611 lines
77 KiB
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611 lines
77 KiB
XML
<document id="BC1DBB547F78021F02A14DBA6EFA75D4" ID-DOI="10.11646/zootaxa.5570.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734877" ID-ZooBank="7177B69B-6132-485D-87C7-4EA807692757" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737737014959" checkinUser="plazi" docAuthor="Rojas-Arias, Laura, Griswold, Terry, Hinojosa-Díaz, Ismael, Morrone, Juan J. & Barajas, Ricardo Ayala" docDate="2025" docId="D838E015FF85B076FF5DC259918E7B9B" docLanguage="en" docName="zootaxa.5570.2.1.pdf" docOrigin="Zootaxa 5570 (2)" docSource="https://doi.org/10.11646/zootaxa.5570.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heriades luisa Rojas-Arias, Griswold & Ayala. 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="238" masterDocId="2401986DFFA4B053FFCAC76393587A50" masterDocTitle="Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species" masterLastPageNumber="259" masterPageNumber="201" pageNumber="234" updateTime="1737737686066" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2AAD2801C7F3431A12548D52D64B7491">Neotropical bees of the genus Heriades Spinola, 1808 (Hymenoptera: Megachilidae: Osmiini) with description of nine new species</mods:title>
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<mods:roleTerm id="7F65BA16CF49B92D86E4C71A24AED67A">Author</mods:roleTerm>
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<mods:namePart id="51C6E27BCB4A1857D9FA941EF2222F66">Rojas-Arias, Laura</mods:namePart>
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<mods:nameIdentifier id="3F65FDAB02DDE44427F90DDE73B77672" type="ORCID">0000-0002-7718-1853</mods:nameIdentifier>
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<mods:affiliation id="65BE5C3B4A7EC8C8A599527FACD2EC5E">Posgrado en Ciencias Biológicas, Universidad Nacional Autónoma de México (Posgrado en Ciencias Biológicas, UNAM. Unidad de</mods:affiliation>
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<mods:name id="4ADA9F973359F002BBFC978AF5942065" type="personal">
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<mods:roleTerm id="C5ACFD7EF2288E327EE9B8433F91F068">Author</mods:roleTerm>
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<mods:namePart id="DE94BD42666151E809CA605F43698280">Griswold, Terry</mods:namePart>
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<mods:nameIdentifier id="273B3CB5CA327C52C8BE57F73BE1339B" type="ORCID">0000-0003-3952-9393</mods:nameIdentifier>
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<mods:namePart id="9A3F9C8698D07550107A3F4189D8FCA5">Hinojosa-Díaz, Ismael</mods:namePart>
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<mods:nameIdentifier id="6BFD93262DC878F5C92CA8D66D9B3A29" type="ORCID">0000-0002-5493-850X</mods:nameIdentifier>
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<mods:namePart id="98B04009E68A00FF23586DDDBC8F894F">Morrone, Juan J.</mods:namePart>
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<mods:affiliation id="A2FF81E4122E5AADBA6FDDEAF4931C0A">Museo de Zoología “ Alfonso L. Herrera ”, Departamento de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional Autónoma</mods:affiliation>
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<mods:namePart id="E8ABDE38A684DC2EC354C2836F4635A7">Barajas, Ricardo Ayala</mods:namePart>
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<mods:nameIdentifier id="C726CDCB177A4A1B52BC7FC6480318C2" type="ORCID">0000-0002-7718-1853</mods:nameIdentifier>
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<mods:title id="F1970F5F1C5633379ECF92B0841FA979">Zootaxa</mods:title>
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<mods:part id="9D83A02C67BB9439704F9BBF95C0F241">
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<mods:date id="3ABEDB6C8FC89686C00120E83BBA682E">2025</mods:date>
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<mods:number id="A8695AEBF2E8CE274C690A4558F51EBF">2025-01-17</mods:number>
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<mods:number id="4589C0CE5E574D8832689BE55D1427B8">5570</mods:number>
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<mods:url id="D35797CC7A39CAE2F24AF15690BD3160">https://doi.org/10.11646/zootaxa.5570.2.1</mods:url>
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<mods:classification id="9027782FB0A9AB70DFBD9940722FD966">journal article</mods:classification>
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<mods:identifier id="631A60D6B8D2123BD6000AC567024900" type="DOI">10.11646/zootaxa.5570.2.1</mods:identifier>
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<treatment id="D838E015FF85B076FF5DC259918E7B9B" ID-DOI="http://doi.org/10.5281/zenodo.14734894" ID-Zenodo-Dep="14734894" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B" httpUri="http://treatment.plazi.org/id/D838E015FF85B076FF5DC259918E7B9B" lastPageId="37" lastPageNumber="238" pageId="33" pageNumber="234">
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<subSubSection id="188B0288FF85B072FF5DC25990677F04" box="[151,831,1338,1365]" pageId="33" pageNumber="234" type="nomenclature">
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<paragraph id="502E5103FF85B072FF5DC25990677F04" blockId="33.[151,831,1338,1400]" box="[151,831,1338,1365]" pageId="33" pageNumber="234">
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<heading id="0B66E66FFF85B072FF5DC25990677F04" bold="true" box="[151,831,1338,1365]" fontSize="11" level="1" pageId="33" pageNumber="234" reason="1">
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<emphasis id="62E58D11FF85B072FF5DC25990677F04" bold="true" box="[151,831,1338,1365]" pageId="33" pageNumber="234">
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<taxonomicName id="97912A80FF85B072FF5DC25991827F04" authority="Rojas-Arias, Griswold & Ayala." authorityName="Rojas-Arias, Griswold & Ayala." authorityYear="2025" box="[151,730,1338,1365]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
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<emphasis id="62E58D11FF85B072FF5DC259921F7F05" bold="true" box="[151,327,1338,1365]" italics="true" pageId="33" pageNumber="234">Heriades luisa</emphasis>
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Rojas-Arias, Griswold & Ayala.
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</taxonomicName>
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<taxonomicNameLabel id="79D6306AFF85B072FD2BC25990677F04" box="[737,831,1338,1364]" pageId="33" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="188B0288FF85B072FF5DC23D92067F28" box="[151,350,1374,1400]" pageId="33" pageNumber="234" type="description">
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<paragraph id="502E5103FF85B072FF5DC23D92067F28" blockId="33.[151,831,1338,1400]" box="[151,350,1374,1400]" pageId="33" pageNumber="234">
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(
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<figureCitation id="C8AA4D86FF85B072FF55C23D925B7F28" box="[159,259,1374,1400]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="33" pageNumber="234">Figs. 2N</figureCitation>
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,
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<figureCitation id="C8AA4D86FF85B072FEC5C23D920F7F28" box="[271,343,1374,1400]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStartId-0="34.[151,250,1098,1122]" captionStartId-1="35.[151,250,1943,1967]" captionTargetBox-0="[159,1428,187,1064]" captionTargetBox-1="[161,1406,179,1888]" captionTargetId-0="figure-22@34.[151,1436,179,1072]" captionTargetId-1="figure-18@35.[161,1426,179,1917]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionText-0="FIGURE 20. Known distribution records for Heriades luisa sp. nov." captionText-1="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14734936" figureDoi-1="http://doi.org/10.5281/zenodo.14734940" httpUri-0="https://zenodo.org/record/14734936/files/figure.png" httpUri-1="https://zenodo.org/record/14734940/files/figure.png" pageId="33" pageNumber="234">20–21</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="188B0288FF85B072FF5DC2C597267CB0" pageId="33" pageNumber="234" type="diagnosis">
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<paragraph id="502E5103FF85B072FF5DC2C597267CB0" blockId="33.[151,1437,1446,2013]" pageId="33" pageNumber="234">
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<emphasis id="62E58D11FF85B072FF5DC2C5924A7F90" bold="true" box="[151,274,1446,1472]" pageId="33" pageNumber="234">Diagnosis.</emphasis>
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Clypeus in profile concave and strongly projected on apical margin; clypeus, supraclypeal area and scutum with small punctures (~0.2 times OD); propodeum with large pits (
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<figureCitation id="C8AA4D86FF85B072FC6AC2A990A87FB4" box="[928,1008,1482,1508]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.A–C ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). D–F scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). G–I propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). J–L Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="33" pageNumber="234">Fig. 3I</figureCitation>
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).
