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<document id="F2AE18F1A5F8EB7ED7672AD0A79B2165" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF4CCD9EFF4CEA57B080ED7C" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cryptacanthophorella manubriata" docType="treatment" docVersion="8" lastPageNumber="161" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="156" updateTime="1698435542289" updateUser="ExternalLinkService">
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<mods:title id="4F976E41920A7E76A90E74FA071F72FB">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
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<mods:name id="A3A0636DA4D16365D0515507E1961D5F" type="personal">
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<mods:namePart id="71A2B241BD9D5D6898CFD2BC9756E187">Antić, Dragan Ž.</mods:namePart>
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<mods:name id="895E1BFB9247FA89C7C6A8EFA2D593FF" type="personal">
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<mods:roleTerm id="1337C9DE44F3E0FAAB6BA5B24CEF98F7">Author</mods:roleTerm>
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<mods:namePart id="CC12B04434EBA874E82256517940D3B2">Makarov, Slobodan E.</mods:namePart>
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<mods:typeOfResource id="55D3085CA27BCF6471838331D5559410">text</mods:typeOfResource>
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<mods:title id="489DE50631480651977E557E753424FC">Zootaxa</mods:title>
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<mods:date id="54424E49FEB7F470382CECAB939D69BE">2016</mods:date>
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<mods:number id="D7CB0DD254E8A8AFE4E43C86607F7D1C">4211</mods:number>
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<mods:identifier id="1E602A3A5D278391A85FCA4125C33EC4" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
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<treatment id="03B7878CFF4CCD9EFF4CEA57B080ED7C" ID-DOI="http://doi.org/10.5281/zenodo.6074476" ID-GBIF-Taxon="127419183" ID-Zenodo-Dep="6074476" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C" httpUri="http://treatment.plazi.org/id/03B7878CFF4CCD9EFF4CEA57B080ED7C" lastPageId="160" lastPageNumber="161" pageId="155" pageNumber="156">
<subSubSection id="C3046511FF4CCDA5FF4CEA57B043EB17" pageId="155" pageNumber="156" type="nomenclature">
<paragraph id="8BA1369AFF4CCDA5FF4CEA57B381EB75" blockId="155.[151,754,546,606]" box="[151,754,546,572]" pageId="155" pageNumber="156">
<heading id="D0E981F6FF4CCDA5FF4CEA57B381EB75" bold="true" box="[151,754,546,572]" fontSize="11" level="1" pageId="155" pageNumber="156" reason="1">
<emphasis id="B96AEA88FF4CCDA5FF4CEA57B381EB75" bold="true" box="[151,754,546,572]" pageId="155" pageNumber="156">
<taxonomicName id="4C1E4D19FF4CCDA5FF4CEA57B347EB75" ID-CoL="847XK" box="[151,564,546,572]" class="Diplopoda" family="Anthroleucosomatidae" genus="Cryptacanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96AEA88FF4CCDA5FF4CEA57B347EB75" bold="true" box="[151,564,546,572]" italics="true" pageId="155" pageNumber="156">Cryptacanthophorella manubriata</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF4CCDA5FF4CEA30B043EB17" blockId="155.[151,754,546,606]" box="[151,304,581,606]" pageId="155" pageNumber="156">
<figureCitation id="13252A1FFF4CCDA5FF4CEA30B043EB17" box="[151,304,581,606]" captionStart-0="FIGURE 129" captionStart-1="FIGURE 130" captionStart-2="FIGURE 131" captionStart-3="FIGURE 132" captionStart-4="FIGURE 133" captionStartId-0="155.[151,250,1935,1957]" captionStartId-1="156.[151,250,1915,1937]" captionStartId-2="157.[151,250,1959,1981]" captionStartId-3="158.[151,250,1186,1208]" captionStartId-4="159.[151,250,1815,1837]" captionTargetBox-0="[188,1398,982,1914]" captionTargetBox-1="[228,1359,193,1894]" captionTargetBox-2="[256,1330,193,1938]" captionTargetBox-3="[291,1278,223,1073]" captionTargetBox-4="[260,1326,193,1794]" captionTargetId-0="figure@155.[188,1398,982,1914]" captionTargetId-1="figure@156.[228,1359,193,1894]" captionTargetId-2="figure@157.[256,1330,193,1938]" captionTargetId-3="figure@158.[240,1347,167,1165]" captionTargetId-4="figure@159.[260,1326,193,1794]" captionTargetPageId-0="155" captionTargetPageId-1="156" captionTargetPageId-2="157" captionTargetPageId-3="158" captionTargetPageId-4="159" captionText-0="FIGURE 129. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia, habitus. Scale line = 1 mm." captionText-1="FIGURE 130. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia. A. Pleurotergite 15, posterior view. B. Pleurotergite 14 and 15, lateral view. C. Prozonite 15, dorsal view. D. Lateral keel 15, dorsal view. E. Pleurotergite 14 and 15, dorsal view." captionText-2="FIGURE 131. Cryptacanthophorella manubriata gen. et sp. nov., holotype male. A. Leg 3. B. Leg 7. C. Leg 4. D. Leg 10. Scale lines = 0.2 mm." captionText-3="FIGURE 132. Cryptacanthophorella manubriata gen. et sp. nov., holotype male anterior gonopods, posterior view. h = horn; hl = hairy lever; lv = lever; wp = nippled part; apr = subapical process; aps = process with minute setae; hap = hidden anterior process. Scale line = 0.2 mm." captionText-4="FIGURE 133. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia, gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, anterolateral view. C. Anterior gonopods, lateral view. D. and E. Anterior gonopods, posterolateral view. F. Anterior gonopods, posterior view. G. Posterior gonopods, posterior view. H. Posterior gonopods, lateral view. c = coxite; h = horn; t = telopodite; al = anterior lateral structure; cp = coxal process; cv = coxal vesicle; hl = hairy structure; lg = medial groove; lp = lateral part; lv = lever; mc = membranous crest; mp = medial part; sp = sternal plate; ss = sternal sac; wp = nippled part; apr = subapical process; aps = process with minute setae; hap = hidden anterior process; scv = syncoxal vesicle." httpUri-0="https://zenodo.org/record/272878/files/figure.png" httpUri-1="https://zenodo.org/record/272879/files/figure.png" httpUri-2="https://zenodo.org/record/272880/files/figure.png" httpUri-3="https://zenodo.org/record/272881/files/figure.png" httpUri-4="https://zenodo.org/record/272882/files/figure.png" pageId="155" pageNumber="156">Figs 129133</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF4CCDA5FF4CEAF8B0ABEBEF" box="[151,472,653,678]" pageId="155" pageNumber="156" type="diagnosis">
<paragraph id="8BA1369AFF4CCDA5FF4CEAF8B0ABEBEF" blockId="155.[151,1436,653,930]" box="[151,472,653,678]" pageId="155" pageNumber="156">
<emphasis id="B96AEA88FF4CCDA5FF4CEAF8B062EBEF" bold="true" box="[151,273,653,678]" pageId="155" pageNumber="156">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF4CCDA5FF1CEAC4B078EBA7" pageId="155" pageNumber="156" type="etymology">
<paragraph id="8BA1369AFF4CCDA5FF1CEAC4B078EBA7" blockId="155.[151,1436,653,930]" pageId="155" pageNumber="156">
<emphasis id="B96AEA88FF4CCDA5FF1CEAC4B03DEB83" bold="true" box="[199,334,689,714]" pageId="155" pageNumber="156">Etymology.</emphasis>
An adjective; to emphasize the presence of a pair of posterior levers (handles) on the anterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF4CCDA0FF1CEA8CB03DEF1F" lastPageId="158" lastPageNumber="159" pageId="155" pageNumber="156" type="materials_examined">
<paragraph id="8BA1369AFF4CCDA5FF1CEA8CB372EA13" blockId="155.[151,1436,653,930]" pageId="155" pageNumber="156">
<materialsCitation id="3B763CC7FF4CCDA5FF1CEA8CB372EA13" ID-GBIF-Occurrence="1424566946" collectingDate="1983-05-16" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" county="Baniskhevi Valley" elevation="850" location="Material" municipality="Borzhomi Nature Reserve" pageId="155" pageNumber="156" specimenCount="17" specimenCount-female="9" specimenCount-male="8" typeStatus="holotype">
<emphasis id="B96AEA88FF4CCDA5FF1CEA8CB0E6EA5B" bold="true" box="[199,405,761,786]" pageId="155" pageNumber="156">
<location id="8EC16041FF4CCDA5FF1CEA8CB05CEA5B" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C:8EC16041FF4CCDA5FF1CEA8CB05CEA5B" box="[199,303,761,786]" country="Georgia" county="Baniskhevi Valley" municipality="Borzhomi Nature Reserve" name="Material" pageId="155" pageNumber="156">Material</location>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FF4CCDA5FE2EEA8FB325EA5B" box="[501,598,761,786]" pageId="155" pageNumber="156" type="male">8 males</specimenCount>
,
<specimenCount id="9D18FD13FF4CCDA5FDB1EA8FB391EA5B" box="[618,738,761,786]" pageId="155" pageNumber="156" type="female">9 females</specimenCount>
).
