248 lines
29 KiB
XML
248 lines
29 KiB
XML
<document id="9D754FFD275C50F0441183FA7165AEC5" ID-CLB-Dataset="25493" ID-DOI="10.11646/zootaxa.4668.1.6" ID-GBIF-Dataset="3996d4b6-345e-4db4-a214-4729d5844552" ID-ISSN="1175-5326" ID-Zenodo-Dep="3449013" ID-ZooBank="4F3088B0-9803-45D3-8ECD-ED88C8141C57" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568897344918" checkinUser="plazi" docAuthor="Chen, Mengzhen, Ma, Zijun & Tian, Mingyi" docDate="2019" docId="03C187C8FF8E8C7DB5B2FD25FDE2FDA3" docLanguage="en" docName="zootaxa.4668.1.6.pdf" docOrigin="Zootaxa 4668 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Qianaphaenops Ueno 2000, n. subgen." docType="treatment" docVersion="8" lastPageNumber="107" masterDocId="FFF8FFB0FF8F8C7FB525FF90FF8BFFBF" masterDocTitle="Discovery of a cavernicolous trechine species from the Sanwang Dong-Erwang Dong cave system, Wulong, Chongqing, southwestern China (Coleoptera Carabidae: Trechinae)" masterLastPageNumber="114" masterPageNumber="105" pageNumber="106" updateTime="1698755406474" updateUser="ExternalLinkService">
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<mods:mods id="AF5F0E882C11A35CF94D9CA2457A52D5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="67F4D2B049030D83F61D4EF7B5FAD720">
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<mods:title id="C2FCBE4DEE80D7EAECBE6FE52142E613">Discovery of a cavernicolous trechine species from the Sanwang Dong-Erwang Dong cave system, Wulong, Chongqing, southwestern China (Coleoptera Carabidae: Trechinae)</mods:title>
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<mods:name id="2B24F659D7D481BF41D687522A605FD9" type="personal">
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<mods:namePart id="4BA1CFA8C7449EECF337DDB699E5B815">Chen, Mengzhen</mods:namePart>
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<mods:namePart id="828B1FB551760CD396A88B9F3B1EB518">Ma, Zijun</mods:namePart>
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<mods:name id="97AC85E541F5C3268C853E7E976A13B5" type="personal">
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<mods:roleTerm id="5E8D2F99AA854111CE3756B0E209FFDD">Author</mods:roleTerm>
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<mods:namePart id="C8B192A8233C9A3B59873383AF788BBB">Tian, Mingyi</mods:namePart>
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<mods:typeOfResource id="8CA44BADB67A0C04280F8B7803BEA630">text</mods:typeOfResource>
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<mods:title id="2F6881092C762EBC51B297C0A02586FD">Zootaxa</mods:title>
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<mods:part id="D8FB6DAD826C04B817A331EBE4E9D278">
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<mods:date id="F7DAC25FD823780EA89B29BDEBDDDE00">2019</mods:date>
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<mods:number id="32F6874443EAF7D2B2AA79DAAACCB139">2019-09-09</mods:number>
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<mods:number id="1CABF6982241C19DE3D02FE8ADAD56F5">4668</mods:number>
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<mods:identifier id="A129A3AB41C406DD26ED7789E8D514ED" type="DOI">10.11646/zootaxa.4668.1.6</mods:identifier>
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<treatment id="03C187C8FF8E8C7DB5B2FD25FDE2FDA3" ID-GBIF-Taxon="159406777" LSID="urn:lsid:plazi:treatment:03C187C8FF8E8C7DB5B2FD25FDE2FDA3" httpUri="http://treatment.plazi.org/id/03C187C8FF8E8C7DB5B2FD25FDE2FDA3" lastPageId="2" lastPageNumber="107" pageId="1" pageNumber="106">
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<subSubSection id="C3726555FF8E8C7EB5B2FD25FDE9FCA8" pageId="1" pageNumber="106" type="nomenclature">
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<paragraph id="8BD736DEFF8E8C7EB5B2FD25FDB4FD70" blockId="1.[151,575,692,720]" box="[151,575,692,720]" pageId="1" pageNumber="106">
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<heading id="D09F81B2FF8E8C7EB5B2FD25FDB4FD70" bold="true" box="[151,575,692,720]" fontSize="11" level="1" pageId="1" pageNumber="106" reason="1">
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<emphasis id="B91CEACCFF8E8C7EB5B2FD25FDB4FD70" bold="true" box="[151,575,692,720]" pageId="1" pageNumber="106">
