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<document id="D940D9241ECDBED3610927AD0834B283" ID-DOI="10.11646/zootaxa.3722.4.1" ID-GBIF-Dataset="c092e547-503a-41ef-a855-4a12b5d776c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="223434" ID-ZooBank="3A461275-7565-4F6E-9785-12C45E12992E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460692161620" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2013" docId="03DE87B0FFD0FFDECADAB197FBC41DA1" docLanguage="en" docName="zt03722p438.pdf" docOrigin="Zootaxa 3722 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malaconothrus liklik Colloff, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="422" masterDocId="FFE7FFC8FFC4FFCBCA4DB328FF871A47" masterDocTitle="Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region" masterLastPageNumber="438" masterPageNumber="401" pageNumber="421" updateTime="1698351897157" updateUser="plazi">
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<mods:title id="06556B5E98CE0BFD0DCDD5EA03B62DAA">Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region</mods:title>
</mods:titleInfo>
<mods:name id="ECE3F2910B9EFC0314F12C1210A51786" type="personal">
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<mods:namePart id="DAFD35164C0679D162532F8844C3812D">Colloff, Matthew J.</mods:namePart>
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<mods:title id="2CFDAF8F35F5E058D208CA6A8A4D833B">Zootaxa</mods:title>
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<mods:date id="23ED9B2094C98416DE3F3A463AF96F5D">2013</mods:date>
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<mods:number id="6CA1E5E3155E31C18322E4264385DB16">3722</mods:number>
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<mods:identifier id="3CC180B92DCAEBA2F20B64A3E54940CD" type="DOI">10.11646/zootaxa.3722.4.1</mods:identifier>
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<treatment id="03DE87B0FFD0FFDECADAB197FBC41DA1" ID-DOI="http://doi.org/10.5281/zenodo.6157860" ID-GBIF-Taxon="119580482" ID-Zenodo-Dep="6157860" LSID="urn:lsid:plazi:treatment:03DE87B0FFD0FFDECADAB197FBC41DA1" httpUri="http://treatment.plazi.org/id/03DE87B0FFD0FFDECADAB197FBC41DA1" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
<subSubSection id="C36D652DFFD0FFDFCADAB197FAA11921" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="8BC836A6FFD0FFDFCADAB197FE7D189E" blockId="20.[151,506,703,762]" box="[151,506,703,729]" pageId="20" pageNumber="421">
<heading id="D08081CAFFD0FFDFCADAB197FE7D189E" bold="true" box="[151,506,703,729]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<emphasis id="B903EAB4FFD0FFDFCADAB197FE7D189E" bold="true" box="[151,506,703,729]" pageId="20" pageNumber="421">
<taxonomicName id="4C774D25FFD0FFDFCADAB197FE11189E" ID-CoL="7VRCW" box="[151,406,703,729]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="liklik" status="sp. nov.">
<emphasis id="B903EAB4FFD0FFDFCADAB197FE11189E" bold="true" box="[151,406,703,729]" italics="true" pageId="20" pageNumber="421">Malaconothrus liklik</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD0FFDFCBD0B197FE7D189E" box="[413,506,703,729]" pageId="20" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFD0FFDFCADAB1C9FF7E18BD" blockId="20.[151,506,703,762]" box="[151,249,737,762]" pageId="20" pageNumber="421">(Fig. 13)</paragraph>
<paragraph id="8BC836A6FFD0FFDFCADAB001FAA11921" blockId="20.[151,1437,809,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCADAB001FEA51905" bold="true" box="[151,290,809,834]" pageId="20" pageNumber="421">Dimensions</emphasis>
.
<typeStatus id="54CC8804FFD0FFDFCB62B001FE101905" box="[303,407,809,834]" pageId="20" pageNumber="421" type="holotype">Holotype</typeStatus>
female length 378 Μm, breadth 187 Μm.