<emphasis id="B96AEA88FF4CCDA5FD26EA8CB202EA5B" bold="true" box="[765,881,761,786]" pageId="155" pageNumber="156">
<typeStatus id="54A58838FF4CCDA5FD26EA8CB21FEA5B" box="[765,876,761,786]" pageId="155" pageNumber="156" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFF4CCDA5FC5BEA8FB570EA5B" box="[896,1027,762,786]" name="Georgia" pageId="155" pageNumber="156">GEORGIA</collectingCountry>
: male,
<collectingMunicipality id="6BC5ACE0FF4CCDA5FBBAEA8CB4EBEA58" box="[1121,1432,761,786]" pageId="155" pageNumber="156">Borzhomi Nature Reserve</collectingMunicipality>
,
<collectingCounty id="62C04E16FF4CCDA5FF4CEB68B016EA7F" box="[151,357,797,822]" pageId="155" pageNumber="156">Baniskhevi Valley</collectingCounty>
,
<elevation id="0033D1A9FF4CCDA5FEABEB6BB083EA7C" box="[368,496,798,822]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="155" pageNumber="156" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<quantity id="4CE69B7FFF4CCDA5FEABEB6BB083EA7C" box="[368,496,798,822]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="155" pageNumber="156" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800900 m</quantity>
</elevation>
asl,
<taxonomicName id="4C1E4D19FF4CCDA5FDFFEB6BB317EA7C" box="[548,612,798,821]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="155" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF4CCDA5FDFFEB6BB317EA7C" box="[548,612,798,821]" italics="true" pageId="155" pageNumber="156">Picea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D19FF4CCDA5FDA8EB6BB3CAEA7C" box="[627,697,798,821]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="155" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF4CCDA5FDA8EB6BB3CAEA7C" box="[627,697,798,821]" italics="true" pageId="155" pageNumber="156">Fagus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D19FF4CCDA5FD06EB6BB236EA7C" box="[733,837,798,821]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="155" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF4CCDA5FD06EB6BB236EA7C" box="[733,837,798,821]" italics="true" pageId="155" pageNumber="156">Carpinus</emphasis>
</taxonomicName>
forest, litter, logs, under stones, 12,
<collectingDate id="EFE4E9B2FF4CCDA5FB3BEB68B40AEA7F" box="[1248,1401,797,822]" pageId="155" pageNumber="156" value="1983-05-16">
<date id="FFA0105AFF4CCDA5FB3BEB68B40AEA7F" box="[1248,1401,797,822]" pageId="155" pageNumber="156" value="1983-05-16">16 May 1983</date>
</collectingDate>
,
<collectorName id="26EB534CFF4CCDA5FA5EEB6BB07FEA13" pageId="155" pageNumber="156">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF4CCDA5FE91EB37B0D0EA13" box="[330,419,834,858]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="155" pageNumber="156">ZMUM</collectionCode>
ρ3405).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF4CCDA5FF1CEB10B19DEAEB" blockId="155.[151,1436,653,930]" pageId="155" pageNumber="156">
<emphasis id="B96AEA88FF4CCDA5FF1CEB10B04CEA37" bold="true" box="[199,319,869,894]" pageId="155" pageNumber="156">Paratypes</emphasis>
(total: 1 male, 6 females).
<materialsCitation id="3B763CC7FF4CCDA5FDB4EB10B19DEAEB" ID-GBIF-Occurrence="1424566928" collectingDate="1983-05-16" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" county="Baniskhevi Valley" elevation="850" location="All" municipality="Borzhomi Nature Reserve" pageId="155" pageNumber="156" specimenCount="7" specimenCount-female="6" specimenCount-male="1">
<location id="8EC16041FF4CCDA5FDB4EB10B3E0EA37" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C:8EC16041FF4CCDA5FDB4EB10B3E0EA37" box="[623,659,869,894]" country="Georgia" name="All" pageId="155" pageNumber="156">All</location>
from
<collectingCountry id="F309760AFF4CCDA5FD02EB13B228EA37" box="[729,859,870,894]" name="Georgia" pageId="155" pageNumber="156">GEORGIA</collectingCountry>
:
<specimenCount id="9D18FD13FF4CCDA5FCBDEB13B2C6EA37" box="[870,949,869,894]" pageId="155" pageNumber="156" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF4CCDA5FC1AEB10B543EA37" box="[961,1072,869,894]" pageId="155" pageNumber="156" type="female">6 females</specimenCount>
, same data as holotype (
<collectionCode id="ED0FAE5FFF4CCDA5FA98EB13B4EFEA37" box="[1347,1436,870,894]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="155" pageNumber="156">ZMUM</collectionCode>
ρ3406).
</materialsCitation>
</paragraph>
<caption id="DF616612FF4CCDA5FF4CEFFAB0DFEE8D" httpUri="https://zenodo.org/record/272878/files/figure.png" pageId="155" pageNumber="156" targetBox="[188,1398,982,1914]" targetPageId="155">
<paragraph id="8BA1369AFF4CCDA5FF4CEFFAB0DFEE8D" blockId="155.[151,1436,1935,1988]" pageId="155" pageNumber="156">
<emphasis id="B96AEA88FF4CCDA5FF4CEFFAB05EEEED" bold="true" box="[151,301,1935,1957]" pageId="155" pageNumber="156">FIGURE 129.</emphasis>
<taxonomicName id="4C1E4D19FF4CCDA5FEEEEFFAB3E0EEEC" box="[309,659,1935,1957]" class="Diplopoda" family="Anthroleucosomatidae" genus="Cryptacanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96AEA88FF4CCDA5FEEEEFFAB3E0EEEC" box="[309,659,1935,1957]" italics="true" pageId="155" pageNumber="156">Cryptacanthophorella manubriata</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF4CCDA5FD41EFE5B248EEEC" bold="true" box="[666,827,1936,1957]" pageId="155" pageNumber="156">gen. et sp. nov.</emphasis>
, male from near Bakhmaro, Chokhatauri District, Georgia, habitus. Scale line = 1 mm.
</paragraph>
</caption>
<caption id="DF616612FF4BCDA2FF4CEF0EB3DAEE86" httpUri="https://zenodo.org/record/272879/files/figure.png" pageId="156" pageNumber="157" targetBox="[228,1359,193,1894]" targetPageId="156">
<paragraph id="8BA1369AFF4BCDA2FF4CEF0EB3DAEE86" blockId="156.[151,1437,1915,1999]" pageId="156" pageNumber="157">
<emphasis id="B96AEA88FF4BCDA2FF4CEF0EB05EEED9" bold="true" box="[151,301,1915,1937]" pageId="156" pageNumber="157">FIGURE 130.</emphasis>
<taxonomicName id="4C1E4D19FF4BCDA2FEEEEF0EB3E0EED8" box="[309,659,1915,1937]" class="Diplopoda" family="Anthroleucosomatidae" genus="Cryptacanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="156" pageNumber="157" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96AEA88FF4BCDA2FEEEEF0EB3E0EED8" box="[309,659,1915,1937]" italics="true" pageId="156" pageNumber="157">Cryptacanthophorella manubriata</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF4BCDA2FD41EF09B248EED8" bold="true" box="[666,827,1916,1937]" pageId="156" pageNumber="157">gen. et sp. nov.</emphasis>
, male from near Bakhmaro, Chokhatauri District, Georgia.
<emphasis id="B96AEA88FF4BCDA2FF4CEFEFB1DCEEE6" bold="true" box="[151,175,1946,1967]" pageId="156" pageNumber="157">A.</emphasis>
Pleurotergite 15, posterior view.
<emphasis id="B96AEA88FF4BCDA2FDDCEFEFB36DEEE6" bold="true" box="[519,542,1946,1967]" pageId="156" pageNumber="157">B.</emphasis>
Pleurotergite 14 and 15, lateral view.
<emphasis id="B96AEA88FF4BCDA2FC73EFEFB2B3EEF9" bold="true" box="[936,960,1946,1968]" pageId="156" pageNumber="157">C.</emphasis>
Prozonite 15, dorsal view.
<emphasis id="B96AEA88FF4BCDA2FB00EFEFB587EEE6" bold="true" box="[1243,1268,1946,1967]" pageId="156" pageNumber="157">D.</emphasis>
Lateral keel 15, dorsal view.
<emphasis id="B96AEA88FF4BCDA2FECCEFCCB05DEE87" bold="true" box="[279,302,1977,1998]" pageId="156" pageNumber="157">E.</emphasis>
Pleurotergite 14 and 15, dorsal view.
</paragraph>
</caption>
<caption id="DF616612FF4ACDA3FF4CEFD2B000EE95" httpUri="https://zenodo.org/record/272880/files/figure.png" pageId="157" pageNumber="158" targetBox="[256,1330,193,1938]" targetPageId="157">
<paragraph id="8BA1369AFF4ACDA3FF4CEFD2B000EE95" blockId="157.[151,1436,1959,2012]" pageId="157" pageNumber="158">
<emphasis id="B96AEA88FF4ACDA3FF4CEFD2B05EEEF5" bold="true" box="[151,301,1959,1981]" pageId="157" pageNumber="158">FIGURE 131.</emphasis>
<taxonomicName id="4C1E4D19FF4ACDA3FEEEEFD2B3E0EEF4" box="[309,659,1959,1981]" class="Diplopoda" family="Anthroleucosomatidae" genus="Cryptacanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96AEA88FF4ACDA3FEEEEFD2B3E0EEF4" box="[309,659,1959,1981]" italics="true" pageId="157" pageNumber="158">Cryptacanthophorella manubriata</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF4ACDA3FD47EFD2B24DEEF5" bold="true" box="[668,830,1959,1980]" pageId="157" pageNumber="158">gen. et sp. nov.</emphasis>
, holotype male.
<emphasis id="B96AEA88FF4ACDA3FC36EFD2B575EEF5" bold="true" box="[1005,1030,1959,1980]" pageId="157" pageNumber="158">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FF4ACDA3FB83EFD2B51CEEF5" bold="true" box="[1112,1135,1959,1980]" pageId="157" pageNumber="158">B.</emphasis>
Leg 7.
<emphasis id="B96AEA88FF4ACDA3FB1AEFD2B5A9EEF4" bold="true" box="[1217,1242,1959,1981]" pageId="157" pageNumber="158">C.</emphasis>
Leg 4.
<emphasis id="B96AEA88FF4ACDA3FAF7EFD2B436EEF5" bold="true" box="[1324,1349,1959,1980]" pageId="157" pageNumber="158">D.</emphasis>
Leg 10. Scale lines = 0.2 mm.
</paragraph>
</caption>
<caption id="DF616612FF49CDA0FF4CECD7B0B1EDBF" httpUri="https://zenodo.org/record/272881/files/figure.png" pageId="158" pageNumber="159" targetBox="[291,1278,223,1073]" targetPageId="158">
<paragraph id="8BA1369AFF49CDA0FF4CECD7B0B1EDBF" blockId="158.[151,1436,1186,1270]" pageId="158" pageNumber="159">
<emphasis id="B96AEA88FF49CDA0FF4CECD7B05EEDFE" bold="true" box="[151,301,1186,1208]" pageId="158" pageNumber="159">FIGURE 132.</emphasis>
<taxonomicName id="4C1E4D19FF49CDA0FEE8ECD7B3E3EDF1" box="[307,656,1186,1208]" class="Diplopoda" family="Anthroleucosomatidae" genus="Cryptacanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="158" pageNumber="159" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96AEA88FF49CDA0FEE8ECD7B3E3EDF1" box="[307,656,1186,1208]" italics="true" pageId="158" pageNumber="159">Cryptacanthophorella manubriata</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF49CDA0FD4CECD6B245EDF1" bold="true" box="[663,822,1187,1208]" pageId="158" pageNumber="159">gen. et sp. nov.</emphasis>
, holotype male anterior gonopods, posterior view.
<emphasis id="B96AEA88FF49CDA0FAE5ECD7B438EDF1" box="[1342,1355,1186,1208]" italics="true" pageId="158" pageNumber="159">h</emphasis>
= horn;
<emphasis id="B96AEA88FF49CDA0FF4CECB4B1D8ED9E" box="[151,171,1217,1239]" italics="true" pageId="158" pageNumber="159">hl</emphasis>
= hairy lever;
<emphasis id="B96AEA88FF49CDA0FE9AECB4B020ED9E" box="[321,339,1217,1239]" italics="true" pageId="158" pageNumber="159">lv</emphasis>
= lever;
<emphasis id="B96AEA88FF49CDA0FE74ECB7B0BFED9E" box="[431,460,1218,1239]" italics="true" pageId="158" pageNumber="159">wp</emphasis>
= nippled part;
<emphasis id="B96AEA88FF49CDA0FDABECB7B3E0ED9E" box="[624,659,1218,1239]" italics="true" pageId="158" pageNumber="159">apr</emphasis>
= subapical process;
<emphasis id="B96AEA88FF49CDA0FCB5ECB7B2E2ED9E" box="[878,913,1218,1239]" italics="true" pageId="158" pageNumber="159">aps</emphasis>
= process with minute setae;
<emphasis id="B96AEA88FF49CDA0FB1BECB4B595ED9E" box="[1216,1254,1217,1239]" italics="true" pageId="158" pageNumber="159">hap</emphasis>
= hidden anterior process. Scale line = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF49CDA0FF1CED68B19DEF47" blockId="158.[151,1436,1309,2018]" pageId="158" pageNumber="159">
<materialsCitation id="3B763CC7FF49CDA0FF1CED68B2D0EC7F" ID-GBIF-Occurrence="1424566937" box="[199,931,1309,1334]" country="Georgia" location="GEORGIA" pageId="158" pageNumber="159" specimenCount="9" specimenCount-female="3" specimenCount-male="6">
<emphasis id="B96AEA88FF49CDA0FF1CED68B008EC7F" bold="true" box="[199,379,1309,1334]" pageId="158" pageNumber="159">Other material</emphasis>
(total:
<specimenCount id="9D18FD13FF49CDA0FE16ED68B355EC7F" box="[461,550,1309,1334]" pageId="158" pageNumber="159" type="male">6 males</specimenCount>
,
<specimenCount id="9D18FD13FF49CDA0FDE9ED6BB3D7EC7F" box="[562,676,1309,1334]" pageId="158" pageNumber="159" type="female">3 females</specimenCount>
). All from
<collectingCountry id="F309760AFF49CDA0FCFBED6BB2D0EC7F" box="[800,931,1310,1334]" name="Georgia" pageId="158" pageNumber="159">GEORGIA</collectingCountry>
</materialsCitation>
: 1 male,
<materialsCitation id="3B763CC7FF49CDA0FBDCED6BB096EC37" ID-GBIF-Occurrence="1424566936" collectingDate="1981-06-08" collectionCode="ZMUM" collectorName="S. Golovatch &amp; J. Martens" country="Georgia" location="Bakhmaro" municipality="Chokhatauri District" pageId="158" pageNumber="159" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D18FD13FF49CDA0FBDCED6BB51FEC7F" box="[1031,1132,1309,1334]" pageId="158" pageNumber="159" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BC5ACE0FF49CDA0FBACED68B413EC7F" box="[1143,1376,1309,1334]" pageId="158" pageNumber="159">Chokhatauri District</collectingMunicipality>
, near
<location id="8EC16041FF49CDA0FF4CED34B07CEC13" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C:8EC16041FF49CDA0FF4CED34B07CEC13" box="[151,271,1345,1370]" country="Georgia" municipality="Chokhatauri District" name="Bakhmaro" pageId="158" pageNumber="159">Bakhmaro</location>
,
<locationDeviation id="9A132FB8FF49CDA0FEC5ED37B354EC13" box="[286,551,1345,1370]" pageId="158" pageNumber="159">
<quantity id="4CE69B7FFF49CDA0FEC5ED37B019EC13" box="[286,362,1345,1370]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="158" pageNumber="159" unit="km" value="40.0">40 km</quantity>
SSE Nabeglavi
</locationDeviation>
, 15501700,
<taxonomicName id="4C1E4D19FF49CDA0FD1CED34B276EC13" box="[711,773,1345,1370]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FD1CED34B276EC13" box="[711,773,1345,1370]" italics="true" pageId="158" pageNumber="159">Abies</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D19FF49CDA0FCC3ED37B22BEC10" box="[792,856,1346,1369]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FCC3ED37B22BEC10" box="[792,856,1346,1369]" italics="true" pageId="158" pageNumber="159">Picea</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D19FF49CDA0FC5FED37B2B9EC10" box="[900,970,1346,1369]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FC5FED37B2B9EC10" box="[900,970,1346,1369]" italics="true" pageId="158" pageNumber="159">Fagus</emphasis>
</taxonomicName>
forest,
<collectingDate id="EFE4E9B2FF49CDA0FBF2ED37B5C5EC13" box="[1065,1206,1346,1370]" pageId="158" pageNumber="159" value="1981-06-08">
<date id="FFA0105AFF49CDA0FBF2ED37B5C5EC13" box="[1065,1206,1346,1370]" pageId="158" pageNumber="159" value="1981-06-08">8 Jun. 1981</date>
</collectingDate>
,
<collectorName id="26EB534CFF49CDA0FB1DED37B42EEC13" box="[1222,1373,1345,1370]" pageId="158" pageNumber="159">S. Golovatch</collectorName>
&amp;
<collectorName id="26EB534CFF49CDA0FA51ED37B180EC34" pageId="158" pageNumber="159">J. Martens</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF49CDA0FEE9ED13B0F8EC37" box="[306,395,1382,1406]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="158" pageNumber="159">ZMUM</collectionCode>
ρ3407)
</materialsCitation>
; 2 males (ZMUM ρ3408),
<materialsCitation id="3B763CC7FF49CDA0FCCEED13B0F5ECEB" ID-GBIF-Occurrence="1424566924" collectingDate="1981-06-08" collectionCode="IZB" collectorName="S. Golovatch &amp; J. Martens" country="Georgia" elevation="1985" location="Bakhmaro" municipality="Chokhatauri District" pageId="158" pageNumber="159" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D18FD13FF49CDA0FCCEED13B212EC37" box="[789,865,1381,1406]" pageId="158" pageNumber="159" type="male">1 male</specimenCount>
(
<collectionCode id="ED0FAE5FFF49CDA0FCAAED13B2D1EC37" box="[881,930,1382,1406]" pageId="158" pageNumber="159">IZB</collectionCode>
), same data, except:
<elevation id="0033D1A9FF49CDA0FB55ED10B45AEC34" box="[1166,1321,1381,1406]" metricMagnitude="3" metricUnit="m" metricValue="1.985" metricValueMax="2.02" metricValueMin="1.95" pageId="158" pageNumber="159" unit="m" value="1985.0" valueMax="2020.0" valueMin="1950.0">
<quantity id="4CE69B7FFF49CDA0FB55ED10B45AEC34" box="[1166,1321,1381,1406]" metricMagnitude="3" metricUnit="m" metricValue="1.985" metricValueMax="2.02" metricValueMin="1.95" pageId="158" pageNumber="159" unit="m" value="1985.0" valueMax="2020.0" valueMin="1950.0">19502020 m</quantity>
</elevation>
asl,
<taxonomicName id="4C1E4D19FF49CDA0FA85ED10B048ECEB" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="species" species="nordmanniana">
<emphasis id="B96AEA88FF49CDA0FA85ED10B048ECEB" italics="true" pageId="158" pageNumber="159">Abies nordmanniana</emphasis>
</taxonomicName>
forest
</materialsCitation>
;
<materialsCitation id="3B763CC7FF49CDA0FE4BEDFFB33DEC8F" ID-GBIF-Occurrence="1424566942" collectingDate="1981-10-13" collectionCode="IZB" collectorName="S. Golovatch" country="Georgia" elevation="800" location="Kintrish Nature Reserve" pageId="158" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Ajaria">
<specimenCount id="9D18FD13FF49CDA0FE4BEDFFB0ADECEB" box="[400,478,1417,1442]" pageId="158" pageNumber="159" type="male">1 male</specimenCount>
,
<collectingRegion id="49DAF878FF49CDA0FE31EDFCB340ECEB" box="[490,563,1417,1442]" country="Georgia" name="Ajaria" pageId="158" pageNumber="159">Ajaria</collectingRegion>
,
<location id="8EC16041FF49CDA0FDE6EDFCB222ECE8" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C:8EC16041FF49CDA0FDE6EDFCB222ECE8" box="[573,849,1417,1442]" country="Georgia" name="Kintrish Nature Reserve" pageId="158" pageNumber="159" stateProvince="Ajaria">Kintrish Nature Reserve</location>
,
<taxonomicName id="4C1E4D19FF49CDA0FC87EDFCB2A6ECEB" box="[860,981,1417,1442]" class="Diplopoda" family="Anthroleucosomatidae" genus="Dentatosoma" kingdom="Animalia" order="Chordeumatida" pageId="158" pageNumber="202" phylum="Arthropoda" rank="species" species="zeraboseli">Zeraboseli</taxonomicName>
,