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<emphasis id="62E58D11FF85B072FBC2C2A9971A7FB4" box="[1032,1090,1482,1508]" italics="true" pageId="33" pageNumber="234">Male</emphasis>
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: with well-defined paraocular pubescence; frons with erected hairs, but clypeus with plumose, thin, short, white hairs, almost transparent in frontal view (
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<figureCitation id="C8AA4D86FF85B072FF15C17192117C7C" box="[223,329,1554,1580]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="33" pageNumber="234">Fig. 21A</figureCitation>
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); posterior margin of S1 produced to median rounded angle, without emargination (
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<figureCitation id="C8AA4D86FF85B072FACDC17196137C7C" box="[1287,1355,1554,1580]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="33" pageNumber="234">Fig. 2</figureCitation>
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Nb), in profile, with soft ventral angle less than one third of distance from apex to base (
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<figureCitation id="C8AA4D86FF85B072FBE9C155973E7C00" box="[1059,1126,1590,1616]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="33" pageNumber="234">Fig. 2</figureCitation>
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Na); posterior margin of S5 with two quadrate lobes, two to three lines of capitate hairs on top of it, and two small tufts of a few simple, thick, short hairs on top of two small, inner, smooth elevations (
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<figureCitation id="C8AA4D86FF85B072FCE3C11D90C87CC8" box="[809,912,1662,1688]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="33" pageNumber="234">Fig. 21E</figureCitation>
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); S6 with one subapical tuft of a few capitate, thick, short erected (
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<figureCitation id="C8AA4D86FF85B072FEB2C1C192837CEC" box="[376,475,1698,1724]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="33" pageNumber="234">Fig. 21F</figureCitation>
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).
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<emphasis id="62E58D11FF85B072FE27C1C191197CEC" box="[493,577,1698,1724]" italics="true" pageId="33" pageNumber="234">Female</emphasis>
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: outer surface of mandible without tubercle; apical margin of clypeus without emargination and five well defined tubercles; apical tooth longer than the rest (
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<figureCitation id="C8AA4D86FF85B072FBC1C1A5972A7CB0" box="[1035,1138,1734,1760]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="33" pageNumber="234">Fig. 21C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="188B0288FF85B077FF0DC18991CF7C25" lastPageId="36" lastPageNumber="237" pageId="33" pageNumber="234" type="description">
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<paragraph id="502E5103FF85B072FF0DC18996207D54" blockId="33.[151,1437,1446,2013]" box="[199,1400,1770,1797]" pageId="33" pageNumber="234">
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<emphasis id="62E58D11FF85B072FF0DC18992FC7D54" bold="true" box="[199,420,1770,1797]" pageId="33" pageNumber="234">
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Description.
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<emphasis id="62E58D11FF85B072FEAAC18992C37D55" bold="true" box="[352,411,1770,1797]" italics="true" pageId="33" pageNumber="234">Male</emphasis>
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:
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</emphasis>
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Body length,
|
||
<quantity id="9769FCE6FF85B072FD89C18891C17D55" box="[579,665,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="33" pageNumber="234" unit="mm" value="7.7">7.7 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF85B072FD6DC189906A7D55" box="[679,818,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="7.7" metricValueMin="6.7" pageId="33" pageNumber="234" unit="mm" value="7.2" valueMax="7.7" valueMin="6.7">
|
||
6.7
|
||
<emphasis id="62E58D11FF85B072FD06C18991827D54" bold="true" box="[716,730,1770,1796]" pageId="33" pageNumber="234">–</emphasis>
|
||
7.7 mm
|
||
</quantity>
|
||
, n=3); forewing length,
|
||
<quantity id="9769FCE6FF85B072FB8BC18997CF7D55" box="[1089,1175,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="33" pageNumber="234" unit="mm" value="4.2">4.2 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF85B072FB6CC18996687D55" box="[1190,1328,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="33" pageNumber="234" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">
|
||
4.1
|
||
<emphasis id="62E58D11FF85B072FB00C18997807D54" bold="true" box="[1226,1240,1770,1796]" pageId="33" pageNumber="234">–</emphasis>
|
||
4.2 mm
|
||
</quantity>
|
||
, n=3).
|
||
</paragraph>
|
||
<paragraph id="502E5103FF85B071FF0DC06D90227FF7" blockId="33.[151,1437,1446,2013]" lastBlockId="34.[151,1437,1168,1987]" lastPageId="34" lastPageNumber="235" pageId="33" pageNumber="234">
|
||
<emphasis id="62E58D11FF85B072FF0DC06D925C7D78" box="[199,260,1806,1832]" italics="true" pageId="33" pageNumber="234">Head</emphasis>
|
||
. Length
|
||
<quantity id="9769FCE6FF85B072FEA3C06D92E77D79" box="[361,447,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="234" unit="mm" value="1.5">1.5 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF85B072FE04C06D91017D79" box="[462,601,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="33" pageNumber="234" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
|
||
, n=3), width
|
||
<quantity id="9769FCE6FF85B072FD3AC06D901E7D79" box="[752,838,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="234" unit="mm" value="1.8">1.8 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF85B072FC9FC06D90B87D79" box="[853,992,1806,1833]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="33" pageNumber="234" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
|
||
, n=3); eyes about 3.3 times longer than broad, 1.7 times wider than gena in profile; lower interocular distance
|
||
<quantity id="9769FCE6FF85B072FC69C05190A07D1D" box="[931,1016,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="234" unit="mm" value="0.9">0.9 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF85B072FBCFC05197D77D1D" box="[1029,1167,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="33" pageNumber="234" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
|
||
, n=3); upper interocular distance
|
||
<quantity id="9769FCE6FF85B072FF30C03592097D21" box="[250,337,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="33" pageNumber="234" unit="mm" value="1.1">1.1 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF85B072FEABC035928E7D21" box="[353,470,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="33" pageNumber="234" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">1–1.1 mm</quantity>
|
||
, n=3); interantennal distance 1.5 times antennocular distance; lateral ocelli large (
|
||
<figureCitation id="C8AA4D86FF85B072FAA5C03593E27DC4" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.A–C ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). D–F scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). G–I propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). J–L Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="33" pageNumber="234">Fig. 3C</figureCitation>
|
||
), 3.2 times DOP in interocellar area; interocellar distance 1.8 times OD, ocellocular distance 1.8 times OD, OPR 1.8 times OD; labrum with basal, medial, wide elevation with one cusp; clypeus 1.7 times broader than long, in profile strongly convex; apical margin without emargination; scape length 0.9 times alveolocellar distance, 3.2 times longer than wide; pedicel longer than broad, about 1.2 times longer than F1 and about 1.1 times longer than F2; flagellum black, F1 longer than broad, remaining flagellomeres 1.36 longer than F1; F11 2.0 times longer than broad.
|
||
<emphasis id="62E58D11FF86B071FF23C3BA92307EA2" box="[233,360,1241,1266]" italics="true" pageId="34" pageNumber="235">Mesosoma.</emphasis>
|
||
Scutum 2.31 times longer than scutellum, 3.4 times longer than tegula, and 4.2 longer than axilla; intertegular distance
|
||
<quantity id="9769FCE6FF86B071FE07C39E917A7F47" box="[461,546,1277,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="34" pageNumber="235" unit="mm" value="1.3">1.3 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF86B071FDFAC39E91ED7F47" box="[560,693,1277,1303]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="34" pageNumber="235" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
|
||
, n=3); metanotum with transverse ridge, in posterior view straight; propodeum with large pits (I),
|
||
<quantity id="9769FCE6FF86B071FE3AC242910C7F6B" box="[496,596,1313,1339]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="235" unit="mm" value="0.08">0.08 mm</quantity>
|
||
long; legs black.
