasis id="B96AEAB4FFB08C427D48FD38FF704E97" italics="true" pageId="5" pageNumber="150">Leptoderoi- des</emphasis>
); lateral sutures of submentum posteriorly broadly separated (posteriorly nearly touching in
<taxonomicName id="4C1E4D25FFB08C427CBDFD1CFA114E97" authorityName="Croissandeau" authorityYear="1898" box="[1260,1419,692,718]" class="Insecta" family="Staphylinidae" genus="Leptoderoides" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CBDFD1CFA114E97" box="[1260,1419,692,718]" italics="true" pageId="5" pageNumber="150">Leptoderoides</emphasis>
</taxonomicName>
); the tempora subequal to eyes, and eyes near middle of head length (tempora very short, vestigial, and eyes in posterior half of head in
<taxonomicName id="4C1E4D25FFB08C427985FD54FDE94F4F" authorityName="Croissandeau" authorityYear="1898" box="[468,627,764,790]" class="Insecta" family="Staphylinidae" genus="Leptoderoides" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427985FD54FDE94F4F" box="[468,627,764,790]" italics="true" pageId="5" pageNumber="150">Leptoderoides</emphasis>
</taxonomicName>
); the pronotum lacking sublateral carinae (present in
<taxonomicName id="4C1E4D25FFB08C427C95FD54FAF94F4F" authorityName="Croissandeau" authorityYear="1898" box="[1220,1379,764,790]" class="Insecta" family="Staphylinidae" genus="Leptoderoides" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427C95FD54FAF94F4F" box="[1220,1379,764,790]" italics="true" pageId="5" pageNumber="150">Leptoderoides</emphasis>
</taxonomicName>
); the hypomeral ridges marked only anteriorly, posteriorly largely obliterated (complete in
<taxonomicName id="4C1E4D25FFB08C427CD3FC88FABB4F63" authorityName="Croissandeau" authorityYear="1898" box="[1154,1313,800,826]" class="Insecta" family="Staphylinidae" genus="Leptoderoides" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CD3FC88FABB4F63" box="[1154,1313,800,826]" italics="true" pageId="5" pageNumber="150">Leptoderoides</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FCEDFCBB4802" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FCEDFE9A4F06" bold="true" box="[151,256,836,863]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427956FCECFDF64F06" authority="Jaloszynski, 2018" authorityName="Jaloszynski" authorityYear="2018" box="[263,620,836,863]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427956FCECFE014F06" bold="true" box="[263,411,836,863]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
<bibRefCitation id="EF8F4B57FFB08C4279F5FCECFDF64F06" author="Jaloszynski, P." box="[420,620,836,863]" pageId="5" pageNumber="150" pagination="111 - 121" refId="ref8335" refString="Jaloszynski, P. (2018) Madagaphes gen. nov. for Madagascan species misplaced in Stenichnaphes Franz (Coleoptera, Staphylinidae, Scydmaeninae). Austral Entomology, 58 (2019), 111 - 121. https: // doi. org / 10.1111 / aen. 12331" type="journal article" year="2018">
Jałoszyński,
<quantity id="4CE69B43FFB08C427A65FCEDFDF64F06" box="[564,620,837,863]" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" pageId="5" pageNumber="150" unit="in" value="2018.0">2018</quantity>
</bibRefCitation>
</taxonomicName>
in the pronotum lacking antebasal pits and groove (present in
<taxonomicName id="4C1E4D25FFB08C427D67FCECFF624FDB" authorityName="Jaloszynski" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427D67FCECFF624FDB" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
); an indistinct, diffuse prosternal intercoxal process (carinate, sharply marked in
<taxonomicName id="4C1E4D25FFB08C427CF1FCC0FAAB4FDB" authorityName="Jaloszynski" authorityYear="2018" box="[1184,1329,872,898]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CF1FCC0FAAB4FDB" box="[1184,1329,872,898]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
); the hypomeral ridges marked only anteriorly, posteriorly largely obliterated (in
<taxonomicName id="4C1E4D25FFB08C427BA6FC24FB124FFF" authorityName="Jaloszynski" authorityYear="2018" box="[1015,1160,908,934]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BA6FC24FB124FFF" box="[1015,1160,908,934]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
marked at middle, obliterated anteriorly and posteriorly); the large setose impressions behind asetose procoxal rests on the mesoventrite (
<taxonomicName id="4C1E4D25FFB08C42789FFC7CFEC54FB7" authorityName="Jaloszynski" authorityYear="2018" box="[206,351,980,1006]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C42789FFC7CFEC54FB7" box="[206,351,980,1006]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
with large asetose procoxal rests and very short setose impressions behind them); the metaventral intercoxal process deeply notched at middle (subtrapezoidal, lacking notch in
<taxonomicName id="4C1E4D25FFB08C427C75FC50FB2F484B" authorityName="Jaloszynski" authorityYear="2018" box="[1060,1205,1016,1042]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427C75FC50FB2F484B" box="[1060,1205,1016,1042]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB08C427A28FBB4FC90486F" authorityName="Jaloszynski" authorityYear="2018" box="[633,778,1052,1078]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A28FBB4FC90486F" box="[633,778,1052,1078]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
exposed); large and deep basal elytral foveae near elytral suture (in
<taxonomicName id="4C1E4D25FFB08C427967FBE8FE5D4803" authorityName="Jaloszynski" authorityYear="2018" box="[310,455,1088,1114]" class="Insecta" family="Staphylinidae" genus="Madagaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427967FBE8FE5D4803" box="[310,455,1088,1114]" italics="true" pageId="5" pageNumber="150">Madagaphes</emphasis>
