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<document id="01C42C438AEAB760F4F547A3593F6CD7" ID-CLB-Dataset="31902" ID-DOI="10.11646/zootaxa.4329.2.1" ID-GBIF-Dataset="7e33ec7d-211b-4042-8d07-344364ee805d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1001915" ID-ZooBank="8226E27D-E3A9-481A-B51B-558643Bbb03A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1507106636447" checkinUser="plazi" docAuthor="Tarasov, Sergei" docDate="2017" docId="03C187E1AE7BB616DD9E4412FB25059F" docLanguage="en" docName="zootaxa.4329.2.1.pdf" docOrigin="Zootaxa 4329 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parachorius semsanganus Tarasov &amp; Keith 2011" docType="treatment" docVersion="11" lastPageNumber="139" masterDocId="FFF8FF99AE5DB630DD09474FFFDB0270" masterDocTitle="A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data" masterLastPageNumber="149" masterPageNumber="101" pageNumber="139" updateTime="1698469310636" updateUser="ExternalLinkService">
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<mods:title id="E671FE82865B8F970803274C73DBAF1F">A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data</mods:title>
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<treatment id="03C187E1AE7BB616DD9E4412FB25059F" ID-DOI="http://doi.org/10.5281/zenodo.3511542" ID-GBIF-Taxon="135106123" ID-Zenodo-Dep="3511542" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F" httpUri="http://treatment.plazi.org/id/03C187E1AE7BB616DD9E4412FB25059F" lastPageNumber="139" pageId="38" pageNumber="139">
<subSubSection id="C372657CAE7BB616DD9E4412FE9901E8" pageId="38" pageNumber="139" type="nomenclature">
<paragraph id="8BD736F7AE7BB616DD9E4412FD2D0107" blockId="38.[151,758,861,920]" box="[151,758,861,887]" pageId="38" pageNumber="139">
<heading id="D09F819BAE7BB616DD9E4412FD2D0107" bold="true" box="[151,758,861,887]" fontSize="11" level="1" pageId="38" pageNumber="139" reason="1">
<taxonomicName id="4C684D74AE7BB616DD9E4412FD2D0107" ID-CoL="6TNM9" authority="Tarasov &amp; Keith, 2011" authorityName="Tarasov &amp; Keith" authorityYear="2011" box="[151,758,861,887]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="139" phylum="Arthropoda" rank="species" species="semsanganus">
<emphasis id="B91CEAE5AE7BB616DD9E4412FD2D0107" bold="true" box="[151,758,861,887]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DD9E4412FE0C0107" bold="true" box="[151,471,861,887]" italics="true" pageId="38" pageNumber="139">Parachorius semsanganus</emphasis>
<bibRefCitation id="EFF94B06AE7BB616DCD74412FD2D0107" author="Tarasov" box="[478,758,861,887]" pageId="38" pageNumber="139" refString="Tarasov, S. &amp; Keith, D. (2011) Parachorius semsanganus sp. n. (Coleoptera, Scarabaeidae, Scarabaeinae) from Laos and its significance in the phylogeny of Oriental Deltochilini. Zookeys, 111, 51 - 57. https: // doi. org / 10.3897 / zookeys. 111.1221" type="journal article" year="2011">Tarasov &amp; Keith, 2011</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BD736F7AE7BB616DD9E4430FE9901E8" blockId="38.[151,758,861,920]" box="[151,322,895,920]" pageId="38" pageNumber="139">
(
<figureCitation id="13532A72AE7BB616DD964430FF2701E8" box="[159,252,895,920]" captionStart="FIGURE 22" captionStartId="39.[151,250,1152,1174]" captionTargetBox="[151,1430,195,1127]" captionTargetId="figure@39.[151,1436,195,1131]" captionTargetPageId="39" captionText="FIGURE 22. Morphology of Parachorius semsanganus. (A) aedeagus, lateral view (paratype). (B) aedeagus, dorsal view (paratype). (C) aedeagus, apical view (paratype). (D) aedeagus lateral view. (E) right metafemur (male, paratype). (F) habitus (male, holotype)." httpUri="https://zenodo.org/record/1001961/files/figure.png" pageId="38" pageNumber="139">Figs. 22</figureCitation>
,