|
||
<emphasis id="62E58D11FF86B071FCD1C24290CF7F6A" box="[795,919,1313,1338]" italics="true" pageId="34" pageNumber="235">Metasoma.</emphasis>
|
||
T1
|
||
<quantity id="9769FCE6FF86B071FC09C24397427F6B" box="[963,1050,1312,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="235" unit="mm" value="1.5">1.5 mm</quantity>
|
||
broad (
|
||
<quantity id="9769FCE6FF86B071FBBAC243965B7F6B" box="[1136,1283,1312,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.615" metricValueMax="1.7" metricValueMin="1.53" pageId="34" pageNumber="235" unit="mm" value="1.615" valueMax="1.7" valueMin="1.53">1.53-1.7 mm</quantity>
|
||
, n=3); T3
|
||
<quantity id="9769FCE6FF86B071FAB3C242939B7F0F" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="34" pageNumber="235" unit="mm" value="1.7">1.7 mm</quantity>
|
||
broad (
|
||
<quantity id="9769FCE6FF86B071FED0C22692F87F0F" box="[282,416,1349,1375]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="34" pageNumber="235" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7-1.9 mm</quantity>
|
||
, n=3); T6 with posterior margin not projected; posterior margin of S1 produced to median rounded angle, without emargination (
|
||
<figureCitation id="C8AA4D86FF86B071FD80C20B91D67FD2" box="[586,654,1384,1410]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="34" pageNumber="235">Fig. 2</figureCitation>
|
||
Nb), in profile, with posterior margin projected and with soft ventral angle less than one third of distance from apex to base (
|
||
<figureCitation id="C8AA4D86FF86B071FCCDC2EF90117FF7" box="[775,841,1420,1447]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1907]" captionTargetBox="[264,1323,378,1874]" captionTargetId="figure-137@3.[264,1323,378,1874]" captionTargetPageId="3" captionText="FIGURE 2. S1 of male, (a) profile with posterior margin of sterna to the right, (b) posterior margin in ventral view, and (c) ventral process of S1 in posterior view with dorsal surface up, when relevant (shown for A and B applies to the reminder) A. H. micheneri B. H. amanda sp. nov. C. H. bruneri D. H. briceida sp. nov. E. H. currani F. H. leavitti G. H. viridiana sp. nov. H. H. texana I. H. cristina sp. nov. J. H. danuncia sp. nov. K. H. vulcanica sp. nov. L. H. mariana sp. nov. M. H. nora sp. nov. N. H. luisa sp. nov. O. H. crucifera." figureDoi="http://doi.org/10.5281/zenodo.14734881" httpUri="https://zenodo.org/record/14734881/files/figure.png" pageId="34" pageNumber="235">Fig. 2</figureCitation>
|
||
Na).
|
||
</paragraph>
|
||
<caption id="04EE018BFF86B071FF5DC32990087E32" ID-DOI="http://doi.org/10.5281/zenodo.14734936" ID-Zenodo-Dep="14734936" box="[151,848,1098,1122]" httpUri="https://zenodo.org/record/14734936/files/figure.png" pageId="34" pageNumber="235" startId="34.[151,250,1098,1122]" targetBox="[159,1428,187,1064]" targetPageId="34" targetType="figure">
|
||
<paragraph id="502E5103FF86B071FF5DC32990087E32" blockId="34.[151,848,1098,1122]" box="[151,848,1098,1122]" pageId="34" pageNumber="235">
|
||
<emphasis id="62E58D11FF86B071FF5DC32992787E32" bold="true" box="[151,288,1098,1122]" pageId="34" pageNumber="235">FIGURE 20.</emphasis>
|
||
Known distribution records for
|
||
<taxonomicName id="97912A80FF86B071FDADC32991A37E32" authorityName="Rojas-Arias & Griswold & Hinojosa-Díaz & Morrone & Barajas" authorityYear="2025" box="[615,763,1098,1122]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
|
||
<emphasis id="62E58D11FF86B071FDADC32991A37E32" box="[615,763,1098,1122]" italics="true" pageId="34" pageNumber="235">Heriades luisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="79D6306AFF86B071FCC8C32890087E32" box="[770,848,1099,1122]" pageId="34" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="502E5103FF86B071FF0DC2D390EC7D92" blockId="34.[151,1437,1168,1987]" pageId="34" pageNumber="235">
|
||
<emphasis id="62E58D11FF86B071FF0DC2D392097F9A" box="[199,337,1456,1482]" italics="true" pageId="34" pageNumber="235">Sculpturing.</emphasis>
|
||
Clypeus with small (at base 0.2 times OD), contiguous, not coarse punctures, smaller towards disc and apical margin; supraclypeal area with smaller, denser punctures than those on base of clypeus; frons with bigger and coarser than those on supraclypeal area, and in frons in front of ocelli even bigger than frons; vertex with bigger, sparser and coarser punctures than on supraclypeal area, without micropunctures between punctures; hypostomal area with punctation slightly smaller and sparser than on clypeus; pronotum anterior surface with punctures sparser than on clypeus, denser on dorsolateral angles; disc of scutum with slightly denser punctures (0.2 times OD) than on vertex, without micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with punctures bigger and sparser; mesepisternum with slightly bigger and slightly sparser punctures, irregularly shaped with micropunctures between punctures; metepisternum with notably smaller punctures, less dense and coarse; T1 with smaller (0.2 times OD), slightly sparser punctures; T2 with punctures smaller and sparser than on T1; T3 with smaller, less coarse punctures than on T1, semicircular shape, separated by half DOP; T4 with slightly bigger, denser and coarser punctures than on T3; T5 with smaller punctures than on T4; dorsal surface of T6 with smaller, denser, irregularly shaped punctures than on T5, on posterior surface with bigger and sparser punctation, DBP one third of DOP; sides of S1 with punctation bigger and sparser than on T1, on disc smaller and irregularly shaped; sides of S2 with punctures as on T1, disc of S2 microreticulate without punctation.
|
||
</paragraph>
|
||
<caption id="04EE018BFF87B070FF5DC0F491A37D83" ID-DOI="http://doi.org/10.5281/zenodo.14734940" ID-Zenodo-Dep="14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="35" pageNumber="236" startId="35.[151,250,1943,1967]" targetBox="[161,1406,179,1888]" targetPageId="35" targetType="figure">
|
||
<paragraph id="502E5103FF87B070FF5DC0F491A37D83" blockId="35.[151,1436,1943,2003]" pageId="35" pageNumber="236">
|
||
<emphasis id="62E58D11FF87B070FF5DC0F492477DFF" bold="true" box="[151,287,1943,1967]" pageId="35" pageNumber="236">FIGURE 21.</emphasis>
|
||
<taxonomicName id="97912A80FF87B070FEEEC0F492E07DFF" authorityName="Rojas-Arias & Griswold & Hinojosa-Díaz & Morrone & Barajas" authorityYear="2025" box="[292,440,1943,1967]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="luisa" status="sp. nov.">
|
||
<emphasis id="62E58D11FF87B070FEEEC0F492E07DFF" box="[292,440,1943,1967]" italics="true" pageId="35" pageNumber="236">Heriades luisa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="79D6306AFF87B070FE74C0FB91547DFF" box="[446,524,1944,1967]" pageId="35" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Male:
|
||
<emphasis id="62E58D11FF87B070FD98C0F491327DFF" bold="true" box="[594,618,1943,1967]" pageId="35" pageNumber="236">A.</emphasis>
|
||
frontal view
|
||
<emphasis id="62E58D11FF87B070FD3AC0F4905F7DFF" bold="true" box="[752,775,1943,1967]" pageId="35" pageNumber="236">B.</emphasis>
|
||
dorsal view. Female:
|
||
<emphasis id="62E58D11FF87B070FC29C0F490A37DFF" bold="true" box="[995,1019,1943,1967]" pageId="35" pageNumber="236">C.</emphasis>
|
||
frontal view
|
||
<emphasis id="62E58D11FF87B070FB4BC0F497C17DFF" bold="true" box="[1153,1177,1943,1967]" pageId="35" pageNumber="236">D.</emphasis>
|
||
dorsal view. Male:
|
||
<emphasis id="62E58D11FF87B070FA94C0F4962E7DFF" bold="true" box="[1374,1398,1943,1967]" pageId="35" pageNumber="236">E.</emphasis>
|
||
S5.
|
||
<emphasis id="62E58D11FF87B070FF5DC0D893F27D83" bold="true" box="[151,170,1979,2003]" pageId="35" pageNumber="236">F.</emphasis>
|
||
S6.
|
||
<emphasis id="62E58D11FF87B070FF12C0D893AB7D83" bold="true" box="[216,243,1979,2003]" pageId="35" pageNumber="236">G.</emphasis>
|
||
genital capsule in dorsal view and
|
||
<emphasis id="62E58D11FF87B070FD9DC0D891297D83" bold="true" box="[599,625,1979,2003]" pageId="35" pageNumber="236">H.</emphasis>
|
||
ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="502E5103FF80B077FF0DC7F4961A7B81" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC7F492687AE0" box="[199,304,151,176]" italics="true" pageId="36" pageNumber="237">Vestiture.</emphasis>
|
||
Sides of clypeus with plumose, white hairs, of similar length, directed ventrally, scattered on middle vertical line; supraclypeal area with few plumose, thin, golden, short hairs; frons with plumose, thin, long, white hairs directed dorsally (
|
||
<figureCitation id="C8AA4D86FF80B077FE6FC7BC91577AA9" box="[421,527,223,249]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21A</figureCitation>
|
||
); pubescence in dorsolateral angles of pronotum not forming continuous, linear band with pubescence on anterior margin of scutum, and without tuft of thick, short hairs; disc of scutum with simple, thin, short, scattered, white hairs on its entire surface; mesepisternum with plumose, thin, short, white hairs, increasing in length towards dorsal surface; disc of metepisternum without characteristic tuft of plumose, thick, short, white hairs; tergal fasciae with rows of thin, long, white hairs (
|
||
<figureCitation id="C8AA4D86FF80B077FC7BC60C97517BD9" box="[945,1033,367,393]" captionStart="FIGURE 3" captionStartId="7.[151,250,1796,1820]" captionTargetBox="[151,1436,270,1769]" captionTargetId="figure-60@7.[151,1436,270,1769]" captionTargetPageId="7" captionText="FIGURE 3. Ocellar size, punctation on scutum, pits on propodeal basal zone, apical terga fasciae and T3 punctation.A–C ocelli: A. small (H. currani) B. medium-sized (H. micheneri) C. large (H. mariana sp. nov.). D–F scutal punctation: D. small (H. luisa sp. nov.) E. medium-sized (H. briceida sp. nov.) F. large (H. cristina sp. nov). G–I propodeal basal zone pits: G and H. small to medium-sized (H. briceida sp. nov. and H. micheneri) I. large (H. cristina. sp. nov.). J–L Terga apical fasciae and T3 punctation: J. Fasciae with row of thick, short, white hairs and T3 with dense punctures (H. briceida sp. nov.) K. Fasciae with rows of thin, long, orangish hairs and T3 with small and sparse punctures (H. bruneri). L. Fasciae with rows of thin, short, white hairs and T3 with large, horseshoe-shaped punctures (H. cristina sp. nov.)" figureDoi="http://doi.org/10.5281/zenodo.14734885" httpUri="https://zenodo.org/record/14734885/files/figure.png" pageId="36" pageNumber="237">Fig. 3L</figureCitation>
|
||
); T1 to T3 with simple, short, thin, white hairs in punctures; disc of T4 to T6 with simple, long, thick, whitish hairs in punctures increasing in length towards posterior terga (
|
||
<figureCitation id="C8AA4D86FF80B077FE62C6D491497B81" box="[424,529,439,465]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21D</figureCitation>
|
||
); posterior surface of S1 with few plumose, scattered, long whitish hairs.
|
||
</paragraph>
|
||
<paragraph id="502E5103FF80B077FF0DC6B89294789D" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC6B892117BA5" box="[199,329,475,501]" italics="true" pageId="36" pageNumber="237">
|
||
<taxonomicName id="97912A80FF80B077FF0DC6B8921D7BA5" box="[199,325,475,501]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="36" pageNumber="237" phylum="Tracheophyta" rank="genus">Terminalia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Posterior margin of S5 sub-quadrate, strongly bilobated, with two quadrate lobes on top, three lines of capitate hairs on top of lobes, and two tufts of a few simple, thick, short hairs on top of two small, inner, smooth elevations (at base of lobes), with transverse, subapical line (
|
||
<figureCitation id="C8AA4D86FF80B077FC9CC54090E6786D" box="[854,958,547,573]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21E</figureCitation>
|
||
); S6 with pointy, acute-angled apex, with very short and thin lamella, with one subapical tuft of a few capitate thick, short hairs erected (
|
||
<figureCitation id="C8AA4D86FF80B077FB04C524966B7831" box="[1230,1331,583,609]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21F</figureCitation>
|
||
); capsule in dorsal view
|
||
<quantity id="9769FCE6FF80B077FE8AC50892FD78D5" box="[320,421,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="36" pageNumber="237" unit="mm" value="1.03">1.03 mm</quantity>
|
||
length, and
|
||
<quantity id="9769FCE6FF80B077FDF8C50991CF78D5" box="[562,663,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="36" pageNumber="237" unit="mm" value="0.52">0.52 mm</quantity>
|
||
width, 1.98 times longer than wide, gonobase
|
||
<quantity id="9769FCE6FF80B077FB60C508965778D5" box="[1194,1295,619,645]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="36" pageNumber="237" unit="mm" value="0.08">0.08 mm</quantity>
|
||
length, 5.75 times wider than long, and gonocoxite
|
||
<quantity id="9769FCE6FF80B077FD9CC5ED91E478F9" box="[598,700,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="36" pageNumber="237" unit="mm" value="0.45">0.45 mm</quantity>
|
||
length, gonostylus
|
||
<quantity id="9769FCE6FF80B077FC56C5ED975A78F9" box="[924,1026,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="36" pageNumber="237" unit="mm" value="0.95">0.95 mm</quantity>
|
||
length with acute apex, penis valve
|
||
<quantity id="9769FCE6FF80B077FF5DC5D093A3789D" box="[151,251,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="36" pageNumber="237" unit="mm" value="0.71">0.71 mm</quantity>
|
||
length (
|
||
<figureCitation id="C8AA4D86FF80B077FE9DC5D09298789D" box="[343,448,691,717]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="502E5103FF80B077FF0DC5B592557E65" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC5B5927F78A1" bold="true" box="[199,295,726,753]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC5B5924678A1" bold="true" box="[199,286,726,753]" italics="true" pageId="36" pageNumber="237">Female</emphasis>
|
||
:
|
||
</emphasis>
|
||
Body length
|
||
<quantity id="9769FCE6FF80B077FE75C5B4914D78A1" box="[447,533,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="36" pageNumber="237" unit="mm" value="8.3">8.3 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF80B077FDEEC5B591F678A1" box="[548,686,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.6" metricValueMin="7.5" pageId="36" pageNumber="237" unit="mm" value="8.05" valueMax="8.6" valueMin="7.5">
|
||
7.5
|
||
<emphasis id="62E58D11FF80B077FD82C5B4910E78A1" bold="true" box="[584,598,727,753]" pageId="36" pageNumber="237">–</emphasis>
|
||
8.6 mm
|
||
</quantity>
|
||
, n=5); forewing length
|
||
<quantity id="9769FCE6FF80B077FC7CC5B4975478A1" box="[950,1036,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="36" pageNumber="237" unit="mm" value="4.6">4.6 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF80B077FBD0C5B497FD78A1" box="[1050,1189,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="36" pageNumber="237" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">
|
||
4.6
|
||
<emphasis id="62E58D11FF80B077FBF5C5B4971578A1" bold="true" box="[1087,1101,727,753]" pageId="36" pageNumber="237">–</emphasis>
|
||
5.1 mm
|
||
</quantity>
|
||
, n=5); similar to male except:
|
||
<emphasis id="62E58D11FF80B077FF24C59892737945" box="[238,299,763,789]" italics="true" pageId="36" pageNumber="237">Head</emphasis>
|
||
. Length
|
||
<quantity id="9769FCE6FF80B077FE44C59892BB7945" box="[398,483,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="237" unit="mm" value="1.9">1.9 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF80B077FE38C59891237945" box="[498,635,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="36" pageNumber="237" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">
|
||
1.9
|
||
<emphasis id="62E58D11FF80B077FDDCC598917C7945" bold="true" box="[534,548,763,789]" pageId="36" pageNumber="237">–</emphasis>
|
||
2.2 mm
|
||
</quantity>
|
||
, n=5), width
|
||
<quantity id="9769FCE6FF80B077FCC5C598903C7945" box="[783,868,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="237" unit="mm" value="1.9">1.9 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF80B077FCB8C59890BF7945" box="[882,999,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="36" pageNumber="237" unit="mm" value="1.95" valueMax="2.0" valueMin="1.9">
|
||
1.9
|
||
<emphasis id="62E58D11FF80B077FC5CC59890FC7945" bold="true" box="[918,932,763,789]" pageId="36" pageNumber="237">–</emphasis>
|
||
2 mm
|
||
</quantity>
|
||
, n=5); eyes about 4.1 times longer than broad, 1.2 times wider than gena in profile; lower interocular distance
|
||
<quantity id="9769FCE6FF80B077FC77C47C975A7969" box="[957,1026,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="237" unit="mm" value="1.0">1 mm</quantity>
|
||
; upper interocular distance
|
||
<quantity id="9769FCE6FF80B077FAF7C47C96C07969" box="[1341,1432,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="36" pageNumber="237" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; interantennal distance 1.6 times antennocular distance; interocellar distance 1.8 times OD, ocellocular distance 1.8 times OD, OPR 1.9 times OD; labrum with one rounded, basal, median tubercle; dorsal margin of mandible with carina; clypeus without emargination and apical margin with five rounded, small tubercles (
|
||
<figureCitation id="C8AA4D86FF80B077FB2BC4E8961179F5" box="[1249,1353,907,933]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21C</figureCitation>
|
||
); scape 3.2 times longer than broad; pedicel 1.3 longer than broad, about 2 times longer than F1 and 1.6 longer than F2; remaining flagellomeres 1.8 longer than F1.
|
||
<emphasis id="62E58D11FF80B077FD0AC4B0906079BC" box="[704,824,979,1004]" italics="true" pageId="36" pageNumber="237">Mesosoma</emphasis>
|
||
. Scutum 2.9 times longer than scutellum; intertegular distance
|
||
<quantity id="9769FCE6FF80B077FF31C495920A7E41" box="[251,338,1014,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="237" unit="mm" value="1.5">1.5 mm</quantity>
|
||
(
|
||
<quantity id="9769FCE6FF80B077FEA8C49492B57E41" box="[354,493,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="36" pageNumber="237" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">
|
||
1.2
|
||
<emphasis id="62E58D11FF80B077FE4CC49492CC7E41" bold="true" box="[390,404,1015,1041]" pageId="36" pageNumber="237">–</emphasis>
|
||
1.6 mm
|
||
</quantity>
|
||
, n=5).
|
||
<emphasis id="62E58D11FF80B077FDF7C49491EA7E40" box="[573,690,1015,1040]" italics="true" pageId="36" pageNumber="237">Metasoma</emphasis>
|
||
. T1
|
||
<quantity id="9769FCE6FF80B077FD22C49490677E41" box="[744,831,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="36" pageNumber="237" unit="mm" value="1.7">1.7 mm</quantity>
|
||
broad (
|
||
<quantity id="9769FCE6FF80B077FC5CC49497797E41" box="[918,1057,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="36" pageNumber="237" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">
|
||
1.7
|
||
<emphasis id="62E58D11FF80B077FC70C49490907E41" bold="true" box="[954,968,1015,1041]" pageId="36" pageNumber="237">–</emphasis>
|
||
1.8 mm
|
||
</quantity>
|
||
, n=5). T3
|
||
<quantity id="9769FCE6FF80B077FB53C49497A87E41" box="[1177,1264,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="237" unit="mm" value="1.9">1.9 mm</quantity>
|
||
broad (
|
||
<quantity id="9769FCE6FF80B077FA8DC494939D7E65" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="36" pageNumber="237" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">
|
||
1.9
|
||
<emphasis id="62E58D11FF80B077FAA1C49496217E41" bold="true" box="[1387,1401,1015,1041]" pageId="36" pageNumber="237">–</emphasis>
|
||
2.1 mm
|
||
</quantity>
|
||
, n=5).
|
||
</paragraph>
|
||
<paragraph id="502E5103FF80B077FF0DC35C90637FB5" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC35C92097E09" box="[199,337,1087,1113]" italics="true" pageId="36" pageNumber="237">Sculpturing.</emphasis>
|
||
Clypeus with small (0.3 times OD), contiguous, coarse punctures becoming smaller on lower two thirds, irregularly shaped, without micropunctures between punctures; supraclypeal area with punctures slightly smaller than base of clypeus; frons with bigger, coarser, and contiguous punctures than on supraclypeal area; on frons in front of ocelli slightly biggerpunctures on sides than on frons; vertex with bigger and sparser than on clypeus, with irregular surface between punctures; juxtantennal carina absent but strongly angulated; gena evenly punctate as on frons in front of ocelli; scutum with smaller and denser punctures than on vertex, with micropunctures between punctures; rest of mesosoma as on scutum except: scutellum with bigger and slightly sparser punctures; mesepisternum with bigger and slightly sparser punctures, with micropuntures between punctures; terga bright, microreticulate between punctures; T1 and T2 with smaller (0.1 times OD) punctures than on scutum, sparser (0.3 times DOP), with rounded shape; T3 with punctures 0.7 times DOP of scutum, rounded, and sparser (2.2 times DOP); dorsal surface of T6 with bigger, coarser and denser punctures than on T5; sterna with punctation smaller and sparser than on scutum, microreticulate on posterior margin.
|
||
</paragraph>
|
||
<paragraph id="502E5103FF80B077FF0DC28C91CF7C25" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC28C92687C58" box="[199,304,1519,1544]" italics="true" pageId="36" pageNumber="237">Vestiture.</emphasis>
|
||
Clypeus with few simple, short, white hairs in punctures; frons with few plumose, thin, white hairs (
|
||
<figureCitation id="C8AA4D86FF80B077FF55C17092507C7D" box="[159,264,1555,1581]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21C</figureCitation>
|
||
); gena with plumose, white hairs increasing in length towards hypostomal area; T4 to T6 with simple, thin, golden hairs in punctures, increasing in length towards posterior terga; subapical margin of T6 with linear tuft of plumose, short, yellowish hairs (
|
||
<figureCitation id="C8AA4D86FF80B077FDE8C13891D37C25" box="[546,651,1627,1653]" captionStart="FIGURE 21" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[161,1406,179,1888]" captionTargetId="figure-18@35.[161,1426,179,1917]" captionTargetPageId="35" captionText="FIGURE 21. Heriades luisa sp. nov. Male: A. frontal view B. dorsal view. Female: C. frontal view D. dorsal view. Male: E. S5. F. S6. G. genital capsule in dorsal view and H. ventral view." figureDoi="http://doi.org/10.5281/zenodo.14734940" httpUri="https://zenodo.org/record/14734940/files/figure.png" pageId="36" pageNumber="237">Fig. 21D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="188B0288FF80B077FF0DC11C92617D8D" pageId="36" pageNumber="237" type="materials_examined">
|
||
<paragraph id="502E5103FF80B077FF0DC11C977D7D1D" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC11C92217CC8" bold="true" box="[199,377,1662,1689]" pageId="36" pageNumber="237">Type material:</emphasis>
|
||
<materialsCitation id="E0F95B5EFF80B077FE4BC11C92AB7CED" collectingDate="2009-01-24" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="5" location="Arriaga" pageId="36" pageNumber="237" specimenCode="ECOAB 46959" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
|
||
<emphasis id="62E58D11FF80B077FE4BC11C97587CC8" bold="true" box="[385,1024,1662,1689]" pageId="36" pageNumber="237">
|
||
<typeStatus id="8F2AEFA1FF80B077FE4BC11C92B67CC9" box="[385,494,1663,1689]" pageId="36" pageNumber="237" type="holotype">Holotype</typeStatus>
|
||
male (
|
||
<specimenCode id="0037F978FF80B077FD8AC11D91A17CC8" box="[576,761,1662,1688]" collectionCode="ECOAB" pageId="36" pageNumber="237">ECOAB 46959</specimenCode>
|
||
):
|
||
<collectingCountry id="28861193FF80B077FCDAC11D90D37CC8" box="[784,907,1662,1688]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="92559FE1FF80B077FC52C11D90A47CC8" box="[920,1020,1662,1688]" country="Mexico" name="Chiapas" pageId="36" pageNumber="237">Chiapas</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<location id="554E07D8FF80B077FBC3C11C97397CC9" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FBC3C11C97397CC9" box="[1033,1121,1663,1689]" country="Mexico" name="Arriaga" pageId="36" pageNumber="237" stateProvince="Chiapas">Arriaga</location>
|
||
, 465,
|
||
<quantity id="9769FCE6FF80B077FB6AC11D97967CC9" box="[1184,1230,1662,1689]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
|
||
<elevation id="DBBCB630FF80B077FB6AC11D97967CC9" box="[1184,1230,1662,1689]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
|
||
</quantity>
|
||
, 16.34207 -93.95,
|
||
<date id="242F77C3FF80B077FF5DC1C0925F7CED" box="[151,263,1699,1725]" pageId="36" pageNumber="237" value="2009-01-24">
|
||
<collectingDate id="346B8E2BFF80B077FF5DC1C0925F7CED" box="[151,263,1699,1725]" pageId="36" pageNumber="237" value="2009-01-24">24/I/2009</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FD6434D5FF80B077FEDCC1C0922C7CED" box="[278,372,1699,1725]" pageId="36" pageNumber="237">P. Sagot</collectorName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FE4CC1C092B37CED" box="[390,491,1699,1725]" pageId="36" pageNumber="237">ECOAB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E0F95B5EFF80B077FE35C1C191367CB1" collectingDate="2009-11-18" collectionCode="ECOAB" country="Mexico" elevation="5" location="Arriaga" pageId="36" pageNumber="237" specimenCode="ECOAB 46961" specimenCount="1" stateProvince="Chiapas" typeStatus="allotype">
|
||
<emphasis id="62E58D11FF80B077FE35C1C197657CEC" bold="true" box="[511,1085,1698,1724]" pageId="36" pageNumber="237">
|
||
<typeStatus id="8F2AEFA1FF80B077FE35C1C1913C7CEC" box="[511,612,1698,1724]" pageId="36" pageNumber="237" type="allotype">Allotype</typeStatus>
|
||
(
|
||
<specimenCode id="0037F978FF80B077FDBDC1C1906A7CEC" box="[631,818,1698,1724]" collectionCode="ECOAB" pageId="36" pageNumber="237">ECOAB 46961</specimenCode>
|
||
):
|
||
<collectingCountry id="28861193FF80B077FC80C1C1909D7CEC" box="[842,965,1698,1724]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="92559FE1FF80B077FC1EC1C197607CEC" box="[980,1080,1698,1724]" country="Mexico" name="Chiapas" pageId="36" pageNumber="237">Chiapas</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<location id="554E07D8FF80B077FB8DC1C097C77CED" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FB8DC1C097C77CED" box="[1095,1183,1699,1725]" country="Mexico" name="Arriaga" pageId="36" pageNumber="237" stateProvince="Chiapas">Arriaga</location>
|
||
, 465,
|
||
<quantity id="9769FCE6FF80B077FB2BC1C196487CED" box="[1249,1296,1698,1725]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
|
||
<elevation id="DBBCB630FF80B077FB2BC1C196487CED" box="[1249,1296,1698,1725]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
|
||
</quantity>
|
||
, 16.34207 - 93.95,
|
||
<date id="242F77C3FF80B077FF22C1A492337CB1" box="[232,363,1735,1761]" pageId="36" pageNumber="237" value="2009-11-18">
|
||
<collectingDate id="346B8E2BFF80B077FF22C1A492337CB1" box="[232,363,1735,1761]" pageId="36" pageNumber="237" value="2009-11-18">18/XI/2009</collectingDate>
|
||
</date>
|
||
, C.