<elevation id="0033D1A9FF49CDA0FC3BEDFFB554ECE8" box="[992,1063,1418,1442]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="158" pageNumber="159" unit="m" value="800.0">
<quantity id="4CE69B7FFF49CDA0FC3BEDFFB554ECE8" box="[992,1063,1418,1442]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="158" pageNumber="159" unit="m" value="800.0">800 m</quantity>
</elevation>
asl,
<taxonomicName id="4C1E4D19FF49CDA0FB80EDFCB473ECEB" box="[1115,1280,1417,1442]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FB80EDFCB473ECEB" box="[1115,1280,1417,1442]" italics="true" pageId="158" pageNumber="159">Rhododendron</emphasis>
</taxonomicName>
thicket, litter,
<collectingDate id="EFE4E9B2FF49CDA0FF4CEDDBB05DEC8F" box="[151,302,1454,1478]" pageId="158" pageNumber="159" value="1981-10-13">
<date id="FFA0105AFF49CDA0FF4CEDDBB05DEC8F" box="[151,302,1454,1478]" pageId="158" pageNumber="159" value="1981-10-13">13 Oct. 1981</date>
</collectingDate>
,
<collectorName id="26EB534CFF49CDA0FEE7EDDBB0A2EC8F" box="[316,465,1453,1478]" pageId="158" pageNumber="159">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF49CDA0FDCFEDDBB335EC8F" box="[532,582,1454,1478]" pageId="158" pageNumber="159">IZB</collectionCode>
)
</materialsCitation>
; 1 male,
<materialsCitation id="3B763CC7FF49CDA0FD62EDDBB19DEF47" ID-GBIF-Occurrence="1424566934" collectingDate="1983-05-17" collectingDateMax="2014-04" collectingDateMin="1983-05-17" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" elevation="1000" location="Pass" municipality="Surami" pageId="158" pageNumber="159" specimenCount="2" specimenCount-female="2" stateProvince="Ajaria">
<specimenCount id="9D18FD13FF49CDA0FD62EDDBB259EC8F" box="[697,810,1453,1478]" pageId="158" pageNumber="159" type="female">2 females</specimenCount>
,
<collectingMunicipality id="6BC5ACE0FF49CDA0FCECEDD8B2FBEC8F" box="[823,904,1453,1478]" pageId="158" pageNumber="159">Surami</collectingMunicipality>
(= Rikoti)
<location id="8EC16041FF49CDA0FBD0EDDBB533EC8C" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C:8EC16041FF49CDA0FBD0EDDBB533EC8C" box="[1035,1088,1454,1477]" country="Georgia" municipality="Surami" name="Pass" pageId="158" pageNumber="159" stateProvince="Ajaria">Pass</location>
,
<emphasis id="B96AEA88FF49CDA0FB96EDDBB51BEC8C" box="[1101,1128,1454,1477]" italics="true" pageId="158" pageNumber="159">ca</emphasis>
<elevation id="0033D1A9FF49CDA0FBABEDDBB5B4EC8C" box="[1136,1223,1454,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="158" pageNumber="159" unit="m" value="1000.0">
<quantity id="4CE69B7FFF49CDA0FBABEDDBB5B4EC8C" box="[1136,1223,1454,1478]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="158" pageNumber="159" unit="m" value="1000.0">1000 m</quantity>
</elevation>
asl,
<taxonomicName id="4C1E4D19FF49CDA0FB24EDDBB436EC8C" box="[1279,1349,1454,1477]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FB24EDDBB436EC8C" box="[1279,1349,1454,1477]" italics="true" pageId="158" pageNumber="159">Fagus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D19FF49CDA0FA8EEDD8B4E6EC8F" box="[1365,1429,1453,1478]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FA8EEDD8B4E6EC8F" box="[1365,1429,1453,1478]" italics="true" pageId="158" pageNumber="159">Alnus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D19FF49CDA0FF4CEDA4B04FECA3" box="[151,316,1489,1514]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FF4CEDA4B04FECA3" box="[151,316,1489,1514]" italics="true" pageId="158" pageNumber="159">Rhododendron</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D19FF49CDA0FE94EDA7B0CAECA0" box="[335,441,1490,1513]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FE94EDA7B0CAECA0" box="[335,441,1490,1513]" italics="true" pageId="158" pageNumber="159">Castanea</emphasis>
</taxonomicName>
forest, litter and under stones,
<collectingDate id="EFE4E9B2FF49CDA0FCF5EDA7B2F4ECA3" box="[814,903,1490,1514]" pageId="158" pageNumber="159" value="2014-04">
<date id="FFA0105AFF49CDA0FCF5EDA7B2F4ECA3" box="[814,903,1490,1514]" pageId="158" pageNumber="159" value="2014-04">14 Apr.</date>
</collectingDate>
&amp;
<collectingDate id="EFE4E9B2FF49CDA0FC6FEDA7B526ECA3" box="[948,1109,1490,1514]" pageId="158" pageNumber="159" value="1983-05-17">
<date id="FFA0105AFF49CDA0FC6FEDA7B526ECA3" box="[948,1109,1490,1514]" pageId="158" pageNumber="159" value="1983-05-17">17 May 1983</date>
</collectingDate>
,
<collectorName id="26EB534CFF49CDA0FBBEEDA7B58FECA3" box="[1125,1276,1489,1514]" pageId="158" pageNumber="159">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF49CDA0FA98EDA7B4EFECA3" box="[1347,1436,1490,1514]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="158" pageNumber="159">ZMUM</collectionCode>
ρ3409).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF49CDA0FF1CEE6CB03DEF1F" blockId="158.[151,1436,1309,2018]" pageId="158" pageNumber="159">
<materialsCitation id="3B763CC7FF49CDA0FF1CEE6CB03DEF1F" ID-GBIF-Occurrence="1424566940" country="Georgia" elevation="850" location="Borzhomi Nature Reserve" municipality="Baniskhevi Valley" pageId="158" pageNumber="159" specimenCount="1" typeStatus="holotype">
<emphasis id="B96AEA88FF49CDA0FF1CEE6CB01AEF7B" bold="true" box="[199,361,1561,1586]" pageId="158" pageNumber="159">
<typeStatus id="54A58838FF49CDA0FF1CEE6CB071EF7B" box="[199,258,1561,1586]" pageId="158" pageNumber="159">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFF49CDA0FEAFEE6FB084EF7B" box="[372,503,1562,1586]" name="Georgia" pageId="158" pageNumber="159">GEORGIA</collectingCountry>
:
<location id="8EC16041FF49CDA0FDDEEE6CB247EF78" LSID="urn:lsid:plazi:treatment:03B7878CFF4CCD9EFF4CEA57B080ED7C:8EC16041FF49CDA0FDDEEE6CB247EF78" box="[517,820,1561,1586]" country="Georgia" municipality="Baniskhevi Valley" name="Borzhomi Nature Reserve" pageId="158" pageNumber="159">Borzhomi Nature Reserve</location>
,
<collectingMunicipality id="6BC5ACE0FF49CDA0FC9FEE6CB565EF7B" box="[836,1046,1561,1586]" pageId="158" pageNumber="159">Baniskhevi Valley</collectingMunicipality>
,
<elevation id="0033D1A9FF49CDA0FBFFEE6FB5D4EF78" box="[1060,1191,1562,1586]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="158" pageNumber="159" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">
<quantity id="4CE69B7FFF49CDA0FBFFEE6FB5D4EF78" box="[1060,1191,1562,1586]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="158" pageNumber="159" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800900 m</quantity>
</elevation>
asl,
<emphasis id="B96AEA88FF49CDA0FB38EE6FB408EF78" box="[1251,1403,1562,1585]" italics="true" pageId="158" pageNumber="159">
<taxonomicName id="4C1E4D19FF49CDA0FB38EE6FB456EF78" box="[1251,1317,1562,1585]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
,
<taxonomicName id="4C1E4D19FF49CDA0FAEEEE6FB408EF78" box="[1333,1403,1562,1585]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
</emphasis>
&amp;
<taxonomicName id="4C1E4D19FF49CDA0FF4CEE4BB18CEF1C" box="[151,255,1598,1621]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="158" pageNumber="159" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF49CDA0FF4CEE4BB18CEF1C" box="[151,255,1598,1621]" italics="true" pageId="158" pageNumber="159">Carpinus</emphasis>
</taxonomicName>
forest.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF49CD9EFF1CEE14B080ED7C" lastPageId="160" lastPageNumber="161" pageId="158" pageNumber="159" type="description">
<paragraph id="8BA1369AFF49CDA0FF1CEE14B2D1EF33" blockId="158.[151,1436,1309,2018]" box="[199,930,1633,1658]" pageId="158" pageNumber="159">
<emphasis id="B96AEA88FF49CDA0FF1CEE14B02AEF33" bold="true" box="[199,345,1633,1658]" pageId="158" pageNumber="159">Description.</emphasis>
Body with 31 segments (including telson) in adults.