<typeStatus id="54CC8804FFD0FFDFC92CB002FC491905" box="[865,974,810,834]" pageId="20" pageNumber="421" type="paratype">Paratypes</typeStatus>
(
<emphasis id="B903EAB4FFD0FFDFC993B003FC6B1905" box="[990,1004,811,834]" italics="true" pageId="20" pageNumber="421">n</emphasis>
= 4) mean length 388 Μm (range 378 397 Μm); mean breadth 184 Μm (range 182189 Μm). Ratio of prodorsum to total length (
<typeStatus id="54CC8804FFD0FFDFCECDB065FB6E1921" box="[1152,1257,845,870]" pageId="20" pageNumber="421" type="holotype">holotype</typeStatus>
): 0.3.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFD0FFDFCA8AB059FDA51C9A" pageId="20" pageNumber="421" type="description">
<paragraph id="8BC836A6FFD0FFDFCA8AB059FB161E79" blockId="20.[151,1437,809,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB059FE9919CD" bold="true" box="[199,286,881,906]" pageId="20" pageNumber="421">Female</emphasis>
.
<emphasis id="B903EAB4FFD0FFDFCB7DB059FE2A19CD" box="[304,429,881,906]" italics="true" pageId="20" pageNumber="421">Prodorsum</emphasis>
. Rostrum broad, sub-rectangular, with transverse inter-rostral ridge above acute naso. Rostral seta (
<emphasis id="B903EAB4FFD0FFDFCB62B0BFFEC119E9" box="[303,326,919,942]" italics="true" pageId="20" pageNumber="421">ro</emphasis>
) smooth, thin, setiform, 26 Μm long, shorter than mutual distance, on apex of carina at junction with inter-rostral ridge on apex of simple, curved carina (Fig. 13a). Lamellar seta (
<emphasis id="B903EAB4FFD0FFDFCE15B091FBEB1995" box="[1112,1132,953,978]" italics="true" pageId="20" pageNumber="421">le</emphasis>
) smooth, setiform, 20 Μm long, shorter than mutual distance, not extending as far as base of seta
<emphasis id="B903EAB4FFD0FFDFC989B0F7FC5C19B1" box="[964,987,991,1014]" italics="true" pageId="20" pageNumber="421">ro</emphasis>
; positioned medial of prodorsal carina. Interlamellar seta (
<emphasis id="B903EAB4FFD0FFDFCB27B72BFE071E5D" box="[362,384,1027,1050]" italics="true" pageId="20" pageNumber="421">in</emphasis>
) smooth, thin, setiform, 28 Μm long, length 0.3 × mutual distance; 2 × length of smooth, thin, setiform exobothridial seta
<emphasis id="B903EAB4FFD0FFDFCB87B70FFE641E79" box="[458,483,1063,1086]" italics="true" pageId="20" pageNumber="421">ex</emphasis>
. Prodorsum smooth, porose. Cerotegument densely granular.
</paragraph>
<paragraph id="8BC836A6FFD0FFDFCA8AB763FBF91F59" blockId="20.[151,1437,809,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB763FEC51E25" box="[199,322,1099,1122]" italics="true" pageId="20" pageNumber="421">Notogaster</emphasis>
. Anterior notogastral margin slightly convex; humeral region rounded (Fig. 13a). Cerotegument of large-broadly-spaced foveolae. Lyrifissure
<emphasis id="B903EAB4FFD0FFDFC833B747FD131EC1" box="[638,660,1135,1158]" italics="true" pageId="20" pageNumber="421">ia</emphasis>
very large: 21 Μm long, 4 Μm broad. With 15 pairs of thin, smooth notogastral setae plus
<emphasis id="B903EAB4FFD0FFDFCBC3B7BAFE111EED" box="[398,406,1170,1194]" italics="true" pageId="20" pageNumber="421">f</emphasis>
1, represented by their alveoli;
<emphasis id="B903EAB4FFD0FFDFC8A7B7BBFD711EED" box="[746,758,1171,1194]" italics="true" pageId="20" pageNumber="421">c</emphasis>
1,
<emphasis id="B903EAB4FFD0FFDFC946B7B9FC9E1EED" box="[779,793,1169,1194]" italics="true" pageId="20" pageNumber="421">d</emphasis>
1 and
<emphasis id="B903EAB4FFD0FFDFC91AB7BBFCE41EED" box="[855,867,1171,1194]" italics="true" pageId="20" pageNumber="421">e</emphasis>
1 much longer (4568 Μm) than others (2033 Μm). Seta
<emphasis id="B903EAB4FFD0FFDFCA82B791FF5C1E97" box="[207,219,1209,1232]" italics="true" pageId="20" pageNumber="421">c</emphasis>
2 positioned closer to
<emphasis id="B903EAB4FFD0FFDFCB9FB791FE591E97" box="[466,478,1209,1232]" italics="true" pageId="20" pageNumber="421">c</emphasis>
3 than to
<emphasis id="B903EAB4FFD0FFDFC808B791FDD61E97" box="[581,593,1209,1232]" italics="true" pageId="20" pageNumber="421">c</emphasis>
1. Notogaster elongated, shield-shaped; margins parallel, broadest between
<emphasis id="B903EAB4FFD0FFDFCADAB7C8FF351EB0" box="[151,178,1248,1271]" italics="true" pageId="20" pageNumber="421">cp</emphasis>
and
<emphasis id="B903EAB4FFD0FFDFCAAAB7C8FF741EB0" box="[231,243,1248,1271]" italics="true" pageId="20" pageNumber="421">e</emphasis>
2. Short, acute, diagonal ridges between
<emphasis id="B903EAB4FFD0FFDFC8FCB7F7FD381EBF" box="[689,703,1247,1272]" italics="true" pageId="20" pageNumber="421">h</emphasis>
2 and
<emphasis id="B903EAB4FFD0FFDFC8B1B7F7FC8D1EBF" box="[764,778,1247,1272]" italics="true" pageId="20" pageNumber="421">h</emphasis>
1. Longitudinal ridges absent. Setae
<emphasis id="B903EAB4FFD0FFDFCED7B7C8FB2F1EB0" box="[1178,1192,1248,1271]" italics="true" pageId="20" pageNumber="421">p</emphasis>
3 markedly anterior of
<emphasis id="B903EAB4FFD0FFDFCADAB62FFF221F59" box="[151,165,1287,1310]" italics="true" pageId="20" pageNumber="421">p</emphasis>
2. Ratio of length to breadth 1.4. Caudal margin U-shaped. Lyrifissurae
<emphasis id="B903EAB4FFD0FFDFC989B62FFC5D1F59" box="[964,986,1287,1310]" italics="true" pageId="20" pageNumber="421">ip</emphasis>
curved, acute.
</paragraph>
<paragraph id="8BC836A6FFD0FFDFCA8AB606FF7D1F92" blockId="20.[151,1437,809,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB606FED11F02" box="[199,342,1326,1349]" italics="true" pageId="20" pageNumber="421">Coxisternum</emphasis>
. Epimeral setal formula 3-1-3-3 (Fig. 13b). Epimeral setae smooth, setiform; seta
<emphasis id="B903EAB4FFD0FFDFCEBCB604FA8A1F02" box="[1265,1293,1324,1349]" italics="true" pageId="20" pageNumber="421">1b</emphasis>
45 Μm long, very much longer than others;
<emphasis id="B903EAB4FFD0FFDFCBA1B679FD8F1F2D" box="[492,520,1361,1386]" italics="true" pageId="20" pageNumber="421">3b</emphasis>
,
<emphasis id="B903EAB4FFD0FFDFC858B679FDB71F2E" box="[533,560,1361,1385]" italics="true" pageId="20" pageNumber="421">3c</emphasis>
,
<emphasis id="B903EAB4FFD0FFDFC873B679FDDE1F2E" box="[574,601,1361,1385]" italics="true" pageId="20" pageNumber="421">4a</emphasis>
and
<emphasis id="B903EAB4FFD0FFDFC8C2B679FD2D1F2E" box="[655,682,1361,1385]" italics="true" pageId="20" pageNumber="421">4c</emphasis>
2230 Μm long; others 514 Μm long;
<emphasis id="B903EAB4FFD0FFDFCE13B679FBFE1F2E" box="[1118,1145,1361,1385]" italics="true" pageId="20" pageNumber="421">1c</emphasis>
in posterolateral position,
<emphasis id="B903EAB4FFD0FFDFCADAB65CFF341FCA" box="[151,179,1396,1421]" italics="true" pageId="20" pageNumber="421">4b</emphasis>
lateral,
<emphasis id="B903EAB4FFD0FFDFCB57B65CFEB11FCA" box="[282,310,1396,1421]" italics="true" pageId="20" pageNumber="421">3b</emphasis>
mediolateral. Epimeral plates all separated medially. Posterolateral margins of plates II acute. Posterior margin of epimere II and anterior margin of epimere III with rows of small tubercles. Apodeme IV concave.
</paragraph>
<paragraph id="8BC836A6FFD0FFDFCA8AB6C9FBBA1CFE" blockId="20.[151,1437,809,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB6C9FE101FBE" box="[199,407,1505,1530]" italics="true" pageId="20" pageNumber="421">Anogenital Region</emphasis>
. Genital plates with five pairs of very short, thin, smooth, setiform setae directed posteriorly (Fig. 13b);
<emphasis id="B903EAB4FFD0FFDFCB5AB52EFEA21C5A" box="[279,293,1542,1565]" italics="true" pageId="20" pageNumber="421">g</emphasis>
1-4 short, sub-equal (69 Μm),
<emphasis id="B903EAB4FFD0FFDFC838B52EFD041C5A" box="[629,643,1542,1565]" italics="true" pageId="20" pageNumber="421">g</emphasis>
5 18 Μm long, positioned midway down plate; distance between
<emphasis id="B903EAB4FFD0FFDFCF1BB52EFAE31C5A" box="[1366,1380,1542,1565]" italics="true" pageId="20" pageNumber="421">g</emphasis>
5 and
<emphasis id="B903EAB4FFD0FFDFCADAB505FF221C03" box="[151,165,1581,1604]" italics="true" pageId="20" pageNumber="421">g</emphasis>
4 2 × that between
<emphasis id="B903EAB4FFD0FFDFCB3EB505FE061C03" box="[371,385,1581,1604]" italics="true" pageId="20" pageNumber="421">g</emphasis>
4 and
<emphasis id="B903EAB4FFD0FFDFCBF3B505FE4B1C03" box="[446,460,1581,1604]" italics="true" pageId="20" pageNumber="421">g</emphasis>
3. Genital plates oval; each plate 79 Μm long, 41 Μm broad; posterior margin transverse. One pair of minute setiform anal setae, 4 Μm long. Each adanal plate 98 Μm long, 28 Μm broad, Adanal setae
<emphasis id="B903EAB4FFD0FFDFCF26B57AFA001C2C" box="[1387,1415,1618,1643]" italics="true" pageId="20" pageNumber="421">ad</emphasis>
2-3 thick, short (6 Μm), spiniform, barbed;
<emphasis id="B903EAB4FFD0FFDFC81CB551FDEA1CD5" box="[593,621,1657,1682]" italics="true" pageId="20" pageNumber="421">ad</emphasis>
1 setiform, smooth, long (24 Μm). Anterior margin of adanal plate acute, overlapped by posterior margin of genital plate, without thickened transverse ridge.
</paragraph>
<paragraph id="8BC836A6FFD0FFDFCA8AB5EDFDA51C9A" blockId="20.[151,1437,809,2009]" box="[199,546,1732,1757]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB5EDFF7B1C9B" box="[199,252,1733,1756]" italics="true" pageId="20" pageNumber="421">Legs</emphasis>
. Pre-tarsi monodactylous.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFD0FFDFCA8AB5C0FC041DF2" pageId="20" pageNumber="421" type="materials_examined">
<paragraph id="8BC836A6FFD0FFDFCA8AB5C0FC041DF2" blockId="20.[151,1437,809,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB5C0FE281D46" bold="true" box="[199,431,1768,1793]" pageId="20" pageNumber="421">Material examined</emphasis>
<typeStatus id="54CC8804FFD0FFDFCBF0B5C0FDAD1D46" box="[445,554,1768,1793]" pageId="20" pageNumber="421" type="holotype">Holotype</typeStatus>
: ANIC 407, litter, rainforest, Hayfield Mission, near Maprik,
<collectingCountry id="F3607636FFD0FFDFCF58B5C1FF6C1D62" name="Papua New Guinea" pageId="20" pageNumber="421">Papua New Guinea</collectingCountry>
,
<geoCoordinate id="EE435061FFD0FFDFCABBB424FEEF1D62" box="[246,360,1804,1829]" direction="south" orientation="latitude" pageId="20" pageNumber="421" precision="15" value="-3.6986113">3°41'55&quot;S</geoCoordinate>
<geoCoordinate id="EE435061FFD0FFDFCB23B424FE751D62" box="[366,498,1804,1829]" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="15" value="143.04945">143°2'58&quot;E</geoCoordinate>
, ca.
<specimenCount id="9D71FD2FFFD0FFDFC869B424FDE11D63" box="[548,614,1804,1829]" pageId="20" pageNumber="421" type="generic">
<quantity id="4C8F9B43FFD0FFDFC869B424FDE41D63" box="[548,611,1804,1829]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="421" unit="m" value="50.0">50 m</quantity>
.
</specimenCount>
, coll. R.W. Taylor,
<date id="FFC91066FFD0FFDFC90DB424FC3B1D62" box="[832,956,1804,1829]" pageId="20" pageNumber="421" value="1972-06-27">27.vi.1972</date>
. One
<typeStatus id="54CC8804FFD0FFDFC9B1B425FBD81D62" box="[1020,1119,1805,1829]" pageId="20" pageNumber="421" type="paratype">paratype</typeStatus>
, ANIC 391, rainforest litter, near Lae,
<collectingCountry id="F3607636FFD0FFDFCB4AB419FE641D0E" box="[263,483,1840,1865]" name="Papua New Guinea" pageId="20" pageNumber="421">Papua New Guinea</collectingCountry>
, ca.
<specimenCount id="9D71FD2FFFD0FFDFC858B418FDDF1D0F" box="[533,600,1840,1865]" pageId="20" pageNumber="421" type="generic">
<quantity id="4C8F9B43FFD0FFDFC858B418FDD31D0F" box="[533,596,1840,1865]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="421" unit="m" value="50.0">50 m</quantity>
.
</specimenCount>
, coll. R.W. Taylor,
<date id="FFC91066FFD0FFDFC97EB418FC2B1D0E" box="[819,940,1840,1865]" pageId="20" pageNumber="421" value="1972-06-11">11.vi.1972</date>
. Two
<typeStatus id="54CC8804FFD0FFDFC9BDB419FBD91D0E" box="[1008,1118,1841,1865]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
, ANIC 395, rainforest litter, “Timber Track,
<quantity id="4C8F9B43FFD0FFDFCB13B47CFE2C1D2A" box="[350,427,1876,1901]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="421" unit="km" value="16.0">16 km</quantity>
north-west of Lae,
<collectingCountry id="F3607636FFD0FFDFC8D1B47DFC041D2A" box="[668,899,1876,1901]" name="Papua New Guinea" pageId="20" pageNumber="421">Papua New Guinea</collectingCountry>
,
<geoCoordinate id="EE435061FFD0FFDFC9DEB47CFC5B1D2A" box="[915,988,1876,1901]" direction="south" orientation="latitude" pageId="20" pageNumber="421" precision="925" value="-6.6">6°36'S</geoCoordinate>
<geoCoordinate id="EE435061FFD0FFDFC9A4B47CFBD41D2A" box="[1001,1107,1876,1901]" direction="east" orientation="longitude" pageId="20" pageNumber="421" precision="925" value="146.86667">146°52'E</geoCoordinate>
,
<specimenCount id="9D71FD2FFFD0FFDFCE29B47DFB3E1D2B" box="[1124,1209,1877,1901]" pageId="20" pageNumber="421" type="generic">
<quantity id="4C8F9B43FFD0FFDFCE29B47DFB321D2B" box="[1124,1205,1877,1901]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="20" pageNumber="421" unit="m" value="220.0">220 m</quantity>
.
</specimenCount>
, coll. R.W. Taylor,
<date id="FFC91066FFD0FFDFCADAB450FE951DD6" box="[151,274,1912,1937]" pageId="20" pageNumber="421" value="1972-06-12">12.vi.1972</date>
. One
<typeStatus id="54CC8804FFD0FFDFCB1EB451FE311DD6" box="[339,438,1913,1937]" pageId="20" pageNumber="421" type="paratype">paratype</typeStatus>
, ANIC 408, same data as ANIC 407.
<typeStatus id="54CC8804FFD0FFDFC910B450FC421DD6" box="[861,965,1912,1937]" pageId="20" pageNumber="421" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54CC8804FFD0FFDFC9B1B451FBE01DD6" box="[1020,1127,1913,1937]" pageId="20" pageNumber="421" type="paratype">paratypes</typeStatus>
deposited in the Australian National Insect Collection, CSIRO Ecosystem Sciences, Canberra.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFD0FFDFCA8AB4E8FCF71D9E" box="[199,880,1984,2009]" pageId="20" pageNumber="421" type="etymology">
<paragraph id="8BC836A6FFD0FFDFCA8AB4E8FCF71D9E" blockId="20.[151,1437,809,2009]" box="[199,880,1984,2009]" pageId="20" pageNumber="421">
<emphasis id="B903EAB4FFD0FFDFCA8AB4E8FEC91D9E" bold="true" box="[199,334,1984,2009]" pageId="20" pageNumber="421">Etymology.</emphasis>
The specific name
<taxonomicName id="4C774D25FFD0FFDFC864B4E8FDE61D9E" box="[553,609,1984,2009]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="liklik">
<emphasis id="B903EAB4FFD0FFDFC864B4E8FDE61D9E" box="[553,609,1984,2009]" italics="true" pageId="20" pageNumber="421">liklik</emphasis>
</taxonomicName>
is Tok Pisin for small.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFD1FFDECA8AB3BFFF341893" pageId="21" pageNumber="422" type="discussion">
<paragraph id="8BC836A6FFD1FFDECA8AB3BFFAF91B2D" blockId="21.[151,1436,151,729]" pageId="21" pageNumber="422">
<emphasis id="B903EAB4FFD1FFDECA8AB3BFFEB31AF7" bold="true" box="[199,308,151,176]" pageId="21" pageNumber="422">Remarks</emphasis>
.
<taxonomicName id="4C774D25FFD1FFDECB06B3BFFDB81AF7" box="[331,575,151,176]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="liklik" status="sp. nov.">
<emphasis id="B903EAB4FFD1FFDECB06B3BFFDB81AF7" box="[331,575,151,176]" italics="true" pageId="21" pageNumber="422">Malaconothrus liklik</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD1FFDEC802B3BFFD351AE8" box="[591,690,151,176]" pageId="21" pageNumber="422" rank="species">
<emphasis id="B903EAB4FFD1FFDEC802B3BFFD2C1AF7" bold="true" box="[591,683,151,176]" pageId="21" pageNumber="422">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from other members of the genus by the following combination of characters: 1) the presence of a well-developed inter-rostral ridge; 2) setae
<emphasis id="B903EAB4FFD1FFDECEECB395FB301A93" box="[1185,1207,189,212]" italics="true" pageId="21" pageNumber="422">in</emphasis>
2 × length of
<emphasis id="B903EAB4FFD1FFDECF11B395FAF21A93" box="[1372,1397,189,212]" italics="true" pageId="21" pageNumber="422">ex</emphasis>
; 3) the foveolate notogastral cerotegument; 4) the shield-shaped notogaster; 5) the short, acute, diagonal notogastral ridges between
<emphasis id="B903EAB4FFD1FFDECB06B22CFEDE1B5A" box="[331,345,260,285]" italics="true" pageId="21" pageNumber="422">h</emphasis>
1 and
<emphasis id="B903EAB4FFD1FFDECBD1B22CFE2D1B5A" box="[412,426,260,285]" italics="true" pageId="21" pageNumber="422">h</emphasis>
2; 6) the extraordinarily long setae
<emphasis id="B903EAB4FFD1FFDEC977B22CFCD11B5A" box="[826,854,260,285]" italics="true" pageId="21" pageNumber="422">3b</emphasis>
; 7) the small tubercles along the posterior edge of epimere II and anterior edge of epimere III; 8) the five pairs of very short genital setae with
<emphasis id="B903EAB4FFD1FFDECEFEB204FB461B04" box="[1203,1217,300,323]" italics="true" pageId="21" pageNumber="422">g</emphasis>
5 halfway along the plate; 9) the singular, heteromorphic adanal setae, with
<emphasis id="B903EAB4FFD1FFDEC949B279FCA71B2D" box="[772,800,337,362]" italics="true" pageId="21" pageNumber="422">ad</emphasis>
2-3, short, barbed, spiniform and
<emphasis id="B903EAB4FFD1FFDECEC8B279FB261B2D" box="[1157,1185,337,362]" italics="true" pageId="21" pageNumber="422">ad</emphasis>
1 long and setiform.
</paragraph>
<paragraph id="8BC836A6FFD1FFDECA8AB250FF341893" blockId="21.[151,1436,151,729]" pageId="21" pageNumber="422">
<taxonomicName id="4C774D25FFD1FFDECA8AB250FE341BD6" box="[199,435,376,401]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="liklik">
<emphasis id="B903EAB4FFD1FFDECA8AB250FE341BD6" box="[199,435,376,401]" italics="true" pageId="21" pageNumber="422">Malaconothrus liklik</emphasis>
</taxonomicName>
belongs to a group of species that are monodactylous, have five or six pairs of genital setae, the centrodorsal setae longer than the others, a shield-shaped notogaster with parallel sides, an inter-rostral ridge and foveolate prodorsal and notogastral cerotegument. The group includes
<taxonomicName id="4C774D25FFD1FFDECE6FB2E9FE211BBA" authority="Palacios-Vargas &amp; Iglesias, 1997" authorityName="Palacios-Vargas &amp; Iglesias" authorityYear="1997" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="chajulensis">
<emphasis id="B903EAB4FFD1FFDECE6FB2E9FB421B9E" box="[1058,1221,448,473]" italics="true" pageId="21" pageNumber="422">M. chajulensis</emphasis>
Palacios-Vargas &amp; Iglesias, 1997 (
<collectingCountry id="F3607636FFD1FFDECB0FB2CCFE191BBA" box="[322,414,484,509]" name="Mexico" pageId="21" pageNumber="422">Mexico</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="4C774D25FFD1FFDECBFCB2CDFC171BBA" authority="Saha &amp; Sanyal, 1996" authorityName="Saha &amp; Sanyal" authorityYear="1996" box="[433,912,484,509]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="dipankari">
<emphasis id="B903EAB4FFD1FFDECBFCB2CDFDC31BBA" box="[433,580,484,509]" italics="true" pageId="21" pageNumber="422">M. dipankari</emphasis>
Saha &amp; Sanyal, 1996 (
<collectingCountry id="F3607636FFD1FFDEC905B2CCFC001BBA" box="[840,903,484,509]" name="India" pageId="21" pageNumber="422">India</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="4C774D25FFD1FFDEC9D1B2CDFAF61BBA" authority="Hammer, 1979 (Java)" authorityName="Hammer, 1979 (Java" box="[924,1393,484,509]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="dorsofoveolatus">
<emphasis id="B903EAB4FFD1FFDEC9D1B2CDFBF31BBA" box="[924,1140,484,509]" italics="true" pageId="21" pageNumber="422">M. dorsofoveolatus</emphasis>
Hammer, 1979 (Java)
</taxonomicName>
;
<taxonomicName id="4C774D25FFD1FFDECF30B2CDFE5D1866" authority="Hammer, 1971" authorityName="Hammer" authorityYear="1971" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="hexasetosus">
<emphasis id="B903EAB4FFD1FFDECF30B2CDFE9A1866" italics="true" pageId="21" pageNumber="422">M. hexasetosus</emphasis>
Hammer, 1971
</taxonomicName>
(
<collectingCountry id="F3607636FFD1FFDECBBCB120FD9B1866" box="[497,540,520,545]" name="Fiji" pageId="21" pageNumber="422">Fiji</collectingCountry>
, Java);
<taxonomicName id="4C774D25FFD1FFDEC8CCB121FBEC1866" authority="Yamamoto, 1998" authorityName="Yamamoto" authorityYear="1998" box="[641,1131,520,545]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B903EAB4FFD1FFDEC8CCB121FCAA1867" box="[641,813,521,544]" italics="true" pageId="21" pageNumber="422">M. marginatus</emphasis>
Yamamoto, 1998 (
<collectingCountry id="F3607636FFD1FFDECE57B120FBE41866" box="[1050,1123,520,545]" name="China" pageId="21" pageNumber="422">China</collectingCountry>
)
</taxonomicName>
;
<taxonomicName id="4C774D25FFD1FFDECE32B121FE381802" authority="Balogh &amp; Mahunka, 1969" authorityName="Balogh &amp; Mahunka" authorityYear="1969" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="pilosellus">
<emphasis id="B903EAB4FFD1FFDECE32B121FA9F1866" box="[1151,1304,520,545]" italics="true" pageId="21" pageNumber="422">M. pilosellus</emphasis>
Balogh &amp; Mahunka, 1969 (
<collectingCountry id="F3607636FFD1FFDECB22B104FE311802" box="[367,438,556,581]" name="Brazil" pageId="21" pageNumber="422">Brazil</collectingCountry>
)
</taxonomicName>
and
<taxonomicName id="4C774D25FFD1FFDEC843B105FBB21802" authority="Willmann, 1932 (Java)" authorityName="Willmann, 1932 (Java" box="[526,1077,556,581]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="pseudolamellatus">
<emphasis id="B903EAB4FFD1FFDEC843B105FC821802" box="[526,773,556,581]" italics="true" pageId="21" pageNumber="422">M. pseudolamellatus</emphasis>
Willmann, 1932 (Java)
</taxonomicName>
. Of these species, only
<taxonomicName id="4C774D25FFD1FFDECF30B105FECD182E" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="dorsofoveolatus">
<emphasis id="B903EAB4FFD1FFDECF30B105FECD182E" italics="true" pageId="21" pageNumber="422">M. dorsofoveolatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D25FFD1FFDECBC9B179FDB6182E" box="[388,561,592,617]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="hexasetosus">
<emphasis id="B903EAB4FFD1FFDECBC9B179FDB6182E" box="[388,561,592,617]" italics="true" pageId="21" pageNumber="422">M. hexasetosus</emphasis>
</taxonomicName>
share the short, acute, diagonal notogastral ridges. The latter species has six pairs of genital setae and
<taxonomicName id="4C774D25FFD1FFDECBF2B15CFD2C18CA" box="[447,683,628,653]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="liklik">
<emphasis id="B903EAB4FFD1FFDECBF2B15CFD2C18CA" box="[447,683,628,653]" italics="true" pageId="21" pageNumber="422">Malaconothrus liklik</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="4C774D25FFD1FFDECE0AB15DFAA518CA" box="[1095,1314,628,653]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="dorsofoveolatus">
<emphasis id="B903EAB4FFD1FFDECE0AB15DFAA518CA" box="[1095,1314,628,653]" italics="true" pageId="21" pageNumber="422">M. dorsofoveolatus</emphasis>
</taxonomicName>
from Java from which it differs by its heteromorphic adanal setae, longer setae
<emphasis id="B903EAB4FFD1FFDEC9DAB1B1FC3718F7" box="[919,944,665,688]" italics="true" pageId="21" pageNumber="422">ex</emphasis>
,
<emphasis id="B903EAB4FFD1FFDEC9F0B1B0FC5E18F6" box="[957,985,664,689]" italics="true" pageId="21" pageNumber="422">3b</emphasis>
and centrodorsal setae and shorter setae
<emphasis id="B903EAB4FFD1FFDECADAB194FF221892" box="[151,165,700,725]" italics="true" pageId="21" pageNumber="422">h</emphasis>
2.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFD1FFDECADAB4F8FBC41DA1" box="[151,1091,2000,2025]" pageId="21" pageNumber="422" type="description">
<paragraph id="8BC836A6FFD1FFDECADAB4F8FBC41DA1" blockId="21.[151,1091,2000,2025]" box="[151,1091,2000,2025]" pageId="21" pageNumber="422">
<emphasis id="B903EAB4FFD1FFDECADAB4F8FEA71DA2" bold="true" box="[151,288,2000,2022]" pageId="21" pageNumber="422">FIGURE 13.</emphasis>
<taxonomicName id="4C774D25FFD1FFDECB68B4F8FE7E1DA1" box="[293,505,2000,2022]" class="Arachnida" family="Malaconothridae" genus="Malaconothrus" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="422" phylum="Arthropoda" rank="species" species="liklik" status="sp. nov.">
<emphasis id="B903EAB4FFD1FFDECB68B4F8FE7E1DA1" box="[293,505,2000,2022]" italics="true" pageId="21" pageNumber="422">Malaconothrus liklik</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFD1FFDECBB2B4F8FDD71DA1" box="[511,592,2000,2022]" pageId="21" pageNumber="422" rank="species">
<emphasis id="B903EAB4FFD1FFDECBB2B4F8FDCE1DA2" bold="true" box="[511,585,2000,2021]" pageId="21" pageNumber="422">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<typeStatus id="54CC8804FFD1FFDEC811B4F8FD321DA1" box="[604,693,2000,2022]" pageId="21" pageNumber="422" type="holotype">holotype</typeStatus>
female; a) dorsal; b) ventral; c) seta
<emphasis id="B903EAB4FFD1FFDECE67B4F8FBB11DA1" box="[1066,1078,2000,2022]" italics="true" pageId="21" pageNumber="422">d</emphasis>
2.
</paragraph>
</subSubSection>
</treatment>
</document>