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<document id="82294FF58F0AB4D45C624605503A4640" ID-DOI="10.11646/zootaxa.4802.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3904978" ID-ZooBank="BA145877-12CB-4C97-A18F-9E4E6F0258CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592923053381" checkinUser="plazi" docAuthor="Obando, Ranulfo González, Gironza, Nancy Carrejo, Panche, Jeferson &amp; Aldrete, Alfonso Neri García" docDate="2020" docId="039787F6D151260BFF7AFD8D4EB7F8BC" docLanguage="en" docName="zootaxa.4802.2.1.pdf" docOrigin="Zootaxa 4802 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loneura carijona Obando &amp; Gironza &amp; Panche &amp; Aldrete 2020, n. sp." docType="treatment" docVersion="4" lastPageNumber="215" masterDocId="FFAEFF8ED15F2605FFEDFFA84C2FFFCF" masterDocTitle="The genus Loneura Navás (Insecta: Psocodea: Psocoptera: Ptiloneuridae) in Colombia, description of new species and key for identification" masterLastPageNumber="249" masterPageNumber="201" pageNumber="215" updateTime="1698776039647" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="260061C005F0E7063FDD4E6F01A92236">The genus Loneura Navás (Insecta: Psocodea: Psocoptera: Ptiloneuridae) in Colombia, description of new species and key for identification</mods:title>
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<mods:namePart id="33B6A5F5261E47710B3ECCA05982A0D1">Obando, Ranulfo González</mods:namePart>
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<mods:namePart id="BB6B3E9D7BEF7B46ABD55E5A0B0F628D">Gironza, Nancy Carrejo</mods:namePart>
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<mods:namePart id="0D87171400C8B90EB500698A8C6058F0">Panche, Jeferson</mods:namePart>
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<mods:namePart id="C8F4161E9038046193D56CB66301F432">Aldrete, Alfonso Neri García</mods:namePart>
<mods:affiliation id="37E2A9FAFEB0F967824ACD40B929E5DF">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
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<mods:title id="50196E017909F92FB546662F419D4600">Zootaxa</mods:title>
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<mods:date id="B09071366E8D5A86E124F66072FA3B98">2020</mods:date>
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<mods:number id="04FEB2F32197077D30E8D3D251798A0A">2020-06-23</mods:number>
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<mods:number id="4A63256D2E3033ADD7FD986089A81482">4802</mods:number>
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<subSubSection id="C324656BD151260BFF7AFD8D4DD2FDF0" box="[151,509,549,575]" pageId="14" pageNumber="215" type="nomenclature">
<paragraph id="8B8136E0D151260BFF7AFD8D4DD2FDF0" blockId="14.[151,509,549,611]" box="[151,509,549,575]" pageId="14" pageNumber="215">
<heading id="D0C9818CD151260BFF7AFD8D4DD2FDF0" bold="true" box="[151,509,549,575]" fontSize="11" level="1" pageId="14" pageNumber="215" reason="1">
<emphasis id="B94AEAF2D151260BFF7AFD8D4DD2FDF0" bold="true" box="[151,509,549,575]" pageId="14" pageNumber="215">
<taxonomicName id="4C3E4D63D151260BFF7AFD8D4D43FDF0" ID-CoL="8NJ5P" authority="Obando &amp; Gironza &amp; Panche &amp; Aldrete, 2020" authorityName="Obando &amp; Gironza &amp; Panche &amp; Aldrete" authorityYear="2020" box="[151,364,549,575]" class="Insecta" family="Ptiloneuridae" genus="Loneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="carijona" status="sp. nov.">
<emphasis id="B94AEAF2D151260BFF7AFD8D4D43FDF0" bold="true" box="[151,364,549,575]" italics="true" pageId="14" pageNumber="215">Loneura carijona</emphasis>
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Male
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C324656BD151260BFF7AFDE14D08FDAC" box="[151,295,585,611]" pageId="14" pageNumber="215" type="description">
<paragraph id="8B8136E0D151260BFF7AFDE14D08FDAC" blockId="14.[151,509,549,611]" box="[151,295,585,611]" pageId="14" pageNumber="215">
(
<figureCitation id="13052A65D151260BFF72FDE14D30FDAC" box="[159,287,585,611]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Figs 3439</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324656BD151260BFF7AFD38492BF99C" pageId="14" pageNumber="215" type="diagnosis">
<paragraph id="8B8136E0D151260BFF7AFD384DF6FCF4" blockId="14.[151,1437,656,1907]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF7AFD384D3DFD64" bold="true" box="[151,274,656,683]" pageId="14" pageNumber="215">Diagnosis.</emphasis>
Forewings with a slender submarginal pigmented band from R4+5 to areola postica; pterostigma hyaline, with broad proximal and distal dark brown bands. Hypandrium deeply cleft in the middle, distal third of each posterior process with a broad lobe extended laterally, apices rounded, each with a subapical macroseta on inner margin; proximally a wide lobe extended outwards (
<figureCitation id="13052A65D151260BFD08FD554F15FCD8" box="[741,826,765,791]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 37</figureCitation>
). Mesal sclerite processes wide based, narrowing posteriorly, acuminate (
<figureCitation id="13052A65D151260BFE95FC894DE2FCF4" box="[376,461,801,827]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="8B8136E0D151260BFF2AFCEC4EA6FAC0" blockId="14.[151,1437,656,1907]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AFCEC4D24FC90" bold="true" box="[199,267,836,863]" pageId="14" pageNumber="215">Color</emphasis>
. Body pale brown. Head pattern (
<figureCitation id="13052A65D151260BFD6EFCED4EF9FCAF" box="[643,726,837,864]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 36</figureCitation>
), vertex light brown; front with two large brown spots; an ochre transverse band between the inner border of each compound eye to the lower edge of the ocellar triangle and to the epistomal sulcus; a cream diagonal band from the inner border of each compound eye to the epistomal sulcus at the level of the antennal fossae. Compound eyes black; ocelli hyaline with ochre centripetal crescents. Labrum, anteclypeus, postclypeus and mandibles dark brown. Genae ochre to light brown. Postgenae light brown. Antennae: scape brown; pedicel and flagella pale brown, flagellomeres 15 with apices cream. Maxillary palps pale brown. Tergal lobes of meso- and metathorax dark brown, with light brown edges. Thoracic pleura pale brown with dark brown areas. Legs: coxae with small ochre spot basally, fore- coxae and trochanters brown; coxae and trochanters of mid and hind- legs light brown; femora light brown, with two brown spots; tibiae and tarsi brown. Wings hyaline (
<figureCitation id="13052A65D151260BFF72FB214D30FB6C" box="[159,287,1160,1187]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Figs 34, 35</figureCitation>
), forewings with brown submarginal band from R4+5 to areola postica, and a dark brown band from the middle of Cu1 to distal end of 1A; dark brown spots on vein ends at wing margin. Hindwings with brown spots on vein ends at wing margin. Abdomen light brown, with subcuticular ochre spots; clunium pale brown. Hypandrium brown. Epiproct and paraprocts light brown.
</paragraph>
<paragraph id="8B8136E0D151260BFF2AFAB1492BF99C" blockId="14.[151,1437,656,1907]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AFAB14D4EFAFC" bold="true" box="[199,353,1305,1331]" pageId="14" pageNumber="215">Morphology.</emphasis>
Head (
<figureCitation id="13052A65D151260BFE59FAB14E26FAFB" box="[436,521,1305,1332]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 36</figureCitation>
): H/MxW: 1.44; H/d: 3.56; IO/MxW: 0.74. Vertex slightly above the level of the upper border of the compound eyes. Outer cusp of lacinial tips broad, with seveneight short denticles. Mx4/Mx2: 1.22. Forewings (
<figureCitation id="13052A65D151260BFE8DFAC94D98FAB4" box="[352,439,1377,1403]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 34</figureCitation>
): L/W: 2.51. Pterostigma: lp/wp: 4.08, areola postica wide, slightly slanted posteriorly, apically rounded: al/ah: 1.68, R4+5 sinuous, M five-branched, M5 forked. Hindwings (
<figureCitation id="13052A65D151260BFB8FFA2D4899FA50" box="[1122,1206,1412,1439]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 35</figureCitation>
): l/w: 2.75; M threebranched. Hypandrium (
<figureCitation id="13052A65D151260BFE44FA014DD3FA0C" box="[425,508,1449,1475]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 37</figureCitation>
), setose as illustrated; phallosome V-shaped anteriorly, side struts slender; external parameres laminar, apices rounded, bearing pores (
<figureCitation id="13052A65D151260BFD3FFA654F09FA28" box="[722,806,1485,1511]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 39</figureCitation>
); three pairs of endophallic sclerites, anterior and lateral pair laminar, fused; mesal pair wide basally, mesal sclerite processes distally acuminate. Paraprocts (
<figureCitation id="13052A65D151260BFB1CFA59496EF9C4" box="[1265,1345,1521,1547]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 38</figureCitation>
) robust, almost triangular, with distal setal field and macrosetae as illustrated; sensory fields with 26 trichobothria on basal rosettes. Epiproct (
<figureCitation id="13052A65D151260BFE87F9914D93F99C" box="[362,444,1593,1619]" captionStart="FIGURES 3439" captionStartId="15.[151,264,1873,1898]" captionTargetBox="[159,1431,193,1839]" captionTargetId="figure@15.[151,1435,181,1848]" captionTargetPageId="15" captionText="FIGURES 3439. Loneura carijona n. sp. Male. 34. Forewing. 35. Hindwing. 36. Front view of head. 37. Hypandrium. 38. Left paraproct and epiproct. 39. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904994" httpUri="https://zenodo.org/record/3904994/files/figure.png" pageId="14" pageNumber="215">Fig. 38</figureCitation>
) semioval, with short and long setae, two lateral macrosetae, as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="C324656BD151260BFF2AF9F54D6EF970" pageId="14" pageNumber="215" type="description">
<paragraph id="8B8136E0D151260BFF2AF9F54D6EF970" blockId="14.[151,1437,656,1907]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AF9F54D59F9B8" bold="true" box="[199,374,1629,1655]" pageId="14" pageNumber="215">Measurements</emphasis>
. FW: 4100, HW: 2850, F: 1032, T: 1750, t1: 750, t2: 100, t3: 133, ctt1: 23, f1: 625, f2: 525, f3: 442, f4: 400, f5: 300, f6: 275, f7: 215, f8: 200, f9: 177, f10: 172, f11: 162, Mx: 426, IO: 490, D: 410, d: 270, IO/d: 1.81, PO: 0.66.
</paragraph>
</subSubSection>
<subSubSection id="C324656BD151260BFF2AF9604DC1F8E4" pageId="14" pageNumber="215" type="materials_examined">
<paragraph id="8B8136E0D151260BFF2AF9604DC1F8E4" blockId="14.[151,1437,656,1907]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AF9604DB8F92C" bold="true" box="[199,407,1736,1763]" pageId="14" pageNumber="215">Material studied.</emphasis>
<materialsCitation id="3B563CBDD151260BFE4CF9614DC1F8E4" collectingDate="2017-04-28" collectingDateMax="2017-04-29" collectingDateMin="2017-04-28" collectorName="J. Panche." country="Colombia" county="San Vicente del Caguan" elevation="917" latitude="2.4639277" location="Reserva Indigena Altamira" longLatPrecision="1" longitude="-74.91724" municipality="Laureles" pageId="14" pageNumber="215" specimenCount="917" stateProvince="Caqueta" typeStatus="holotype">
<typeStatus id="54858842D151260BFE4CF9614E26F92C" box="[417,521,1737,1763]" pageId="14" pageNumber="215" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F3297670D151260BFDB7F9614EDCF92C" box="[602,755,1737,1763]" name="Colombia" pageId="14" pageNumber="215">COLOMBIA</collectingCountry>
.
<collectingRegion id="49FAF802D151260BFCEDF9614F70F92C" box="[768,863,1737,1763]" country="Colombia" name="Caqueta" pageId="14" pageNumber="215">Caquetá</collectingRegion>
.
<collectingCounty id="62E04E6CD151260BFC81F96148ACF92C" box="[876,1155,1737,1763]" pageId="14" pageNumber="215">San Vicente del Caguán</collectingCounty>
,
<collectingMunicipality id="6BE5AC9AD151260BFB7CF96148DBF92C" box="[1169,1268,1737,1763]" pageId="14" pageNumber="215">Laureles</collectingMunicipality>
,
<location id="8EE1603BD151260BFAEFF9614D15F8C8" LSID="urn:lsid:plazi:treatment:039787F6D151260BFF7AFD8D4EB7F8BC:8EE1603BD151260BFAEFF9614D15F8C8" country="Colombia" county="San Vicente del Caguan" latitude="2.4639277" longLatPrecision="1" longitude="-74.91724" municipality="Laureles" name="Reserva Indigena Altamira" pageId="14" pageNumber="215" stateProvince="Caqueta">Reserva Indígena Altamira</location>
,
<geoCoordinate id="EE0A5027D151260BFEA4F9444DC7F8C8" box="[329,488,1772,1799]" degrees="2" direction="north" minutes="27" orientation="latitude" pageId="14" pageNumber="215" precision="1" seconds="50.14" value="2.4639277">2º2750.14”N</geoCoordinate>
:
<geoCoordinate id="EE0A5027D151260BFE1BF9444EB7F8C8" box="[502,664,1772,1799]" degrees="74" direction="west" minutes="55" orientation="longitude" pageId="14" pageNumber="215" precision="1" seconds="2.06" value="-74.91724">74º552.06”W</geoCoordinate>
,
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<elevation id="0013D1D3D151260BFD4BF9454EDCF8C8" box="[678,755,1773,1799]" metricMagnitude="2" metricUnit="m" metricValue="9.17" pageId="14" pageNumber="215" unit="m" value="917.0">917 m</elevation>
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.
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<date id="FF801020D151260BFCECF9454F85F8C8" box="[769,938,1773,1799]" pageId="14" pageNumber="215" value="2017-04-28" valueMax="2017-04-29" valueMin="2017-04-28">
<collectingDate id="EFC4E9C8D151260BFCECF9454F85F8C8" box="[769,938,1773,1799]" pageId="14" pageNumber="215" value="2017-04-28" valueMax="2017-04-29" valueMin="2017-04-28">2829.IV.2017</collectingDate>
</date>
.
<collectingMethod id="527F4EF7D151260BFC5AF9454879F8C8" box="[951,1110,1773,1799]" pageId="14" pageNumber="215">Led light trap</collectingMethod>
in forest canopy.
<collectorName id="26CB5336D151260BFAC4F94549B2F8C8" box="[1321,1437,1773,1799]" pageId="14" pageNumber="215">J. Panche.</collectorName>
MUSENUV slide code 29312.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324656BD151260BFF2AF89D4EB7F8BC" pageId="14" pageNumber="215" type="etymology">
<paragraph id="8B8136E0D151260BFF2AF89D4EB7F8BC" blockId="14.[151,1437,656,1907]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AF89D4D61F880" bold="true" box="[199,334,1845,1871]" pageId="14" pageNumber="215">Etymology.</emphasis>
This species is dedicated to the Carijona indigenous people, who inhabited the Departments of Amazonas,
<collectingRegion id="49FAF802D151260BFEF6F8F14D58F8BC" box="[283,375,1881,1907]" country="Colombia" name="Caqueta" pageId="14" pageNumber="215">Caquetá</collectingRegion>
and
<collectingRegion id="49FAF802D151260BFE43F8F14E38F8BC" box="[430,535,1881,1907]" country="Colombia" name="Guaviare" pageId="14" pageNumber="215">Guaviare</collectingRegion>
,
<collectingCountry id="F3297670D151260BFDCFF8F14EBBF8BC" box="[546,660,1881,1907]" name="Colombia" pageId="14" pageNumber="215">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>