</paragraph>
<paragraph id="8BA1369AFF49CDA0FF1CEEF3B2CAEF88" blockId="158.[151,1436,1309,2018]" pageId="158" pageNumber="159">
MEASUREMENTS. Males
<quantity id="4CE69B7FFF49CDA0FE07EEF3B326EFD4" box="[476,597,1670,1694]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="158" pageNumber="159" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF49CDA0FB55EEF0B462EFD4" box="[1166,1297,1669,1694]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="158" pageNumber="159" unit="mm" value="0.925" valueMax="1.0" valueMin="0.85">0.851 mm</quantity>
. Females
<quantity id="4CE69B7FFF49CDA0FA5BEEF3B189EF88" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.15" metricValueMin="0.9" pageId="158" pageNumber="159" unit="mm" value="10.25" valueMax="11.5" valueMin="9.0">9 11.5 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF49CDA0FCE9EEDCB2C6EF88" box="[818,949,1705,1730]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="158" pageNumber="159" unit="mm" value="0.925" valueMax="1.0" valueMin="0.85">0.851 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFF49CDA0FF1CEEBBB5E1EFAF" blockId="158.[151,1436,1309,2018]" box="[199,1170,1741,1766]" pageId="158" pageNumber="159">
COLORATION (
<figureCitation id="13252A1FFF49CDA0FEB7EEB8B0BDEFAF" box="[364,462,1741,1766]" captionStart="FIGURE 129" captionStartId="155.[151,250,1935,1957]" captionTargetBox="[188,1398,982,1914]" captionTargetId="figure@155.[188,1398,982,1914]" captionTargetPageId="155" captionText="FIGURE 129. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia, habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272878/files/figure.png" pageId="158" pageNumber="159">Fig. 129</figureCitation>
). Yellowish, with traces of greyish and brownish pigmentation.
</paragraph>
<paragraph id="8BA1369AFF49CDA0FF1CEE87B067EEA8" blockId="158.[151,1436,1309,2018]" pageId="158" pageNumber="159">
HEAD. Without frontal depression in males. Labrum with three medial teeth and 5+4 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 6+5 setae, on each plate arranged in 12 irregular rows. Stipites with
<emphasis id="B96AEA88FF49CDA0FEABEF4FB0F8EE18" box="[368,395,1850,1873]" italics="true" pageId="158" pageNumber="159">ca</emphasis>
20+20 setae. Antennae
<quantity id="4CE69B7FFF49CDA0FD42EF4CB383EE18" box="[665,752,1849,1874]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="158" pageNumber="159" unit="mm" value="1.6">1.6 mm</quantity>
long in
<typeStatus id="54A58838FF49CDA0FC96EF4CB2C0EE1B" box="[845,947,1849,1874]" pageId="158" pageNumber="159" type="holotype">holotype</typeStatus>
. Length of antennomeres (in mm): I (0.08), II (0.18), III (0.42), IV (0.23), V (0.38), VI (0.15), VII (0.12) and VIII (0.04). Length/breadth ratios of antennomeres IVII: I (1), II (2), III (5), IV (2.5), V (3.5), VI (1.3) and VII (1.2). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 1518, arranged in 46 rows in both sexes.
</paragraph>
<caption id="DF616612FF48CDA1FF4CEF62B35DEEAC" httpUri="https://zenodo.org/record/272882/files/figure.png" pageId="159" pageNumber="160" targetBox="[260,1326,193,1794]" targetPageId="159">
<paragraph id="8BA1369AFF48CDA1FF4CEF62B35DEEAC" blockId="159.[151,1437,1815,2022]" pageId="159" pageNumber="160">
<emphasis id="B96AEA88FF48CDA1FF4CEF62B05EEE65" bold="true" box="[151,301,1815,1837]" pageId="159" pageNumber="160">FIGURE 133.</emphasis>
<taxonomicName id="4C1E4D19FF48CDA1FEEEEF62B3E0EE64" box="[309,659,1815,1837]" class="Diplopoda" family="Anthroleucosomatidae" genus="Cryptacanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="159" pageNumber="160" phylum="Arthropoda" rank="species" species="manubriata">
<emphasis id="B96AEA88FF48CDA1FEEEEF62B3E0EE64" box="[309,659,1815,1837]" italics="true" pageId="159" pageNumber="160">Cryptacanthophorella manubriata</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF48CDA1FD41EF6DB248EE64" bold="true" box="[666,827,1816,1837]" pageId="159" pageNumber="160">gen. et sp. nov.</emphasis>
, male from near Bakhmaro, Chokhatauri District, Georgia, gonopods.
<emphasis id="B96AEA88FF48CDA1FEDCEF43B06CEE02" bold="true" box="[263,287,1846,1867]" pageId="159" pageNumber="160">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FF48CDA1FD5DEF42B3EEEE05" bold="true" box="[646,669,1847,1868]" pageId="159" pageNumber="160">B.</emphasis>
Anterior gonopods, anterolateral view.
<emphasis id="B96AEA88FF48CDA1FBE8EF43B53FEE05" bold="true" box="[1075,1100,1846,1868]" pageId="159" pageNumber="160">C.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FF48CDA1FF4CEF20B1DCEE23" bold="true" box="[151,175,1877,1898]" pageId="159" pageNumber="160">D.</emphasis>
and
<emphasis id="B96AEA88FF48CDA1FF3AEF20B18BEE23" bold="true" box="[225,248,1877,1898]" pageId="159" pageNumber="160">E.</emphasis>
Anterior gonopods, posterolateral view.
<emphasis id="B96AEA88FF48CDA1FD4DEF20B3D4EE23" bold="true" box="[662,679,1877,1898]" pageId="159" pageNumber="160">F.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FF48CDA1FBCCEF20B55FEE22" bold="true" box="[1047,1068,1877,1899]" pageId="159" pageNumber="160">G.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B96AEA88FF48CDA1FF4CEF01B1C2EEC0" bold="true" box="[151,177,1908,1929]" pageId="159" pageNumber="160">H.</emphasis>
Posterior gonopods, lateral view.
