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<document id="DA4D64CB06EECB5D783F8152C924DDC2" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D5852AFF985991FCEB5666FC88" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) longiceps Paulian 1938" docType="treatment" docVersion="10" lastPageNumber="476" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="475" updateTime="1698565278120" updateUser="plazi">
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<mods:title id="F6B320186C1DA08FDCF43F3B84810203">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="CBCDFEE98D9D4322760AB6B1DDE1AB1C">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="22155C4D53FD844F5D9F70EF02E954D4">Louzada, Júlio</mods:namePart>
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<mods:namePart id="838F9EE5D59666859EAF9F479FB42912">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:typeOfResource id="DD7B962FF292F63D6855BEBAAA8D7138">text</mods:typeOfResource>
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<mods:title id="C794241AAF877AE7C5A77B372D2F01E3">Zootaxa</mods:title>
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<mods:date id="B7EA0BE3BF7D4C4804D2EBA90FE63F47">2015</mods:date>
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<treatment id="03DE87D5852AFF985991FCEB5666FC88" ID-DOI="http://doi.org/10.5281/zenodo.6115726" ID-GBIF-Taxon="119598943" ID-Zenodo-Dep="6115726" LSID="urn:lsid:plazi:treatment:03DE87D5852AFF985991FCEB5666FC88" httpUri="http://treatment.plazi.org/id/03DE87D5852AFF985991FCEB5666FC88" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
<subSubSection id="C36D6548852AFF995991FCEB577FFC9A" pageId="24" pageNumber="475" type="nomenclature">
<paragraph id="8BC836C3852AFF995991FCEB5053FCF8" blockId="24.[151,1308,786,846]" box="[151,1308,786,812]" pageId="24" pageNumber="475">
<heading id="D08081AF852AFF995991FCEB5053FCF8" bold="true" box="[151,1308,786,812]" fontSize="11" level="1" pageId="24" pageNumber="475" reason="1">
<emphasis id="B903EAD1852AFF995991FCEB5053FCF8" bold="true" box="[151,1308,786,812]" pageId="24" pageNumber="475">
<taxonomicName id="4C774D40852AFF995991FCEB5643FCF8" ID-CoL="6CJ5K" authority="Paulian 1938" authorityName="Paulian" authorityYear="1938" box="[151,780,786,812]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="longiceps" subGenus="Aganhyboma">
<emphasis id="B903EAD1852AFF995991FCEB5462FCF8" bold="true" box="[151,301,786,812]" italics="true" pageId="24" pageNumber="475">Deltochilum</emphasis>
(
<emphasis id="B903EAD1852AFF99583BFCEB5491FCF8" bold="true" box="[317,478,786,812]" italics="true" pageId="24" pageNumber="475">Aganhyboma</emphasis>
)
<emphasis id="B903EAD1852AFF9958F6FCEB572DFCF8" bold="true" box="[496,610,786,812]" italics="true" pageId="24" pageNumber="475">longiceps</emphasis>
<bibRefCitation id="EFE64B32852AFF995B6FFCEB5643FCF8" author="Paulian" box="[617,780,786,812]" pageId="24" pageNumber="475" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
</taxonomicName>
new subgeneric placement and new status
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C3852AFF995991FCCC577FFC9A" blockId="24.[151,1308,786,846]" box="[151,560,821,846]" pageId="24" pageNumber="475">
(
<figureCitation id="134C2A46852AFF995999FCCC55BAFC9A" box="[159,245,821,846]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="24" pageNumber="475">Figs 16</figureCitation>
,
<figureCitation id="134C2A46852AFF995807FCCF546FFC9A" box="[257,288,822,846]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="24" pageNumber="475">37</figureCitation>
, 61, 80, 83, 84, 91, 110)
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852AFF995991FC8451ACFBF8" pageId="24" pageNumber="475" type="reference_group">
<paragraph id="8BC836C3852AFF995991FC8457A9FC66" blockId="24.[151,1304,893,1069]" pageId="24" pageNumber="475">
<treatmentCitationGroup id="AB6711ED852AFF995991FC8457A9FC66" pageId="24" pageNumber="475">
<taxonomicName id="4C774D40852AFF995991FC8454B7FC47" ID-CoL="34KB6" box="[151,504,893,915]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="valgum" subGenus="Deltochilum">
<emphasis id="B903EAD1852AFF995991FC84545BFC47" box="[151,276,893,915]" italics="true" pageId="24" pageNumber="475">Deltochilum</emphasis>
(
<emphasis id="B903EAD1852AFF995825FC8454EFFC47" box="[291,416,893,915]" italics="true" pageId="24" pageNumber="475">Deltochilum</emphasis>
)
<emphasis id="B903EAD1852AFF9958A8FC8454B7FC47" box="[430,504,893,915]" italics="true" pageId="24" pageNumber="475">valgum</emphasis>
</taxonomicName>
<taxonomicName id="4C774D40852AFF9958F9FC875602FC47" ID-CoL="6CJ5K" authority="Paulian 1938: 269" authorityName="Paulian" authorityPageNumber="269" authorityYear="1938" box="[511,845,893,915]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="variety" species="valgum" variety="longiceps">
var.
<emphasis id="B903EAD1852AFF995B2DFC8457C3FC47" box="[555,652,893,915]" italics="true" pageId="24" pageNumber="475">longiceps</emphasis>
<treatmentCitation id="0AD610D2852AFF995B94FC845602FC47" author="Paulian" box="[658,845,893,915]" page="269" pageId="24" pageNumber="475" year="1938">
<bibRefCitation id="EFE64B32852AFF995B94FC845654FC47" author="Paulian" box="[658,795,893,915]" pageId="24" pageNumber="475" refString="Paulian, R. (1938) Contribution a lʼetude des canthonides americains. Annales de la societe entomologique de France, 107, 213 - 296." type="journal article" year="1938">Paulian 1938</bibRefCitation>
: 269
</treatmentCitation>
</taxonomicName>
, 273 (original description, identification key)
<taxonomicName id="4C774D40852AFF995991FC65573AFC66" ID-CoL="6CJ5K" authority="Blackwelder 1944: 203" authorityName="Blackwelder" authorityPageNumber="203" authorityYear="1944" box="[151,629,924,946]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="B903EAD1852AFF995991FC655434FC66" box="[151,379,924,946]" italics="true" pageId="24" pageNumber="475">Deltochilum longiceps</emphasis>
:
<treatmentCitation id="0AD610D2852AFF99588FFC65573AFC66" author="Blackwelder" box="[393,629,924,946]" page="203" pageId="24" pageNumber="475" year="1944">
<bibRefCitation id="EFE64B32852AFF99588FFC655709FC66" author="Blackwelder" box="[393,582,924,946]" pageId="24" pageNumber="475" refString="Blackwelder, R. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum, 185, 189 - 265." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 203
</treatmentCitation>
</taxonomicName>
(checklist)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C3852AFF995991FC435492FC04" blockId="24.[151,1304,893,1069]" box="[151,477,954,976]" pageId="24" pageNumber="475">
<bibRefCitation id="EFE64B32852AFF995991FC435479FC04" author="Contreras" box="[151,310,954,976]" pageId="24" pageNumber="475" refString="Contreras, G. (1951) Catalogo de coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 8, 220 - 229." type="journal article" year="1951">Contreras 1951</bibRefCitation>
: 222 (catalogue)
</paragraph>
<paragraph id="8BC836C3852AFF995991FC20568EFBDA" blockId="24.[151,1304,893,1069]" pageId="24" pageNumber="475">
<treatmentCitationGroup id="AB6711ED852AFF995991FC20568EFBDA" pageId="24" pageNumber="475">
<taxonomicName id="4C774D40852AFF995991FC2056EEFC3B" ID-CoL="6CJ5K" authority="Vulcano &amp; Pereira 1967: 557" authorityName="Vulcano &amp; Pereira" authorityPageNumber="557" authorityYear="1967" box="[151,929,985,1007]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="subSpecies" species="valgum" subGenus="Deltohyboma" subSpecies="longiceps">
<emphasis id="B903EAD1852AFF995991FC205723FC3B" box="[151,620,985,1007]" italics="true" pageId="24" pageNumber="475">Deltochilum (Deltohyboma) valgum longiceps</emphasis>
:
<treatmentCitation id="0AD610D2852AFF995B7FFC2056EEFC3B" author="Vulcano" box="[633,929,985,1007]" page="557" pageId="24" pageNumber="475" year="1967">
<bibRefCitation id="EFE64B32852AFF995B7FFC20563EFC3B" author="Vulcano" box="[633,881,985,1007]" pageId="24" pageNumber="475" refString="Vulcano, M. A. &amp; Pereira, F. S. (1967) Sinopse dos Passalidae e Scarabaeidae s. str. da regiao Amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica (Zoologia), 5, 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967</bibRefCitation>
: 557
</treatmentCitation>
</taxonomicName>
(identification key)
<taxonomicName id="4C774D40852AFF995991FC015734FBDA" ID-CoL="6CJ5K" authority="Cano 1998: 177" authorityName="Cano" authorityPageNumber="177" authorityYear="1998" box="[151,635,1016,1038]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="subSpecies" species="valgum" subSpecies="longiceps">
<emphasis id="B903EAD1852AFF995991FC015483FBDA" box="[151,460,1016,1038]" italics="true" pageId="24" pageNumber="475">Deltochilum valgum longiceps</emphasis>
:
<treatmentCitation id="0AD610D2852AFF9958DCFC015734FBDA" author="Cano" box="[474,635,1016,1038]" page="177" pageId="24" pageNumber="475" year="1998">
<bibRefCitation id="EFE64B32852AFF9958DCFC015703FBDA" author="Cano" box="[474,588,1016,1038]" pageId="24" pageNumber="475" refString="Cano, E. B. (1998) Deltochilum valgum acropyge Bates (Coleoptera: Scarabaeidae: Scarabaeinae): habitats and distribution. The Coleopterists Bulletin, 52, 174 - 178." type="journal article" year="1998">Cano 1998</bibRefCitation>
: 177
</treatmentCitation>
</taxonomicName>
(comment on feeding behavior)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836C3852AFF995991FBEF51ACFBF8" blockId="24.[151,1304,893,1069]" box="[151,1251,1046,1069]" pageId="24" pageNumber="475">
<bibRefCitation id="EFE64B32852AFF995991FBEF546AFBF8" author="Escobar" box="[151,293,1046,1068]" pageId="24" pageNumber="475" refString="Escobar, F. (2003) Feeding habitats and distributional records of 11 species of Neotropical Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 57, 131 - 132. http: // dx. doi. org / 10.1649 / 642" type="journal article" year="2003">Escobar 2003</bibRefCitation>
: 131 (observations of feeding behavior, referred as
<taxonomicName id="4C774D40852AFF995A2BFBEF5142FBF9" ID-CoL="6CJ5R" box="[813,1037,1046,1069]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD1852AFF995A2BFBEF5142FBF9" box="[813,1037,1046,1069]" italics="true" pageId="24" pageNumber="475">Deltochilum acropyge</emphasis>
</taxonomicName>
, &quot;misidentification&quot;)
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852AFF995991FBA0506FF822" pageId="24" pageNumber="475" type="description">
<paragraph id="8BC836C3852AFF995991FBA0506FF822" blockId="24.[151,1437,1113,2038]" pageId="24" pageNumber="475">
<emphasis id="B903EAD1852AFF995991FBA0543CFBA6" bold="true" box="[151,371,1113,1138]" pageId="24" pageNumber="475">Description. Body.</emphasis>
Head, pronotum, elytra and ventral surface black or dark brown, with an evident sheen. Length&gt;
<quantity id="4C8F9B26852AFF9959B5FB875444FB41" box="[179,267,1150,1174]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="24" pageNumber="475" unit="mm" value="13.0">13 mm</quantity>
.
<emphasis id="B903EAD1852AFF99581DFB84542DFB42" bold="true" box="[283,354,1149,1174]" pageId="24" pageNumber="475">Head.</emphasis>
Punctures dense, but not touching each other; foveiform punctures separated at the dorsal interocular surface by approximately one diameter (Fig. 61). Punctures around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae or with almost inconspicuous setae. Dorsal interocular distance approximately three times eye width (Fig. 61).
<emphasis id="B903EAD1852AFF995991FAF455B9FAF2" bold="true" box="[151,246,1293,1318]" pageId="24" pageNumber="475">Thorax.</emphasis>
Surface of pronotum with dense foveiform punctures, not touching each other. Punctures on the sides and posterior angles as close as those on anterior angles. Discal punctures smaller and more dispersed than those of lateral portion, separated by approximately two or three times their diameter. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by at least one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles straight, with approximately the same length of margin between anterior and lateral angles. Hypomera with dense foveiform punctures at anterior portion and absent at posteromedial portion (
<figureCitation id="134C2A46852AFF995ABCFA1C515BFA2A" box="[954,1044,1509,1534]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="24" pageNumber="475">Fig. 83</figureCitation>
); anteromedial punctures usually separated by less than one diameter. Diagonal band of hypomera from posteromedial portion to middle of lateral margin lacking punctures or with one or two isolated punctures on otherwise smooth surface (
<figureCitation id="134C2A46852AFF995DDDF9D4507DF992" box="[1243,1330,1581,1606]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="24" pageNumber="475">Fig. 83</figureCitation>
).
<emphasis id="B903EAD1852AFF995C4FF9D450D3F992" bold="true" box="[1353,1436,1581,1606]" pageId="24" pageNumber="475">Elytra.</emphasis>
Lateral margins slightly curved outward (
<figureCitation id="134C2A46852AFF995B87F9A857AAF9BE" box="[641,741,1617,1642]" captionStart="FIGURES 103 – 110" captionStartId="49.[151,264,1951,1974]" captionTargetBox="[197,1421,212,1926]" captionTargetId="figure@49.[165,1421,193,1931]" captionTargetPageId="49" captionText="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244808/files/figure.png" pageId="24" pageNumber="475">Fig. 110</figureCitation>
) and surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated; strial punctures often elliptical, separated by irregular spacing (Fig. 84). Interstriae flattened, with foveiform punctures surrounded by approximately five well-defined shiny microtubercles. Foveiform punctures separated on elytral disc by approximately half diameter. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length (Fig. 80). Apical carina or tubercle of second interstria inconspicuous. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria noticeably smaller than the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina incomplete, effaced medially (Fig. 80). Parameres symmetric and elongate, length equal to or longer than phallobase; in lateral view forming an angle greater than 60° with phallobase; apex rounded (
<figureCitation id="134C2A46852AFF995DD5F8405066F806" box="[1235,1321,1977,2002]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="24" pageNumber="475">Fig. 16</figureCitation>
). Lamella copulatrix (LC): subrectangular sclerites (
<figureCitation id="134C2A46852AFF995B6AF824578FF822" box="[620,704,2013,2038]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="24" pageNumber="475">Fig. 37</figureCitation>
). Fronto-lateral peripheral sclerite (FLP) as in Fig. 54.
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852BFF9859C1FF6E515DFE5C" pageId="25" pageNumber="476" type="materials_examined">
<paragraph id="8BC836C3852BFF9859C1FF6E5796FEC9" blockId="25.[151,1437,151,861]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF9859C1FF6E576EFF64" bold="true" box="[199,545,151,176]" pageId="25" pageNumber="476">
Material studied.
<typeStatus id="54CC8861852BFF9858B2FF6E576EFF64" box="[436,545,151,176]" pageId="25" pageNumber="476" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="ED66AE06852BFF985B3AFF6157DCFF64" box="[572,659,152,176]" pageId="25" pageNumber="476">BMNH</collectionCode>
)
<emphasis id="B903EAD1852BFF985BA8FF6E567BFF64" bold="true" box="[686,820,151,176]" pageId="25" pageNumber="476">[examined]</emphasis>
♀:
<emphasis id="B903EAD1852BFF985A7EFF6E5158FF64" bold="true" box="[888,1047,151,176]" pageId="25" pageNumber="476">
<collectingCountry id="F3607653852BFF985A7EFF6E5158FF64" box="[888,1047,151,176]" name="Colombia" pageId="25" pageNumber="476">COLOMBIA</collectingCountry>
</emphasis>
(no locality given in original description). 1. [white label, handwritten]. 416; 2. [white label, printed in black]. 6745; 3. [white label, bordered in red, round].
<typeStatus id="54CC8861852BFF98582BFF195427FF2C" box="[301,360,224,248]" pageId="25" pageNumber="476">Type</typeStatus>
; 4. [white label, handwritten].
<taxonomicName id="4C774D40852BFF985BD9FF265606FF2C" box="[735,841,223,248]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="longiceps">longiceps</taxonomicName>
/ Reiche/
<collectingCountry id="F3607653852BFF985AC7FF26517CFF2C" box="[961,1075,223,248]" name="Colombia" pageId="25" pageNumber="476">Colombie</collectingCountry>
; 5. [white label, handwritten].
<taxonomicName id="4C774D40852BFF985991FEFD546CFEC9" box="[151,291,260,285]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">Deltochilum</taxonomicName>
/
<taxonomicName id="4C774D40852BFF985834FEFD5768FEC9" box="[306,551,260,285]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="variety" species="valgum" variety="longiceps">valgum var. longiceps</taxonomicName>
/ Paulian/
<typeStatus id="54CC8861852BFF985B9CFEFC579AFEC9" box="[666,725,261,285]" pageId="25" pageNumber="476">Type</typeStatus>
.
</paragraph>
<paragraph id="8BC836C3852BFF9859C1FEDE51B1FE94" blockId="25.[151,1437,151,861]" box="[199,1278,295,320]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF9859C1FEDE5738FE94" bold="true" box="[199,631,295,320]" pageId="25" pageNumber="476">
Non-type material [1]:
<collectingCountry id="F3607653852BFF9858DFFEDE5738FE94" box="[473,631,295,320]" name="Colombia" pageId="25" pageNumber="476">COLOMBIA</collectingCountry>
</emphasis>
: TOLIMA: Mariquita,
<date id="FFC91003852BFF985A78FEDE5692FE94" box="[894,989,295,320]" pageId="25" pageNumber="476" value="1995-11">XI.1995</date>
, F. Escobar (
<specimenCount id="9D71FD4A852BFF985D76FED151D6FEEB" box="[1136,1177,296,320]" pageId="25" pageNumber="476" type="generic">1 ♂</specimenCount>
<collectionCode id="ED66AE06852BFF985DA6FED151BCFE94" box="[1184,1267,296,320]" pageId="25" pageNumber="476">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C3852BFF9859C1FEB55788FEB0" blockId="25.[151,1437,151,861]" box="[199,711,332,357]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF9859C1FEB554E3FEB1" bold="true" box="[199,428,332,357]" pageId="25" pageNumber="476">Sampling methods.</emphasis>
un-baited pitfall trap [1].
</paragraph>
<paragraph id="8BC836C3852BFF9859C1FE96515DFE5C" blockId="25.[151,1437,151,861]" box="[199,1042,367,392]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF9859C1FE965465FE5C" bold="true" box="[199,298,367,392]" pageId="25" pageNumber="476">Habitat.</emphasis>
<collectingCountry id="F3607653852BFF985837FE9654D0FE5C" box="[305,415,367,392]" name="Colombia" pageId="25" pageNumber="476">Colombia</collectingCountry>
(
<figureCitation id="134C2A46852BFF9858A8FE96574DFE5C" box="[430,514,367,392]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="25" pageNumber="476">Fig. 91</figureCitation>
), one specimen with label data &quot;Bosque Seco&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C36D6548852BFF9859C1FE6D5666FC88" pageId="25" pageNumber="476" type="diagnosis">
<paragraph id="8BC836C3852BFF9859C1FE6D543CFD19" blockId="25.[151,1437,151,861]" pageId="25" pageNumber="476">
<emphasis id="B903EAD1852BFF9859C1FE6D54A2FE79" bold="true" box="[199,493,404,429]" pageId="25" pageNumber="476">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D40852BFF985B96FE6C57B8FE78" box="[656,759,405,428]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD1852BFF985B96FE6C57B8FE78" box="[656,759,405,428]" italics="true" pageId="25" pageNumber="476">acropyge</emphasis>
</taxonomicName>
complex): foveiform punctures separated at the dorsal interocular surface by approximately one diameter (Fig. 61); foveiform punctures of pronotal disc separated by approximately two or three times their diameter; punctures of elytral striae often elliptical, separated by irregular spacing (Fig. 84); apical carina or tubercle of second elytral interstria inconspicuous; pseudepipleural carina incomplete, effaced medially (Fig. 80). The pattern of hypomera punctures on individuals of this species is also characteristic (
<figureCitation id="134C2A46852BFF98583AFDBE54C0FDB4" box="[316,399,583,608]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="25" pageNumber="476">Fig. 83</figureCitation>
). The following aedeagus characters are also diagnostic: parameres symmetric and elongate, length equal to or longer than phallobase; in lateral view forming an angle greater than 60° with phallobase; apex rounded (
<figureCitation id="134C2A46852BFF985803FD765413FD7C" box="[261,348,655,680]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="25" pageNumber="476">Fig. 16</figureCitation>
). Lamella copulatrix (LC): subrectangular sclerites (
<figureCitation id="134C2A46852BFF985ABAFD76515BFD7C" box="[956,1044,655,680]" captionStart="FIGURES 37 – 41" captionStartId="42.[151,264,1811,1834]" captionTargetBox="[218,1374,212,1759]" captionTargetId="figure@42.[197,1374,193,1771]" captionTargetPageId="42" captionText="FIGURES 37 – 41. Lamella copulatrix sclerites in D. (Aganhyboma). 37. D. longiceps new status. 38. D. valgum. 39. D. icaroides. 40. D. icariforme. 41. D. subrubrum new species. Figures 42 – 49. Superior right peripheral sclerite (SRP) in D. (Aganhyboma). 42. D. titovidaurrei new species. 43. D. viridescens new status. 44. D. trisignatum. 45. D. finestriatum new species. 46. D. acropyge new status. 47. D. valgum. 48. D. icaroides. 49. D. kolleri new species. Figures 50 – 54. Fronto-lateral peripheral sclerite (FLP) in D. (Aganhyboma). 50. D. cupreicolle. 51. D. viridescens new status. 52. D. trisignatum. 53. D. finestriatum new species. 54. D. schefflerorum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244801/files/figure.png" pageId="25" pageNumber="476">Fig. 37</figureCitation>
). Fronto-lateral peripheral sclerite (FLP) as in Fig. 54.
</paragraph>
<paragraph id="8BC836C3852BFF9859C1FD2E5666FC88" blockId="25.[151,1437,151,861]" pageId="25" pageNumber="476">
<bibRefCitation id="EFE64B32852BFF9859C1FD2E5436FD24" author="Escobar" box="[199,377,727,752]" pageId="25" pageNumber="476" refString="Escobar, F. (2003) Feeding habitats and distributional records of 11 species of Neotropical Scarabaeinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin, 57, 131 - 132. http: // dx. doi. org / 10.1649 / 642" type="journal article" year="2003">Escobar (2003)</bibRefCitation>
reported the feeding behavior of seven specimens from
<collectingCountry id="F3607653852BFF985D2AFD2E51D5FD24" box="[1068,1178,727,752]" name="Colombia" pageId="25" pageNumber="476">Colombia</collectingCountry>
(Tolima), supposedly identified as
<taxonomicName id="4C774D40852BFF985820FD045736FCC0" authority="Bates. As" authorityName="Bates. As" box="[294,633,764,788]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="subSpecies" species="valgum" subSpecies="acropyge">
<emphasis id="B903EAD1852BFF985820FD045747FCC0" box="[294,520,764,788]" italics="true" pageId="25" pageNumber="476">D. valgum acropyge</emphasis>
Bates.
<collectingCountry id="F3607653852BFF985B5CFD055736FCC0" box="[602,633,764,788]" name="American Samoa" pageId="25" pageNumber="476">As</collectingCountry>
</taxonomicName>
far as we know,
<taxonomicName id="4C774D40852BFF985A33FD04514DFCC0" authority="Bates" authorityName="Bates" box="[821,1026,765,788]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD1852BFF985A33FD0456F1FCC0" box="[821,958,765,788]" italics="true" pageId="25" pageNumber="476">D. acropyge</emphasis>
Bates
</taxonomicName>
inhabits only the Caribbean slope of Central
<collectingCountry id="F3607653852BFF9859F4FCE6541DFCEC" box="[242,338,799,824]" name="United States of America" pageId="25" pageNumber="476">America</collectingCountry>
to
<collectingCountry id="F3607653852BFF98587FFCE6549DFCEC" box="[377,466,799,824]" name="Mexico" pageId="25" pageNumber="476">Mexico</collectingCountry>
. From
<collectingCountry id="F3607653852BFF985B25FCE657DEFCEC" box="[547,657,799,824]" name="Colombia" pageId="25" pageNumber="476">Colombia</collectingCountry>
we know only specimens of
<taxonomicName id="4C774D40852BFF985AE6FCD85120FCEC" box="[992,1135,800,824]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="B903EAD1852BFF985AE6FCD85120FCEC" box="[992,1135,800,824]" italics="true" pageId="25" pageNumber="476">D. longiceps</emphasis>
</taxonomicName>
. Therefore, we belive that those specimens reported from
<collectingCountry id="F3607653852BFF9858F0FCBD572AFC89" box="[502,613,836,861]" name="Colombia" pageId="25" pageNumber="476">Colombia</collectingCountry>
are
<taxonomicName id="4C774D40852BFF985B93FCBC566DFC88" box="[661,802,836,860]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="B903EAD1852BFF985B93FCBC566DFC88" box="[661,802,836,860]" italics="true" pageId="25" pageNumber="476">D. longiceps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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