<figureCitation id="13532A72AE7BB616DC0144CFFEE101E8" box="[264,314,896,920]" captionStart="FIGURE 27" captionStartId="44.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1748]" captionTargetId="figure@44.[151,1436,193,1748]" captionTargetPageId="44" captionText="FIGURE 27. Distribution maps of Parachorius species. (A) Parachorius peninsularis. (B) Parachorius pseudojavanus. (C) Parachorius schuelkei. (D) Parachorius semsanganus. (E) Parachorius solodovnikovi. (F) Parachorius thomsoni. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study. Literature data — localities retrieved from literature." httpUri="https://zenodo.org/record/1001971/files/figure.png" pageId="38" pageNumber="139">27D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7BB616DD9E4487FD6D01AE" box="[151,694,968,990]" pageId="38" pageNumber="139" type="reference_group">
<paragraph id="8BD736F7AE7BB616DD9E4487FD6D01AE" blockId="38.[151,694,968,990]" box="[151,694,968,990]" pageId="38" pageNumber="139">
<treatmentCitationGroup id="AB7811D9AE7BB616DD9E4487FD6D01AE" box="[151,694,968,990]" pageId="38" pageNumber="139">
<taxonomicName id="4C684D74AE7BB616DD9E4487FD6D01AE" ID-CoL="6TNM9" authority="Tarasov &amp; Keith, 2011: 51" authorityName="Tarasov &amp; Keith" authorityPageNumber="51" authorityYear="2011" box="[151,694,968,990]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="139" phylum="Arthropoda" rank="species" species="semsanganus">
<emphasis id="B91CEAE5AE7BB616DD9E4487FE7801AE" box="[151,419,968,990]" italics="true" pageId="38" pageNumber="139">Parachorius semsanganus</emphasis>
<treatmentCitation id="0AC910E6AE7BB616DCA04486FD6D01AE" ID-DOI="http://doi.org/10.5281/zenodo.3504928" ID-Zenodo-Dep="3504928" LSID="urn:lsid:zoobank.org:act:E79FA444-725E-40E6-986A-9FB48B7D232A" author="Tarasov" box="[425,694,968,990]" httpUri="http://treatment.plazi.org/id/5DC603CD659041AEA44DDF1C9BD81C2D" page="51" pageId="38" pageNumber="139" year="2011">
<bibRefCitation id="EFF94B06AE7BB616DCA04486FD4801AE" author="Tarasov" box="[425,659,968,990]" pageId="38" pageNumber="139" refString="Tarasov, S. &amp; Keith, D. (2011) Parachorius semsanganus sp. n. (Coleoptera, Scarabaeidae, Scarabaeinae) from Laos and its significance in the phylogeny of Oriental Deltochilini. Zookeys, 111, 51 - 57. https: // doi. org / 10.3897 / zookeys. 111.1221" type="journal article" year="2011">Tarasov &amp; Keith, 2011</bibRefCitation>
: 51
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</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7BB616DD9E4345FD030652" box="[151,728,1034,1059]" pageId="38" pageNumber="139" type="materials_examined">
<paragraph id="8BD736F7AE7BB616DD9E4345FD030652" blockId="38.[151,1436,1034,2031]" box="[151,728,1034,1059]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DD9E4345FEF60653" bold="true" box="[151,301,1034,1059]" pageId="38" pageNumber="139">Type locality</emphasis>
. Phou Sane Mountain, northern Laos.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7BB616DDCE4361FCD10637" box="[199,778,1070,1095]" pageId="38" pageNumber="139" type="distribution">
<paragraph id="8BD736F7AE7BB616DDCE4361FCD10637" blockId="38.[151,1436,1034,2031]" box="[199,778,1070,1095]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4361FEBB0637" bold="true" box="[199,352,1070,1095]" pageId="38" pageNumber="139">Distribution.</emphasis>
Northern
<collectingCountry id="F37F7667AE7BB616DCDA4360FDD70636" box="[467,524,1071,1094]" name="Laos" pageId="38" pageNumber="139">Laos</collectingCountry>
, Phou Sane Mountain.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7BB616DDCE431DFD7C06C3" pageId="38" pageNumber="139" type="diagnosis">
<paragraph id="8BD736F7AE7BB616DDCE431DFD7C06C3" blockId="38.[151,1436,1034,2031]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE431DFE9A061B" bold="true" box="[199,321,1106,1131]" pageId="38" pageNumber="139">Diagnosis.</emphasis>
This species stands out in being the largest representative of