<emphasis id="B96AEA88FF48CDA1FDD0EF01B365EEC0" box="[523,534,1908,1929]" italics="true" pageId="159" pageNumber="160">c</emphasis>
= coxite;
<emphasis id="B96AEA88FF48CDA1FDA6EF06B3F9EEC0" box="[637,650,1907,1929]" italics="true" pageId="159" pageNumber="160">h</emphasis>
= horn;
<emphasis id="B96AEA88FF48CDA1FD3AEF01B39BEEC0" box="[737,744,1908,1929]" italics="true" pageId="159" pageNumber="160">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FF48CDA1FCADEF06B2F9EEC0" box="[886,906,1907,1929]" italics="true" pageId="159" pageNumber="160">al</emphasis>
= anterior lateral structure;
<emphasis id="B96AEA88FF48CDA1FB7DEF01B5CDEEC0" box="[1190,1214,1908,1929]" italics="true" pageId="159" pageNumber="160">cp</emphasis>
= coxal process;
<emphasis id="B96AEA88FF48CDA1FAAAEF01B4F4EEC0" box="[1393,1415,1908,1929]" italics="true" pageId="159" pageNumber="160">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FF48CDA1FEFDEFE7B049EEE1" box="[294,314,1938,1960]" italics="true" pageId="159" pageNumber="160">hl</emphasis>
= hairy structure;
<emphasis id="B96AEA88FF48CDA1FE28EFE7B374EEE1" box="[499,519,1938,1960]" italics="true" pageId="159" pageNumber="160">lg</emphasis>
= medial groove;
<emphasis id="B96AEA88FF48CDA1FD64EFE7B3A0EEE1" box="[703,723,1938,1960]" italics="true" pageId="159" pageNumber="160">lp</emphasis>
= lateral part;
<emphasis id="B96AEA88FF48CDA1FCBEEFE7B204EEE1" box="[869,887,1938,1960]" italics="true" pageId="159" pageNumber="160">lv</emphasis>
= lever;
<emphasis id="B96AEA88FF48CDA1FC0AEFE6B29CEEE1" box="[977,1007,1939,1960]" italics="true" pageId="159" pageNumber="160">mc</emphasis>
= membranous crest;
<emphasis id="B96AEA88FF48CDA1FB15EFE6B59EEEE1" box="[1230,1261,1939,1960]" italics="true" pageId="159" pageNumber="160">mp</emphasis>
= medial part;
<emphasis id="B96AEA88FF48CDA1FA5EEFE6B4EFEEE1" box="[1413,1436,1939,1960]" italics="true" pageId="159" pageNumber="160">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF48CDA1FEE1EFC7B03DEE8E" box="[314,334,1970,1991]" italics="true" pageId="159" pageNumber="160">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF48CDA1FE33EFC7B376EE8E" box="[488,517,1970,1991]" italics="true" pageId="159" pageNumber="160">wp</emphasis>
= nippled part;
<emphasis id="B96AEA88FF48CDA1FD6BEFC7B3A0EE8E" box="[688,723,1970,1991]" italics="true" pageId="159" pageNumber="160">apr</emphasis>
= subapical process;
<emphasis id="B96AEA88FF48CDA1FC6EEFC7B2ABEE8E" box="[949,984,1970,1991]" italics="true" pageId="159" pageNumber="160">aps</emphasis>
= process with minute setae;
<emphasis id="B96AEA88FF48CDA1FAC9EFC4B44BEE8E" box="[1298,1336,1969,1991]" italics="true" pageId="159" pageNumber="160">hap</emphasis>
= hidden anterior process;
<emphasis id="B96AEA88FF48CDA1FE9EEFA4B016EEAF" box="[325,357,2001,2022]" italics="true" pageId="159" pageNumber="160">scv</emphasis>
= syncoxal vesicle.
</paragraph>
</caption>
<paragraph id="8BA1369AFF77CD9EFF1CE8EDB189E99D" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BA1369AFF77CD9EFF1CE895B0C1E809" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">
BODY SEGMENTS (
<figureCitation id="13252A1FFF77CD9EFE7FE8AAB363E9B1" box="[420,528,223,248]" captionStart="FIGURE 129" captionStartId="155.[151,250,1935,1957]" captionTargetBox="[188,1398,982,1914]" captionTargetId="figure@155.[188,1398,982,1914]" captionTargetPageId="155" captionText="FIGURE 129. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia, habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272878/files/figure.png" pageId="160" pageNumber="161">Figs 129</figureCitation>
,
<figureCitation id="13252A1FFF77CD9EFDF8E895B322E9B1" box="[547,593,224,248]" captionStart="FIGURE 130" captionStartId="156.[151,250,1915,1937]" captionTargetBox="[228,1359,193,1894]" captionTargetId="figure@156.[228,1359,193,1894]" captionTargetPageId="156" captionText="FIGURE 130. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia. A. Pleurotergite 15, posterior view. B. Pleurotergite 14 and 15, lateral view. C. Prozonite 15, dorsal view. D. Lateral keel 15, dorsal view. E. Pleurotergite 14 and 15, dorsal view." httpUri="https://zenodo.org/record/272879/files/figure.png" pageId="160" pageNumber="161">130</figureCitation>
). Lateral keels like lateral swellings. Macrochaetae medium-sized and trichoid. CIX (pleurotergite 15) ~ 1.4; MIX (pleurotergite 15) ~ 0.5; PIX (pleurotergite 15) ~ 0.7; MA (pleurotergite 15) ~ 110˚.
</paragraph>
<paragraph id="8BA1369AFF77CD9EFF1CE938B39FE8CE" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<paragraph id="8BA1369AFF77CD9EFF1CE9E0B295E886" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">WALKING LEGS. In both sexes, leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae.</paragraph>
<paragraph id="8BA1369AFF77CD9EFF1CE9A8B2D4EBCC" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFF77CD9EFDCDE9A9B304E8BD" box="[534,631,476,501]" captionStart="FIGURE 131" captionStartId="157.[151,250,1959,1981]" captionTargetBox="[256,1330,193,1938]" captionTargetId="figure@157.[256,1330,193,1938]" captionTargetPageId="157" captionText="FIGURE 131. Cryptacanthophorella manubriata gen. et sp. nov., holotype male. A. Leg 3. B. Leg 7. C. Leg 4. D. Leg 10. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272880/files/figure.png" pageId="160" pageNumber="161">Fig. 131</figureCitation>
). Leg-pairs 37 enlarged, especially so leg-pairs 3, 4 and 7. Leg-pairs 3 and 4 each with a basal external protrusion on prefemur; prefemur and femur thickened. Leg-pair 5 with a basal, subtriangular, oral protrusion on femur. Leg-pair 6 without peculiarities. Leg-pair 7 with a well-developed, spatulated, lamellar, posterior, coxal process; prefemora with a bump. Leg-pair 10 with coxal glands and a posterior coxal horn. Leg-pair 11 with coxal glands, without other peculiarities.
</paragraph>
<paragraph id="8BA1369AFF77CD9EFF1CEAE5B0D7EA81" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFF77CD9EFE0BEAFAB345EBE1" box="[464,566,655,680]" captionStart="FIGURE 132" captionStartId="158.[151,250,1186,1208]" captionTargetBox="[291,1278,223,1073]" captionTargetId="figure@158.[240,1347,167,1165]" captionTargetPageId="158" captionText="FIGURE 132. Cryptacanthophorella manubriata gen. et sp. nov., holotype male anterior gonopods, posterior view. h = horn; hl = hairy lever; lv = lever; wp = nippled part; apr = subapical process; aps = process with minute setae; hap = hidden anterior process. Scale line = 0.2 mm." httpUri="https://zenodo.org/record/272881/files/figure.png" pageId="160" pageNumber="161">Figs 132</figureCitation>
,
<figureCitation id="13252A1FFF77CD9EFD9FEAE5B31DEBE1" box="[580,622,656,680]" captionStart="FIGURE 133" captionStartId="159.[151,250,1815,1837]" captionTargetBox="[260,1326,193,1794]" captionTargetId="figure@159.[260,1326,193,1794]" captionTargetPageId="159" captionText="FIGURE 133. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia, gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, anterolateral view. C. Anterior gonopods, lateral view. D. and E. Anterior gonopods, posterolateral view. F. Anterior gonopods, posterior view. G. Posterior gonopods, posterior view. H. Posterior gonopods, lateral view. c = coxite; h = horn; t = telopodite; al = anterior lateral structure; cp = coxal process; cv = coxal vesicle; hl = hairy structure; lg = medial groove; lp = lateral part; lv = lever; mc = membranous crest; mp = medial part; sp = sternal plate; ss = sternal sac; wp = nippled part; apr = subapical process; aps = process with minute setae; hap = hidden anterior process; scv = syncoxal vesicle." httpUri="https://zenodo.org/record/272882/files/figure.png" pageId="160" pageNumber="161">133</figureCitation>
AF). Sternal plate (
<emphasis id="B96AEA88FF77CD9EFC81EAE4B200EBE1" box="[858,883,657,680]" italics="true" pageId="160" pageNumber="161">sp</emphasis>
) with an anterior, medial, hairy, sternal sac (
<emphasis id="B96AEA88FF77CD9EFAADEAE4B4FFEBE1" box="[1398,1420,657,680]" italics="true" pageId="160" pageNumber="161">ss</emphasis>
). Coxal processes completely fused basally; consisting of a medial part (
<emphasis id="B96AEA88FF77CD9EFC65EAC0B293EB85" box="[958,992,693,716]" italics="true" pageId="160" pageNumber="161">mp</emphasis>
) and two lateral parts (
<emphasis id="B96AEA88FF77CD9EFB30EAC1B472EB84" box="[1259,1281,692,717]" italics="true" pageId="160" pageNumber="161">lp</emphasis>
). Medial part with a longitudinal medial groove (
<emphasis id="B96AEA88FF77CD9EFDFDEAA2B34FEBB9" box="[550,572,727,752]" italics="true" pageId="160" pageNumber="161">lg</emphasis>
); apically, with a pair of posteriorly curved horns (
<emphasis id="B96AEA88FF77CD9EFBA4EAA2B5FEEBB9" box="[1151,1165,727,752]" italics="true" pageId="160" pageNumber="161">h</emphasis>
); subapically, on lateral side with a U-shaped rift; posteriorly, with a nippled part (
<emphasis id="B96AEA88FF77CD9EFCC4EA88B233EA5D" box="[799,832,765,788]" italics="true" pageId="160" pageNumber="161">wp</emphasis>
). Lateral parts with an anterior lamellar structure (
<emphasis id="B96AEA88FF77CD9EFAAEEA89B4F9EA5C" box="[1397,1418,764,789]" italics="true" pageId="160" pageNumber="161">al</emphasis>
); posteriorly with a C-shaped lever (
<emphasis id="B96AEA88FF77CD9EFDF2EB6AB34EEA71" box="[553,573,799,824]" italics="true" pageId="160" pageNumber="161">lv</emphasis>
), this latter being orientated anteriorly; levers with an additional subapical process (
<emphasis id="B96AEA88FF77CD9EFF27EB30B050EA15" box="[252,291,837,860]" italics="true" pageId="160" pageNumber="161">apr</emphasis>
). Gonopods are characterized by the presence of a pair of hidden anterior processes (
<emphasis id="B96AEA88FF77CD9EFB3DEB31B463EA14" box="[1254,1296,836,861]" italics="true" pageId="160" pageNumber="161">hap</emphasis>
) situated on both sides of the medial part, only their upper third visible, passing through the U-shaped rift of medial part. Posteriorly, with a medial syncoxal vesicle (
<emphasis id="B96AEA88FF77CD9EFD59EBF8B3D5EAED" box="[642,678,909,932]" italics="true" pageId="160" pageNumber="161">scv</emphasis>
), as well as with a pair of hairy levers (
<emphasis id="B96AEA88FF77CD9EFB87EBF9B501EAEC" box="[1116,1138,908,933]" italics="true" pageId="160" pageNumber="161">hl</emphasis>
) and an additional process with minute setae (
<emphasis id="B96AEA88FF77CD9EFEB7EBC4B0E0EA81" box="[364,403,945,968]" italics="true" pageId="160" pageNumber="161">aps</emphasis>
).
</paragraph>
<paragraph id="8BA1369AFF77CD9EFF1CEBA0B080ED7C" blockId="160.[151,1437,151,1077]" pageId="160" pageNumber="161">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFF77CD9EFE0EEBA1B341EAA5" box="[469,562,980,1005]" captionStart="FIGURE 133" captionStartId="159.[151,250,1815,1837]" captionTargetBox="[260,1326,193,1794]" captionTargetId="figure@159.[260,1326,193,1794]" captionTargetPageId="159" captionText="FIGURE 133. Cryptacanthophorella manubriata gen. et sp. nov., male from near Bakhmaro, Chokhatauri District, Georgia, gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, anterolateral view. C. Anterior gonopods, lateral view. D. and E. Anterior gonopods, posterolateral view. F. Anterior gonopods, posterior view. G. Posterior gonopods, posterior view. H. Posterior gonopods, lateral view. c = coxite; h = horn; t = telopodite; al = anterior lateral structure; cp = coxal process; cv = coxal vesicle; hl = hairy structure; lg = medial groove; lp = lateral part; lv = lever; mc = membranous crest; mp = medial part; sp = sternal plate; ss = sternal sac; wp = nippled part; apr = subapical process; aps = process with minute setae; hap = hidden anterior process; scv = syncoxal vesicle." httpUri="https://zenodo.org/record/272882/files/figure.png" pageId="160" pageNumber="161">Fig. 133</figureCitation>
G, H). Coxited (
<emphasis id="B96AEA88FF77CD9EFD3CEBA0B380EAA5" box="[743,755,981,1004]" italics="true" pageId="160" pageNumber="161">c</emphasis>
) divided. Telpodites (
<emphasis id="B96AEA88FF77CD9EFC30EBA0B280EAA5" box="[1003,1011,981,1004]" italics="true" pageId="160" pageNumber="161">t</emphasis>
) present on posterolateral side. Coxal processes (
<emphasis id="B96AEA88FF77CD9EFECDEB8CB043ED59" box="[278,304,1017,1040]" italics="true" pageId="160" pageNumber="161">cp</emphasis>
) simple, curved posteriorly. Anterior side carrying a membranous crest (
<emphasis id="B96AEA88FF77CD9EFB5FEB8CB5D6ED59" box="[1156,1189,1017,1040]" italics="true" pageId="160" pageNumber="161">mc</emphasis>
). Coxal vesicles (
<emphasis id="B96AEA88FF77CD9EFAA1EB8CB4E0ED59" box="[1402,1427,1017,1040]" italics="true" pageId="160" pageNumber="161">cv</emphasis>
) placed basally on anterior side.
</paragraph>
</subSubSection>
</treatment>
</document>