</taxonomicName>
vestigial fovea near humerus);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FBCDFCFE496A" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FBCDFE9A4826" bold="true" box="[151,256,1124,1151]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427958FBCCFDE34827" authority="Jaloszynski, 2013 b" authorityName="Jaloszynski" authorityYear="2013" box="[265,633,1124,1150]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427958FBCCFE024827" bold="true" box="[265,408,1124,1150]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
Jałoszyński, 2013b
</taxonomicName>
in lack of the antebasal pronotal groove (present in
<taxonomicName id="4C1E4D25FFB08C427C89FBCDFAC54827" authorityName="Jaloszynski" authorityYear="2013" box="[1240,1375,1125,1150]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427C89FBCDFAC54827" box="[1240,1375,1125,1150]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
</taxonomicName>
); the hypomeral ridges marked only anteriorly, posteriorly largely obliterated (complete in
<taxonomicName id="4C1E4D25FFB08C427CCEFB21FABC48FB" authorityName="Jaloszynski" authorityYear="2013" box="[1183,1318,1161,1186]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CCEFB21FABC48FB" box="[1183,1318,1161,1186]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
</taxonomicName>
); the prothorax lacking thick bristles (present in
<taxonomicName id="4C1E4D25FFB08C427AD4FB05FC96489F" authorityName="Jaloszynski" authorityYear="2013" box="[645,780,1197,1222]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427AD4FB05FC96489F" box="[645,780,1197,1222]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
</taxonomicName>
); the basisternal part of prosternum much shorter than the coxal part (subequal in length in
<taxonomicName id="4C1E4D25FFB08C427A10FB79FD5248B3" authorityName="Jaloszynski" authorityYear="2013" box="[577,712,1233,1258]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A10FB79FD5248B3" box="[577,712,1233,1258]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
</taxonomicName>
); the large setose impressions behind asetose procoxal rests on the mesoventrite (absent in
<taxonomicName id="4C1E4D25FFB08C4279A8FB5DFD1A4957" authorityName="Jaloszynski" authorityYear="2013" box="[505,640,1269,1294]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C4279A8FB5DFD1A4957" box="[505,640,1269,1294]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
</taxonomicName>
); the protarsi in male unmodified (strongly broadened at base and with conspicuously long and dense ventral setae in
<taxonomicName id="4C1E4D25FFB08C427A9AFAB1FCC8496B" authorityName="Jaloszynski" authorityYear="2013" box="[715,850,1305,1330]" class="Insecta" family="Staphylinidae" genus="Mexiconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A9AFAB1FCC8496B" box="[715,850,1305,1330]" italics="true" pageId="5" pageNumber="150">Mexiconnus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FA95FCC049BE" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FA95FEF6490E" bold="true" box="[151,364,1340,1367]" pageId="5" pageNumber="150">
- from
<taxonomicName id="4C1E4D25FFB08C427954FA94FEF6490E" authorityName="Casey" authorityYear="1897" box="[261,364,1340,1367]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427954FA94FEF6490E" bold="true" box="[261,364,1340,1367]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
</emphasis>
in the tempora subequal to eyes (much longer than eyes in
<taxonomicName id="4C1E4D25FFB08C427BACFA94FBFB490F" authorityName="Casey" authorityYear="1897" box="[1021,1121,1340,1366]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BACFA94FBFB490F" box="[1021,1121,1340,1366]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
); the basisternal part of prosternum much shorter than coxal part (subequal in length in
<taxonomicName id="4C1E4D25FFB08C427B30FAC8FC5F4923" authorityName="Casey" authorityYear="1897" box="[865,965,1376,1402]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427B30FAC8FC5F4923" box="[865,965,1376,1402]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB08C42792DFA2CFE7A49C7" authorityName="Casey" authorityYear="1897" box="[380,480,1412,1438]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C42792DFA2CFE7A49C7" box="[380,480,1412,1438]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
exposed); large and deep basal elytral foveae (vestigial in
<taxonomicName id="4C1E4D25FFB08C427C3EFA2CFB4949C7" authorityName="Casey" authorityYear="1897" box="[1135,1235,1412,1438]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427C3EFA2CFB4949C7" box="[1135,1235,1412,1438]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
); the mesoventral intercoxal process present (absent in
<taxonomicName id="4C1E4D25FFB08C427A3EFA00FD49499B" authorityName="Casey" authorityYear="1897" box="[623,723,1448,1474]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A3EFA00FD49499B" box="[623,723,1448,1474]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
); metacoxae laterally reaching lateral margins of metaventrite (broadly separated from sides of metaventrite in