<taxonomicName id="4C684D74AE7BB616DEFB431DFBA5061B" box="[1010,1150,1106,1131]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE7BB616DEFB431DFBA5061B" box="[1010,1150,1106,1131]" italics="true" pageId="38" pageNumber="139">Parachorius</emphasis>
</taxonomicName>
. It can be separated from all other species of
<taxonomicName id="4C684D74AE7BB616DC784339FE2606FF" box="[369,509,1142,1167]" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAE5AE7BB616DC784339FE2606FF" box="[369,509,1142,1167]" italics="true" pageId="38" pageNumber="139">Parachorius</emphasis>
</taxonomicName>
by metatibia with large teeth in males and inner margin denticulate and parameres with apex bent upward and membranous lobes.
</paragraph>
</subSubSection>
<subSubSection id="C372657CAE7BB616DDCE43F1FB0E04F7" pageId="38" pageNumber="139" type="description">
<paragraph id="8BD736F7AE7BB616DDCE43F1FB9A06A7" blockId="38.[151,1436,1034,2031]" box="[199,1089,1214,1239]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE43F1FE8206A7" bold="true" box="[199,345,1214,1239]" pageId="38" pageNumber="139">Description.</emphasis>
<emphasis id="B91CEAE5AE7BB616DC5643F1FE4406A7" box="[351,415,1214,1239]" italics="true" pageId="38" pageNumber="139">Body.</emphasis>
Body black, length
<quantity id="4C909B12AE7BB616DF8843F1FCC206A6" box="[641,793,1214,1239]" metricMagnitude="-3" metricUnit="m" metricValue="9.399999999999999" metricValueMax="10.6" metricValueMin="8.2" pageId="38" pageNumber="139" unit="mm" value="9.399999999999999" valueMax="10.6" valueMin="8.2">8.210.6 mm</quantity>
. Elytra uniformly colored.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE43ADFB41068B" blockId="38.[151,1436,1034,2031]" box="[199,1178,1250,1275]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE43ADFED0068B" box="[199,267,1250,1275]" italics="true" pageId="38" pageNumber="139">Head.</emphasis>
Clypeus with two teeth, outer margin of each tooth very slightly notched basally.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE4248FB580733" blockId="38.[151,1436,1034,2031]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4248FEE6076E" box="[199,317,1287,1310]" italics="true" pageId="38" pageNumber="139">Pronotum.</emphasis>
Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 12 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE4201FD1B0716" blockId="38.[151,1436,1034,2031]" box="[199,704,1358,1383]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4201FECE0717" box="[199,277,1358,1383]" italics="true" pageId="38" pageNumber="139">Elytra.</emphasis>
Elytra covered with sparse punctures.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE423BFD9407FB" blockId="38.[151,1436,1034,2031]" box="[199,591,1394,1419]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE423BFEC907FB" box="[199,274,1396,1419]" italics="true" pageId="38" pageNumber="139">Wings.</emphasis>
Wings normally developed.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE42D8FB1E044F" blockId="38.[151,1436,1034,2031]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE42D8FED807DE" box="[199,259,1431,1454]" italics="true" pageId="38" pageNumber="139">Legs.</emphasis>
Profemur with almost straight or sinuate anterior margin. Protibia with apical spur reaching protarsomere 2. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin serrate (sometimes slightly), simple, not modified. Metatibia slightly sinuate, inner margin denticulate with large teeth on inner margin, produced with a small angle apically.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE4103FBBD0413" blockId="38.[151,1436,1034,2031]" box="[199,1126,1610,1635]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4103FEB90413" box="[199,354,1612,1635]" italics="true" pageId="38" pageNumber="139">Metasternum.</emphasis>
Transverse basal ridge of metasternum reduced and extremely short.