|
||
<taxonomicName id="97912A80FF80B077FE55C1A491367CB1" authority="(ECOAB)" baseAuthorityName="ECOAB" box="[415,622,1735,1761]" class="Magnoliopsida" family="Fabaceae" genus="Balboa" kingdom="Plantae" order="Fabales" pageId="36" pageNumber="237" phylum="Tracheophyta" rank="genus">Balboa (ECOAB)</taxonomicName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E0F95B5EFF80B077FDB6C1A497E47D55" collectingDate="2009-11-18" collectionCode="ECOAB" country="Mexico" elevation="5" location="Arriaga" pageId="36" pageNumber="237" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Chiapas" typeStatus="paratype">
|
||
<emphasis id="62E58D11FF80B077FDB6C1A490A07CB0" bold="true" box="[636,1016,1734,1761]" pageId="36" pageNumber="237">
|
||
<typeStatus id="8F2AEFA1FF80B077FDB6C1A491AE7CB1" box="[636,758,1735,1761]" pageId="36" pageNumber="237" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<collectingCountry id="28861193FF80B077FCCFC1A590D87CB0" box="[773,896,1734,1760]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="92559FE1FF80B077FC45C1A590AB7CB0" box="[911,1011,1734,1760]" country="Mexico" name="Chiapas" pageId="36" pageNumber="237">Chiapas</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<location id="554E07D8FF80B077FBCBC1A497017CB1" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FBCBC1A497017CB1" box="[1025,1113,1735,1761]" country="Mexico" name="Arriaga" pageId="36" pageNumber="237" stateProvince="Chiapas">Arriaga</location>
|
||
, 465,
|
||
<quantity id="9769FCE6FF80B077FB50C1A597927CB1" box="[1178,1226,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
|
||
<elevation id="DBBCB630FF80B077FB50C1A597927CB1" box="[1178,1226,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
|
||
</quantity>
|
||
, 16.34207 -93.95, 24, 24/I, 14/III, 17,
|
||
<date id="242F77C3FF80B077FEBCC18892A27D55" box="[374,506,1771,1797]" pageId="36" pageNumber="237" value="2009-11-18">
|
||
<collectingDate id="346B8E2BFF80B077FEBCC18892A27D55" box="[374,506,1771,1797]" pageId="36" pageNumber="237" value="2009-11-18">18/XI/2009</collectingDate>
|
||
</date>
|
||
, C.
|
||
<taxonomicName id="97912A80FF80B077FDEDC18890B47D55" authority=", J. Merida, M. Guzman, P. Sagot" authorityName="J. Merida, M. Guzman, P. Sagot" box="[551,1004,1771,1797]" class="Magnoliopsida" family="Fabaceae" genus="Balboa" kingdom="Plantae" order="Fabales" pageId="36" pageNumber="237" phylum="Tracheophyta" rank="genus">Balboa, J. Mérida, M. Guzmán, P. Sagot</taxonomicName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FC31C18897047D55" box="[1019,1116,1771,1797]" pageId="36" pageNumber="237">ECOAB</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FBA9C18897DE7D55" box="[1123,1158,1771,1797]" count="6" pageId="36" pageNumber="237" type="female">6♀</specimenCount>
|
||
<specimenCount id="46979A8AFF80B077FB44C18997EA7D55" box="[1166,1202,1770,1797]" count="5" pageId="36" pageNumber="237" type="male">5♂</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="E0F95B5EFF80B077FB03C189912B7D79" collectingDate="2009-01-24" collectionCode="ECOAB" collectorName="Sagot" country="Mexico" elevation="5" pageId="36" pageNumber="237" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
|
||
465,
|
||
<quantity id="9769FCE6FF80B077FB31C189967F7D55" box="[1275,1319,1770,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">
|
||
<elevation id="DBBCB630FF80B077FB31C189967F7D55" box="[1275,1319,1770,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="237" unit="m" value="5.0">5 m</elevation>
|
||
</quantity>
|
||
, 16.34207 -93.95,
|
||
<date id="242F77C3FF80B077FF3AC06C92077D79" box="[240,351,1807,1833]" pageId="36" pageNumber="237" value="2009-01-24">
|
||
<collectingDate id="346B8E2BFF80B077FF3AC06C92077D79" box="[240,351,1807,1833]" pageId="36" pageNumber="237" value="2009-01-24">24/I/2009</collectingDate>
|
||
</date>
|
||
, P.