<taxonomicName id="4C1E4D25FFB08C427AB7FA64FCD049BF" authorityName="Casey" authorityYear="1897" box="[742,842,1484,1510]" class="Insecta" family="Staphylinidae" genus="Neladius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427AB7FA64FCD049BF" box="[742,842,1484,1510]" italics="true" pageId="5" pageNumber="150">Neladius</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6FA59FEF64AC2" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6FA59FE1F4A52" bold="true" box="[151,389,1520,1547]" pageId="5" pageNumber="150">
- from
<taxonomicName id="4C1E4D25FFB08C427956FA58FE1F4A52" authorityName="Thomson" authorityYear="1859" box="[263,389,1520,1547]" class="Insecta" family="Staphylinidae" genus="Neuraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427956FA58FE1F4A52" bold="true" box="[263,389,1520,1547]" italics="true" pageId="5" pageNumber="150">Neuraphes</emphasis>
</taxonomicName>
</emphasis>
in tempora subequal to eyes, and eyes near middle of head length (tempora very short, vestigial, and eyes in posterior half of head in
<taxonomicName id="4C1E4D25FFB08C427AEBF9BCFCA94A77" authorityName="Thomson" authorityYear="1859" box="[698,819,1556,1582]" class="Insecta" family="Staphylinidae" genus="Neuraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427AEBF9BCFCA94A77" box="[698,819,1556,1582]" italics="true" pageId="5" pageNumber="150">Neuraphes</emphasis>
</taxonomicName>
); the pronotum lacking lateral carinae, antebasal pits, groove and median longitudinal wrinkle or carina (all present in
<taxonomicName id="4C1E4D25FFB08C427BC9F990FB8B4A0B" authorityName="Thomson" authorityYear="1859" box="[920,1041,1592,1618]" class="Insecta" family="Staphylinidae" genus="Neuraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BC9F990FB8B4A0B" box="[920,1041,1592,1618]" italics="true" pageId="5" pageNumber="150">Neuraphes</emphasis>
</taxonomicName>
); the aedeagus with elongate sclerites in the endophallus, broadest in basal half (lacking elongate sclerites and most often broadest in distal half in
<taxonomicName id="4C1E4D25FFB08C4278B0F928FEC04AC3" authorityName="Thomson" authorityYear="1859" box="[225,346,1664,1690]" class="Insecta" family="Staphylinidae" genus="Neuraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C4278B0F928FEC04AC3" box="[225,346,1664,1690]" italics="true" pageId="5" pageNumber="150">Neuraphes</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6F90DFB3C4B2A" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6F90DFE9A4AE6" bold="true" box="[151,256,1700,1727]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427959F90CFDED4AE6" authority="Jaloszynski, 2014" authorityName="Jaloszynski" authorityYear="2014" box="[264,631,1700,1727]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427959F90CFE3E4AE6" bold="true" box="[264,420,1700,1727]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
<bibRefCitation id="EF8F4B57FFB08C4279FCF90CFDED4AE6" author="Jaloszynski, P." box="[429,631,1700,1727]" pageId="5" pageNumber="150" pagination="481 - 491" refId="ref8126" refString="Jaloszynski, P. (2014) Obesoconnus gen. n., a new bizarre Neotropical genus of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3794 (3), 481 - 491. https: // doi. org / 10.11646 / zootaxa. 3794.3.9" type="journal article" year="2014">
Jałoszyński,
<quantity id="4CE69B43FFB08C427A6EF90DFDED4AE6" box="[575,631,1701,1727]" metricMagnitude="1" metricUnit="m" metricValue="5.11556" pageId="5" pageNumber="150" unit="in" value="2014.0">2014</quantity>
</bibRefCitation>
</taxonomicName>
in the ant-like body form (compactly oval in
<taxonomicName id="4C1E4D25FFB08C427CDAF90CFAB84AE7" authorityName="Jaloszynski" authorityYear="2014" box="[1163,1314,1700,1726]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CDAF90CFAB84AE7" box="[1163,1314,1700,1726]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
); the head with a distinct occipital constriction, with small eyes (lacking constriction and with conspicuously large eyes in
<taxonomicName id="4C1E4D25FFB08C427894F944FEC64B5F" authorityName="Jaloszynski" authorityYear="2014" box="[197,348,1772,1798]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427894F944FEC64B5F" box="[197,348,1772,1798]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
); the hypostomal ridges complete (incomplete in
<taxonomicName id="4C1E4D25FFB08C427BDCF944FBBE4B5F" authorityName="Jaloszynski" authorityYear="2014" box="[909,1060,1772,1798]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BDCF944FBBE4B5F" box="[909,1060,1772,1798]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
); the pronotum lacking antebasal pits and groove (present in
<taxonomicName id="4C1E4D25FFB08C4279AFF8B8FD0F4B73" authorityName="Jaloszynski" authorityYear="2014" box="[510,661,1808,1834]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C4279AFF8B8FD0F4B73" box="[510,661,1808,1834]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
); the prosternum strongly convex posteromedially (deeply emarginate posteromedially in
<taxonomicName id="4C1E4D25FFB08C427999F89CFDC54B17" authorityName="Jaloszynski" authorityYear="2014" box="[456,607,1844,1870]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427999F89CFDC54B17" box="[456,607,1844,1870]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB08C427D54F89CFA064B17" authorityName="Jaloszynski" authorityYear="2014" box="[1285,1436,1844,1870]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427D54F89CFA064B17" box="[1285,1436,1844,1870]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
exposed); the aedeagus lacking the basal ‘pumping apparatus’ (present in
<taxonomicName id="4C1E4D25FFB08C427BAFF8F0FB0F4B2B" authorityName="Jaloszynski" authorityYear="2014" box="[1022,1173,1880,1906]" class="Insecta" family="Staphylinidae" genus="Obesoconnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427BAFF8F0FB0F4B2B" box="[1022,1173,1880,1906]" italics="true" pageId="5" pageNumber="150">Obesoconnus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB08C4278C6F8D5FCD24B86" blockId="5.[151,1437,152,2015]" pageId="5" pageNumber="150">
<emphasis id="B96AEAB4FFB08C4278C6F8D5FE9A4BCE" bold="true" box="[151,256,1916,1943]" pageId="5" pageNumber="150">- from</emphasis>
<taxonomicName id="4C1E4D25FFB08C427959F8D4FDEE4BCE" authority="Franz, 1975" authorityName="Franz" authorityYear="1975" box="[264,628,1916,1943]" class="Insecta" family="Staphylinidae" genus="Palaeoscydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427959F8D4FE794BCE" bold="true" box="[264,483,1916,1943]" italics="true" pageId="5" pageNumber="150">Palaeoscydmaenus</emphasis>
<bibRefCitation id="EF8F4B57FFB08C4279BCF8D5FDEE4BCE" author="Franz, H." box="[493,628,1916,1943]" pageId="5" pageNumber="150" pagination="1 - 312" refId="ref7759" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Os- terreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">
Franz,
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</bibRefCitation>
</taxonomicName>
in the pronotum lacking antebasal groove (present in
<taxonomicName id="4C1E4D25FFB08C427CB2F8D4FF764BE3" authorityName="Franz" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Palaeoscydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427CB2F8D4FF764BE3" italics="true" pageId="5" pageNumber="150">Palaeoscydmaenus</emphasis>
</taxonomicName>
); the elytra with deep basal foveae (absent in
<taxonomicName id="4C1E4D25FFB08C427AA6F808FC544BE3" authorityName="Franz" authorityYear="1975" box="[759,974,1952,1978]" class="Insecta" family="Staphylinidae" genus="Palaeoscydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427AA6F808FC544BE3" box="[759,974,1952,1978]" italics="true" pageId="5" pageNumber="150">Palaeoscydmaenus</emphasis>
</taxonomicName>
); the mesoventrite with a carinate mesoventral intercoxal process (absent in
<taxonomicName id="4C1E4D25FFB08C427A31F86CFCAD4B87" authorityName="Franz" authorityYear="1975" box="[608,823,1988,2014]" class="Insecta" family="Staphylinidae" genus="Palaeoscydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB08C427A31F86CFCAD4B87" box="[608,823,1988,2014]" italics="true" pageId="5" pageNumber="150">Palaeoscydmaenus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FF31FD554DD2" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FF31FE3E4CEA" bold="true" box="[151,420,152,179]" pageId="6" pageNumber="151">
- from
<taxonomicName id="4C1E4D25FFB38C41795BFF30FE3E4CEA" authorityName="Casey" authorityYear="1897" box="[266,420,152,179]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C41795BFF30FE3E4CEA" bold="true" box="[266,420,152,179]" italics="true" pageId="6" pageNumber="151">Parascydmus</emphasis>
</taxonomicName>
</emphasis>
in the frons lacking deep supraantennal notches and not impressed at middle (notches present and frons with a transverse impression in
<taxonomicName id="4C1E4D25FFB38C417AA8FF14FC0A4C8F" authorityName="Casey" authorityYear="1897" box="[761,912,188,214]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417AA8FF14FC0A4C8F" box="[761,912,188,214]" italics="true" pageId="6" pageNumber="151">Parascydmus</emphasis>
</taxonomicName>
); the lateral sutures of submentum posteriorly broadly separated (nearly touching in
<taxonomicName id="4C1E4D25FFB38C417A28FF48FC8A4CA3" authorityName="Casey" authorityYear="1897" box="[633,784,224,250]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417A28FF48FC8A4CA3" box="[633,784,224,250]" italics="true" pageId="6" pageNumber="151">Parascydmus</emphasis>
</taxonomicName>
); the prothorax lacking lateral carinae and antebasal pits (present in
<taxonomicName id="4C1E4D25FFB38C417916FEACFE444D47" authorityName="Casey" authorityYear="1897" box="[327,478,260,286]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417916FEACFE444D47" box="[327,478,260,286]" italics="true" pageId="6" pageNumber="151">Parascydmus</emphasis>
</taxonomicName>
); the metaventral intercoxal process with a pair of short lateral subtriangular, blunt projections (with a pair of long pointed lateral spines in
<taxonomicName id="4C1E4D25FFB38C417B03FE80FC734D1B" authorityName="Casey" authorityYear="1897" box="[850,1001,296,322]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B03FE80FC734D1B" box="[850,1001,296,322]" italics="true" pageId="6" pageNumber="151">Parascydmus</emphasis>
</taxonomicName>
); the aedeagus broadest in basal half, presumably with parameres and with long endophallic sclerites (broadest in distal half, lacking parameres and lacking endophallic sclerites in
<taxonomicName id="4C1E4D25FFB38C417A76FED8FD244DD3" authorityName="Casey" authorityYear="1897" box="[551,702,368,394]" class="Insecta" family="Staphylinidae" genus="Parascydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417A76FED8FD244DD3" box="[551,702,368,394]" italics="true" pageId="6" pageNumber="151">Parascydmus</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FE3DFAC64E3A" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FE3DFE9A4DF6" bold="true" box="[151,256,404,431]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417955FE3CFDCF4DF6" authority="Jaloszynski, 2015" authorityName="Jaloszynski" authorityYear="2015" box="[260,597,404,431]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417955FE3CFE104DF6" bold="true" box="[260,394,404,431]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
<bibRefCitation id="EF8F4B57FFB38C4179DEFE3CFDCF4DF6" author="Jaloszynski, P." box="[399,597,404,431]" pageId="6" pageNumber="151" pagination="330 - 339" refId="ref8177" refString="Jaloszynski, P. (2015) A new genus of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae) from Japan. Entomological Science, 18 (3), 330 - 339. https: // doi. org / 10.1111 / ens. 12136" type="journal article" year="2015">
Jałoszyński,
<quantity id="4CE69B43FFB38C417A4CFE3CFDCF4DF6" box="[541,597,404,431]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="6" pageNumber="151" unit="in" value="2015.0">2015</quantity>
</bibRefCitation>
</taxonomicName>
in the frons lacking deep supraantennal notches (present in
<taxonomicName id="4C1E4D25FFB38C417CB2FE3CFAFE4DF7" authorityName="Jaloszynski" authorityYear="2015" box="[1251,1380,404,430]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417CB2FE3CFAFE4DF7" box="[1251,1380,404,430]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
</taxonomicName>
); the lateral sutures of submentum posteriorly broadly separated (nearly touching in
<taxonomicName id="4C1E4D25FFB38C417C60FE10FB284D8B" authorityName="Jaloszynski" authorityYear="2015" box="[1073,1202,440,466]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417C60FE10FB284D8B" box="[1073,1202,440,466]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
</taxonomicName>
); the pronotum lacking lateral carinae and antebasal pits (present in
<taxonomicName id="4C1E4D25FFB38C417AB8FE74FCF04DAF" authorityName="Jaloszynski" authorityYear="2015" box="[745,874,476,502]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417AB8FE74FCF04DAF" box="[745,874,476,502]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
</taxonomicName>
); the hypomeral ridges marked anteriorly (absent in
<taxonomicName id="4C1E4D25FFB38C4178B2FDA8FEFE4E43" authorityName="Jaloszynski" authorityYear="2015" box="[227,356,512,538]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C4178B2FDA8FEFE4E43" box="[227,356,512,538]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
</taxonomicName>
); the metaventral intercoxal process with a pair of short lateral subtriangular, blunt projections (with a pair of long pointed lateral spines in
<taxonomicName id="4C1E4D25FFB38C417AE4FD8CFCAC4E67" authorityName="Jaloszynski" authorityYear="2015" box="[693,822,548,574]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417AE4FD8CFCAC4E67" box="[693,822,548,574]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
</taxonomicName>
); the aedeagus presumably with parameres, lacking an elongate inner apodeme (lacking parameres and with a conspicuous sclerotized apodeme in
<taxonomicName id="4C1E4D25FFB38C417C9BFDE0FAD14E3B" authorityName="Jaloszynski" authorityYear="2015" box="[1226,1355,584,610]" class="Insecta" family="Staphylinidae" genus="Rutaraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417C9BFDE0FAD14E3B" box="[1226,1355,584,610]" italics="true" pageId="6" pageNumber="151">Rutaraphes</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FDC5FCA54F4E" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FDC5FE9A4EDE" bold="true" box="[151,256,620,647]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417957FDC4FDBF4EDF" authority="King, 1864" authorityName="King" authorityYear="1864" box="[262,549,620,647]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417957FDC4FE384EDE" bold="true" box="[262,418,620,647]" italics="true" pageId="6" pageNumber="151">Scydmaenilla</emphasis>
<bibRefCitation id="EF8F4B57FFB38C4179F9FDC4FDBF4EDF" author="King, R. L." box="[424,549,620,646]" pageId="6" pageNumber="151" pagination="91 - 99" refId="ref8386" refString="King, R. L. (1864) On the Scydmaenides of New South Wales. Transactions of the Entomological Society of New South Wales, 1, 91 - 99." type="journal article" year="1864">
King,
<quantity id="4CE69B43FFB38C4179BCFDC4FDBF4EDF" box="[493,549,620,646]" metricMagnitude="1" metricUnit="m" metricValue="4.73456" pageId="6" pageNumber="151" unit="in" value="1864.0">1864</quantity>
</bibRefCitation>
</taxonomicName>
in gradually thickened antennae (with distinct trimerous club in
<taxonomicName id="4C1E4D25FFB38C417CA5FDC4FA114EDF" authorityName="King" authorityYear="1864" box="[1268,1419,620,646]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417CA5FDC4FA114EDF" box="[1268,1419,620,646]" italics="true" pageId="6" pageNumber="151">Scydmaenilla</emphasis>
</taxonomicName>
); the pronotum lacking antebasal pits and groove (present in
<taxonomicName id="4C1E4D25FFB38C417B35FD38FC614EF3" authorityName="King" authorityYear="1864" box="[868,1019,656,682]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B35FD38FC614EF3" box="[868,1019,656,682]" italics="true" pageId="6" pageNumber="151">Scydmaenilla</emphasis>
</taxonomicName>
); large and deep basal elytral foveae (vestigial in
<taxonomicName id="4C1E4D25FFB38C417908FD1CFE6A4E97" authorityName="King" authorityYear="1864" box="[345,496,692,718]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417908FD1CFE6A4E97" box="[345,496,692,718]" italics="true" pageId="6" pageNumber="151">Scydmaenilla</emphasis>
</taxonomicName>
); the mesoventral intercoxal process on entire length fused with the ventrite (in
<taxonomicName id="4C1E4D25FFB38C417894FD70FEC64EAB" authorityName="King" authorityYear="1864" box="[197,348,728,754]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417894FD70FEC64EAB" box="[197,348,728,754]" italics="true" pageId="6" pageNumber="151">Scydmaenilla</emphasis>
</taxonomicName>
the posterior 1/3 of mesoventral carina separated from the ventrite and in lateral view forming a subtriangular projection with its apex directed posterad);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FC89FB7E4FFE" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FC89FE9A4F62" bold="true" box="[151,256,800,827]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417956FC88FDDD4F62" authority="Reitter, 1891" authorityName="Reitter" authorityYear="1891" box="[263,583,800,827]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417956FC88FE354F62" bold="true" box="[263,431,800,827]" italics="true" pageId="6" pageNumber="151">Scydmoraphes</emphasis>
<bibRefCitation id="EF8F4B57FFB38C4179E7FC88FDDD4F62" author="Reitter, E." box="[438,583,800,827]" pageId="6" pageNumber="151" pagination="1 - 420" refId="ref8542" refString="Reitter, E. (1891) [new names]. In: von Heyden, L., Reitter, E. &amp; Weise, J. (Eds.), Catalogus Coleopterorum Europae, Caucasi et Armeniae rossicae. Friedlander &amp; Sohn, Berlin, pp. i-vii + 1 - 420." type="book chapter" year="1891">
Reitter,
<quantity id="4CE69B43FFB38C417A5EFC89FDDD4F62" box="[527,583,801,827]" metricMagnitude="1" metricUnit="m" metricValue="4.80314" pageId="6" pageNumber="151" unit="in" value="1891.0">1891</quantity>
</bibRefCitation>
</taxonomicName>
in the tempora subequal to eyes, and eyes near middle of head length (tempora very short, vestigial, and eyes in posterior half of head in
<taxonomicName id="4C1E4D25FFB38C417BCDFCECFBA54F07" authorityName="Reitter" authorityYear="1891" box="[924,1087,836,862]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417BCDFCECFBA54F07" box="[924,1087,836,862]" italics="true" pageId="6" pageNumber="151">Scydmoraphes</emphasis>
</taxonomicName>
); the pronotum lacking lateral carinae and antebasal groove (present in
<taxonomicName id="4C1E4D25FFB38C417AC3FCC0FCAF4FDB" authorityName="Reitter" authorityYear="1891" box="[658,821,872,898]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417AC3FCC0FCAF4FDB" box="[658,821,872,898]" italics="true" pageId="6" pageNumber="151">Scydmoraphes</emphasis>
</taxonomicName>
); the hypomeral ridges incomplete (complete in
<taxonomicName id="4C1E4D25FFB38C417D0FFCC0FEA94FFF" authorityName="Reitter" authorityYear="1891" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417D0FFCC0FEA94FFF" italics="true" pageId="6" pageNumber="151">Scydmoraphes</emphasis>
</taxonomicName>
); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB38C417B9CFC24FBEA4FFF" authorityName="Reitter" authorityYear="1891" box="[973,1136,908,934]" class="Insecta" family="Staphylinidae" genus="Scydmoraphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B9CFC24FBEA4FFF" box="[973,1136,908,934]" italics="true" pageId="6" pageNumber="151">Scydmoraphes</emphasis>
</taxonomicName>
exposed);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FC19FA834802" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FC19FE9A4F92" bold="true" box="[151,256,944,971]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417957FC18FE6E4F92" authority="Franz, 1989" authorityName="Franz" authorityYear="1989" box="[262,500,944,971]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417957FC18FEF24F93" bold="true" box="[262,360,944,970]" italics="true" pageId="6" pageNumber="151">Siamites</emphasis>
<bibRefCitation id="EF8F4B57FFB38C41793FFC19FE6E4F92" author="Franz, H." box="[366,500,945,971]" pageId="6" pageNumber="151" pagination="33 - 80" refId="ref7910" refString="Franz, H. (1989) 4. Beitrag zur Scydmaenidenfauna von Thailand (Die Scydmaenidenausbeute von D. H. Burckhardt u. I. Lobl). Revue suisse de Zoologie, 96, 33 - 80. https: // doi. org / 10.5962 / bhl. part. 82027" type="journal article" year="1989">
Franz,
<quantity id="4CE69B43FFB38C4179EDFC19FE6E4F92" box="[444,500,945,971]" metricMagnitude="1" metricUnit="m" metricValue="5.05206" pageId="6" pageNumber="151" unit="in" value="1989.0">1989</quantity>
</bibRefCitation>
</taxonomicName>
in the head, ventral thorax and abdomen lacking rosettes of thick bristles, also sides of head and pronotum lacking bristles (all present in
<taxonomicName id="4C1E4D25FFB38C417B78FC7DFC124FB6" authorityName="Franz" authorityYear="1989" box="[809,904,981,1007]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B78FC7DFC124FB6" box="[809,904,981,1007]" italics="true" pageId="6" pageNumber="151">Siamites</emphasis>
</taxonomicName>
); the lateral sutures of submentum posteriorly broadly separated (touching in
<taxonomicName id="4C1E4D25FFB38C417A76FC51FD1C484A" authorityName="Franz" authorityYear="1989" box="[551,646,1017,1043]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417A76FC51FD1C484A" box="[551,646,1017,1043]" italics="true" pageId="6" pageNumber="151">Siamites</emphasis>
</taxonomicName>
); the pronotum lacking the antebasal groove and pits (present in
<taxonomicName id="4C1E4D25FFB38C417D3EFC51FE9A486F" authorityName="Franz" authorityYear="1989" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417D3EFC51FE9A486F" italics="true" pageId="6" pageNumber="151">Siamites</emphasis>
</taxonomicName>
); the hypomeral ridges marked anteriorly and obliterated posteriorly (nearly complete, obliterated only anteriorly in
<taxonomicName id="4C1E4D25FFB38C417904FBE9FE2E4802" authorityName="Franz" authorityYear="1989" box="[341,436,1089,1115]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417904FBE9FE2E4802" box="[341,436,1089,1115]" italics="true" pageId="6" pageNumber="151">Siamites</emphasis>
</taxonomicName>
); the aedeagus narrowing toward apex (in
<taxonomicName id="4C1E4D25FFB38C417BC5FBE9FC694802" authorityName="Franz" authorityYear="1989" box="[916,1011,1089,1115]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417BC5FBE9FC694802" box="[916,1011,1089,1115]" italics="true" pageId="6" pageNumber="151">Siamites</emphasis>
</taxonomicName>
broadening toward apex);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FBCDFE7648B2" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FBCDFE9A4826" bold="true" box="[151,256,1124,1151]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417957FBCCFDD14827" authority="Franz, 1980 b" authorityName="Franz" authorityYear="1980" box="[262,587,1124,1151]" class="Insecta" family="Staphylinidae" genus="Stenichnaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417957FBCCFE2A4826" bold="true" box="[262,432,1124,1151]" italics="true" pageId="6" pageNumber="151">Stenichnaphes</emphasis>
Franz, 1980b
</taxonomicName>
in tempora subequal to eyes, and eyes near middle of head length (tempora very short, vestigial, and eyes in posterior half of head in
<taxonomicName id="4C1E4D25FFB38C417B04FB20FC6D48FB" authorityName="Franz" authorityYear="1980" box="[853,1015,1160,1186]" class="Insecta" family="Staphylinidae" genus="Stenichnaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B04FB20FC6D48FB" box="[853,1015,1160,1186]" italics="true" pageId="6" pageNumber="151">Stenichnaphes</emphasis>
</taxonomicName>
); the pronotum lacking the antebasal groove and pits (present in
<taxonomicName id="4C1E4D25FFB38C417A55FB04FD3C489F" authorityName="Franz" authorityYear="1980" box="[516,678,1196,1222]" class="Insecta" family="Staphylinidae" genus="Stenichnaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417A55FB04FD3C489F" box="[516,678,1196,1222]" italics="true" pageId="6" pageNumber="151">Stenichnaphes</emphasis>
</taxonomicName>
); the mesoventrite with a carinate mesoventral intercoxal process (absent in
<taxonomicName id="4C1E4D25FFB38C417968FB78FE4148B3" authorityName="Franz" authorityYear="1980" box="[313,475,1232,1258]" class="Insecta" family="Staphylinidae" genus="Stenichnaphes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417968FB78FE4148B3" box="[313,475,1232,1258]" italics="true" pageId="6" pageNumber="151">Stenichnaphes</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FB5DFB894922" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FB5DFE9A4956" bold="true" box="[151,256,1268,1295]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417958FB5CFDAC4957" authority="Franz, 1966" authorityName="Franz" authorityYear="1966" box="[265,566,1268,1295]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417958FB5CFE394956" bold="true" box="[265,419,1268,1295]" italics="true" pageId="6" pageNumber="151">Stenichnodes</emphasis>
<bibRefCitation id="EF8F4B57FFB38C4179FCFB5DFDAC4957" author="Franz, H." box="[429,566,1268,1295]" pageId="6" pageNumber="151" pagination="613 - 628" refId="ref7686" refString="Franz, H. (1966) Scydmaeniden aus Nigeria, Senegal, der Tschadrepublik und Ostafrika (Coleoptera, Scydmaenidae). The An- nals and Magazine of Natural History, Series 13, 8 (95), 613 - 628." type="journal article" year="1966">
Franz,
<quantity id="4CE69B43FFB38C4179AFFB5CFDAC4957" box="[510,566,1268,1294]" metricMagnitude="1" metricUnit="m" metricValue="4.99364" pageId="6" pageNumber="151" unit="in" value="1966.0">1966</quantity>
</bibRefCitation>
</taxonomicName>
in the pronotum lacking the antebasal groove and pits (present in
<taxonomicName id="4C1E4D25FFB38C417D11FB5CFE9C496B" authorityName="Franz" authorityYear="1966" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417D11FB5CFE9C496B" italics="true" pageId="6" pageNumber="151">Stenichnodes</emphasis>
</taxonomicName>
); largely obliterated hypomeral ridges (complete in
<taxonomicName id="4C1E4D25FFB38C417B06FAB0FC71496B" authorityName="Franz" authorityYear="1966" box="[855,1003,1304,1330]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B06FAB0FC71496B" box="[855,1003,1304,1330]" italics="true" pageId="6" pageNumber="151">Stenichnodes</emphasis>
</taxonomicName>
); the aedeagus presumably with parameres, with elongate endophallic sclerites and without the basal pumping apparatus (in
<taxonomicName id="4C1E4D25FFB38C417CFBFA94FAA4490F" authorityName="Franz" authorityYear="1966" box="[1194,1342,1340,1366]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417CFBFA94FAA4490F" box="[1194,1342,1340,1366]" italics="true" pageId="6" pageNumber="151">Stenichnodes</emphasis>
</taxonomicName>
without parameres and endophallic sclerites and with the basal pumping apparatus);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FA2DFCAB49BE" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FA2DFE9A49C6" bold="true" box="[151,256,1412,1439]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417954FA2CFDA349C6" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[261,569,1412,1439]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417954FA2CFE1C49C6" bold="true" box="[261,390,1412,1439]" italics="true" pageId="6" pageNumber="151">Stenichnus</emphasis>
<bibRefCitation id="EF8F4B57FFB38C4179DAFA2CFDA349C6" author="Thomson, C. G." box="[395,569,1412,1439]" pageId="6" pageNumber="151" refId="ref8599" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Vol. 1. Berlingska Boktryckeriet, Lund, 290 pp. https: // doi. org / 10.5962 / bhl. title. 138677" type="book" year="1859">
Thomson,
<quantity id="4CE69B43FFB38C417A50FA2CFDA349C6" box="[513,569,1412,1439]" metricMagnitude="1" metricUnit="m" metricValue="4.7218599999999995" pageId="6" pageNumber="151" unit="in" value="1859.0">1859</quantity>
</bibRefCitation>
</taxonomicName>
in subtriangular, flat mandibles (narrow, falciform, nearly cylindrical in crosssection in
<taxonomicName id="4C1E4D25FFB38C417968FA00FE28499B" authorityName="Thomson" authorityYear="1859" box="[313,434,1448,1474]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417968FA00FE28499B" box="[313,434,1448,1474]" italics="true" pageId="6" pageNumber="151">Stenichnus</emphasis>
</taxonomicName>
); the pronotum lacking the antebasal pits (present in
<taxonomicName id="4C1E4D25FFB38C417C5CFA00FB1C499B" authorityName="Thomson" authorityYear="1859" box="[1037,1158,1448,1474]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417C5CFA00FB1C499B" box="[1037,1158,1448,1474]" italics="true" pageId="6" pageNumber="151">Stenichnus</emphasis>
</taxonomicName>
); the mesoscutellum not exposed between elytral bases (in
<taxonomicName id="4C1E4D25FFB38C417A15FA64FD2749BF" authorityName="Thomson" authorityYear="1859" box="[580,701,1484,1510]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417A15FA64FD2749BF" box="[580,701,1484,1510]" italics="true" pageId="6" pageNumber="151">Stenichnus</emphasis>
</taxonomicName>
exposed);
</paragraph>
<paragraph id="8BA136A6FFB38C4178C6FA59FB1E4A0A" blockId="6.[151,1437,152,1619]" pageId="6" pageNumber="151">
<emphasis id="B96AEAB4FFB38C4178C6FA59FE9A4A52" bold="true" box="[151,256,1520,1547]" pageId="6" pageNumber="151">- from</emphasis>
<taxonomicName id="4C1E4D25FFB38C417957FA58FDF24A52" authority="Jaloszynski, 2013 a" authorityName="Jaloszynski" authorityYear="2013" box="[262,616,1520,1547]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417957FA58FE154A52" bold="true" box="[262,399,1520,1547]" italics="true" pageId="6" pageNumber="151">Zeanichnus</emphasis>
Jałoszyński, 2013a
</taxonomicName>
in the lateral sutures of submentum straight and posteriorly broadly sepa- rated (sinuate in
<taxonomicName id="4C1E4D25FFB38C41792FF9BCFE654A77" authorityName="Jaloszynski" authorityYear="2013" box="[382,511,1556,1582]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C41792FF9BCFE654A77" box="[382,511,1556,1582]" italics="true" pageId="6" pageNumber="151">Zeanichnus</emphasis>
</taxonomicName>
, their posterior ends approximate); the pronotum lacking antebasal pits and groove (present in
<taxonomicName id="4C1E4D25FFB38C417913F990FE594A0B" authorityName="Jaloszynski" authorityYear="2013" box="[322,451,1592,1618]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417913F990FE594A0B" box="[322,451,1592,1618]" italics="true" pageId="6" pageNumber="151">Zeanichnus</emphasis>
</taxonomicName>
); large and deep basal elytral foveae (vestigial in
<taxonomicName id="4C1E4D25FFB38C417BA3F990FBE94A0B" authorityName="Jaloszynski" authorityYear="2013" box="[1010,1139,1592,1618]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417BA3F990FBE94A0B" box="[1010,1139,1592,1618]" italics="true" pageId="6" pageNumber="151">Zeanichnus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA136A6FFB38C417896F929FBCF4ABA" blockId="6.[151,1437,1664,1763]" pageId="6" pageNumber="151">
In a preliminary phylogenetic analysis of
<taxonomicName id="4C1E4D25FFB38C417AC4F928FCB74AC3" box="[661,813,1664,1690]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
,
<taxonomicName id="4C1E4D25FFB38C417B69F928FC3F4AC3" authorityName="Casey" authorityYear="1897" box="[824,933,1664,1690]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B69F928FC3F4AC3" box="[824,933,1664,1690]" italics="true" pageId="6" pageNumber="151">Catalinus</emphasis>
</taxonomicName>
takes place as a sister genus to the Australian
<taxonomicName id="4C1E4D25FFB38C4178C6F90CFEF44AE7" authorityName="Franz" authorityYear="1975" box="[151,366,1700,1726]" class="Insecta" family="Staphylinidae" genus="Palaeoscydmaenus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C4178C6F90CFEF44AE7" box="[151,366,1700,1726]" italics="true" pageId="6" pageNumber="151">Palaeoscydmaenus</emphasis>
</taxonomicName>
(Jałoszyński, unpublished), but this placement is weakly supported and requires verification, as morphologically
<taxonomicName id="4C1E4D25FFB38C417906F960FE5E4ABB" authorityName="Casey" authorityYear="1897" box="[343,452,1736,1762]" class="Insecta" family="Staphylinidae" genus="Catalinus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417906F960FE5E4ABB" box="[343,452,1736,1762]" italics="true" pageId="6" pageNumber="151">Catalinus</emphasis>
</taxonomicName>
seems to be more similar to the
<collectingCountry id="F3097636FFB38C417B61F961FC5C4ABB" box="[816,966,1736,1763]" name="New Zealand" pageId="6" pageNumber="151">New Zealand</collectingCountry>
<taxonomicName id="4C1E4D25FFB38C417B9CF960FBD44ABB" authorityName="Jaloszynski" authorityYear="2013" box="[973,1102,1736,1762]" class="Insecta" family="Staphylinidae" genus="Zeanichnus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAB4FFB38C417B9CF960FBD44ABB" box="[973,1102,1736,1762]" italics="true" pageId="6" pageNumber="151">Zeanichnus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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