</paragraph>
<paragraph id="8BD736F7AE7BB616DDCE4121FB0E04F7" blockId="38.[151,1436,1034,2031]" box="[199,1237,1646,1671]" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DDCE4121FEE004F7" box="[199,315,1646,1671]" italics="true" pageId="38" pageNumber="139">Aedeagus.</emphasis>
Parameres symmetrical, bearing membranous lobes and bending upward apically.
</paragraph>
</subSubSection>
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<paragraph id="8BD736F7AE7BB616DDCE41DDFB25059F" blockId="38.[151,1436,1034,2031]" pageId="38" pageNumber="139">
<materialsCitation id="3B003CAAAE7BB616DDCE41DDFE7C04BF" ID-GBIF-Occurrence="1658411524" collectionCode="LAOS-NE" country="Laos" latitude="19.636667" location="Material" longLatPrecision="12" longitude="103.33667" municipality="Phonsavan" pageId="38" pageNumber="139" specimenCount="1" specimenCount-male="1" stateProvince="Xieng Khouang" typeStatus="holotype">
<emphasis id="B91CEAE5AE7BB616DDCE41DDFDF204DB" bold="true" box="[199,553,1682,1707]" pageId="38" pageNumber="139">
<location id="8EB7602CAE7BB616DDCE41DDFEF404DB" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DDCE41DDFEF404DB" box="[199,303,1682,1707]" country="Laos" latitude="19.636667" longLatPrecision="12" longitude="103.33667" municipality="Phonsavan" name="Material" pageId="38" pageNumber="139" stateProvince="Xieng Khouang">Material</location>
examined.
<typeStatus id="54D38855AE7BB616DCB541DDFDF204DB" box="[444,553,1682,1707]" pageId="38" pageNumber="139" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F37F7667AE7BB616DF3341DDFDAF04DB" box="[570,628,1682,1707]" name="Laos" pageId="38" pageNumber="139">
<emphasis id="B91CEAE5AE7BB616DF3341DDFDAF04DB" bold="true" box="[570,628,1682,1707]" pageId="38" pageNumber="139">Laos</emphasis>
</collectingCountry>
:
<specimenCount id="9D6EFD7EAE7BB616DF8C41DDFD0D04DB" box="[645,726,1682,1707]" pageId="38" pageNumber="139" type="male">
<emphasis id="B91CEAE5AE7BB616DF8C41DDFD0D04DB" bold="true" box="[645,726,1682,1707]" pageId="38" pageNumber="139">1 male</emphasis>
</specimenCount>
, &quot;
<collectionCode id="ED79AE32AE7BB616DFE641DCFCB604DB" box="[751,877,1683,1707]" pageId="38" pageNumber="139">LAOS-NE</collectionCode>
,
<collectingRegion id="49ACF815AE7BB616DE7241DDFBF404DB" box="[891,1071,1682,1707]" country="Laos" name="Xiangkhouang" pageId="38" pageNumber="139">Xieng Khouang</collectingRegion>
prov.,
<geoCoordinate id="EE5C5030AE7BB616D98B41DCFAD904DB" box="[1154,1282,1683,1707]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="9" value="19.636667">19°38.20'N</geoCoordinate>
<geoCoordinate id="EE5C5030AE7BB616D80241DCFA4C04DB" box="[1291,1431,1683,1707]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="9" value="103.33667">103°20.20'E</geoCoordinate>
,
<collectingMunicipality id="6BB3AC8DAE7BB616DD9E41F9FECB04BF" box="[151,272,1718,1743]" pageId="38" pageNumber="139">Phonsavan</collectingMunicipality>
(
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NE)
</materialsCitation>
:
<materialsCitation id="3B003CAAAE7BB616DCBD41F8FB3204BF" ID-GBIF-Occurrence="1658411527" box="[436,1257,1718,1743]" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="PHOU, SANE" collectorName="D. Hauck" country="Laos" elevation="1420" location="Mt." pageId="38" pageNumber="139" specimenCount="1" stateProvince="Xieng Khouang" typeStatus="holotype">
<collectionCode id="ED79AE32AE7BB616DCBD41F8FD8904BF" box="[436,594,1719,1743]" pageId="38" pageNumber="139">PHOU SANE</collectionCode>
<location id="8EB7602CAE7BB616DF5541F8FD5D04BE" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DF5541F8FD5D04BE" box="[604,646,1719,1742]" country="Laos" name="Mt." pageId="38" pageNumber="139" stateProvince="Xieng Khouang">Mt.</location>
,
<elevation id="0045D1C4AE7BB616DF9A41F8FD3604BE" box="[659,749,1719,1743]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="38" pageNumber="139" unit="m" value="1420.0">
<quantity id="4C909B12AE7BB616DF9A41F8FD3604BE" box="[659,749,1719,1743]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="38" pageNumber="139" unit="m" value="1420.0">1420 m</quantity>
</elevation>
,
<collectingDate id="EF92E9DFAE7BB616DFF341F8FC4304BF" box="[762,920,1719,1743]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<date id="FFD61037AE7BB616DFF341F8FC4304BF" box="[762,920,1719,1743]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10.-30.v.2009</date>
</collectingDate>
,
<collectorName id="269D5321AE7BB616DEAC41F8FBC904BF" box="[933,1042,1718,1743]" pageId="38" pageNumber="139">D. Hauck</collectorName>
leg. / HMB Basel
</materialsCitation>
,
<materialsCitation id="3B003CAAAE7BB616D9FF41F8FBF20567" ID-GBIF-Occurrence="1658411541" collectionCode="NMPC, NHMB" country="Laos" county="Kuban" latitude="19.636667" location="Expedition" longLatPrecision="127" longitude="103.33667" municipality="Hauck" pageId="38" pageNumber="139" specimenCount="1" stateProvince="Prague" typeStatus="holotype">
<collectionCode id="ED79AE32AE7BB616D9FF41F8FA9E04BF" box="[1270,1349,1719,1743]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="38" pageNumber="139">NMPC</collectionCode>
<collectingRegion id="49ACF815AE7BB616D84741F8FA4704BF" box="[1358,1436,1719,1743]" country="Czech Republic" name="Praha" pageId="38" pageNumber="139">Prague</collectingRegion>
<collectingCountry id="F37F7667AE7BB616DD9E4194FF160482" box="[151,205,1755,1778]" name="Laos" pageId="38" pageNumber="139">Laos</collectingCountry>
2009
<location id="8EB7602CAE7BB616DC174195FE450483" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DC174195FE450483" box="[286,414,1754,1779]" country="Laos" county="Kuban" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Hauck" name="Expedition" pageId="38" pageNumber="139" stateProvince="Prague">Expedition</location>
: M. Brancucci, M. Geiser, Z.
<location id="8EB7602CAE7BB616DE044194FC890482" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DE044194FC890482" box="[781,850,1755,1778]" country="Laos" county="Kuban" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Hauck" name="Kraus" pageId="38" pageNumber="139" stateProvince="Prague">Kraus</location>
, D.
<collectingMunicipality id="6BB3AC8DAE7BB616DE824195FC0C0483" box="[907,983,1754,1779]" pageId="38" pageNumber="139">Hauck</collectingMunicipality>
, V.
<collectingCounty id="62B64E7BAE7BB616D9054195FB8C0483" box="[1036,1111,1754,1779]" pageId="38" pageNumber="139">Kubán</collectingCounty>
/
<typeStatus id="54D38855AE7BB616D9714194FAD70483" box="[1144,1292,1755,1779]" pageId="38" pageNumber="139" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C684D74AE7BB616D8114195FDEB0566" authority="D. Keith &amp; S. Tarasov" authorityName="D. Keith &amp; S. Tarasov" class="Insecta" family="Scarabaeidae" genus="Parachorius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="139" phylum="Arthropoda" rank="species" species="semsanganus">Parachorius semsanganus D. Keith &amp; S. Tarasov</taxonomicName>
det. 2011&quot;,
<geoCoordinate id="EE5C5030AE7BB616DFB341B0FCF50567" box="[698,814,1791,1815]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="92" value="19.636667">19°38.2'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DE3241B0FC6C0567" box="[827,951,1791,1815]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="92" value="103.33667">103°20.2'E</geoCoordinate>
(
<collectionCode id="ED79AE32AE7BB616DECE41B0FBFA0567" box="[967,1057,1791,1815]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="139">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE7BB616D93D41B1FD03052F" ID-GBIF-Occurrence="1658411534" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB" collectorName="M. Brancucci" country="Laos" county="Lam Phou Sane Mountain" elevation="1400" latitude="19.633333" location="Phonsavan" longLatPrecision="1271" longitude="103.333336" municipality="Ban Na" pageId="38" pageNumber="139" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Xieng Khouang" typeStatus="paratype">
<typeStatus id="54D38855AE7BB616D93D41B1FB770567" box="[1076,1196,1790,1815]" pageId="38" pageNumber="139" type="paratype">
<emphasis id="B91CEAE5AE7BB616D93D41B1FB770567" bold="true" box="[1076,1196,1790,1815]" pageId="38" pageNumber="139">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D6EFD7EAE7BB616D9B541B1FACD0567" box="[1212,1302,1790,1815]" pageId="38" pageNumber="139" type="male">
<emphasis id="B91CEAE5AE7BB616D9B541B1FACD0567" bold="true" box="[1212,1302,1790,1815]" pageId="38" pageNumber="139">4 males</emphasis>
</specimenCount>
,
<specimenCount id="9D6EFD7EAE7BB616D82C41B1FA4E0567" box="[1317,1429,1790,1815]" pageId="38" pageNumber="139" type="female">
<emphasis id="B91CEAE5AE7BB616D82C41B1FA4E0567" bold="true" box="[1317,1429,1790,1815]" pageId="38" pageNumber="139">3 females</emphasis>
</specimenCount>
,
<collectingRegion id="49ACF815AE7BB616DD9E406DFE90054B" box="[151,331,1826,1851]" country="Laos" name="Xiangkhouang" pageId="38" pageNumber="139">Xieng Khouang</collectingRegion>
Province,
<quantity id="4C909B12AE7BB616DCCE406CFDCB054B" box="[455,528,1826,1851]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="139" unit="km" value="30.0">30 km</quantity>
NE
<location id="8EB7602CAE7BB616DF4E406DFD1E054B" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DF4E406DFD1E054B" box="[583,709,1826,1851]" country="Laos" county="Lam Phou Sane Mountain" latitude="19.633333" longLatPrecision="1271" longitude="103.333336" municipality="Ban Na" name="Phonsavan" pageId="38" pageNumber="139" stateProvince="Xieng Khouang">Phonsavan</location>
:
<collectingMunicipality id="6BB3AC8DAE7BB616DFD8406CFCF1054A" box="[721,810,1827,1850]" pageId="38" pageNumber="139">Ban Na</collectingMunicipality>
,
<collectingCounty id="62B64E7BAE7BB616DE31406CFBBD054B" box="[824,1126,1826,1851]" pageId="38" pageNumber="139">Lam Phou Sane Mountain</collectingCounty>
,
<geoCoordinate id="EE5C5030AE7BB616D97A406CFB0A054B" box="[1139,1233,1827,1851]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="925" value="19.633333">19°38'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616D9D6406CFA92054B" box="[1247,1353,1827,1851]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="925" value="103.333336">103°20'E</geoCoordinate>
,
<elevation id="0045D1C4AE7BB616D85F406CFF36052E" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="38" pageNumber="139" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<quantity id="4C909B12AE7BB616D85F406CFF36052E" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="38" pageNumber="139" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300 1500 m</quantity>
</elevation>
, on
<collectingDate id="EF92E9DFAE7BB616DC124008FE6C052F" box="[283,439,1863,1887]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<date id="FFD61037AE7BB616DC124008FE6C052F" box="[283,439,1863,1887]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">1030/v/2009</date>
</collectingDate>
(
<collectorName id="269D5321AE7BB616DCCF4008FDBB052F" box="[454,608,1862,1887]" pageId="38" pageNumber="139">M. Brancucci</collectorName>
) (
<collectionCode id="ED79AE32AE7BB616DF7F4008FD0B052F" box="[630,720,1863,1887]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="139">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE7BB616DFEB4009FC1105D7" ID-GBIF-Occurrence="1658411530" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB" collectorName="Z. Kraus" country="Laos" elevation="1420" latitude="19.636667" location="Phou Sane Mountain" longLatPrecision="127" longitude="103.33667" municipality="Expedition" pageId="38" pageNumber="139" specimenCount="2" specimenCount-male="2" stateProvince="Xieng Khouang" typeStatus="paratype">
<typeStatus id="54D38855AE7BB616DFEB4009FC81052F" box="[738,858,1862,1887]" pageId="38" pageNumber="139" type="paratype">
<emphasis id="B91CEAE5AE7BB616DFEB4009FC81052F" bold="true" box="[738,858,1862,1887]" pageId="38" pageNumber="139">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D6EFD7EAE7BB616DE614009FC1B052F" box="[872,960,1862,1887]" pageId="38" pageNumber="139" type="male">
<emphasis id="B91CEAE5AE7BB616DE614009FC1B052F" bold="true" box="[872,960,1862,1887]" pageId="38" pageNumber="139">2 males</emphasis>
</specimenCount>
,
<collectingRegion id="49ACF815AE7BB616DEC74009FBA4052F" box="[974,1151,1862,1887]" country="Laos" name="Xiangkhouang" pageId="38" pageNumber="139">Xieng Khouang</collectingRegion>
Province, Phonsavan (
<quantity id="4C909B12AE7BB616D8764008FF6005F3" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="139" unit="km" value="30.0">30 km</quantity>
NE):
<location id="8EB7602CAE7BB616DC0A4025FE2E05F3" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DC0A4025FE2E05F3" box="[259,501,1898,1923]" country="Laos" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Expedition" name="Phou Sane Mountain" pageId="38" pageNumber="139" stateProvince="Xieng Khouang">Phou Sane Mountain</location>
,
<collectingCountry id="F37F7667AE7BB616DF0B4024FDE305F2" box="[514,568,1899,1922]" name="Laos" pageId="38" pageNumber="139">Laos</collectingCountry>
2009
<collectingMunicipality id="6BB3AC8DAE7BB616DF8B4025FCDA05F3" box="[642,769,1898,1923]" pageId="38" pageNumber="139">Expedition</collectingMunicipality>
: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubán,
<geoCoordinate id="EE5C5030AE7BB616DD9E40C0FED105D7" box="[151,266,1935,1959]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="92" value="19.636667">19°38.2'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DC1340C0FE4205D7" box="[282,409,1935,1959]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="92" value="103.33667">103°20.2'E</geoCoordinate>
,
<elevation id="0045D1C4AE7BB616DCAE40C0FDD905D6" box="[423,514,1935,1959]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="38" pageNumber="139" unit="m" value="1420.0">
<quantity id="4C909B12AE7BB616DCAE40C0FDD905D6" box="[423,514,1935,1959]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="38" pageNumber="139" unit="m" value="1420.0">1420 m</quantity>
</elevation>
, on
<collectingDate id="EF92E9DFAE7BB616DF3140C0FD0805D7" box="[568,723,1935,1959]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<date id="FFD61037AE7BB616DF3140C0FD0805D7" box="[568,723,1935,1959]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">1030/v/2009</date>
</collectingDate>
(
<collectorName id="269D5321AE7BB616DFEE40C0FC9405D7" box="[743,847,1935,1959]" pageId="38" pageNumber="139">Z. Kraus</collectorName>
) (
<collectionCode id="ED79AE32AE7BB616DE6140C0FC1905D7" box="[872,962,1935,1959]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="139">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B003CAAAE7BB616DED140C1FB21059F" ID-GBIF-Occurrence="1658411532" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB" collectorName="D. Hauck" country="Laos" elevation="1420" latitude="19.636667" location="Phou Sane Mountain" longLatPrecision="127" longitude="103.33667" municipality="Expedition" pageId="38" pageNumber="139" specimenCount="9" specimenCount-female="9" stateProvince="Xieng Khouang" typeStatus="paratype">
<typeStatus id="54D38855AE7BB616DED140C1FB8B05D7" box="[984,1104,1934,1959]" pageId="38" pageNumber="139" type="paratype">
<emphasis id="B91CEAE5AE7BB616DED140C1FB8B05D7" bold="true" box="[984,1104,1934,1959]" pageId="38" pageNumber="139">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D6EFD7EAE7BB616D96B40C1FB0E05D7" box="[1122,1237,1934,1959]" pageId="38" pageNumber="139" type="female">
<emphasis id="B91CEAE5AE7BB616D96B40C1FB0E05D7" bold="true" box="[1122,1237,1934,1959]" pageId="38" pageNumber="139">9 females</emphasis>
</specimenCount>
,
<collectingRegion id="49ACF815AE7BB616D9EF40C1FA4705D7" box="[1254,1436,1934,1959]" country="Laos" name="Xiangkhouang" pageId="38" pageNumber="139">Xieng Khouang</collectingRegion>
Province, Phonsavan (
<quantity id="4C909B12AE7BB616DC9340FCFE3C05BB" box="[410,487,1970,1995]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="38" pageNumber="139" unit="km" value="30.0">30 km</quantity>
NE):
<location id="8EB7602CAE7BB616DF3D40FDFCF005BB" LSID="urn:lsid:plazi:treatment:03C187E1AE7BB616DD9E4412FB25059F:8EB7602CAE7BB616DF3D40FDFCF005BB" box="[564,811,1970,1995]" country="Laos" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Expedition" name="Phou Sane Mountain" pageId="38" pageNumber="139" stateProvince="Xieng Khouang">Phou Sane Mountain</location>
,
<collectingCountry id="F37F7667AE7BB616DE3240FCFCAA05BA" box="[827,881,1971,1994]" name="Laos" pageId="38" pageNumber="139">Laos</collectingCountry>
2009
<collectingMunicipality id="6BB3AC8DAE7BB616DEC940FDFB9B05BB" box="[960,1088,1970,1995]" pageId="38" pageNumber="139">Expedition</collectingMunicipality>
: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubán,
<geoCoordinate id="EE5C5030AE7BB616DCD14098FD97059F" box="[472,588,2007,2031]" direction="north" orientation="latitude" pageId="38" pageNumber="139" precision="92" value="19.636667">19°38.2'N</geoCoordinate>
,
<geoCoordinate id="EE5C5030AE7BB616DF514098FD0C059F" box="[600,727,2007,2031]" direction="east" orientation="longitude" pageId="38" pageNumber="139" precision="92" value="103.33667">103°20.2'E</geoCoordinate>
,
<elevation id="0045D1C4AE7BB616DFEB4098FCE2059E" box="[738,825,2007,2031]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="38" pageNumber="139" unit="m" value="1420.0">
<quantity id="4C909B12AE7BB616DFEB4098FCE2059E" box="[738,825,2007,2031]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="38" pageNumber="139" unit="m" value="1420.0">1420 m</quantity>
</elevation>
, on
<collectingDate id="EF92E9DFAE7BB616DE614098FBDF059F" box="[872,1028,2007,2031]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<date id="FFD61037AE7BB616DE614098FBDF059F" box="[872,1028,2007,2031]" pageId="38" pageNumber="139" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">1030/v/2009</date>
</collectingDate>
(
<collectorName id="269D5321AE7BB616D91A4098FB5A059F" box="[1043,1153,2006,2031]" pageId="38" pageNumber="139">D. Hauck</collectorName>
) (
<collectionCode id="ED79AE32AE7BB616D99E4098FB2A059F" box="[1175,1265,2007,2031]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="139">NHMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>