|
||
<collectorName id="FD6434D5FF80B077FE40C06C92927D79" box="[394,458,1807,1833]" pageId="36" pageNumber="237">Sagot</collectorName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FE11C06C91647D79" box="[475,572,1807,1833]" pageId="36" pageNumber="237">ECOAB</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FD8FC06C91317D79" box="[581,617,1807,1833]" count="1" pageId="36" pageNumber="237" type="male">1♂</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E0F95B5EFF80B077FD48C06C96CF7D79" box="[642,1431,1807,1833]" collectingDate="2016-03-24" collectionCode="ECOAB" collectorName="P. Sagot" country="Mexico" elevation="412" location="Bochil" pageId="36" pageNumber="237" specimenCount="4" specimenCount-female="4" stateProvince="Chiapas" typeStatus="paratype">
|
||
<location id="554E07D8FF80B077FD48C06C91977D79" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FD48C06C91977D79" box="[642,719,1807,1833]" country="Mexico" name="Bochil" pageId="36" pageNumber="237" stateProvince="Chiapas">Bochil</location>
|
||
,
|
||
<quantity id="9769FCE6FF80B077FD17C06C90707D79" box="[733,808,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="36" pageNumber="237" unit="m" value="412.0">
|
||
<elevation id="DBBCB630FF80B077FD17C06C90707D79" box="[733,808,1807,1833]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="36" pageNumber="237" unit="m" value="412.0">412 m</elevation>
|
||
</quantity>
|
||
, 16.97647 -93.06,
|
||
<date id="242F77C3FF80B077FBCBC06C97DC7D79" box="[1025,1156,1807,1833]" pageId="36" pageNumber="237" value="2016-03-24">
|
||
<collectingDate id="346B8E2BFF80B077FBCBC06C97DC7D79" box="[1025,1156,1807,1833]" pageId="36" pageNumber="237" value="2016-03-24">24/III/2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FD6434D5FF80B077FB5BC06C97B67D79" box="[1169,1262,1807,1833]" pageId="36" pageNumber="237">P. Sagot</collectorName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FB35C06C96387D79" box="[1279,1376,1807,1833]" pageId="36" pageNumber="237">ECOAB</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FAA3C06C96D67D79" box="[1385,1422,1807,1833]" count="4" pageId="36" pageNumber="237" type="female">4♀</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E0F95B5EFF80B077FF5DC05097797D1D" box="[151,1057,1842,1869]" collectingDate="2021-01-25" collectionCode="ECOAB" collectorName="Sagot" country="Mexico" elevation="553" location="Chiapa de Corzo" pageId="36" pageNumber="237" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Chiapas" typeStatus="paratype">
|
||
<location id="554E07D8FF80B077FF5DC050920E7D1D" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FF5DC050920E7D1D" box="[151,342,1843,1869]" country="Mexico" name="Chiapa de Corzo" pageId="36" pageNumber="237" stateProvince="Chiapas">Chiapa de Corzo</location>
|
||
,
|
||
<quantity id="9769FCE6FF80B077FEABC05192F27D1D" box="[353,426,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" pageId="36" pageNumber="237" unit="m" value="553.0">
|
||
<elevation id="DBBCB630FF80B077FEABC05192F27D1D" box="[353,426,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" pageId="36" pageNumber="237" unit="m" value="553.0">553 m</elevation>
|
||
</quantity>
|
||
, 16.46994 -92.99,
|
||
<date id="242F77C3FF80B077FDB6C05191B47D1D" box="[636,748,1842,1869]" pageId="36" pageNumber="237" value="2021-01-25">
|
||
<collectingDate id="346B8E2BFF80B077FDB6C05191B47D1D" box="[636,748,1842,1869]" pageId="36" pageNumber="237" value="2021-01-25">25/I/2021</collectingDate>
|
||
</date>
|
||
, P.
|
||
<collectorName id="FD6434D5FF80B077FCD8C050900A7D1D" box="[786,850,1843,1869]" pageId="36" pageNumber="237">Sagot</collectorName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FCAAC05090997D1D" box="[864,961,1843,1869]" pageId="36" pageNumber="237">ECOAB</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FC02C05090B37D1D" box="[968,1003,1843,1869]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
|
||
<specimenCount id="46979A8AFF80B077FC38C050974F7D1D" box="[1010,1047,1843,1869]" count="3" pageId="36" pageNumber="237" type="male">3♂</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="502E5103FF80B077FF0DC03592617D8D" blockId="36.[151,1437,151,2013]" pageId="36" pageNumber="237">
|
||
<emphasis id="62E58D11FF80B077FF0DC035916C7D20" bold="true" box="[199,564,1878,1904]" pageId="36" pageNumber="237">Additional material examined:</emphasis>
|
||
<materialsCitation id="E0F95B5EFF80B077FDF3C03591867DC5" collectingDate="1987-11-02" collectionCode="BBSL" collectorName="T. Griswold" country="Mexico" elevation="250" latitude="17.989445" location="Los Amates Playa Azul" longLatPrecision="21" longitude="-102.365" pageId="36" pageNumber="237" specimenCount="1" specimenCount-female="1" stateProvince="Michoacan de Ocampo">
|
||
<emphasis id="62E58D11FF80B077FDF3C03590117D21" bold="true" box="[569,841,1878,1905]" pageId="36" pageNumber="237">
|
||
<collectingCountry id="28861193FF80B077FDF3C03591EC7D20" box="[569,692,1878,1904]" name="Mexico" pageId="36" pageNumber="237">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion id="92559FE1FF80B077FD74C035901C7D21" box="[702,836,1878,1905]" country="Mexico" name="Michoacan de Ocampo" pageId="36" pageNumber="237">Michoacán</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<location id="554E07D8FF80B077FC85C03497097D21" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FC85C03497097D21" box="[847,1105,1879,1905]" country="Mexico" latitude="17.989445" longLatPrecision="21" longitude="-102.365" name="Los Amates Playa Azul" pageId="36" pageNumber="237" stateProvince="Michoacan de Ocampo">Los Amates Playa Azul</location>
|
||
26kmN, [
|
||
<quantity id="9769FCE6FF80B077FB74C03596517D21" box="[1214,1289,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="237" unit="m" value="250.0">
|
||
<elevation id="DBBCB630FF80B077FB74C03596517D21" box="[1214,1289,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="237" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="35A537C4FF80B077FAD9C03596C47D21" box="[1299,1436,1878,1905]" degrees="17" direction="north" minutes="59" orientation="latitude" pageId="36" pageNumber="237" precision="15" seconds="22" value="17.989445">17°59’22”N</geoCoordinate>
|
||
<geoCoordinate id="35A537C4FF80B077FF5DC01992617DC5" box="[151,313,1914,1941]" degrees="102" direction="west" minutes="21" orientation="longitude" pageId="36" pageNumber="237" precision="15" seconds="54" value="-102.365">102°21’54”W</geoCoordinate>
|
||
],
|
||
<date id="242F77C3FF80B077FE81C01892997DC5" box="[331,449,1915,1941]" pageId="36" pageNumber="237" value="1987-11-02">
|
||
<collectingDate id="346B8E2BFF80B077FE81C01892997DC5" box="[331,449,1915,1941]" pageId="36" pageNumber="237" value="1987-11-02">2/XI/1987</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FD6434D5FF80B077FE01C018910A7DC5" box="[459,594,1915,1941]" pageId="36" pageNumber="237">T. Griswold</collectorName>
|
||
, (
|
||
<collectionCode id="3680C9C6FF80B077FDAEC01891F37DC5" box="[612,683,1915,1941]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="36" pageNumber="237" type="Museum">BBSL</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FD7BC018918A7DC5" box="[689,722,1915,1941]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E0F95B5EFF80B077FD20C019911B7DE9" collectingDate="1993-04-07" collectionCode="BBSL" collectorName="La Ventosa & A. Rodriguez" country="Mexico" elevation="29" latitude="16.551666" location="La Ventosa" longLatPrecision="21" longitude="-94.94583" pageId="36" pageNumber="237" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
|
||
<emphasis id="62E58D11FF80B077FD20C01990137DC4" bold="true" box="[746,843,1914,1940]" pageId="36" pageNumber="237">
|
||
<collectingRegion id="92559FE1FF80B077FD20C019901E7DC4" box="[746,838,1914,1940]" country="Mexico" name="Oaxaca" pageId="36" pageNumber="237">Oaxaca</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<collectorName id="FD6434D5FF80B077FC9BC01890957DC5" box="[849,973,1915,1941]" pageId="36" pageNumber="237">
|
||
<location id="554E07D8FF80B077FC9BC01890957DC5" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FC9BC01890957DC5" box="[849,973,1915,1941]" country="Mexico" latitude="16.551666" longLatPrecision="21" longitude="-94.94583" name="La Ventosa" pageId="36" pageNumber="237" stateProvince="Oaxaca">La Ventosa</location>
|
||
</collectorName>
|
||
10kmN,
|
||
<quantity id="9769FCE6FF80B077FBFEC01897367DC5" box="[1076,1134,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="36" pageNumber="237" unit="m" value="29.0">
|
||
<elevation id="DBBCB630FF80B077FBFEC01897367DC5" box="[1076,1134,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="36" pageNumber="237" unit="m" value="29.0">29 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="35A537C4FF80B077FBB3C018965A7DC5" box="[1145,1282,1915,1941]" degrees="16" direction="north" minutes="33" orientation="latitude" pageId="36" pageNumber="237" precision="15" seconds="06" value="16.551666">16°33’06”N</geoCoordinate>
|
||
<geoCoordinate id="35A537C4FF80B077FAC2C01996C07DC5" box="[1288,1432,1914,1941]" degrees="94" direction="west" minutes="56" orientation="longitude" pageId="36" pageNumber="237" precision="15" seconds="45" value="-94.94583">94°56’45”W</geoCoordinate>
|
||
,
|
||
<date id="242F77C3FF80B077FF5DC0FC92557DE9" box="[151,269,1951,1977]" pageId="36" pageNumber="237" value="1993-04-07">
|
||
<collectingDate id="346B8E2BFF80B077FF5DC0FC92557DE9" box="[151,269,1951,1977]" pageId="36" pageNumber="237" value="1993-04-07">7/IV/1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FD6434D5FF80B077FED0C0FC92EC7DE9" box="[282,436,1951,1977]" pageId="36" pageNumber="237">A. Rodriguez</collectorName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FE0FC0FC91547DE9" box="[453,524,1951,1977]" country="USA" name="USDA, Agriculture Research Service, Pollinating Insects-- Biology, Management and Systematics Research" pageId="36" pageNumber="237" type="Museum">BBSL</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FDDFC0FC916E7DE9" box="[533,566,1951,1977]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="E0F95B5EFF80B077FD98C0FC926C7D8D" collectingDate="2018-11-19" collectionCode="ECOAB" collectorName="N. Arnold" country="Mexico" elevation="1154" location="Santa Maria Zoquitlan" pageId="36" pageNumber="237" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
|
||
<location id="554E07D8FF80B077FD98C0FC900F7DE9" LSID="urn:lsid:plazi:treatment:D838E015FF85B076FF5DC259918E7B9B:554E07D8FF80B077FD98C0FC900F7DE9" box="[594,855,1951,1977]" country="Mexico" name="Santa Maria Zoquitlan" pageId="36" pageNumber="237" stateProvince="Oaxaca">Santa Maria Zoquitlan</location>
|
||
,
|
||
<quantity id="9769FCE6FF80B077FCAEC0FD90E57DE9" box="[868,957,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="36" pageNumber="237" unit="m" value="1154.0">
|
||
<elevation id="DBBCB630FF80B077FCAEC0FD90E57DE9" box="[868,957,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="36" pageNumber="237" unit="m" value="1154.0">1154 m</elevation>
|
||
</quantity>
|
||
, 16.51338 -96.34,
|
||
<date id="242F77C3FF80B077FB52C0FC96447DE9" box="[1176,1308,1951,1977]" pageId="36" pageNumber="237" value="2018-11-19">
|
||
<collectingDate id="346B8E2BFF80B077FB52C0FC96447DE9" box="[1176,1308,1951,1977]" pageId="36" pageNumber="237" value="2018-11-19">19/XI/2018</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FD6434D5FF80B077FAE3C0FC96C47DE9" box="[1321,1436,1951,1977]" pageId="36" pageNumber="237">N. Arnold</collectorName>
|
||
(
|
||
<collectionCode id="3680C9C6FF80B077FF55C0A092587D8D" box="[159,256,1987,2013]" pageId="36" pageNumber="237">ECOAB</collectionCode>
|
||
<specimenCount id="46979A8AFF80B077FECCC0A092707D8D" box="[262,296,1987,2013]" count="1" pageId="36" pageNumber="237" type="female">1♀</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="188B0288FF81B076FF0DC7F5966A7AA8" pageId="37" pageNumber="238" type="distribution">
|
||
<paragraph id="502E5103FF81B076FF0DC7F5966A7AA8" blockId="37.[151,1437,150,460]" pageId="37" pageNumber="238">
|
||
<emphasis id="62E58D11FF81B076FF0DC7F592397AE0" bold="true" box="[199,353,150,176]" pageId="37" pageNumber="238">Distribution.</emphasis>
|
||
This species is present along the Pacific slope of southern
|
||
<collectingCountry id="28861193FF81B076FBD9C7F497337AE1" box="[1043,1131,151,177]" name="Mexico" pageId="37" pageNumber="238">Mexico</collectingCountry>
|
||
, in areas with tropical dry forest, and is present in the states of
|
||
<collectingRegion id="92559FE1FF81B076FD8FC7D8919C7A85" box="[581,708,187,213]" country="Mexico" name="Michoacan de Ocampo" pageId="37" pageNumber="238">Michoacán</collectingRegion>
|
||
,
|
||
<collectingRegion id="92559FE1FF81B076FD18C7D8907E7A85" box="[722,806,187,213]" country="Mexico" name="Oaxaca" pageId="37" pageNumber="238">Oaxaca</collectingRegion>
|
||
and
|
||
<collectingRegion id="92559FE1FF81B076FCA8C7D890E77A85" box="[866,959,187,213]" country="Mexico" name="Chiapas" pageId="37" pageNumber="238">Chiapas</collectingRegion>
|
||
, being endemic to
|
||
<collectingCountry id="28861193FF81B076FB57C7D897AD7A85" box="[1181,1269,187,213]" name="Mexico" pageId="37" pageNumber="238">Mexico</collectingCountry>
|
||
, in the Pacific Lowlands biogeographic province (
|
||
<bibRefCitation id="34002CF2FF81B076FDEEC7BD90507AA8" author="Morrone, J. J. & Escalante, T. & Rodriguez-Tapia, G. & Carmona, A. & Arana, M. & Mercado-Gomez, J. D." box="[548,776,222,248]" pageId="37" pageNumber="238" refId="ref43590" refString="Morrone, J. J., Escalante, T., Rodriguez-Tapia, G., Carmona, A., Arana, M. & Mercado-Gomez, J. D. (2022) Biogeographic regionalization of the Neotropical region: New map and shapefile. Anais da Academia Brasileira de Ciencias, 94, e 20211167. https: // doi. org / 10.1590 / 0001 - 3765202220211167" type="journal volume" year="2022">
|
||
Morrone
|
||
<emphasis id="62E58D11FF81B076FD45C7BC91987AA8" box="[655,704,222,248]" italics="true" pageId="37" pageNumber="238">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
) (
|
||
<figureCitation id="C8AA4D86FF81B076FCD5C7BD902B7AA8" box="[799,883,222,248]" captionStart="FIGURE 20" captionStartId="34.[151,250,1098,1122]" captionTargetBox="[159,1428,187,1064]" captionTargetId="figure-22@34.[151,1436,179,1072]" captionTargetPageId="34" captionText="FIGURE 20. Known distribution records for Heriades luisa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14734936" httpUri="https://zenodo.org/record/14734936/files/figure.png" pageId="37" pageNumber="238">Fig. 20</figureCitation>
|
||
). Altitudinal range: sea level to
|
||
<quantity id="9769FCE6FF81B076FB1DC7BD96767AA8" box="[1239,1326,222,249]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="37" pageNumber="238" unit="m" value="1154.0">1154 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="188B0288FF81B076FF0DC66296277B6E" pageId="37" pageNumber="238" type="etymology">
|
||
<paragraph id="502E5103FF81B076FF0DC66296277B6E" blockId="37.[151,1437,150,460]" pageId="37" pageNumber="238">
|
||
<emphasis id="62E58D11FF81B076FF0DC66292167B4B" bold="true" box="[199,334,257,283]" pageId="37" pageNumber="238">Etymology.</emphasis>
|
||
This species is dedicated to Dr. Luisa Ruz, a Chilean taxonomist and melitologist who has contributed greatly to the knowledge of native bees in South America, especially bees of the family
|
||
<taxonomicName id="97912A80FF81B076FB3FC64796237B6E" authorityName="Latreille" authorityYear="1802" box="[1269,1403,292,318]" class="Insecta" family="Andrenidae" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="family">Andrenidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="188B0288FF81B076FF0DC624918E7B9B" pageId="37" pageNumber="238" type="discussion">
|
||
<paragraph id="502E5103FF81B076FF0DC624918E7B9B" blockId="37.[151,1437,150,460]" pageId="37" pageNumber="238">
|
||
<emphasis id="62E58D11FF81B076FF0DC62492177B31" bold="true" box="[199,335,327,353]" pageId="37" pageNumber="238">Comments.</emphasis>
|
||
Females show variation in tubercles on the apical margin of clypeus in frontal view only, from well-defined rounded tubercles to crenulated margin. Although body size is large for the genus, some small females can be found in
|
||
<collectingRegion id="92559FE1FF81B076FE85C6ED92967BF8" box="[335,462,398,424]" country="Mexico" name="Michoacan de Ocampo" pageId="37" pageNumber="238">Michoacán</collectingRegion>
|
||
, Los Amates, and
|
||
<collectingRegion id="92559FE1FF81B076FD6AC6ED91A57BF8" box="[672,765,398,424]" country="Mexico" name="Chiapas" pageId="37" pageNumber="238">Chiapas</collectingRegion>
|
||
, Arriaga,
|
||
<collectingCountry id="28861193FF81B076FCA6C6ED909D7BF8" box="[876,965,398,424]" name="Mexico" pageId="37" pageNumber="238">Mexico</collectingCountry>
|
||
. Males seem to vary on S6 with posterior tuft in a V-line or a rounded shape of capitate hairs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |