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<document ID-DOI="10.5733/afin.053.0119" ID-ISSN="2305-2562" ID-Zenodo-Dep="7917198" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683666793957" checkinUser="felipe" docAuthor="Wagner, Rüdiger &amp; Stuckenberg, Brian R." docDate="2012" docId="03DE879F2331FFE662F7FDA9C06F810C" docLanguage="en" docName="AfrInvertebr.53.1.355-367.pdf" docOrigin="African Invertebrates 53 (1)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.053.0119" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Nemopalpus inexpectatus Wagner &amp; Stuckenberg 2012, sp. n." docType="treatment" docVersion="2" lastPageNumber="358" masterDocId="FFE7FFE72335FFE2635DFFF8C435842D" masterDocTitle="New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae: Bruchomyiinae)" masterLastPageNumber="355" masterPageNumber="355" pageNumber="358" updateTime="1683694472035" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>New fossil and extant species of Nemopalpus Macquart (Diptera: Psychodidae: Bruchomyiinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wagner, Rüdiger</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Stuckenberg, Brian R.</mods:namePart>
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<mods:title>African Invertebrates</mods:title>
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<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-06-30</mods:number>
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<mods:detail type="volume">
<mods:number>53</mods:number>
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<mods:number>1</mods:number>
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<mods:start>355</mods:start>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.053.0119</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.053.0119</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7917198</mods:identifier>
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<taxonomicName ID-CoL="8CYT7" authority="Wagner" authorityName="Wagner &amp; Stuckenberg" authorityYear="2012" box="[426,826,593,620]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. n.">
<emphasis bold="true" box="[426,731,593,619]" pageId="4" pageNumber="358">Nemopalpus inexpectatus</emphasis>
Wagner
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,
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<paragraph blockId="4.[181,1142,593,1313]" box="[629,694,633,659]" pageId="4" pageNumber="358">
<figureCitation box="[629,694,633,659]" captionStart="Figs 14" captionStartId="2.[181,220,1582,1603]" captionTargetBox="[202,1125,976,1556]" captionTargetId="figure-371@2.[181,1141,972,1560]" captionTargetPageId="2" captionText="Figs 14. New fossil species of Nemopalpus: (13) N. velteni Wagner, sp. n., male: (1) habitus in amber; (2) head with mouthparts and basal antennomeres; (3) terminalia, dorsal view; (4) N. inexpectatus:DJQHU, VS. Q., PDOH WHUPLQDOLD, ODWHUDO YLHZ. 6RXUFHV RI ¿JXUHV:)LJV Ɨ± (5:);)LJ. 4 (%6 ZLWK additions by RW). Scale bars: Fig. 1 = 0.5 mm; Figs 2, 3 = 0.1 mm; Fig. 4 = scale unknown." figureDoi="http://doi.org/10.5281/zenodo.7917200" httpUri="https://zenodo.org/record/7917200/files/figure.png" pageId="4" pageNumber="358">Fig. 4</figureCitation>
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<emphasis bold="true" box="[181,1095,671,700]" pageId="4" pageNumber="358">(W\PRORJ\: 7KH VSHFL¿F HSLWKHW UHÀHFWV WKH XQH[SHFWHG GLVFRYHU\ RI DQ DGGLWLRQDO</emphasis>
<taxonomicName pageId="4" pageNumber="358">
<emphasis italics="true" pageId="4" pageNumber="358">Nemopalpus</emphasis>
species
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from Baltic amber.
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<paragraph blockId="4.[181,1142,593,1313]" box="[181,321,745,771]" pageId="4" pageNumber="358">Description:</paragraph>
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<emphasis box="[181,239,787,811]" italics="true" pageId="4" pageNumber="358">Male</emphasis>
.
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<emphasis box="[181,242,827,851]" italics="true" pageId="4" pageNumber="358">Head</emphasis>
&amp;
<emphasis box="[278,351,827,851]" italics="true" pageId="4" pageNumber="358">thorax</emphasis>
: Typical of the genus.
</paragraph>
<paragraph blockId="4.[181,1142,593,1313]" pageId="4" pageNumber="358">
<emphasis box="[181,287,867,891]" italics="true" pageId="4" pageNumber="358">Abdomen</emphasis>
: Terminalia with large gonocoxite in lateral view, increasing in width distally. Gonostylus small, elongate, blade-like, S-shaped, with fringe of setae, between which lies well sclerotised tip of aedeagus. Tergum 9 cone-shaped, cerci small, just aside cerci with
<emphasis bold="true" box="[181,1127,959,988]" pageId="4" pageNumber="358">D SDLU RI VPDOO WUL¿G DSSHQGDJHV DORQJ YHQWUDO PDUJLQ RI WHUJLWH (IXQFWLRQ XQNQRZQ).</emphasis>
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<paragraph blockId="4.[181,1142,593,1313]" pageId="4" pageNumber="358">
<emphasis bold="true" box="[181,1141,1001,1025]" pageId="4" pageNumber="358">+RORW\SH: ƃ (= 9ƟƟ) %DOWLF $PEHU; /DWH (RFHQH ($PEHU FROOHFWLRQ RI WKH *HRZLVVHQVFKDIWOLFKHV =HQWUXP</emphasis>
der Universität Göttingen).
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<paragraph blockId="4.[181,1142,593,1313]" pageId="4" pageNumber="358">
Notes: This specimen was repeatedly examined (
<emphasis box="[729,741,1064,1088]" italics="true" pageId="4" pageNumber="358">e</emphasis>
.
<emphasis box="[749,763,1064,1088]" italics="true" pageId="4" pageNumber="358">g</emphasis>
.,
<bibRefCitation author="HENNIG, W." box="[784,932,1062,1088]" pageId="4" pageNumber="358" pagination="1 - 69" refId="ref6537" refString="HENNIG, W. 1972. Insektenfossilien aus der unteren Kreide IV. Psychodidae (Phlebotominae), mit einer kritischen Ubersicht uber das phylogenetische System der Familie und die bisher beschriebenen Fossilien (Diptera). Stuttgarter Beitrage zur Naturkunde 241: 1 - 69." type="journal article" year="1972">Hennig 1972</bibRefCitation>
; B.R. Stuckenberg pers. comm., 2002), but never recognised as a separate species.After the recovery and redescription of other
<taxonomicName authority="(Wagner 2006)" baseAuthorityName="Wagner" baseAuthorityYear="2006" box="[399,710,1126,1152]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="genus">
<emphasis box="[399,538,1128,1152]" italics="true" pageId="4" pageNumber="358">Nemopalpus</emphasis>
(
<bibRefCitation author="WAGNER, R." box="[550,701,1126,1152]" pageId="4" pageNumber="358" pagination="83 - 95" refId="ref6936" refString="WAGNER, R. 2006. Amber Bruchomyiinae - descriptions of already known and new species, and the position RI WKH µVXEIDPLO \ ¶ ZLWKLQ 3 V \ FKRGLGDH (sensu lato) (Diptera). Studia dipterologica 13: 83 - 95." type="journal article" year="2006">Wagner 2006</bibRefCitation>
)
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, this species is now easily distinguished from other Baltic amber taxa. Although only visible in lateral view, the terminalia are unique. Gonocoxites of
<taxonomicName authority="Wagner, 2006" authorityName="Wagner" authorityYear="2006" box="[449,739,1190,1217]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="hoffeinsi">
<emphasis box="[449,468,1192,1216]" italics="true" pageId="4" pageNumber="358">N</emphasis>
.
<emphasis box="[482,578,1192,1216]" italics="true" pageId="4" pageNumber="358">hoffeinsi</emphasis>
<bibRefCitation author="WAGNER, R." box="[584,739,1190,1217]" pageId="4" pageNumber="358" pagination="83 - 95" refId="ref6936" refString="WAGNER, R. 2006. Amber Bruchomyiinae - descriptions of already known and new species, and the position RI WKH µVXEIDPLO \ ¶ ZLWKLQ 3 V \ FKRGLGDH (sensu lato) (Diptera). Studia dipterologica 13: 83 - 95." type="journal article" year="2006">Wagner, 2006</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Meunier, 1905" authorityName="Meunier" authorityYear="1905" box="[793,1101,1190,1217]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="tertiariae">
<emphasis box="[793,812,1192,1216]" italics="true" pageId="4" pageNumber="358">N</emphasis>
.
<emphasis box="[825,931,1192,1216]" italics="true" pageId="4" pageNumber="358">tertiariae</emphasis>
Meunier, 1905
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are elongate, tube-shaped; those of
<taxonomicName authority="Edwards, 1921" authorityName="Edwards" authorityYear="1921" box="[528,876,1222,1249]" class="Insecta" family="Psychodidae" genus="Nemopalpus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="molophilinus">
<emphasis box="[528,547,1224,1248]" italics="true" pageId="4" pageNumber="358">N</emphasis>
.
<emphasis box="[559,705,1224,1248]" italics="true" pageId="4" pageNumber="358">molophilinus</emphasis>
<bibRefCitation author="EDWARDS, F. W." box="[710,876,1222,1248]" pageId="4" pageNumber="358" pagination="437 - 439" refId="ref6470" refString="EDWARDS, F. W. 1921. A note on the subfamily Bruchomyiinae (Diptera Nematocera). Annals and Magazine of Natural History, Ser. 9 7: 437 - 439." type="journal article" year="1921">Edwards, 1921</bibRefCitation>
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are similar in shape, but the gonostylus is bipartite and larger than the gonocoxite. In addition, the new species has
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