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Genus
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<taxonomicName id="4C684D5DFF8E8C7EB5CBFD25FDB4FD70" ID-CoL="8KYP3" authority="Ueno, 2000" authorityName="Ueno" authorityYear="2000" box="[238,575,692,720]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEACCFF8E8C7EB5CBFD25FE26FD6F" bold="true" box="[238,429,693,720]" italics="true" pageId="1" pageNumber="106">Qianaphaenops</emphasis>
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<bibRefCitation id="EFF94B2FFF8E8C7EB491FD24FDB4FD70" author="Ueno, S. I." box="[436,575,692,719]" pageId="1" pageNumber="106" pagination="1 - 38" refId="ref4211" refString="Ueno, S. I. (2000) New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China. Journal of the Speleological Society of Japan, 25, 1 - 38." type="journal article" year="2000">Uéno, 2000</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD736DEFF8E8C7EB5B2FD6DFDE9FCA8" blockId="1.[151,610,765,791]" box="[151,610,765,791]" pageId="1" pageNumber="106">
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<heading id="D09F81B2FF8E8C7EB5B2FD6DFDE9FCA8" bold="true" box="[151,610,765,791]" fontSize="11" level="1" pageId="1" pageNumber="106" reason="1">
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<emphasis id="B91CEACCFF8E8C7EB5B2FD6DFDE9FCA8" bold="true" box="[151,610,765,791]" pageId="1" pageNumber="106">
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<emphasis id="B91CEACCFF8E8C7EB5B2FD6DFEA0FCA8" bold="true" box="[151,299,765,791]" italics="true" pageId="1" pageNumber="106">Sanwangius</emphasis>
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Tian et Chen,
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<taxonomicNameLabel id="A22F57B7FF8E8C7EB4C6FD6DFDE9FCA8" box="[483,610,765,791]" pageId="1" pageNumber="106" rank="subGenus">n. subgen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3726555FF8E8C7EB5B2FCD5FB88FCE0" box="[151,1027,837,863]" pageId="1" pageNumber="106" type="type_taxon">
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<paragraph id="8BD736DEFF8E8C7EB5B2FCD5FB88FCE0" blockId="1.[151,1027,837,863]" box="[151,1027,837,863]" pageId="1" pageNumber="106">
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<emphasis id="B91CEACCFF8E8C7EB5B2FCD5FEB2FCE0" bold="true" box="[151,313,837,863]" pageId="1" pageNumber="106">
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<typeStatus id="54D3887CFF8E8C7EB5B2FCD5FF5FFCE0" box="[151,212,837,863]" pageId="1" pageNumber="106">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C684D5DFF8E8C7EB464FCD5FCC2FCE0" ID-CoL="8M678" authority="Tian" authorityName="Tian" box="[321,841,837,863]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="species" species="rowselli" subGenus="Sanwangius">
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<emphasis id="B91CEACCFF8E8C7EB464FCD5FC83FCE0" box="[321,776,837,863]" italics="true" pageId="1" pageNumber="106">Qianaphaenops (Sanwangius) rowselli</emphasis>
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<emphasis id="B91CEACCFF8E8C7EB62AFCD5FCC2FCE0" bold="true" box="[783,841,837,863]" pageId="1" pageNumber="106">Tian</emphasis>
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</taxonomicName>
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<emphasis id="B91CEACCFF8E8C7EB674FCD5FB88FCE0" bold="true" box="[849,1027,837,863]" pageId="1" pageNumber="106">
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et Chen,
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<taxonomicNameLabel id="A22F57B7FF8E8C7EB6E4FCD5FB88FCE0" box="[961,1027,837,863]" pageId="1" pageNumber="106" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3726555FF8E8C7EB5B2FC1CFCF1F9EC" pageId="1" pageNumber="106" type="diagnosis">
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<paragraph id="8BD736DEFF8E8C7EB5B2FC1CFCF1F9EC" blockId="1.[151,1437,908,2015]" pageId="1" pageNumber="106">
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<emphasis id="B91CEACCFF8E8C7EB5B2FC1CFE99FC18" bold="true" box="[151,274,908,935]" pageId="1" pageNumber="106">Diagnosis.</emphasis>
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Semi-aphaenopsian and medium to large sized cave beetles, with moderately elongate body and appendages, eyeless and depigmented (
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<figureCitation id="13532A5BFF8E8C7EB4DCFC21FDB2FC74" box="[505,569,945,971]" captionStart="FIGURE 1" captionStartId="3.[151,250,1946,1971]" captionTargetBox="[323,1251,179,1918]" captionTargetId="figure@3.[323,1263,179,1919]" captionTargetPageId="3" captionText="FIGURE 1 Habitus of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3449017" httpUri="https://zenodo.org/record/3449017/files/figure.png" pageId="1" pageNumber="106">Fig. 1</figureCitation>
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); fore part (head plus prothorax) evidently shorter than elytra, surface smooth; head much longer than wide, genae subparallel-sided, frontal furrows long and deep, divergent posteriorly, two pairs of supraorbital setiferous pores present; mandibles moderately developed, feebly curved apically, shorter than width of head, right mandible tridentate though the median tooth reduced in some individuals; labial suture clearly visible; mentum bisetose on either side of tooth, base concave, tooth short and bifid; submentum 6-setose; ligula thick and short, bisetose at apex; suborbital setae between base of head and submentum (
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<figureCitation id="13532A5BFF8E8C7EB105FBF5FBF7FBC0" box="[1056,1148,1125,1151]" captionStart="FIGURE 2" captionStartId="4.[151,250,1303,1328]" captionTargetBox="[345,1243,181,1261]" captionTargetId="figure@4.[345,1243,181,1277]" captionTargetPageId="4" captionText="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." figureDoi="http://doi.org/10.5281/zenodo.3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="1" pageNumber="106">Fig. 2A</figureCitation>
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); antennae thin and long, but not extending over elytral apices. Propleura not tumid and so invisible from above; pronotum barrel-shaped; front slightly wider than base, two pairs of latero-marginal setae present, hind angles reflexed and blunt. Elytra rather stout and strongly convex, widest at about 3/7 of elytra from base, side margins narrowly bordered and ciliate throughout, prehumeral part widely rounded; whole disc densely punctate and pubescent; striae well-defined and punctate; two dorsal and the preapical setiferous pores present; all marginal umbilicate pores distant from (except the 2
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<superScript id="7C1D9B96FF8E8C7EB5F5FAACFF6BFAF5" attach="left" box="[208,224,1340,1354]" fontSize="6" pageId="1" pageNumber="106">nd</superScript>
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and apical pores which are close to) the marginal gutter, humeral group not aggregated, the 1
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<superScript id="7C1D9B96FF8E8C7EB02AFAACFA91FAF5" attach="left" box="[1295,1306,1340,1354]" fontSize="6" pageId="1" pageNumber="106">st</superScript>
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pore transversely and backwardly shifted onto the 7
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<superScript id="7C1D9B96FF8E8C7EB748FAF0FDF1FAD1" attach="left" box="[621,634,1376,1390]" fontSize="6" pageId="1" pageNumber="106">th</superScript>
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stria, behind the level of the 2
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<superScript id="7C1D9B96FF8E8C7EB6F5FAF0FC6BFAD1" attach="left" box="[976,992,1376,1390]" fontSize="6" pageId="1" pageNumber="106">nd</superScript>
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pore; the 5
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<superScript id="7C1D9B96FF8E8C7EB178FAF0FBE1FAD1" attach="left" box="[1117,1130,1376,1390]" fontSize="6" pageId="1" pageNumber="106">th</superScript>
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and 6
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<superScript id="7C1D9B96FF8E8C7EB18AFAF0FB37FAD1" attach="left" box="[1199,1212,1376,1390]" fontSize="6" pageId="1" pageNumber="106">th</superScript>
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pores of the middle group closely spaced (
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<figureCitation id="13532A5BFF8E8C7EB4B1FA15FE67FA20" box="[404,492,1413,1439]" captionStart="FIGURE 2" captionStartId="4.[151,250,1303,1328]" captionTargetBox="[345,1243,181,1261]" captionTargetId="figure@4.[345,1243,181,1277]" captionTargetPageId="4" captionText="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." figureDoi="http://doi.org/10.5281/zenodo.3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="1" pageNumber="106">Fig. 2B</figureCitation>
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). Protibia with a longitudinal sulcus on dorsal surface; the 1
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<superScript id="7C1D9B96FF8E8C7EB1B0FA14FB2BFA2D" attach="left" box="[1173,1184,1412,1426]" fontSize="6" pageId="1" pageNumber="106">st</superScript>
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and 2
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<superScript id="7C1D9B96FF8E8C7EB1C3FA14FB7DFA2D" attach="left" box="[1254,1270,1412,1426]" fontSize="6" pageId="1" pageNumber="106">nd</superScript>
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protarsomeres modified in male, which are strongly dilated, inwardly denticulate at apices. Ventrites VII bisetose apically in male (
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<figureCitation id="13532A5BFF8E8C7EB585FA5DFF72FA58" box="[160,249,1485,1511]" captionStart="FIGURE 2" captionStartId="4.[151,250,1303,1328]" captionTargetBox="[345,1243,181,1261]" captionTargetId="figure@4.[345,1243,181,1277]" captionTargetPageId="4" captionText="FIGURE 2 Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., A head (ventral), male, paratype B left elytron (chaetotaxy indicated by white points), male, paratype C ventrite VII, male, paratype." figureDoi="http://doi.org/10.5281/zenodo.3449019" httpUri="https://zenodo.org/record/3449019/files/figure.png" pageId="1" pageNumber="106">Fig. 2C</figureCitation>
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), but quadrisetose in female. Male genitalia moderately sclerotized (
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<figureCitation id="13532A5BFF8E8C7EB6DAFA5DFBC8FA58" box="[1023,1091,1485,1511]" captionStart="FIGURE 3" captionStartId="4.[151,250,1957,1982]" captionTargetBox="[321,1265,1386,1932]" captionTargetId="figure@4.[321,1265,1386,1932]" captionTargetPageId="4" captionText="FIGURE 3 Male genitalia of Qianaphaenops (Sanwangius) rowselli Tian et Chen, n. subgen., n. sp., holotype A median lobe and parameres, lateral view B apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3449023" httpUri="https://zenodo.org/record/3449023/files/figure.png" pageId="1" pageNumber="106">Fig. 3</figureCitation>
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), shaped between
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<emphasis id="B91CEACCFF8E8C7EB030FA5DFA17FA58" box="[1301,1436,1485,1511]" italics="true" pageId="1" pageNumber="106">Tiankengius</emphasis>
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<bibRefCitation id="EFF94B2FFF8E8C7EB5B2FA61FE0BF9B4" author="Tian, M. Y. & Huang, S. B. & Wang, D. M." box="[151,384,1521,1547]" pageId="1" pageNumber="106" pagination="81 - 87" refId="ref4060" refString="Tian, M. Y., Huang, S. B. & Wang, D. M. (2018) Occurrence of hypogean trechine beetles in Hanzhong Tiankeng Cluster, southwestern Shaanxi, China (Coleoptera: Carabidae: Trechinae). Annales de la Societe entomologique de France, New Series, 54 (1), 81 - 87. http: // doi. org / 10.1080 / 00379271.2017.1417056" type="journal article" year="2018">Tian et Huang, 2018</bibRefCitation>
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and
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<emphasis id="B91CEACCFF8E8C7EB49CFA61FD4AF9B4" box="[441,705,1521,1547]" italics="true" pageId="1" pageNumber="106">
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<taxonomicName id="4C684D5DFF8E8C7EB49CFA61FDE1F9B4" authorityName="Ueno" authorityYear="2000" box="[441,618,1521,1547]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">Qianaphaenops</taxonomicName>
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(s. str.)
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</emphasis>
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<bibRefCitation id="EFF94B2FFF8E8C7EB7ECFA61FCDBF9B4" author="Ueno, S. I." box="[713,848,1521,1547]" pageId="1" pageNumber="106" pagination="1 - 38" refId="ref4211" refString="Ueno, S. I. (2000) New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China. Journal of the Speleological Society of Japan, 25, 1 - 38." type="journal article" year="2000">Uéno, 2000</bibRefCitation>
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; rather small and short, strongly curved ventrally in lateral view, with a thin and small sagittal aileron; apical lobe elongated in dorsal view, much longer than wide; parameres well developed though much shorter than median lobe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3726555FF8E8C7DB5E2F9CDFBAFFE10" lastPageId="2" lastPageNumber="107" pageId="1" pageNumber="106" type="discussion">
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<paragraph id="8BD736DEFF8E8C7EB5E2F9CDFBFAF860" blockId="1.[151,1437,908,2015]" pageId="1" pageNumber="106">
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<emphasis id="B91CEACCFF8E8C7EB5E2F9CDFEB0F9C8" bold="true" box="[199,315,1629,1655]" pageId="1" pageNumber="106">Remarks.</emphasis>
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Instead of being allied to the genus
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<emphasis id="B91CEACCFF8E8C7EB7D5F9CDFC3AF9C8" box="[752,945,1629,1655]" italics="true" pageId="1" pageNumber="106">Deuveaphaenops</emphasis>
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Tian et
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<bibRefCitation id="EFF94B2FFF8E8C7EB13DF9CDFB26F9C8" author="Huang, S. B. & Cen, Y. J. & Tian, M. Y." box="[1048,1197,1629,1655]" pageId="1" pageNumber="106" pagination="286 - 295" refId="ref3916" refString="Huang, S. B., Cen, Y. J. & Tian, M. Y. (2017) A new genus and a new subgenus of cavernicolous beetles from Furong Jiang valley, southwestern China (Coleoptera: Carabidae: Trechinae). Annales de la Societe entomologique de France, New Series, 53 (4), 286 - 295. http: // doi. org / 10.1080 / 00379271.2017.1344566" type="journal article" year="2017">Huang, 2017</bibRefCitation>
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described also from Wulong (
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<bibRefCitation id="EFF94B2FFF8E8C7EB5DAF911FE44F924" author="Huang, S. B. & Cen, Y. J. & Tian, M. Y." box="[255,463,1665,1691]" pageId="1" pageNumber="106" pagination="286 - 295" refId="ref3916" refString="Huang, S. B., Cen, Y. J. & Tian, M. Y. (2017) A new genus and a new subgenus of cavernicolous beetles from Furong Jiang valley, southwestern China (Coleoptera: Carabidae: Trechinae). Annales de la Societe entomologique de France, New Series, 53 (4), 286 - 295. http: // doi. org / 10.1080 / 00379271.2017.1344566" type="journal article" year="2017">
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Huang
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<emphasis id="B91CEACCFF8E8C7EB476F912FE07F924" box="[339,396,1665,1691]" italics="true" pageId="1" pageNumber="106">et al.</emphasis>
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2017
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</bibRefCitation>
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),
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<emphasis id="B91CEACCFF8E8C7EB4C6F911FDE1F924" box="[483,618,1665,1691]" italics="true" pageId="1" pageNumber="106">Sanwangius</emphasis>
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is more closely allied to the genus
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<taxonomicName id="4C684D5DFF8E8C7EB6DFF911FABDF924" authority="Ueno, 2000" authorityName="Ueno" authorityYear="2000" box="[1018,1334,1665,1691]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">
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<emphasis id="B91CEACCFF8E8C7EB6DFF911FB20F924" box="[1018,1195,1665,1691]" italics="true" pageId="1" pageNumber="106">Qianaphaenops</emphasis>
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<bibRefCitation id="EFF94B2FFF8E8C7EB196F911FABDF924" author="Ueno, S. I." box="[1203,1334,1665,1691]" pageId="1" pageNumber="106" pagination="1 - 38" refId="ref4211" refString="Ueno, S. I. (2000) New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China. Journal of the Speleological Society of Japan, 25, 1 - 38." type="journal article" year="2000">Uéno, 2000</bibRefCitation>
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</taxonomicName>
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which is distributed in northeastern
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<collectingRegion id="49ACF83CFF8E8C7EB4ECF935FDA2F900" box="[457,553,1701,1727]" country="China" name="Guizhou" pageId="1" pageNumber="106">Guizhou</collectingRegion>
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and southern
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<collectingRegion id="49ACF83CFF8E8C7EB7E9F935FCADF900" box="[716,806,1701,1727]" country="China" name="Shaanxi" pageId="1" pageNumber="106">Shaanxi</collectingRegion>
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(
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<bibRefCitation id="EFF94B2FFF8E8C7EB612F935FC31F900" author="Ueno, S. I." box="[823,954,1701,1727]" pageId="1" pageNumber="106" pagination="1 - 38" refId="ref4211" refString="Ueno, S. I. (2000) New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China. Journal of the Speleological Society of Japan, 25, 1 - 38." type="journal article" year="2000">Uéno 2000</bibRefCitation>
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;
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<bibRefCitation id="EFF94B2FFF8E8C7EB6E3F935FB0BF900" author="Tian, M. Y. & Huang, S. B. & Wang, D. M." box="[966,1152,1701,1727]" pageId="1" pageNumber="106" pagination="81 - 87" refId="ref4060" refString="Tian, M. Y., Huang, S. B. & Wang, D. M. (2018) Occurrence of hypogean trechine beetles in Hanzhong Tiankeng Cluster, southwestern Shaanxi, China (Coleoptera: Carabidae: Trechinae). Annales de la Societe entomologique de France, New Series, 54 (1), 81 - 87. http: // doi. org / 10.1080 / 00379271.2017.1417056" type="journal article" year="2018">
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Tian
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<emphasis id="B91CEACCFF8E8C7EB124F936FBB0F900" box="[1025,1083,1701,1727]" italics="true" pageId="1" pageNumber="106">et al.</emphasis>
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2018
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</bibRefCitation>
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).
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<emphasis id="B91CEACCFF8E8C7EB1B1F935FA90F900" box="[1172,1307,1701,1727]" italics="true" pageId="1" pageNumber="106">Sanwangius</emphasis>
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is allied to the nominate subgenus
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<emphasis id="B91CEACCFF8E8C7EB4B9F959FD2AF95C" box="[412,673,1737,1763]" italics="true" pageId="1" pageNumber="106">
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|
<taxonomicName id="4C684D5DFF8E8C7EB4B9F959FDC6F95C" authorityName="Ueno" authorityYear="2000" box="[412,589,1737,1763]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">Qianaphaenops</taxonomicName>
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(s. str.)
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</emphasis>
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, since both subgenera share the following character states: tridentate right mandible, barrel-shaped pronotum, presence of labial suture, presence of posterior latero-marginal setae on pronotum, presence of two dorsal and the preapical pores on elytra, and 1
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<superScript id="7C1D9B96FF8E8C7EB6EEF880FC5DF8A1" attach="left" box="[971,982,1808,1822]" fontSize="6" pageId="1" pageNumber="106">st</superScript>
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and 2
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<superScript id="7C1D9B96FF8E8C7EB13FF880FBA1F8A1" attach="left" box="[1050,1066,1808,1822]" fontSize="6" pageId="1" pageNumber="106">nd</superScript>
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protarsomeres modified in male. However, both of them can be easily distinguished from each other by: (1) the 1
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<superScript id="7C1D9B96FF8E8C7EB100F8A4FBBBF8FD" attach="left" box="[1061,1072,1844,1858]" fontSize="6" pageId="1" pageNumber="106">st</superScript>
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pore of the marginal umbilicate series on elytra transversely and backwardly shifted onto site of the 7
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<superScript id="7C1D9B96FF8E8C7EB69CF8C8FC4DF8D9" attach="left" box="[953,966,1880,1894]" fontSize="6" pageId="1" pageNumber="106">th</superScript>
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stria in
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<emphasis id="B91CEACCFF8E8C7EB10DF8C9FB24F8CC" box="[1064,1199,1881,1907]" italics="true" pageId="1" pageNumber="106">Sanwangius</emphasis>
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, but not backwardly shifted in
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<emphasis id="B91CEACCFF8E8C7EB42DF8EDFD85F828" box="[264,526,1917,1943]" italics="true" pageId="1" pageNumber="106">
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|
<taxonomicName id="4C684D5DFF8E8C7EB42DF8EDFE32F828" authorityName="Ueno" authorityYear="2000" box="[264,441,1917,1943]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">Qianaphaenops</taxonomicName>
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(s. str.)
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</emphasis>
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; (2) elytra stout and strongly convex, wholly pubescent and punctate in
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<emphasis id="B91CEACCFF8E8C7EB01FF8EDFF4DF804" italics="true" pageId="1" pageNumber="106">Sanwangius</emphasis>
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|
, whereas more elongate, less convex and impunctate in
|
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<emphasis id="B91CEACCFF8E8C7EB66BF831FBD3F804" box="[846,1112,1953,1979]" italics="true" pageId="1" pageNumber="106">
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|
<taxonomicName id="4C684D5DFF8E8C7EB66BF831FC74F804" authorityName="Ueno" authorityYear="2000" box="[846,1023,1953,1979]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">Qianaphaenops</taxonomicName>
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(s. str.)
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</emphasis>
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|
; and (3) aedeagus short and evidently arcuate in
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<emphasis id="B91CEACCFF8E8C7EB45EF855FD89F860" box="[379,514,1989,2015]" italics="true" pageId="1" pageNumber="106">Sanwangius</emphasis>
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|
, instead of long and straight in
|
|
<emphasis id="B91CEACCFF8E8C7EB640F855FBE0F860" box="[869,1131,1989,2015]" italics="true" pageId="1" pageNumber="106">
|
|
<taxonomicName id="4C684D5DFF8E8C7EB640F855FB9DF860" authorityName="Ueno" authorityYear="2000" box="[869,1046,1989,2015]" class="Insecta" family="Carabidae" genus="Qianaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="106" phylum="Arthropoda" rank="genus">Qianaphaenops</taxonomicName>
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(s. str.)
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</emphasis>
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|
.
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</paragraph>
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<paragraph id="8BD736DEFF8D8C7DB5BAFF09FBAFFE10" blockId="2.[151,1437,153,540]" pageId="2" pageNumber="107">
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<emphasis id="B91CEACCFF8D8C7DB5BAFF09FEADFF0C" box="[159,294,153,179]" italics="true" pageId="2" pageNumber="107">Sanwangius</emphasis>
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|
is different from the subgenus
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|
<emphasis id="B91CEACCFF8D8C7DB7AEFF09FC99FF0C" box="[651,786,153,179]" italics="true" pageId="2" pageNumber="107">Tiankengius</emphasis>
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|
<bibRefCitation id="EFF94B2FFF8D8C7DB63FFF09FB88FF0C" author="Tian, M. Y. & Huang, S. B. & Wang, D. M." box="[794,1027,153,179]" pageId="2" pageNumber="107" pagination="81 - 87" refId="ref4060" refString="Tian, M. Y., Huang, S. B. & Wang, D. M. (2018) Occurrence of hypogean trechine beetles in Hanzhong Tiankeng Cluster, southwestern Shaanxi, China (Coleoptera: Carabidae: Trechinae). Annales de la Societe entomologique de France, New Series, 54 (1), 81 - 87. http: // doi. org / 10.1080 / 00379271.2017.1417056" type="journal article" year="2018">Tian et Huang, 2018</bibRefCitation>
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|
from southern
|
|
<collectingRegion id="49ACF83CFF8D8C7DB197FF09FAF2FF0C" box="[1202,1401,153,179]" country="China" name="Shaanxi" pageId="2" pageNumber="107">Shaanxi Province</collectingRegion>
|
|
by the following characteristics: (1) propleura invisible from above in
|
|
<emphasis id="B91CEACCFF8D8C7DB6AAFF2DFB9DFF68" box="[911,1046,189,215]" italics="true" pageId="2" pageNumber="107">Sanwangius</emphasis>
|
|
, versus visible in
|
|
<emphasis id="B91CEACCFF8D8C7DB1C0FF2DFAE7FF68" box="[1253,1388,189,215]" italics="true" pageId="2" pageNumber="107">Tiankengius</emphasis>
|
|
; (2) pronotum barrel-like, with obtuse and reflexed hind angles in
|
|
<emphasis id="B91CEACCFF8D8C7DB646FF71FC61FF44" box="[867,1002,225,251]" italics="true" pageId="2" pageNumber="107">Sanwangius</emphasis>
|
|
, versus subquadrate, with rectangular hind angles in
|
|
<emphasis id="B91CEACCFF8D8C7DB461FE95FE40FEA0" box="[324,459,261,287]" italics="true" pageId="2" pageNumber="107">Tiankengius</emphasis>
|
|
; (3) elytra shorter and more convex, wholly pubescent and punctate, lateral margins ciliate throughout, and the 1
|
|
<superScript id="7C1D9B96FF8D8C7DB4F5FEB8FE50FE89" attach="left" box="[464,475,296,310]" fontSize="6" pageId="2" pageNumber="107">st</superScript>
|
|
marginal umbilicate pore backwardly shifted in
|
|
<emphasis id="B91CEACCFF8D8C7DB6DCFEB9FB0BFEFC" box="[1017,1152,297,323]" italics="true" pageId="2" pageNumber="107">Sanwangius</emphasis>
|
|
, versus elytra more elongate, moderately convex, glabrous and impunctate, lateral margins smooth, and the 1
|
|
<superScript id="7C1D9B96FF8D8C7DB14FFEDCFBFEFEE5" attach="left" box="[1130,1141,332,346]" fontSize="6" pageId="2" pageNumber="107">st</superScript>
|
|
marginal umbilicate pore not backwardly shifted in
|
|
<emphasis id="B91CEACCFF8D8C7DB499FEE1FDC8FE34" box="[444,579,369,395]" italics="true" pageId="2" pageNumber="107">Tiankengius</emphasis>
|
|
; and (4) aedeagus longer and more elongate, moderately curved ventrally in
|
|
<emphasis id="B91CEACCFF8D8C7DB5B2FE05FE95FE10" box="[151,286,405,431]" italics="true" pageId="2" pageNumber="107">Sanwangius</emphasis>
|
|
, versus shorter and stouter, strongly curved ventrally in
|
|
<emphasis id="B91CEACCFF8D8C7DB6B3FE05FB96FE10" box="[918,1053,405,431]" italics="true" pageId="2" pageNumber="107">Tiankengius</emphasis>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3726555FF8D8C7DB5E2FE29FDE2FDA3" pageId="2" pageNumber="107" type="etymology">
|
|
<paragraph id="8BD736DEFF8D8C7DB5E2FE29FE4DFE48" blockId="2.[151,1437,153,540]" pageId="2" pageNumber="107">
|
|
<emphasis id="B91CEACCFF8D8C7DB5E2FE29FEC5FE6C" bold="true" box="[199,334,441,467]" pageId="2" pageNumber="107">Etymology.</emphasis>
|
|
“Sanwang + -ius”. To indicate that the beetles are living in Sanwang Dong-Erwang Dong cave system. Gender masculine.
|
|
</paragraph>
|
|
<paragraph id="8BD736DEFF8D8C7DB5E2FD91FDE2FDA3" blockId="2.[151,1437,153,540]" box="[199,617,513,540]" pageId="2" pageNumber="107">
|
|
<emphasis id="B91CEACCFF8D8C7DB5E2FD91FE91FDA4" bold="true" box="[199,282,513,539]" pageId="2" pageNumber="107">Range.</emphasis>
|
|
<collectingCountry id="F37F764EFF8D8C7DB404FD91FEEFFDA4" box="[289,356,513,539]" name="China" pageId="2" pageNumber="107">China</collectingCountry>
|
|
(southern
|
|
<collectingRegion id="49ACF83CFF8D8C7DB4F9FD91FDD5FDA3" box="[476,606,513,540]" country="China" name="Chongqing" pageId="2" pageNumber="107">Chongqing</collectingRegion>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |