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<document id="F2E759C07105C5FD64AA76CA0F0E51CF" ID-DOI="10.5281/zenodo.155753" ID-GBIF-Dataset="591f48a8-6ce9-453f-83c4-2e939759c052" ID-ISSN="1175­5326" ID-Zenodo-Dep="155753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459442263743" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="F207683E6E3CC40D8C38869CFAE0FB14" docLanguage="en" docName="zt00114.pdf" docOrigin="Zootaxa 114" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tropimenelytron Pace 1983" docType="treatment" docVersion="14" lastPageNumber="11" masterDocId="0E3E10466E3EC4078D308720FFEDFFB2" masterDocTitle="A revision of Nearctic species of the genus Tropimenelytron Pace, 1983 (Coleoptera: Staphylinidae: Aleocharinae), a new genus for North America" masterLastPageNumber="24" masterPageNumber="1" pageNumber="3" updateTime="1698190169484" updateUser="plazi">
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<mods:title id="2A844AFA815428D77DE570697913ED01">A revision of Nearctic species of the genus Tropimenelytron Pace, 1983 (Coleoptera: Staphylinidae: Aleocharinae), a new genus for North America</mods:title>
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<mods:date id="114E2A11112B6940A3925112C6E0C5FF">2002</mods:date>
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<treatment id="F207683E6E3CC40D8C38869CFAE0FB14" ID-DOI="http://doi.org/10.5281/zenodo.6277326" ID-GBIF-Taxon="119330575" ID-Zenodo-Dep="6277326" LSID="urn:lsid:plazi:treatment:F207683E6E3CC40D8C38869CFAE0FB14" httpUri="http://treatment.plazi.org/id/F207683E6E3CC40D8C38869CFAE0FB14" lastPageId="10" lastPageNumber="11" pageId="2" pageNumber="3">
<subSubSection id="32B48AA36E3CC4058C38869CFCA6FE67" box="[264,843,443,470]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="7A11D9286E3CC4058C38869CFCA6FE67" blockId="2.[264,843,443,470]" box="[264,843,443,470]" pageId="2" pageNumber="3">
<heading id="21596E446E3CC4058C38869CFCA6FE67" bold="true" box="[264,843,443,470]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<taxonomicName id="BDAEA2AB6E3CC4058C38869CFDB3FE67" ID-CoL="8VSDG" authority="Pace, 1983" authorityName="Pace" authorityYear="1983" box="[264,606,443,470]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3CC4058C38869CFE3FFE64" bold="true" box="[264,466,444,470]" italics="true" pageId="2" pageNumber="3">Tropimenelytron</emphasis>
<bibRefCitation id="1E3FA4D96E3CC4058CEA869BFDB3FE67" author="Pace" box="[474,606,443,469]" pageId="2" pageNumber="23" refString="Pace, R. (1983) Il genere Tropimenelytron Scheerpeltz (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie, 13 (2), 185 - 190." type="journal article" year="1983">Pace, 1983</bibRefCitation>
</taxonomicName>
(
<figureCitation id="E295C5AD6E3CC4058F5D869BFD1DFE67" box="[621,752,443,469]" captionStart-0="FIGURES 1 ­ 5" captionStart-1="FIGURES 6 ­ 11" captionStart-2="FIGURES 12 ­ 15" captionStart-3="FIGURES 16 ­ 21" captionStartId-0="2.[264,382,1787,1811]" captionStartId-1="3.[264,383,1648,1672]" captionStartId-2="5.[264,383,1635,1659]" captionStartId-3="6.[264,383,1667,1691]" captionTargetBox-0="[299,1289,735,1765]" captionTargetBox-1="[288,1296,741,1614]" captionTargetBox-2="[297,1291,428,1595]" captionTargetBox-3="[301,1282,802,1631]" captionTargetId-0="figure@2.[286,1302,724,1777]" captionTargetId-1="figure@3.[278,1309,729,1621]" captionTargetId-2="figure@5.[280,1306,409,1608]" captionTargetId-3="figure@6.[290,1298,793,1649]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 1 ­ 5. Mouthparts of Tropimenelytron tuberiventre (Eppelsheim) (Polikesh, Azerbaijan). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." captionText-1="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." captionText-2="FIGURES 12 ­ 15. Details of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12 ­ 14), 0.2 mm (15)." captionText-3="FIGURES 16 ­ 21. Male genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rensselaerville, New York (16 ­ 18) and Morgan Monroe State Park, Indiana (21 )) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (19 ­ 20 )). 16 – everted internal sac of aedeagus, lateral view; 17 – medial lamellae, parameral view; 18 – copulatory piece, parameral view; 19 – apex of everted internal sac, lateral view; 20 – apex of everted internal sac, parameral view; 21 – internal sac retracted in median lobe, parameral view. CP ­ copulatory piece; ML ­ medial lamellae. Scale bar 0.1 mm." httpUri-0="https://zenodo.org/record/155754/files/figure.png" httpUri-1="https://zenodo.org/record/155755/files/figure.png" httpUri-2="https://zenodo.org/record/155756/files/figure.png" httpUri-3="https://zenodo.org/record/155757/files/figure.png" pageId="2" pageNumber="3">Figs. 1­21</figureCitation>
,
<figureCitation id="E295C5AD6E3CC4058FCD869BFCAEFE67" box="[765,835,443,469]" captionStart-0="FIGURES 35 ­ 38" captionStart-1="FIGURES 39 ­ 44" captionStart-2="FIGURES 45 ­ 48" captionStart-3="FIGURES 49 ­ 54" captionStart-4="FIGURES 55 ­ 58" captionStart-5="FIGURES 59 ­ 64" captionStartId-0="12.[264,383,1248,1272]" captionStartId-1="13.[264,383,1231,1255]" captionStartId-2="14.[264,383,1346,1370]" captionStartId-3="15.[264,382,1742,1766]" captionStartId-4="17.[264,383,1668,1692]" captionStartId-5="18.[264,383,1139,1163]" captionTargetBox-0="[325,1262,417,1213]" captionTargetBox-1="[294,1263,291,1184]" captionTargetBox-2="[298,1291,463,1313]" captionTargetBox-3="[306,1282,761,1723]" captionTargetBox-4="[317,1275,833,1634]" captionTargetBox-5="[304,1295,297,1090]" captionTargetId-0="figure@12.[318,1269,409,1222]" captionTargetId-1="figure@13.[280,1276,284,1195]" captionTargetId-2="figure@14.[287,1299,449,1320]" captionTargetId-3="figure@15.[287,1300,753,1731]" captionTargetId-4="figure@17.[299,1288,809,1641]" captionTargetId-5="figure@18.[280,1313,284,1107]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionTargetPageId-4="17" captionTargetPageId-5="18" captionText-0="FIGURES 35 ­ 38. Details of abdominal segment 8 of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Morgan Monroe State Park (35 ­ 36) and Rockville, Indiana (37 ­ 38 )). 35 – male tergum 8; 36 – male sternum 8; 37 – female tergum 8; 38 – female sternum 8. Scale bar 0.5 mm." captionText-1="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." captionText-2="FIGURES 45 ­ 48. Details of abdominal segment 8 of Tropimenelytron robustum Gusarov, sp. n. (holotype (45 ­ 46), paratype from Hope, British Columbia (47 ­ 48 )). 45 – male tergum 8; 46 – male sternum 8; 47 – female tergum 8; 48 – female sternum 8. Scale bar 0.5 mm." captionText-3="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." captionText-4="FIGURES 55 ­ 58. Details of abdominal segment 8 of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 55 – male tergum 8; 56 – male sternum 8; 57 – female tergum 8; 58 – female sternum 8. Scale bar 0.5 mm." captionText-5="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri-0="https://zenodo.org/record/155760/files/figure.png" httpUri-1="https://zenodo.org/record/155761/files/figure.png" httpUri-2="https://zenodo.org/record/155762/files/figure.png" httpUri-3="https://zenodo.org/record/155763/files/figure.png" httpUri-4="https://zenodo.org/record/155764/files/figure.png" httpUri-5="https://zenodo.org/record/155765/files/figure.png" pageId="2" pageNumber="3">35­64</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="32B48AA36E3CC4058C388527FD70FD2C" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="7A11D9286E3CC4058C388527FD95FD8C" blockId="2.[264,1097,519,670]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="5ABEFE066E3CC4058C388527FD95FD8C" pageId="2" pageNumber="3">
<taxonomicName id="BDAEA2AB6E3CC4058C388527FD24FDAC" ID-CoL="8VSDG" authority="Scheerpeltz, 1955: 171" authorityName="Scheerpeltz" authorityPageNumber="171" authorityYear="1955" box="[264,713,519,542]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3CC4058C388527FE5AFDAC" box="[264,439,519,542]" italics="true" pageId="2" pageNumber="3">Tropimenelytron</emphasis>
:
<treatmentCitation id="FB0FFF396E3CC4058CFD8527FD24FDAC" author="Scheerpeltz" box="[461,713,519,542]" page="171" pageId="2" pageNumber="3" year="1955">
<bibRefCitation id="1E3FA4D96E3CC4058CFD8527FD7DFDAC" author="Scheerpeltz" box="[461,656,519,542]" pageId="2" pageNumber="24" refString="Scheerpeltz, O. (1955) Die neue Systematik der Gattung Sipalia Muls. Rey (Col. Staphylinidae). Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien, 92, 166 - 180." type="journal article" year="1955">Scheerpeltz, 1955</bibRefCitation>
: 171
</treatmentCitation>
</taxonomicName>
(first record but unavailable name).
<taxonomicName id="BDAEA2AB6E3CC4058C388507FD99FD8C" ID-CoL="8VSDG" authority="Pace, 1983: 187" authorityName="Pace" authorityPageNumber="187" authorityYear="1983" box="[264,628,551,574]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3CC4058C388507FE5AFD8C" box="[264,439,551,574]" italics="true" pageId="2" pageNumber="3">Tropimenelytron</emphasis>
<treatmentCitation id="FB0FFF396E3CC4058C8F8507FD99FD8C" author="Pace" box="[447,628,551,574]" page="187" pageId="2" pageNumber="3" year="1983">
<bibRefCitation id="1E3FA4D96E3CC4058C8F8507FDD5FD8C" author="Pace" box="[447,568,551,574]" pageId="2" pageNumber="23" refString="Pace, R. (1983) Il genere Tropimenelytron Scheerpeltz (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie, 13 (2), 185 - 190." type="journal article" year="1983">Pace, 1983</bibRefCitation>
: 187
</treatmentCitation>
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.
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</paragraph>
<paragraph id="7A11D9286E3CC4058C388567FCE2FDEC" blockId="2.[264,1097,519,670]" box="[264,783,583,606]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="5ABEFE066E3CC4058C388567FCE2FDEC" box="[264,783,583,606]" pageId="2" pageNumber="3">
<taxonomicName id="BDAEA2AB6E3CC4058C388567FDAEFDEC" ID-CoL="8VRYC" authority="(Tropimenelytron)" baseAuthorityName="Tropimenelytron" box="[264,579,583,606]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3CC4058C388567FE95FDEC" box="[264,376,583,606]" italics="true" pageId="2" pageNumber="3">Pelioptera</emphasis>
(
<emphasis id="48DA053A6E3CC4058CB88567FDDAFDEC" box="[392,567,583,606]" italics="true" pageId="2" pageNumber="3">Tropimenelytron</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="FB0FFF396E3CC4058F668567FCE6FDEC" author="Pace" box="[598,779,583,606]" page="839" pageId="2" pageNumber="3" year="1991">
<bibRefCitation id="1E3FA4D96E3CC4058F668567FD22FDEC" author="Pace" box="[598,719,583,606]" pageId="2" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace, 1991</bibRefCitation>
: 839
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7A11D9286E3CC4058C388547FD70FD2C" blockId="2.[264,1097,519,670]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="5ABEFE066E3CC4058C388547FD70FD2C" pageId="2" pageNumber="3">
<taxonomicName id="BDAEA2AB6E3CC4058C388547FDAEFDCC" ID-CoL="8VRYC" authority="(Tropimenelytron)" baseAuthorityName="Tropimenelytron" box="[264,579,615,638]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3CC4058C388547FE95FDCC" box="[264,376,615,638]" italics="true" pageId="2" pageNumber="3">Pelioptera</emphasis>
(
<emphasis id="48DA053A6E3CC4058CB88547FDDAFDCC" box="[392,567,615,638]" italics="true" pageId="2" pageNumber="3">Tropimenelytron</emphasis>
)
</taxonomicName>
:
<treatmentCitation id="FB0FFF396E3CC4058F668547FCC2FDCC" author="Assing" box="[598,815,615,638]" page="1000" pageId="2" pageNumber="3" year="2000">
<bibRefCitation id="1E3FA4D96E3CC4058F668547FD0BFDCC" author="Assing" box="[598,742,615,638]" pageId="2" pageNumber="23" refString="Assing, V. (2000) Three new brachypterous Athetini from North Korea (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 32 (2), 999 - 1006." type="journal article" year="2000">Assing, 2000</bibRefCitation>
: 1000
</treatmentCitation>
.
<taxonomicName id="BDAEA2AB6E3CC4058C3885A7FD74FD2C" ID-CoL="8VSDG" authority="Assing, 2001: 168" authorityName="Assing" authorityPageNumber="168" authorityYear="2001" box="[264,665,647,670]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3CC4058C3885A7FE5AFD2C" box="[264,439,647,670]" italics="true" pageId="2" pageNumber="3">Tropimenelytron</emphasis>
:
<treatmentCitation id="FB0FFF396E3CC4058CFD85A7FD74FD2C" author="Assing" box="[461,665,647,670]" page="168" pageId="2" pageNumber="3" year="2001">
<bibRefCitation id="1E3FA4D96E3CC4058CFD85A7FDB0FD2C" author="Assing" box="[461,605,647,670]" pageId="2" pageNumber="23" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing, 2001</bibRefCitation>
: 168
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="2ED189A06E3CC4058C3881DBFD25F8E0" httpUri="https://zenodo.org/record/155754/files/figure.png" pageId="2" pageNumber="3" targetBox="[299,1289,735,1765]" targetPageId="2">
<paragraph id="7A11D9286E3CC4058C3881DBFD25F8E0" blockId="2.[264,1323,1787,1874]" pageId="2" pageNumber="3">
<emphasis id="48DA053A6E3CC4058C3881DBFE5DF8A1" bold="true" box="[264,432,1787,1811]" pageId="2" pageNumber="3">FIGURES 1­5.</emphasis>
Mouthparts of
<taxonomicName id="BDAEA2AB6E3CC4058F6E81DCFBC1F8A0" authority="Eppelsheim" authorityName="Eppelsheim" box="[606,1068,1787,1811]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tuberiventre">
<emphasis id="48DA053A6E3CC4058F6E81DCFC78F8A1" box="[606,917,1788,1811]" italics="true" pageId="2" pageNumber="3">Tropimenelytron tuberiventre</emphasis>
(Eppelsheim)
</taxonomicName>
(Polikesh, Azerbaijan). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm.
</paragraph>
</caption>
<subSubSection id="32B48AA36E3DC4048C38863BFC61FDC4" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="7A11D9286E3DC4048C38863BFC61FDC4" blockId="3.[264,1324,283,630]" pageId="3" pageNumber="4">
<emphasis id="48DA053A6E3DC4048C38863BFE65FE87" bold="true" box="[264,392,283,309]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="BDAEA2AB6E3DC4048CAD863CFDB3FE87" box="[413,606,284,309]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3DC4048CAD863CFDB3FE87" box="[413,606,284,309]" italics="true" pageId="3" pageNumber="4">Tropimenelytron</emphasis>
</taxonomicName>
can be distinguished from other aleocharine genera by the combination of the following characters: parallel­sided body; ligula with wide base, split apically (
<figureCitation id="E295C5AD6E3DC4048C49864CFE29FE34" box="[377,452,364,390]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
); pronotum subquadrate, with microsetae directed posteriorly along the midline of the disc (
<typeStatus id="A515678A6E3DC4048F3F86B4FDA6FE1C" box="[527,587,404,430]" pageId="3" pageNumber="4">Type</typeStatus>
V,
<bibRefCitation id="1E3FA4D96E3DC4048FB286B4FC76FE1C" author="Benick" box="[642,923,404,430]" pageId="3" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Krefeld: Goecke &amp; Evers Verlag, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
) (
<figureCitation id="E295C5AD6E3DC4048E8B86B4FBF4FE1C" box="[955,1049,404,430]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1635,1659]" captionTargetBox="[297,1291,428,1595]" captionTargetId="figure@5.[280,1306,409,1608]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12 ­ 14), 0.2 mm (15)." httpUri="https://zenodo.org/record/155756/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
); pronotal macrosetae moderately long; pronotal hypomera fully visible in lateral view; mesocoxae separated; medial macroseta of mesotibia as long as tibial width; tarsal formula 4­5­5; metatarsal segment 1 about as long as segment 2; one empodial seta; apical process of median lobe of aedeagus straight or slightly bent ventrally at the very apex (in lateral view;
<figureCitation id="E295C5AD6E3DC40489A68514FB14FDFC" box="[1174,1273,564,590]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="3" pageNumber="4">Figs. 41</figureCitation>
,
<figureCitation id="E295C5AD6E3DC40488368514FACBFDFC" box="[1286,1318,564,590]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="3" pageNumber="4">51</figureCitation>
,
<figureCitation id="E295C5AD6E3DC4048C38857CFEC5FDC4" box="[264,296,604,630]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="3" pageNumber="4">61</figureCitation>
); spermatheca without umbiculus (
<figureCitation id="E295C5AD6E3DC4048FF4857CFCCBFDC4" box="[708,806,604,630]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="3" pageNumber="4">Figs. 39</figureCitation>
,
<figureCitation id="E295C5AD6E3DC4048E02857CFCBFFDC4" box="[818,850,604,630]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="3" pageNumber="4">49</figureCitation>
,
<figureCitation id="E295C5AD6E3DC4048E6E857CFC92FDC4" box="[862,895,604,630]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="3" pageNumber="4">59</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="2ED189A06E3DC4048C388150FC2CF95D" httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="3" pageNumber="4" targetBox="[288,1296,741,1614]" targetPageId="3">
<paragraph id="7A11D9286E3DC4048C388150FC2CF95D" blockId="3.[264,1323,1648,1775]" pageId="3" pageNumber="4">
<emphasis id="48DA053A6E3DC4048C388150FE51F93A" bold="true" box="[264,444,1648,1672]" pageId="3" pageNumber="4">FIGURES 6­11.</emphasis>
Mouthparts of
<taxonomicName id="BDAEA2AB6E3DC4048F5B8150FBE6F935" authority="Gusarov" authorityName="Gusarov" box="[619,1035,1648,1671]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="americanum" status="sp. nov.">
<emphasis id="48DA053A6E3DC4048F5B8150FC4AF935" box="[619,935,1648,1671]" italics="true" pageId="3" pageNumber="4">Tropimenelytron americanum</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="53E9B8416E3DC40489268150FBBFF93A" box="[1046,1106,1648,1672]" pageId="3" pageNumber="4" rank="species">
<emphasis id="48DA053A6E3DC40489268150FBBFF93A" bold="true" box="[1046,1106,1648,1672]" pageId="3" pageNumber="4">sp. n.</emphasis>
</taxonomicNameLabel>
(paratype from Eau Claire, Wisconsin (6­8, 10­11) and
<taxonomicName id="BDAEA2AB6E3DC4048FBF81B3FC2AF918" authority="Eppelsheim" authorityName="Eppelsheim" box="[655,967,1683,1706]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tuberiventre">
<emphasis id="48DA053A6E3DC4048FBF81B3FCC3F918" box="[655,814,1683,1706]" italics="true" pageId="3" pageNumber="4">T. tuberiventre</emphasis>
(Eppelsheim)
</taxonomicName>
(Polikesh, Azerbaijan (9)). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm.
</paragraph>
</caption>
<subSubSection id="32B48AA36E3DC4038C088003FE51FCA4" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="7A11D9286E3DC4048C088003FB7DF8D7" blockId="3.[264,1323,1827,1893]" pageId="3" pageNumber="4">
<taxonomicName id="BDAEA2AB6E3DC4048C088003FE14F88E" ID-CoL="8VSDG" box="[312,505,1827,1852]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3DC4048C088003FE14F88E" box="[312,505,1827,1852]" italics="true" pageId="3" pageNumber="4">Tropimenelytron</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BDAEA2AB6E3DC4048F938003FC3AF88F" ID-CoL="8VRPP" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[675,983,1827,1853]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3DC4048F938003FCE0F88E" box="[675,781,1827,1852]" italics="true" pageId="3" pageNumber="4">Geostiba</emphasis>
<bibRefCitation id="1E3FA4D96E3DC4048E298003FC3AF88F" author="Thomson" box="[793,983,1827,1853]" pageId="3" pageNumber="24" refString="Thomson, C. G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">
Thomson,
<quantity id="BD5674CD6E3DC4048EAC8003FC3AF88F" box="[924,983,1827,1853]" metricMagnitude="1" metricUnit="m" metricValue="4.71932" pageId="3" pageNumber="4" unit="in" value="1858.0">1858</quantity>
</bibRefCitation>
</taxonomicName>
in having ligula with wide base, separated mesocoxae and different shape of aedeagus and spermatheca.
</paragraph>
<paragraph id="7A11D9286E3AC4038C08863CFB64FE64" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
<taxonomicName id="BDAEA2AB6E3AC4038C08863CFE14FE87" ID-CoL="8VSDG" box="[312,505,284,309]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3AC4038C08863CFE14FE87" box="[312,505,284,309]" italics="true" pageId="4" pageNumber="5">Tropimenelytron</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BDAEA2AB6E3AC4038F95863CFBC6FE84" ID-CoL="75ZRX" authority="Kraatz, 1857" authorityName="Kraatz" authorityYear="1857" box="[677,1067,284,310]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="micans">
<emphasis id="48DA053A6E3AC4038F95863CFC6DFE87" box="[677,896,284,309]" italics="true" pageId="4" pageNumber="5">Pelioptera micans</emphasis>
<bibRefCitation id="1E3FA4D96E3AC4038EBD863CFBC6FE84" author="Kraatz" box="[909,1067,284,310]" pageId="4" pageNumber="23" refString="Kraatz, G. (1857) Beitrage zur Kenntniss der Termitophilen. Linnaea Entomologica, 11, 44 - 56." type="journal article" year="1857">Kraatz, 1857</bibRefCitation>
</taxonomicName>
(the
<typeStatus id="A515678A6E3AC4038942863CFB49FE84" box="[1138,1188,284,310]" pageId="4" pageNumber="5">type</typeStatus>
species of
<taxonomicName id="BDAEA2AB6E3AC4038C388664FDC5FEEC" ID-CoL="8VRYC" authority="Kraatz, 1857" authorityName="Kraatz" authorityYear="1857" box="[264,552,324,350]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3AC4038C388664FE69FEEF" box="[264,388,324,349]" italics="true" pageId="4" pageNumber="5">Pelioptera</emphasis>
<bibRefCitation id="1E3FA4D96E3AC4038CBD8664FDC5FEEC" author="Kraatz" box="[397,552,324,350]" pageId="4" pageNumber="23" refString="Kraatz, G. (1857) Beitrage zur Kenntniss der Termitophilen. Linnaea Entomologica, 11, 44 - 56." type="journal article" year="1857">Kraatz, 1857</bibRefCitation>
</taxonomicName>
) in the following characters: first segment of labial palpus with seta absent (present in
<emphasis id="48DA053A6E3AC4038F78864CFD56FE37" box="[584,699,364,389]" italics="true" pageId="4" pageNumber="5">P. m i c a n s</emphasis>
); prementum with three asetose pores (two in
<taxonomicName id="BDAEA2AB6E3AC4038825864CFEB6FE1F" ID-CoL="75ZRX" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="micans">
<emphasis id="48DA053A6E3AC4038825864CFEB6FE1F" italics="true" pageId="4" pageNumber="5">P. micans</emphasis>
</taxonomicName>
), spermatheca without umbiculus; copulatory piece with short apical process (with very long flagellum­like apical process in
<emphasis id="48DA053A6E3AC4038FC7869CFC8EFE67" box="[759,867,444,469]" italics="true" pageId="4" pageNumber="5">P. m i c a n s</emphasis>
) (see discussion below).
</paragraph>
<paragraph id="7A11D9286E3AC4038C0886C4FE4EFDC4" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
<taxonomicName id="BDAEA2AB6E3AC4038C0886C4FE14FE4F" ID-CoL="8VSDG" box="[312,505,484,509]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3AC4038C0886C4FE14FE4F" box="[312,505,484,509]" italics="true" pageId="4" pageNumber="5">Tropimenelytron</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BDAEA2AB6E3AC4038FAC86C4FC16FE4C" ID-CoL="8VRN4" authority="Mulsant &amp; Rey, 1874" authorityName="Mulsant &amp; Rey" authorityYear="1874" box="[668,1019,484,510]" class="Insecta" family="Staphylinidae" genus="Earota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3AC4038FAC86C4FD01FE4F" box="[668,748,484,509]" italics="true" pageId="4" pageNumber="5">Earota</emphasis>
<bibRefCitation id="1E3FA4D96E3AC4038FC686C4FC16FE4C" author="Mulsant" box="[758,1019,484,510]" pageId="4" pageNumber="23" refString="Mulsant, E. &amp; Rey, C. (1874) Tribu des Brevipennes. Famille des Aleochariens. Septieme branche: Myrmedoniaires. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts utiles de Lyon, serie 4. 6 [1873], 33 - 738." type="journal article" year="1874">Mulsant &amp; Rey, 1874</bibRefCitation>
</taxonomicName>
by smaller and narrower body; first segment of labial palpus lacking seta; shorter mesosternal process; incomplete infraorbital carina, less transverse pronotum and different shape of aedeagus and spermatheca.
</paragraph>
<paragraph id="7A11D9286E3AC4038C0885A4FE51FCA4" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
<taxonomicName id="BDAEA2AB6E3AC4038C0885A4FE14FD2F" ID-CoL="8VSDG" box="[312,505,644,669]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3AC4038C0885A4FE14FD2F" box="[312,505,644,669]" italics="true" pageId="4" pageNumber="5">Tropimenelytron</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BDAEA2AB6E3AC4038FA985A4FC5AFD2C" ID-CoL="8VS3L" authority="Ashe, 1986" authorityName="Ashe" authorityYear="1986" box="[665,951,644,670]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E3AC4038FA985A4FCCAFD2F" box="[665,807,644,669]" italics="true" pageId="4" pageNumber="5">Seeversiella</emphasis>
<bibRefCitation id="1E3FA4D96E3AC4038E0085A4FC5AFD2C" author="Ashe" box="[816,951,644,670]" lastPageId="4" lastPageNumber="23" pageId="4" pageNumber="22" refString="Ashe, J. S. (1986) Seeversiella bispinosa, a new genus and species of athetine Aleocharinae (Coleoptera: Staphylinidae) from North America. Journal of the New York Entomological Society, 94 (4), 500 - 511." type="journal article" year="1986">
Ashe,
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</bibRefCitation>
</taxonomicName>
in having pronotal pubescence of
<typeStatus id="A515678A6E3AC4038C19858CFEB1FD74" box="[297,348,684,710]" pageId="4" pageNumber="5">type</typeStatus>
V (
<bibRefCitation id="1E3FA4D96E3AC4038CB9858CFD78FD74" author="Benick" box="[393,661,684,710]" pageId="4" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Krefeld: Goecke &amp; Evers Verlag, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
); ligula with wide base; separated mesocoxae; posterior angles of male tergum 3 not extended as spines, and copulatory piece of aedeagus without long flagellum.
</paragraph>
</subSubSection>
<subSubSection id="32B48AA36E3AC4018C088403FEC2FD5C" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="7A11D9286E3AC4038C088403FE75FC3C" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
<emphasis id="48DA053A6E3AC4038C088403FE3DFC8F" bold="true" box="[312,464,803,829]" pageId="4" pageNumber="5">Description.</emphasis>
Length
<quantity id="BD5674CD6E3AC4038F798404FD34FC8C" box="[585,729,804,830]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.7" metricValueMin="2.5" pageId="4" pageNumber="5" unit="mm" value="3.1" valueMax="3.7" valueMin="2.5">2.5­3.7 mm</quantity>
. Body parallel­sided, from uniformly brownish yellow to reddish brown with dark brown head and abdominal segment 6, and brown appendages.
</paragraph>
<paragraph id="7A11D9286E3AC4038C0884BCFDEBFA0C" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
Head slightly longer than wide; eyes small, temples 1.5­3 times as long as eyes; infraorbital carina short, not reaching posterior margin of eye. Antennal article 2 longer than article 3, articles 4­10 transverse (
<figureCitation id="E295C5AD6E3AC4038FDE84CCFCA1FBB4" box="[750,844,1004,1030]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1635,1659]" captionTargetBox="[297,1291,428,1595]" captionTargetId="figure@5.[280,1306,409,1608]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12 ­ 14), 0.2 mm (15)." httpUri="https://zenodo.org/record/155756/files/figure.png" pageId="4" pageNumber="5">Fig. 14</figureCitation>
). Labrum (
<figureCitation id="E295C5AD6E3AC4038ED484CCFBC0FBB4" box="[996,1069,1004,1030]" captionStart="FIGURES 1 ­ 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[299,1289,735,1765]" captionTargetId="figure@2.[286,1302,724,1777]" captionTargetPageId="2" captionText="FIGURES 1 ­ 5. Mouthparts of Tropimenelytron tuberiventre (Eppelsheim) (Polikesh, Azerbaijan). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155754/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
) transverse, anterior margin concave. Adoral surface of labrum (epipharynx) as in
<figureCitation id="E295C5AD6E3AC4038ED88334FBDDFB9C" box="[1000,1072,1044,1070]" captionStart="FIGURES 1 ­ 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[299,1289,735,1765]" captionTargetId="figure@2.[286,1302,724,1777]" captionTargetPageId="2" captionText="FIGURES 1 ­ 5. Mouthparts of Tropimenelytron tuberiventre (Eppelsheim) (Polikesh, Azerbaijan). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155754/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
. Mandibles (
<figureCitation id="E295C5AD6E3AC40389E08334FEFAFBE4" captionStart="FIGURES 1 ­ 5" captionStartId="2.[264,382,1787,1811]" captionTargetBox="[299,1289,735,1765]" captionTargetId="figure@2.[286,1302,724,1777]" captionTargetPageId="2" captionText="FIGURES 1 ­ 5. Mouthparts of Tropimenelytron tuberiventre (Eppelsheim) (Polikesh, Azerbaijan). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155754/files/figure.png" pageId="4" pageNumber="5">Figs. 3­ 5</figureCitation>
) broad, right mandible with very small medial tooth; dorsal molar area with velvety patch consisting of tiny denticles (visible at 400x). Maxilla (
<figureCitation id="E295C5AD6E3AC40389108344FB7CFBCC" box="[1056,1169,1124,1150]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Figs. 6­8</figureCitation>
) with galea extending slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; apex of lacinia with row of closely spaced spines, middle portion covered with numerous setae. Maxillary palpus with four segments (
<figureCitation id="E295C5AD6E3AC4038EFE83FCFBFBFB44" box="[974,1046,1244,1270]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
). Labium as in
<figureCitation id="E295C5AD6E3AC40389E383FCFECAFAAC" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Figs. 9­ 11</figureCitation>
; labial palpi with three segments (
<figureCitation id="E295C5AD6E3AC4038FF38224FCE0FAAC" box="[707,781,1284,1310]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
), first segment of labial palpus lacking seta (
<figureCitation id="E295C5AD6E3AC4038C21820CFEB7FAF4" box="[273,346,1324,1350]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Fig. 9</figureCitation>
); ligula with wide base, split apically; medial area of prementum without pores or pseudopores, lateral areas with 3 asetose pores and single setose pore. Hypopharyngeal lobes as in
<figureCitation id="E295C5AD6E3AC4038CBE825CFE0BFA24" box="[398,486,1404,1430]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
. Mentum (
<figureCitation id="E295C5AD6E3AC4038F40825CFD28FA24" box="[624,709,1404,1430]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
) with slightly concave anterior margin, medial area with numerous pores.
</paragraph>
<paragraph id="7A11D9286E3AC4028C0882ECFBCEFEEC" blockId="4.[264,1325,284,1870]" lastBlockId="5.[264,1323,284,350]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Pronotum (
<figureCitation id="E295C5AD6E3AC4038C8D82ECFDF9FA54" box="[445,532,1484,1510]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1635,1659]" captionTargetBox="[297,1291,428,1595]" captionTargetId="figure@5.[280,1306,409,1608]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12 ­ 14), 0.2 mm (15)." httpUri="https://zenodo.org/record/155756/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
) subquadrate, broadest at apical third, narrows posteriorly or with subparallel lateral margins; anterior margin straight, posterior margin convex; surface covered with microsetae directed posteriorly in midline, posteriorly and obliquely laterally in lateral areas (
<typeStatus id="A515678A6E3AC4038CF98164FDE8F9EC" box="[457,517,1604,1630]" pageId="4" pageNumber="5">Type</typeStatus>
V,
<bibRefCitation id="1E3FA4D96E3AC4038F018164FCD3F9EC" author="Benick" box="[561,830,1604,1630]" pageId="4" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Krefeld: Goecke &amp; Evers Verlag, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
); macrosetae moderately long; hypomera fully visible in lateral view. Meso­ and metasternum as in
<figureCitation id="E295C5AD6E3AC4038EE6814CFBC3F934" box="[982,1070,1644,1670]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1635,1659]" captionTargetBox="[297,1291,428,1595]" captionTargetId="figure@5.[280,1306,409,1608]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12 ­ 14), 0.2 mm (15)." httpUri="https://zenodo.org/record/155756/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
, mesosternal process short and wide, extended about 1/4 length of mesocoxal cavities, metasternal process about 1/3 length of mesocoxal cavities; mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 3:4:4; mesocoxal cavities margined posteriorly; mesocoxae separated. Medial macroseta of mesotibia as long as tibial width. Tarsal segmentation 4­5­5; metatarsal segment 1 about as long as segment 2 (
<figureCitation id="E295C5AD6E3BC4028FE9863CFCDCFE84" box="[729,817,284,310]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1635,1659]" captionTargetBox="[297,1291,428,1595]" captionTargetId="figure@5.[280,1306,409,1608]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12 ­ 14), 0.2 mm (15)." httpUri="https://zenodo.org/record/155756/files/figure.png" pageId="5" pageNumber="6">Fig. 15</figureCitation>
). One empodial seta present. Wings fully developed (in Nearctic species). Posterior margin of elytra straight.
</paragraph>
<caption id="2ED189A06E3BC4028C388143FC9CF90D" httpUri="https://zenodo.org/record/155756/files/figure.png" pageId="5" pageNumber="6" targetBox="[297,1291,428,1595]" targetPageId="5">
<paragraph id="7A11D9286E3BC4028C388143FC9CF90D" blockId="5.[264,1323,1635,1727]" pageId="5" pageNumber="6">
<emphasis id="48DA053A6E3BC4028C388143FE20F9C9" bold="true" box="[264,461,1635,1659]" pageId="5" pageNumber="6">FIGURES 12­15.</emphasis>
Details of
<taxonomicName id="BDAEA2AB6E3BC4028F688144FC11F9C8" authority="Gusarov" authorityName="Gusarov" box="[600,1020,1635,1659]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="americanum" status="sp. nov.">
<emphasis id="48DA053A6E3BC4028F688144FC7AF9C9" box="[600,919,1636,1659]" italics="true" pageId="5" pageNumber="6">Tropimenelytron americanum</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="53E9B8416E3BC402893B8143FBA7F9C9" box="[1035,1098,1635,1659]" pageId="5" pageNumber="6" rank="species">
<emphasis id="48DA053A6E3BC402893B8143FBA7F9C9" bold="true" box="[1035,1098,1635,1659]" pageId="5" pageNumber="6">sp. n.</emphasis>
</taxonomicNameLabel>
(paratype from Eau Claire, Wisconsin). 12 – pronotum; 13 – mesometathorax, ventral view; 14 – right antenna; 15 – right metatarsus. Scale bar 0.4 mm (12­14), 0.2 mm (15).
</paragraph>
</caption>
<paragraph id="7A11D9286E3BC4028C0881D3FD55F8EF" blockId="5.[264,1321,1779,1885]" pageId="5" pageNumber="6">Abdominal terga 3­5 with moderately transverse basal impression. Tergum 7 ­ 1.1 times as long as tergum 6. Punctation on terga 6­7 slightly sparser than on terga 3­5. Tergum 7 with white palisade fringe.</paragraph>
<paragraph id="7A11D9286E38C4018C08863CFEC2FD5C" blockId="6.[264,1325,284,750]" pageId="6" pageNumber="7">
Male secondary sexual characters include longitudinal carina along anterior half of sutural margin of each elytron, tiny medial knob at posterior margin of tergum 3, small medial knob at apical third of tergum 4, short medial carina in front of anterior margin of tergum 7 and uneven posterior margin of tergum 8 (
<figureCitation id="E295C5AD6E38C4018E4586B4FC23FE1C" box="[885,974,404,430]" captionStart="FIGURES 35 ­ 38" captionStartId="12.[264,383,1248,1272]" captionTargetBox="[325,1262,417,1213]" captionTargetId="figure@12.[318,1269,409,1222]" captionTargetPageId="12" captionText="FIGURES 35 ­ 38. Details of abdominal segment 8 of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Morgan Monroe State Park (35 ­ 36) and Rockville, Indiana (37 ­ 38 )). 35 – male tergum 8; 36 – male sternum 8; 37 – female tergum 8; 38 – female sternum 8. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/155760/files/figure.png" pageId="6" pageNumber="7">Fig. 35</figureCitation>
). Some (smaller) males may lack these features altogether and externally look like females. Median lobe of aedeagus narrows apically (in parameral view;
<figureCitation id="E295C5AD6E38C4018FEC86C4FCA9FE4C" box="[732,836,484,510]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="6" pageNumber="7">Figs. 40</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018E6686C4FC9BFE4C" box="[854,886,484,510]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="6" pageNumber="7">50</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018EB886C4FC44FE4C" box="[904,937,484,510]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="6" pageNumber="7">60</figureCitation>
), apex straight or slightly bent ventrally (in lateral view;
<figureCitation id="E295C5AD6E38C4018F7E852CFD59FD94" box="[590,692,524,550]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="6" pageNumber="7">Figs. 41</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018FF6852CFD0BFD94" box="[710,742,524,550]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="6" pageNumber="7">51</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018FC8852CFCF4FD94" box="[760,793,524,550]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="6" pageNumber="7">61</figureCitation>
). Parameres with two long and two short macrosetae (
<figureCitation id="E295C5AD6E38C4018CAD8514FDEFFDFC" box="[413,514,564,590]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="6" pageNumber="7">Figs. 44</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018F3F8514FDC2FDFC" box="[527,559,564,590]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="6" pageNumber="7">54</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018F0D8514FDB3FDFC" box="[573,606,564,590]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="6" pageNumber="7">64</figureCitation>
). Copulatory piece of internal sac with short apical process and lateral denticles near the base of process (CP;
<figureCitation id="E295C5AD6E38C4018E4F857CFBFAFDC4" box="[895,1047,604,630]" captionStart="FIGURES 16 ­ 21" captionStartId="6.[264,383,1667,1691]" captionTargetBox="[301,1282,802,1631]" captionTargetId="figure@6.[290,1298,793,1649]" captionTargetPageId="6" captionText="FIGURES 16 ­ 21. Male genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rensselaerville, New York (16 ­ 18) and Morgan Monroe State Park, Indiana (21 )) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (19 ­ 20 )). 16 – everted internal sac of aedeagus, lateral view; 17 – medial lamellae, parameral view; 18 – copulatory piece, parameral view; 19 – apex of everted internal sac, lateral view; 20 – apex of everted internal sac, parameral view; 21 – internal sac retracted in median lobe, parameral view. CP ­ copulatory piece; ML ­ medial lamellae. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155757/files/figure.png" pageId="6" pageNumber="7">Figs. 18, 20</figureCitation>
). Medial lamellae (in parameral view) broad (ML;
<figureCitation id="E295C5AD6E38C4018F6D85A4FCF9FD2C" box="[605,788,644,670]" captionStart="FIGURES 16 ­ 21" captionStartId="6.[264,383,1667,1691]" captionTargetBox="[301,1282,802,1631]" captionTargetId="figure@6.[290,1298,793,1649]" captionTargetPageId="6" captionText="FIGURES 16 ­ 21. Male genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rensselaerville, New York (16 ­ 18) and Morgan Monroe State Park, Indiana (21 )) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (19 ­ 20 )). 16 – everted internal sac of aedeagus, lateral view; 17 – medial lamellae, parameral view; 18 – copulatory piece, parameral view; 19 – apex of everted internal sac, lateral view; 20 – apex of everted internal sac, parameral view; 21 – internal sac retracted in median lobe, parameral view. CP ­ copulatory piece; ML ­ medial lamellae. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155757/files/figure.png" pageId="6" pageNumber="7">Figs. 17, 20­21</figureCitation>
). Spermatheca without umbiculus, with thick C­shaped distal portion and thin proximal portion which may form 1­2 coils (
<figureCitation id="E295C5AD6E38C40189A7858CFB17FD74" box="[1175,1274,684,710]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="6" pageNumber="7">Figs. 39</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018836858CFACBFD74" box="[1286,1318,684,710]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="6" pageNumber="7">49</figureCitation>
,
<figureCitation id="E295C5AD6E38C4018C3885F4FECAFD5C" box="[264,295,724,750]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="6" pageNumber="7">59</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="2ED189A06E38C4018C3881A3FE27F8D9" httpUri="https://zenodo.org/record/155757/files/figure.png" pageId="6" pageNumber="7" targetBox="[301,1282,802,1631]" targetPageId="6">
<paragraph id="7A11D9286E38C4018C3881A3FE27F8D9" blockId="6.[264,1325,1667,1899]" pageId="6" pageNumber="7">
<emphasis id="48DA053A6E38C4018C3881A3FE29F929" bold="true" box="[264,452,1667,1691]" pageId="6" pageNumber="7">FIGURES 16­21</emphasis>
. Male genitalia of
<taxonomicName id="BDAEA2AB6E38C4018FA581A4FBDEF929" authority="Gusarov" authorityName="Gusarov" box="[661,1075,1668,1691]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="americanum" status="sp. nov.">
<emphasis id="48DA053A6E38C4018FA581A4FC3CF929" box="[661,977,1668,1691]" italics="true" pageId="6" pageNumber="7">Tropimenelytron americanum</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="53E9B8416E38C401890E81A3FB97F929" box="[1086,1146,1667,1691]" pageId="6" pageNumber="7" rank="species">
<emphasis id="48DA053A6E38C401890E81A3FB97F929" bold="true" box="[1086,1146,1667,1691]" pageId="6" pageNumber="7">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Rensselaerville, New York (16­18) and Morgan Monroe State Park, Indiana (21)) and
<taxonomicName id="BDAEA2AB6E38C40188278186FDC9F952" authority="Eppelsheim" authorityName="Eppelsheim" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tuberiventre">
<emphasis id="48DA053A6E38C40188278186FAC9F90F" box="[1303,1316,1702,1725]" italics="true" pageId="6" pageNumber="7">T</emphasis>
.
<emphasis id="48DA053A6E38C4018C3881E9FE64F952" box="[264,393,1737,1760]" italics="true" pageId="6" pageNumber="7">tuberiventre</emphasis>
(Eppelsheim)
</taxonomicName>
(Polikesh, Azerbaijan (19­20)). 16 – everted internal sac of aedeagus, lateral view; 17 – medial lamellae, parameral view; 18 – copulatory piece, parameral view; 19 – apex of everted internal sac, lateral view; 20 – apex of everted internal sac, parameral view; 21 – internal sac retracted in median lobe, parameral view. CP ­ copulatory piece; ML ­ medial lamellae. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption id="2ED189A06E39C4008C3882E0FED6F9D0" httpUri="https://zenodo.org/record/155758/files/figure.png" pageId="7" pageNumber="8" targetBox="[285,1312,288,1432]" targetPageId="7">
<paragraph id="7A11D9286E39C4008C3882E0FED6F9D0" blockId="7.[264,1323,1472,1634]" pageId="7" pageNumber="8">
<emphasis id="48DA053A6E39C4008C3882E0FE21FA6A" bold="true" box="[264,460,1472,1496]" pageId="7" pageNumber="8">FIGURES 22­26.</emphasis>
Details of
<taxonomicName id="BDAEA2AB6E39C4008F6282E1FBEAFA65" authority="Motschulsky" authorityName="Motschulsky" box="[594,1031,1472,1496]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="testaceipennis">
<emphasis id="48DA053A6E39C4008F6282E1FC8EFA6A" box="[594,867,1473,1496]" italics="true" pageId="7" pageNumber="8">Pelioptera testaceipennis</emphasis>
(Motschulsky)
</taxonomicName>
(syntype of
<taxonomicName id="BDAEA2AB6E39C40089A582E1FE87FA48" authority="Cameron" authorityName="Cameron" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="48DA053A6E39C40089A582E1FAC7FA6A" box="[1173,1322,1473,1496]" italics="true" pageId="7" pageNumber="8">P. longicornis</emphasis>
Cameron
</taxonomicName>
from Sumatra, Indonesia (22, 25)),
<emphasis id="48DA053A6E39C4008E2D82C3FC6AFA48" box="[797,903,1507,1530]" italics="true" pageId="7" pageNumber="8">P. m i c a n s</emphasis>
Kraatz (Sri Lanka (23­24)) and
<taxonomicName id="BDAEA2AB6E39C400882782C3FE08F9AF" authority="Kraatz" authorityName="Kraatz" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="exasperata">
<emphasis id="48DA053A6E39C400882782C3FE90F9AF" italics="true" pageId="7" pageNumber="8">P. exasperata</emphasis>
(Kraatz)
</taxonomicName>
(Dehra Dun, India (26)). 22­23 – prementum, ventral view; 24­26 – mesometathorax, ventral view. Scale bar 0.15 mm (23), 0.2 mm (22), 0.4 mm (24, 26), 0.8 mm (25).
</paragraph>
</caption>
<subSubSection id="32B48AA36E39C40F8C0881B5FC6BFEEC" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="7A11D9286E39C4008C0881B5FE7AF96A" blockId="7.[264,1324,1685,1872]" pageId="7" pageNumber="8">
<emphasis id="48DA053A6E39C4008C0881B5FE08F91D" bold="true" box="[312,485,1685,1711]" pageId="7" pageNumber="8">
<typeStatus id="A515678A6E39C4008C0881B5FE98F91D" box="[312,373,1685,1711]" pageId="7" pageNumber="8">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="BDAEA2AB6E39C4008F3A81B6FC54F902" authority="Eppelsheim" authorityName="Eppelsheim" box="[522,953,1686,1712]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tuberiventris">
<emphasis id="48DA053A6E39C4008F3A81B6FCF0F91D" box="[522,797,1686,1711]" italics="true" pageId="7" pageNumber="8">Geostiba tuberiventris</emphasis>
Eppelsheim
</taxonomicName>
<emphasis id="48DA053A6E39C4008EFC81B6FC0EF91D" box="[972,995,1686,1711]" italics="true" pageId="7" pageNumber="8">in</emphasis>
<bibRefCitation id="1E3FA4D96E39C4008EC581B6FB60F902" author="Leder" box="[1013,1165,1686,1712]" pageId="7" pageNumber="23" refString="Leder, H. (1879) Beitrag zur kaukasischen Kafer-Fauna. Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft in Wien, 29, 451 - 488." type="journal article" year="1879">Leder, 1879</bibRefCitation>
by original designation.
</paragraph>
<paragraph id="7A11D9286E39C40F8C0881C5FC6BFEEC" blockId="7.[264,1324,1685,1872]" lastBlockId="8.[264,1324,284,1870]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="48DA053A6E39C4008C0881C5FE76F94D" bold="true" box="[312,411,1765,1791]" pageId="7" pageNumber="8">Gender.</emphasis>
The name
<taxonomicName id="BDAEA2AB6E39C4008F1781C6FD04F94D" box="[551,745,1766,1791]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E39C4008F1781C6FD04F94D" box="[551,745,1766,1791]" italics="true" pageId="7" pageNumber="8">Tropimenelytron</emphasis>
</taxonomicName>
ends in a Greek word “ ” transliterated into Latin without other changes and therefore takes the neutral gender (Article 30.1.2 of the Code;
<bibRefCitation id="1E3FA4D96E39C4008CB68016FDF9F8E2" author="ICZN" box="[390,532,1846,1872]" pageId="7" pageNumber="23" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. London: The International Trust for Zoological Nomenclature. xxix + 306 pp." type="book" year="1999">
<collectionCode id="1CBF41ED6E39C4008CB68016FE27F8E2" box="[390,458,1846,1872]" pageId="7" pageNumber="8">ICZN</collectionCode>
1999
</bibRefCitation>
). Because a species name must agree in gender with the generic name with which it is combined (Article 31.2.1) when
<taxonomicName id="BDAEA2AB6E36C40F8EF8863CFB64FE87" box="[968,1161,284,309]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tuberiventris">
<emphasis id="48DA053A6E36C40F8EF8863CFB64FE87" box="[968,1161,284,309]" italics="true" pageId="8" pageNumber="9">G. tuberiventris</emphasis>
</taxonomicName>
is placed in
<taxonomicName id="BDAEA2AB6E36C40F8C388664FE24FEEF" box="[264,457,324,349]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C388664FE24FEEF" box="[264,457,324,349]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
the correct spelling is
<taxonomicName id="BDAEA2AB6E36C40F8FE48664FC93FEEF" box="[724,894,324,349]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tuberiventre">
<emphasis id="48DA053A6E36C40F8FE48664FC93FEEF" box="[724,894,324,349]" italics="true" pageId="8" pageNumber="9">T. tuberiventre</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="32B48AA36E36C40F8C08864BFD22FDFC" pageId="8" pageNumber="9" type="discussion">
<paragraph id="7A11D9286E36C40F8C08864BFD22FDFC" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
<emphasis id="48DA053A6E36C40F8C08864BFE51FE37" bold="true" box="[312,444,363,389]" pageId="8" pageNumber="9">Discussion</emphasis>
. Although originally described as a separate genus (
<bibRefCitation id="1E3FA4D96E36C40F89A8864CFACCFE34" author="Pace" box="[1176,1313,364,390]" pageId="8" pageNumber="23" refString="Pace, R. (1983) Il genere Tropimenelytron Scheerpeltz (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie, 13 (2), 185 - 190." type="journal article" year="1983">Pace 1983</bibRefCitation>
)
<taxonomicName id="BDAEA2AB6E36C40F8C3886B4FE24FE1F" box="[264,457,404,429]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C3886B4FE24FE1F" box="[264,457,404,429]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
was subsequently (
<bibRefCitation id="1E3FA4D96E36C40F8F9F86B4FCC1FE1C" author="Pace" box="[687,812,404,430]" pageId="8" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
) downgraded to subgeneric rank within the genus
<taxonomicName id="BDAEA2AB6E36C40F8C67869CFE3EFE67" box="[343,467,444,469]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C67869CFE3EFE67" box="[343,467,444,469]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
.
<bibRefCitation id="1E3FA4D96E36C40F8CC0869CFD6CFE64" author="Pace" box="[496,641,444,470]" pageId="8" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace (1991)</bibRefCitation>
argued that
<emphasis id="48DA053A6E36C40F8E2B869CFB87FE67" box="[795,1130,444,469]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="BDAEA2AB6E36C40F8E2B869CFC77FE67" box="[795,922,444,469]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Pelioptera</taxonomicName>
,
<taxonomicName id="BDAEA2AB6E36C40F8E99869CFB87FE67" box="[937,1130,444,469]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Tropimenelytron</taxonomicName>
</emphasis>
and
<taxonomicName id="BDAEA2AB6E36C40F899A869CFAC6FE67" box="[1194,1323,444,469]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F899A869CFAC6FE67" box="[1194,1323,444,469]" italics="true" pageId="8" pageNumber="9">Geostibida</emphasis>
</taxonomicName>
have “almost identical” ligula, but in
<taxonomicName id="BDAEA2AB6E36C40F8FE286C4FCA3FE4F" box="[722,846,484,509]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8FE286C4FCA3FE4F" box="[722,846,484,509]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
mesocoxae are more widely separated. Pace considered this similarity in ligula sufficient to include
<taxonomicName id="BDAEA2AB6E36C40F891D852CFB03FD97" box="[1069,1262,524,549]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F891D852CFB03FD97" box="[1069,1262,524,549]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
and
<taxonomicName id="BDAEA2AB6E36C40F8C388514FE64FDFF" box="[264,393,564,589]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C388514FE64FDFF" box="[264,393,564,589]" italics="true" pageId="8" pageNumber="9">Geostibida</emphasis>
</taxonomicName>
in
<taxonomicName id="BDAEA2AB6E36C40F8C9E8514FDC7FDFF" box="[430,554,564,589]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C9E8514FDC7FDFF" box="[430,554,564,589]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
as subgenera.
</paragraph>
</subSubSection>
<subSubSection id="32B48AA36E36C40F8C08857CFE6CFA24" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="7A11D9286E36C40F8C08857CFB42FBE4" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
<taxonomicName id="BDAEA2AB6E36C40F8C08857CFD43FDC4" ID-CoL="75ZRX" authority="Kraatz, 1857" authorityName="Kraatz" authorityYear="1857" box="[312,686,604,630]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="micans">
<emphasis id="48DA053A6E36C40F8C08857CFDE3FDC7" box="[312,526,604,629]" italics="true" pageId="8" pageNumber="9">Pelioptera micans</emphasis>
<bibRefCitation id="1E3FA4D96E36C40F8F26857CFD43FDC4" author="Kraatz" box="[534,686,604,630]" pageId="8" pageNumber="23" refString="Kraatz, G. (1857) Beitrage zur Kenntniss der Termitophilen. Linnaea Entomologica, 11, 44 - 56." type="journal article" year="1857">Kraatz, 1857</bibRefCitation>
</taxonomicName>
(
<figureCitation id="E295C5AD6E36C40F8F8E857CFCA5FDC4" box="[702,840,604,630]" captionStart="FIGURES 22 ­ 26" captionStartId="7.[264,383,1472,1496]" captionTargetBox="[285,1312,288,1432]" captionTargetId="figure@7.[280,1323,284,1440]" captionTargetPageId="7" captionText="FIGURES 22 ­ 26. Details of Pelioptera testaceipennis (Motschulsky) (syntype of P. longicornis Cameron from Sumatra, Indonesia (22, 25 )), P. m i c a n s Kraatz (Sri Lanka (23 ­ 24 )) and P. exasperata (Kraatz) (Dehra Dun, India (26 )). 22 ­ 23 – prementum, ventral view; 24 ­ 26 – mesometathorax, ventral view. Scale bar 0.15 mm (23), 0.2 mm (22), 0.4 mm (24, 26), 0.8 mm (25)." httpUri="https://zenodo.org/record/155758/files/figure.png" pageId="8" pageNumber="9">Figs. 23­24</figureCitation>
,
<figureCitation id="E295C5AD6E36C40F8E65857CFC70FDC4" box="[853,925,604,630]" captionStart="FIGURES 27 ­ 34" captionStartId="9.[264,383,1746,1770]" captionTargetBox="[316,1284,476,1734]" captionTargetId="figure@9.[305,1296,473,1736]" captionTargetPageId="9" captionText="FIGURES 27 ­ 34. Male genitalia of Pelioptera micans Kraatz (syntype from Sri Lanka (27 ­ 29 )), P. exasperata (Kraatz) (Dehra Dun, India (30 ­ 32 )) and P. testaceipennis (Motschulsky) (Java, Indonesia (33 ­ 34 )). 27, 30, 33 – median lobe of aedeagus, parameral view; 28 – apex of median lobe, parameral view; 29 – copulatory piece of internal sac, lateral view; 31, 34 – median lobe of aedeagus, lateral view; 32 – copulatory piece of internal sac, parameral view. Scale bar 0.2 mm (28 ­ 29, 32), 0.4 mm (27, 30 ­ 31, 33 ­ 34)." httpUri="https://zenodo.org/record/155759/files/figure.png" pageId="8" pageNumber="9">27­29</figureCitation>
;
<figureCitation id="E295C5AD6E36C40F8E99857CFC03FDC4" box="[937,1006,604,630]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="8" pageNumber="9">
Fig.
<quantity id="BD5674CD6E36C40F8ED0857CFC03FDC4" box="[992,1006,604,630]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="9" unit="in" value="9.0">9</quantity>
</figureCitation>
in
<bibRefCitation id="1E3FA4D96E36C40F8925857CFB5BFDC4" author="Sawada" box="[1045,1206,604,630]" pageId="8" pageNumber="24" refString="Sawada, K. (1980) Atheta and Its Allies of Southeast Asia (Coleoptera; Staphylinidae). II. Reexamination of the Species Mainly from Borneo. Contributions from the Biological Laboratory, Kyoto University, 26, 23 - 66." type="journal article" year="1980">Sawada 1980</bibRefCitation>
), the
<typeStatus id="A515678A6E36C40F89C7857CFAC4FDC4" box="[1271,1321,604,630]" pageId="8" pageNumber="9">type</typeStatus>
species of
<taxonomicName id="BDAEA2AB6E36C40F8CB885A4FDE9FD2F" ID-CoL="8VRYC" box="[392,516,644,669]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8CB885A4FDE9FD2F" box="[392,516,644,669]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
, differs from
<taxonomicName id="BDAEA2AB6E36C40F8F9E85A4FCB6FD2F" ID-CoL="59CR2" box="[686,859,644,669]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tuberiventre">
<emphasis id="48DA053A6E36C40F8F9E85A4FCB6FD2F" box="[686,859,644,669]" italics="true" pageId="8" pageNumber="9">T. tuberiventre</emphasis>
</taxonomicName>
and
<taxonomicName id="BDAEA2AB6E36C40F8EA885A4FB55FD2C" ID-CoL="59CPY" authority="Gusarov" authorityName="Gusarov" box="[920,1208,644,670]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="americanum" status="sp. nov.">
<emphasis id="48DA053A6E36C40F8EA885A4FBA7FD2F" box="[920,1098,644,669]" italics="true" pageId="8" pageNumber="9">T. americanum</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="53E9B8416E36C40F89F685A3FAE7FD2F" box="[1222,1290,643,669]" pageId="8" pageNumber="9" rank="species">
<emphasis id="48DA053A6E36C40F89F685A3FAE7FD2F" bold="true" box="[1222,1290,643,669]" pageId="8" pageNumber="9">sp. n.</emphasis>
</taxonomicNameLabel>
in the following characters: first segment of labial palpus with seta present (
<figureCitation id="E295C5AD6E36C40F89F7858CFACFFD74" box="[1223,1314,684,710]" captionStart="FIGURES 22 ­ 26" captionStartId="7.[264,383,1472,1496]" captionTargetBox="[285,1312,288,1432]" captionTargetId="figure@7.[280,1323,284,1440]" captionTargetPageId="7" captionText="FIGURES 22 ­ 26. Details of Pelioptera testaceipennis (Motschulsky) (syntype of P. longicornis Cameron from Sumatra, Indonesia (22, 25 )), P. m i c a n s Kraatz (Sri Lanka (23 ­ 24 )) and P. exasperata (Kraatz) (Dehra Dun, India (26 )). 22 ­ 23 – prementum, ventral view; 24 ­ 26 – mesometathorax, ventral view. Scale bar 0.15 mm (23), 0.2 mm (22), 0.4 mm (24, 26), 0.8 mm (25)." httpUri="https://zenodo.org/record/155758/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
) (absent in
<taxonomicName id="BDAEA2AB6E36C40F8CB585F4FDABFD5F" ID-CoL="8VSDG" box="[389,582,724,749]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8CB585F4FDABFD5F" box="[389,582,724,749]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
:
<figureCitation id="E295C5AD6E36C40F8F5085F4FD4AFD5C" box="[608,679,724,750]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
) and equidistant from setae b and; prementum with two asetose pores (
<figureCitation id="E295C5AD6E36C40F8F3385DCFD8FFCA4" box="[515,610,764,790]" captionStart="FIGURES 22 ­ 26" captionStartId="7.[264,383,1472,1496]" captionTargetBox="[285,1312,288,1432]" captionTargetId="figure@7.[280,1323,284,1440]" captionTargetPageId="7" captionText="FIGURES 22 ­ 26. Details of Pelioptera testaceipennis (Motschulsky) (syntype of P. longicornis Cameron from Sumatra, Indonesia (22, 25 )), P. m i c a n s Kraatz (Sri Lanka (23 ­ 24 )) and P. exasperata (Kraatz) (Dehra Dun, India (26 )). 22 ­ 23 – prementum, ventral view; 24 ­ 26 – mesometathorax, ventral view. Scale bar 0.15 mm (23), 0.2 mm (22), 0.4 mm (24, 26), 0.8 mm (25)." httpUri="https://zenodo.org/record/155758/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
) (three in
<taxonomicName id="BDAEA2AB6E36C40F8FC885DCFC54FCA7" ID-CoL="8VSDG" box="[760,953,764,789]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8FC885DCFC54FCA7" box="[760,953,764,789]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
:
<figureCitation id="E295C5AD6E36C40F8ED385DCFBD9FCA4" box="[995,1076,764,790]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
), spermatheca with umbiculus (
<figureCitation id="E295C5AD6E36C40F8CA48404FE31FC8C" box="[404,476,804,830]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
, O in
<bibRefCitation id="1E3FA4D96E36C40F8F198404FD24FC8C" author="Sawada" box="[553,713,804,830]" pageId="8" pageNumber="24" refString="Sawada, K. (1980) Atheta and Its Allies of Southeast Asia (Coleoptera; Staphylinidae). II. Reexamination of the Species Mainly from Borneo. Contributions from the Biological Laboratory, Kyoto University, 26, 23 - 66." type="journal article" year="1980">Sawada 1980</bibRefCitation>
) (without umbiculus in
<taxonomicName id="BDAEA2AB6E36C40F8ED88404FB44FC8F" ID-CoL="8VSDG" box="[1000,1193,804,829]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8ED88404FB44FC8F" box="[1000,1193,804,829]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
:
<figureCitation id="E295C5AD6E36C40F89F38404FACBFC8C" box="[1219,1318,804,830]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="8" pageNumber="9">Figs. 39</figureCitation>
,
<figureCitation id="E295C5AD6E36C40F8C38846CFEC5FCD4" box="[264,296,844,870]" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="8" pageNumber="9">49</figureCitation>
,
<figureCitation id="E295C5AD6E36C40F8C09846CFEB7FCD4" box="[313,346,844,870]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="8" pageNumber="9">59</figureCitation>
), copulatory piece with very long apical process (flagellum­like;
<figureCitation id="E295C5AD6E36C40F894F846CFB34FCD4" box="[1151,1241,844,870]" captionStart="FIGURES 27 ­ 34" captionStartId="9.[264,383,1746,1770]" captionTargetBox="[316,1284,476,1734]" captionTargetId="figure@9.[305,1296,473,1736]" captionTargetPageId="9" captionText="FIGURES 27 ­ 34. Male genitalia of Pelioptera micans Kraatz (syntype from Sri Lanka (27 ­ 29 )), P. exasperata (Kraatz) (Dehra Dun, India (30 ­ 32 )) and P. testaceipennis (Motschulsky) (Java, Indonesia (33 ­ 34 )). 27, 30, 33 – median lobe of aedeagus, parameral view; 28 – apex of median lobe, parameral view; 29 – copulatory piece of internal sac, lateral view; 31, 34 – median lobe of aedeagus, lateral view; 32 – copulatory piece of internal sac, parameral view. Scale bar 0.2 mm (28 ­ 29, 32), 0.4 mm (27, 30 ­ 31, 33 ­ 34)." httpUri="https://zenodo.org/record/155759/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
) (with short apical process in
<taxonomicName id="BDAEA2AB6E36C40F8F168454FD0AFC3F" ID-CoL="8VSDG" box="[550,743,884,909]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8F168454FD0AFC3F" box="[550,743,884,909]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
:
<figureCitation id="E295C5AD6E36C40F8E378454FC1BFC3C" box="[775,1014,884,910]" captionStart="FIGURES 16 ­ 21" captionStartId="6.[264,383,1667,1691]" captionTargetBox="[301,1282,802,1631]" captionTargetId="figure@6.[290,1298,793,1649]" captionTargetPageId="6" captionText="FIGURES 16 ­ 21. Male genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rensselaerville, New York (16 ­ 18) and Morgan Monroe State Park, Indiana (21 )) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (19 ­ 20 )). 16 – everted internal sac of aedeagus, lateral view; 17 – medial lamellae, parameral view; 18 – copulatory piece, parameral view; 19 – apex of everted internal sac, lateral view; 20 – apex of everted internal sac, parameral view; 21 – internal sac retracted in median lobe, parameral view. CP ­ copulatory piece; ML ­ medial lamellae. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155757/files/figure.png" pageId="8" pageNumber="9">Figs. 16, 18, 19­20</figureCitation>
). Considering the above differences between the examined species of
<taxonomicName id="BDAEA2AB6E36C40F8E1384BCFC09FC07" ID-CoL="8VSDG" box="[803,996,924,949]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8E1384BCFC09FC07" box="[803,996,924,949]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
(including the
<typeStatus id="A515678A6E36C40F89AB84BCFB23FC04" box="[1179,1230,924,950]" pageId="8" pageNumber="9">type</typeStatus>
species of the genus) and the
<typeStatus id="A515678A6E36C40F8F3384E4FDD8FC6C" box="[515,565,964,990]" pageId="8" pageNumber="9">type</typeStatus>
species of
<taxonomicName id="BDAEA2AB6E36C40F8F8784E4FCDEFC6F" ID-CoL="8VRYC" box="[695,819,964,989]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8F8784E4FCDEFC6F" box="[695,819,964,989]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
I do not regard their similarity in the shape of ligula to be sufficient to include
<taxonomicName id="BDAEA2AB6E36C40F8F9484CCFC88FBB7" ID-CoL="8VSDG" box="[676,869,1004,1029]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8F9484CCFC88FBB7" box="[676,869,1004,1029]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
in
<taxonomicName id="BDAEA2AB6E36C40F8EBB84CCFBEAFBB7" ID-CoL="8VRYC" box="[907,1031,1004,1029]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8EBB84CCFBEAFBB7" box="[907,1031,1004,1029]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
. In this paper I consider
<taxonomicName id="BDAEA2AB6E36C40F8C388334FE24FB9F" ID-CoL="8VSDG" box="[264,457,1044,1069]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C388334FE24FB9F" box="[264,457,1044,1069]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
to represent a genus separate from
<taxonomicName id="BDAEA2AB6E36C40F8E9D8334FBC4FB9F" ID-CoL="8VRYC" box="[941,1065,1044,1069]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8E9D8334FBC4FB9F" box="[941,1065,1044,1069]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
, pending a detailed examination of related genera and subgenera and analysis of their relationships.
</paragraph>
<paragraph id="7A11D9286E36C40F8C088344FE6CFA24" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
My examination of three species of
<taxonomicName id="BDAEA2AB6E36C40F8FDF8344FC86FBCF" ID-CoL="8VRYC" box="[751,875,1124,1149]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8FDF8344FC86FBCF" box="[751,875,1124,1149]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
(
<figureCitation id="E295C5AD6E36C40F8E4F8344FBE7FBCC" box="[895,1034,1124,1150]" captionStart-0="FIGURES 22 ­ 26" captionStart-1="FIGURES 27 ­ 34" captionStartId-0="7.[264,383,1472,1496]" captionStartId-1="9.[264,383,1746,1770]" captionTargetBox-0="[285,1312,288,1432]" captionTargetBox-1="[316,1284,476,1734]" captionTargetId-0="figure@7.[280,1323,284,1440]" captionTargetId-1="figure@9.[305,1296,473,1736]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURES 22 ­ 26. Details of Pelioptera testaceipennis (Motschulsky) (syntype of P. longicornis Cameron from Sumatra, Indonesia (22, 25 )), P. m i c a n s Kraatz (Sri Lanka (23 ­ 24 )) and P. exasperata (Kraatz) (Dehra Dun, India (26 )). 22 ­ 23 – prementum, ventral view; 24 ­ 26 – mesometathorax, ventral view. Scale bar 0.15 mm (23), 0.2 mm (22), 0.4 mm (24, 26), 0.8 mm (25)." captionText-1="FIGURES 27 ­ 34. Male genitalia of Pelioptera micans Kraatz (syntype from Sri Lanka (27 ­ 29 )), P. exasperata (Kraatz) (Dehra Dun, India (30 ­ 32 )) and P. testaceipennis (Motschulsky) (Java, Indonesia (33 ­ 34 )). 27, 30, 33 – median lobe of aedeagus, parameral view; 28 – apex of median lobe, parameral view; 29 – copulatory piece of internal sac, lateral view; 31, 34 – median lobe of aedeagus, lateral view; 32 – copulatory piece of internal sac, parameral view. Scale bar 0.2 mm (28 ­ 29, 32), 0.4 mm (27, 30 ­ 31, 33 ­ 34)." httpUri-0="https://zenodo.org/record/155758/files/figure.png" httpUri-1="https://zenodo.org/record/155759/files/figure.png" pageId="8" pageNumber="9">Figs. 22­34</figureCitation>
) and comparison of the drawings and descriptions published by
<bibRefCitation id="1E3FA4D96E36C40F8E2A83ACFC3DFB14" author="Sawada" box="[794,976,1164,1190]" pageId="8" pageNumber="24" refString="Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera; Staphylinidae). III. Japanese Species described by the previous Authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222." type="journal article" year="1977">Sawada (1977</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E36C40F8ED783ACFBC9FB14" author="Sawada" box="[999,1060,1164,1190]" pageId="8" pageNumber="24" refString="Sawada, K. (1980) Atheta and Its Allies of Southeast Asia (Coleoptera; Staphylinidae). II. Reexamination of the Species Mainly from Borneo. Contributions from the Biological Laboratory, Kyoto University, 26, 23 - 66." type="journal article" year="1980">1980</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E36C40F890C83ACFB94FB14" author="Sawada" box="[1084,1145,1164,1190]" pageId="8" pageNumber="24" refString="Sawada, K. (1982) Atheta and Its Allies of Southeast Asia (Coleoptera; Staphylinidae). III. Oriental Species Described by V. von Motschulsky and G. Kraatz. Contributions from the Biological Laboratory, Kyoto University, 26 (2), 141 - 187." type="journal article" year="1982">1982</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E36C40F89A283ACFB22FB14" author="Sawada" box="[1170,1231,1164,1190]" pageId="8" pageNumber="24" refString="Sawada, K. (1987) Atheta and Its Allies of Southeast Asia (Coleoptera: Staphylinidae). V. Singaporean Species Described in Cameron, 1920. Contributions from the Biological Laboratory, Kyoto University, 27 (2), 137 - 150." type="journal article" year="1987">1987</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E36C40F89D783ACFACEFB14" author="Sawada" box="[1255,1315,1164,1190]" pageId="8" pageNumber="24" refString="Sawada, K. (1989) New Species of Aleocharinae from Japan, I (Staphylinidae; Coleoptera). Contributions from the Biological Laboratory, Kyoto University, 27 (3), 273 - 307." type="journal article" year="1989">1989</bibRefCitation>
) demonstrate that the species currently assigned to
<taxonomicName id="BDAEA2AB6E36C40F8E448394FC1DFB7F" ID-CoL="8VRYC" box="[884,1008,1204,1229]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8E448394FC1DFB7F" box="[884,1008,1204,1229]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
often have very different structures of internal sac (cf.
<figureCitation id="E295C5AD6E36C40F8F6F83FCFCFBFB44" box="[607,790,1244,1270]" captionStart="FIGURES 27 ­ 34" captionStartId="9.[264,383,1746,1770]" captionTargetBox="[316,1284,476,1734]" captionTargetId="figure@9.[305,1296,473,1736]" captionTargetPageId="9" captionText="FIGURES 27 ­ 34. Male genitalia of Pelioptera micans Kraatz (syntype from Sri Lanka (27 ­ 29 )), P. exasperata (Kraatz) (Dehra Dun, India (30 ­ 32 )) and P. testaceipennis (Motschulsky) (Java, Indonesia (33 ­ 34 )). 27, 30, 33 – median lobe of aedeagus, parameral view; 28 – apex of median lobe, parameral view; 29 – copulatory piece of internal sac, lateral view; 31, 34 – median lobe of aedeagus, lateral view; 32 – copulatory piece of internal sac, parameral view. Scale bar 0.2 mm (28 ­ 29, 32), 0.4 mm (27, 30 ­ 31, 33 ­ 34)." httpUri="https://zenodo.org/record/155759/files/figure.png" pageId="8" pageNumber="9">Figs. 29, 32, 34</figureCitation>
and
<figureCitation id="E295C5AD6E36C40F8E6083FCFC5CFB44" box="[848,945,1244,1270]" captionStart="FIGURES 6 ­ 11" captionStartId="3.[264,383,1648,1672]" captionTargetBox="[288,1296,741,1614]" captionTargetId="figure@3.[278,1309,729,1621]" captionTargetPageId="3" captionText="FIGURES 6 ­ 11. Mouthparts of Tropimenelytron americanum Gusarov, sp. n. (paratype from Eau Claire, Wisconsin (6 ­ 8, 10 ­ 11) and T. tuberiventre (Eppelsheim) (Polikesh, Azerbaijan (9 )). 6 – right maxilla, ventral view; 7 – right lacinia, dorsal view; 8 – right galea, dorsal view; 9 – prementum; 10 – hypopharynx; 11 – mentum. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155755/files/figure.png" pageId="8" pageNumber="9">Figs. 10</figureCitation>
, K; 11, I; 12, J in
<bibRefCitation id="1E3FA4D96E36C40F89B483FCFACEFB44" author="Sawada" box="[1156,1315,1244,1270]" pageId="8" pageNumber="24" refString="Sawada, K. (1980) Atheta and Its Allies of Southeast Asia (Coleoptera; Staphylinidae). II. Reexamination of the Species Mainly from Borneo. Contributions from the Biological Laboratory, Kyoto University, 26, 23 - 66." type="journal article" year="1980">Sawada 1980</bibRefCitation>
) and mesometathorax (
<figureCitation id="E295C5AD6E36C40F8F2D8224FD5CFAAC" box="[541,689,1284,1310]" captionStart="FIGURES 22 ­ 26" captionStartId="7.[264,383,1472,1496]" captionTargetBox="[285,1312,288,1432]" captionTargetId="figure@7.[280,1323,284,1440]" captionTargetPageId="7" captionText="FIGURES 22 ­ 26. Details of Pelioptera testaceipennis (Motschulsky) (syntype of P. longicornis Cameron from Sumatra, Indonesia (22, 25 )), P. m i c a n s Kraatz (Sri Lanka (23 ­ 24 )) and P. exasperata (Kraatz) (Dehra Dun, India (26 )). 22 ­ 23 – prementum, ventral view; 24 ­ 26 – mesometathorax, ventral view. Scale bar 0.15 mm (23), 0.2 mm (22), 0.4 mm (24, 26), 0.8 mm (25)." httpUri="https://zenodo.org/record/155758/files/figure.png" pageId="8" pageNumber="9">Figs. 24­26</figureCitation>
). This may indicate that some species currently assigned to
<taxonomicName id="BDAEA2AB6E36C40F8CA5820CFDFCFAF7" ID-CoL="8VRYC" box="[405,529,1324,1349]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8CA5820CFDFCFAF7" box="[405,529,1324,1349]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
</taxonomicName>
are not related and should not be congeneric with the
<typeStatus id="A515678A6E36C40F89AB820CFB20FAF4" box="[1179,1229,1324,1350]" pageId="8" pageNumber="9">type</typeStatus>
species of the genus. This problem requires further study and it is briefly discussed at the end of this paper.
</paragraph>
</subSubSection>
<subSubSection id="32B48AA36E36C40D8C088284FAE0FB14" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="7A11D9286E36C40E8C088284FBD4FE1C" blockId="8.[264,1324,284,1870]" lastBlockId="9.[264,1324,284,430]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
In his key to subgenera of
<taxonomicName id="BDAEA2AB6E36C40F8FBF8284FC5CFA0C" authority="Pace (1991)" authorityName="Pace" authorityYear="1991" box="[655,945,1444,1470]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8FBF8284FCE6FA0F" box="[655,779,1444,1469]" italics="true" pageId="8" pageNumber="9">Pelioptera</emphasis>
,
<bibRefCitation id="1E3FA4D96E36C40F8E2F8284FC5CFA0C" author="Pace" box="[799,945,1444,1470]" pageId="8" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace (1991)</bibRefCitation>
</taxonomicName>
listed two characters to allow separation between
<taxonomicName id="BDAEA2AB6E36C40F8F3082ECFD2CFA57" box="[512,705,1484,1509]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8F3082ECFD2CFA57" box="[512,705,1484,1509]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
and
<taxonomicName id="BDAEA2AB6E36C40F8E3982ECFC67FA57" box="[777,906,1484,1509]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8E3982ECFC67FA57" box="[777,906,1484,1509]" italics="true" pageId="8" pageNumber="9">Geostibida</emphasis>
</taxonomicName>
: the difference in the shape of spermatheca (S­shaped in
<taxonomicName id="BDAEA2AB6E36C40F8F0F82D4FD2DF9BF" box="[575,704,1524,1549]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8F0F82D4FD2DF9BF" box="[575,704,1524,1549]" italics="true" pageId="8" pageNumber="9">Geostibida</emphasis>
</taxonomicName>
and semicircular in
<taxonomicName id="BDAEA2AB6E36C40F8E8082D4FB9CF9BF" box="[944,1137,1524,1549]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8E8082D4FB9CF9BF" box="[944,1137,1524,1549]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
) and the length of elytra (shorter than pronotum in
<taxonomicName id="BDAEA2AB6E36C40F8E35813CFC6BF987" box="[773,902,1564,1589]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8E35813CFC6BF987" box="[773,902,1564,1589]" italics="true" pageId="8" pageNumber="9">Geostibida</emphasis>
</taxonomicName>
and longer than pronotum in
<taxonomicName id="BDAEA2AB6E36C40F8C388164FE24F9EF" box="[264,457,1604,1629]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E36C40F8C388164FE24F9EF" box="[264,457,1604,1629]" italics="true" pageId="8" pageNumber="9">Tropimenelytron</emphasis>
</taxonomicName>
). Unfortunately, these characters are not reliable. The shape of the proximal portion of spermatheca varies even among closely related species. For example, in
<taxonomicName id="BDAEA2AB6E36C40F8C1781B4FE3AF91F" box="[295,471,1684,1709]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="americanum">
<emphasis id="48DA053A6E36C40F8C1781B4FE3AF91F" box="[295,471,1684,1709]" italics="true" pageId="8" pageNumber="9">T. americanum</emphasis>
</taxonomicName>
(
<figureCitation id="E295C5AD6E36C40F8CD881B4FDD2F91C" box="[488,575,1684,1710]" captionStart="FIGURES 39 ­ 44" captionStartId="13.[264,383,1231,1255]" captionTargetBox="[294,1263,291,1184]" captionTargetId="figure@13.[280,1276,284,1195]" captionTargetPageId="13" captionText="FIGURES 39 ­ 44. Genitalia of Tropimenelytron americanum Gusarov, sp. n. (paratypes from Rockville (39) and Morgan Monroe State Park, Indiana (40 ­ 44 )). 39 – spermatheca; 40 – median lobe of aedeagus, parameral view; 41 – median lobe of aedeagus, lateral view; 42 – apex of median lobe, parameral view; 43 – apex of median lobe, lateral view; 44 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (39, 42 ­ 44), 0.2 mm (40 ­ 41)." httpUri="https://zenodo.org/record/155761/files/figure.png" pageId="8" pageNumber="9">Fig. 39</figureCitation>
) this portion is longer than in two other Nearctic species (
<figureCitation id="E295C5AD6E36C40F89C181B4FEC5F964" captionStart="FIGURES 49 ­ 54" captionStartId="15.[264,382,1742,1766]" captionTargetBox="[306,1282,761,1723]" captionTargetId="figure@15.[287,1300,753,1731]" captionTargetPageId="15" captionText="FIGURES 49 ­ 54. Genitalia of Tropimenelytron robustum Gusarov, sp. n. (paratype from Hope, British Columbia (49) and holotype (50 ­ 54 )). 49 – spermatheca; 50 – median lobe of aedeagus, parameral view; 51 – median lobe of aedeagus, lateral view; 52 – apex of median lobe, parameral view; 53 – apex of median lobe, lateral view; 54 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (49, 52 ­ 54), 0.2 mm (50 ­ 51)." httpUri="https://zenodo.org/record/155763/files/figure.png" pageId="8" pageNumber="9">Figs. 49</figureCitation>
,
<figureCitation id="E295C5AD6E36C40F8C0E819CFEB0F964" box="[318,349,1724,1750]" captionStart="FIGURES 59 ­ 64" captionStartId="18.[264,383,1139,1163]" captionTargetBox="[304,1295,297,1090]" captionTargetId="figure@18.[280,1313,284,1107]" captionTargetPageId="18" captionText="FIGURES 59 ­ 64. Genitalia of Tropimenelytron californicum Gusarov, sp. n. (paratypes from Mineral, California). 59 – spermatheca; 60 – median lobe of aedeagus, parameral view; 61 – median lobe of aedeagus, lateral view; 62 – apex of median lobe, parameral view; 63 – apex of median lobe, lateral view; 64 – apical lobe of left paramere, side facing median lobe. Scale bar 0.1 mm (59, 62 ­ 64), 0.2 mm (60 ­ 61)." httpUri="https://zenodo.org/record/155765/files/figure.png" pageId="8" pageNumber="9">59</figureCitation>
) and the spermatheca can be described as S­shaped. In many groups of
<taxonomicName id="BDAEA2AB6E36C40F8C3881C4FE49F94C" box="[264,420,1764,1790]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
the length of elytra may be subject to variation even within the same species (see for example
<bibRefCitation id="1E3FA4D96E36C40F8CE6802CFD9DF894" author="Muona" box="[470,624,1804,1830]" pageId="8" pageNumber="23" refString="Muona, J. (1991) The North European and British species of the genus Meotica Mulsant et Rey (Coleoptera, Staphylinidae). Deutsche entomologische Zeitschrift, N. F., 38 (1 - 3), 225 - 246." type="journal article" year="1991">Muona 1991</bibRefCitation>
;
<bibRefCitation id="1E3FA4D96E36C40F8F4D802CFCFBF894" author="Assing" box="[637,790,1804,1830]" pageId="8" pageNumber="23" refString="Assing, V. (1999) A revision of the species of Geostiba Thomson 1858 from Greece and Cyprus (Coleoptera, Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 31 (2), 845 - 928." type="journal article" year="1999">Assing 1999</bibRefCitation>
). The specimens with well developed wings have longer elytra, while in wingless specimens the elytra are shorter. In many groups of aleocharines the species or populations restricted in their distribution to limited areas in the mountains often loose the ability to fly, have reduced wings and shorter elytra in comparison to their more widespread relatives in the plain. Clearly the length of elytra alone is not a sufficient character to separate subgenera of
<taxonomicName id="BDAEA2AB6E37C40E8E8786B4FBDEFE1F" box="[951,1075,404,429]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E37C40E8E8786B4FBDEFE1F" box="[951,1075,404,429]" italics="true" pageId="9" pageNumber="10">Pelioptera</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="2ED189A06E37C40E8C3881F2FD5DF8DD" httpUri="https://zenodo.org/record/155759/files/figure.png" pageId="9" pageNumber="10" targetBox="[316,1284,476,1734]" targetPageId="9">
<paragraph id="7A11D9286E37C40E8C3881F2FD5DF8DD" blockId="9.[264,1323,1746,1903]" pageId="9" pageNumber="10">
<emphasis id="48DA053A6E37C40E8C3881F2FE24F958" bold="true" box="[264,457,1746,1770]" pageId="9" pageNumber="10">FIGURES 27­34.</emphasis>
Male genitalia of
<taxonomicName id="BDAEA2AB6E37C40E8FA181F3FC4CF958" authority="Kraatz" authorityName="Kraatz" box="[657,929,1747,1770]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="micans">
<emphasis id="48DA053A6E37C40E8FA181F3FCBEF958" box="[657,851,1747,1770]" italics="true" pageId="9" pageNumber="10">Pelioptera micans</emphasis>
Kraatz
</taxonomicName>
(syntype from Sri Lanka (27­29)),
<taxonomicName id="BDAEA2AB6E37C40E882781F3FE08F8B6" authority="Kraatz" authorityName="Kraatz" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="exasperata">
<emphasis id="48DA053A6E37C40E882781F3FE90F8B6" italics="true" pageId="9" pageNumber="10">P. exasperata</emphasis>
(Kraatz)
</taxonomicName>
(Dehra Dun, India (30­32)) and
<taxonomicName id="BDAEA2AB6E37C40E8E4781CDFB31F8B6" authority="Motschulsky" authorityName="Motschulsky" box="[887,1244,1773,1796]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="testaceipennis">
<emphasis id="48DA053A6E37C40E8E4781CDFBDCF8B6" box="[887,1073,1773,1796]" italics="true" pageId="9" pageNumber="10">P. testaceipennis</emphasis>
(Motschulsky)
</taxonomicName>
(Java, Indonesia (33­34)). 27, 30, 33 – median lobe of aedeagus, parameral view; 28 – apex of median lobe, parameral view; 29 – copulatory piece of internal sac, lateral view; 31, 34 – median lobe of aedeagus, lateral view; 32 – copulatory piece of internal sac, parameral view. Scale bar 0.2 mm (28­29, 32), 0.4 mm (27, 30­31, 33­34).
</paragraph>
</caption>
<paragraph id="7A11D9286E34C40D8C08863CFB28FCA4" blockId="10.[264,1324,284,1190]" pageId="10" pageNumber="11">
Pace himself is not consistent in applying these two characters to separate
<taxonomicName id="BDAEA2AB6E34C40D8C388664FE24FEEF" box="[264,457,324,349]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8C388664FE24FEEF" box="[264,457,324,349]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
</taxonomicName>
and
<taxonomicName id="BDAEA2AB6E34C40D8F388664FD64FEEF" box="[520,649,324,349]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8F388664FD64FEEF" box="[520,649,324,349]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
. For example, in
<taxonomicName id="BDAEA2AB6E34C40D8E5F8664FB7EFEEC" authority="Pace, 1985" authorityName="Pace" authorityYear="1985" box="[879,1171,324,350]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nepalense">
<emphasis id="48DA053A6E34C40D8E5F8664FBE9FEEF" box="[879,1028,324,349]" italics="true" pageId="10" pageNumber="11">T. nepalense</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D893E8664FB7EFEEC" author="Pace" box="[1038,1171,324,350]" pageId="10" pageNumber="24" refString="Pace, R. (1985 a) Aleocharinae riportate dall'Himalaya dal Prof. Franz. Parte I (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (Nouvelle Serie), 2 (1), 91 - 105." type="journal article" year="1985" yearSuffix="a">Pace, 1985</bibRefCitation>
</taxonomicName>
a (
<figureCitation id="E295C5AD6E34C40D89828664FAE4FEEC" box="[1202,1289,324,350]" captionStart="FIGURES 45 ­ 48" captionStartId="14.[264,383,1346,1370]" captionTargetBox="[298,1291,463,1313]" captionTargetId="figure@14.[287,1299,449,1320]" captionTargetPageId="14" captionText="FIGURES 45 ­ 48. Details of abdominal segment 8 of Tropimenelytron robustum Gusarov, sp. n. (holotype (45 ­ 46), paratype from Hope, British Columbia (47 ­ 48 )). 45 – male tergum 8; 46 – male sternum 8; 47 – female tergum 8; 48 – female sternum 8. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/155762/files/figure.png" pageId="10" pageNumber="11">
Fig.
<quantity id="BD5674CD6E34C40D89DC8664FAE4FEEC" box="[1260,1289,324,350]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="10" pageNumber="11" unit="in" value="45.0">45</quantity>
</figureCitation>
in
<bibRefCitation id="1E3FA4D96E34C40D8C38864CFE78FE34" author="Pace" box="[264,405,364,390]" pageId="10" pageNumber="24" refString="Pace, R. (1985 a) Aleocharinae riportate dall'Himalaya dal Prof. Franz. Parte I (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (Nouvelle Serie), 2 (1), 91 - 105." type="journal article" year="1985" yearSuffix="a">Pace 1985a</bibRefCitation>
) and
<taxonomicName id="BDAEA2AB6E34C40D8CD1864CFCE6FE34" authority="Pace,1987" authorityName="Pace" authorityYear="1987" box="[481,779,364,390]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="parbatense">
<emphasis id="48DA053A6E34C40D8CD1864CFD68FE37" box="[481,645,364,389]" italics="true" pageId="10" pageNumber="11">T. parbatense</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8FA2864CFCE6FE34" author="Pace" box="[658,779,364,390]" pageId="10" pageNumber="24" refString="Pace, R. (1987 a) Staphylinidae dell'Himalaya Nepalese. Aleocharinae raccolte dal Prof. Dr. J. Martens (Insecta: Coleoptera). Courier Forschungsinstitut Senckenberg, 93, 383 - 441." type="journal article" year="1987" yearSuffix="a">Pace,1987</bibRefCitation>
</taxonomicName>
a (
<figureCitation id="E295C5AD6E34C40D8E1D864CFC6BFE34" box="[813,902,364,390]" captionStart="FIGURES 45 ­ 48" captionStartId="14.[264,383,1346,1370]" captionTargetBox="[298,1291,463,1313]" captionTargetId="figure@14.[287,1299,449,1320]" captionTargetPageId="14" captionText="FIGURES 45 ­ 48. Details of abdominal segment 8 of Tropimenelytron robustum Gusarov, sp. n. (holotype (45 ­ 46), paratype from Hope, British Columbia (47 ­ 48 )). 45 – male tergum 8; 46 – male sternum 8; 47 – female tergum 8; 48 – female sternum 8. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/155762/files/figure.png" pageId="10" pageNumber="11">
Fig.
<quantity id="BD5674CD6E34C40D8E59864CFC6BFE34" box="[873,902,364,390]" metricMagnitude="0" metricUnit="m" metricValue="1.1938" pageId="10" pageNumber="11" unit="in" value="47.0">47</quantity>
</figureCitation>
in
<bibRefCitation id="1E3FA4D96E34C40D8E87864CFBAEFE34" author="Pace" box="[951,1091,364,390]" pageId="10" pageNumber="24" refString="Pace, R. (1987 a) Staphylinidae dell'Himalaya Nepalese. Aleocharinae raccolte dal Prof. Dr. J. Martens (Insecta: Coleoptera). Courier Forschungsinstitut Senckenberg, 93, 383 - 441." type="journal article" year="1987" yearSuffix="a">Pace 1987a</bibRefCitation>
) (both assigned to
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<emphasis id="48DA053A6E34C40D8C3886B4FE69FE1F" box="[264,388,404,429]" italics="true" pageId="10" pageNumber="11">Pelioptera</emphasis>
(
<emphasis id="48DA053A6E34C40D8CA586B4FDBBFE1F" box="[405,598,404,429]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
)
</taxonomicName>
in
<bibRefCitation id="1E3FA4D96E34C40D8FB586B4FD12FE1C" author="Pace" box="[645,767,404,430]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
) the spermatheca is no less S­shaped than in
<taxonomicName id="BDAEA2AB6E34C40D883E86B4FDDBFE64" authority="Pace, 1984" authorityName="Pace" authorityYear="1984" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="himalayiensis">
<emphasis id="48DA053A6E34C40D883E86B4FE44FE67" italics="true" pageId="10" pageNumber="11">G. himalayiensis</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8C83869CFDDBFE64" author="Pace" box="[435,566,444,470]" pageId="10" pageNumber="23" refString="Pace, R. (1984) Aleocharinae dell'Himalya [sic!]. LI. Contributo alla conoscenza delle Aleocharinae (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France (Nouvelle Serie), 20 (3), 309 - 339." type="journal article" year="1984">Pace, 1984</bibRefCitation>
</taxonomicName>
(Fig.
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<bibRefCitation id="1E3FA4D96E34C40D8FE6869CFCB8FE64" author="Pace" box="[726,853,444,470]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
; assigned to
<taxonomicName id="BDAEA2AB6E34C40D8EC1869CFAE6FE64" authority="(Geostibida)" baseAuthorityName="Geostibida" box="[1009,1291,444,470]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8EC1869CFB80FE67" box="[1009,1133,444,469]" italics="true" pageId="10" pageNumber="11">Pelioptera</emphasis>
(
<emphasis id="48DA053A6E34C40D89B0869CFAECFE67" box="[1152,1281,444,469]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
)
</taxonomicName>
in
<bibRefCitation id="1E3FA4D96E34C40D8C3886C4FE65FE4C" author="Pace" box="[264,392,484,510]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
). In
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<emphasis id="48DA053A6E34C40D8CFB86C4FE33FE4F" box="[459,478,484,509]" italics="true" pageId="10" pageNumber="11">P.</emphasis>
(
<emphasis id="48DA053A6E34C40D8CC486C4FDFDFE4F" box="[500,528,484,509]" italics="true" pageId="10" pageNumber="11">G.</emphasis>
)
<emphasis id="48DA053A6E34C40D8F1486C4FD52FE4F" box="[548,703,484,509]" italics="true" pageId="10" pageNumber="11">problematica</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8FF886C4FCA1FE4C" author="Pace" box="[712,844,484,510]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace, 1991</bibRefCitation>
</taxonomicName>
the elytra are both described (p.
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<bibRefCitation id="1E3FA4D96E34C40D8C38852CFE6BFD94" author="Pace" box="[264,390,524,550]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
) and illustrated (Fig.
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<bibRefCitation id="1E3FA4D96E34C40D8FDB852CFC87FD94" author="Pace" box="[747,874,524,550]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
) as being longer than pronotum, but nevertheless the species is placed in the subgenus
<taxonomicName id="BDAEA2AB6E34C40D8E648514FC38FDFF" box="[852,981,564,589]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8E648514FC38FDFF" box="[852,981,564,589]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
and not
<taxonomicName id="BDAEA2AB6E34C40D890B8514FB11FDFF" box="[1083,1276,564,589]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D890B8514FB11FDFF" box="[1083,1276,564,589]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
</taxonomicName>
. In
<taxonomicName id="BDAEA2AB6E34C40D8C38857CFE4AFDC7" box="[264,423,604,629]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="parbatense">
<emphasis id="48DA053A6E34C40D8C38857CFE4AFDC7" box="[264,423,604,629]" italics="true" pageId="10" pageNumber="11">T. parbatense</emphasis>
</taxonomicName>
(
<figureCitation id="E295C5AD6E34C40D8C87857CFDE7FDC4" box="[439,522,604,630]" captionStart="FIGURES 45 ­ 48" captionStartId="14.[264,383,1346,1370]" captionTargetBox="[298,1291,463,1313]" captionTargetId="figure@14.[287,1299,449,1320]" captionTargetPageId="14" captionText="FIGURES 45 ­ 48. Details of abdominal segment 8 of Tropimenelytron robustum Gusarov, sp. n. (holotype (45 ­ 46), paratype from Hope, British Columbia (47 ­ 48 )). 45 – male tergum 8; 46 – male sternum 8; 47 – female tergum 8; 48 – female sternum 8. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/155762/files/figure.png" pageId="10" pageNumber="11">
Fig.
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</figureCitation>
in
<bibRefCitation id="1E3FA4D96E34C40D8F00857CFD5AFDC4" author="Pace" box="[560,695,604,630]" pageId="10" pageNumber="24" refString="Pace, R. (1987 a) Staphylinidae dell'Himalaya Nepalese. Aleocharinae raccolte dal Prof. Dr. J. Martens (Insecta: Coleoptera). Courier Forschungsinstitut Senckenberg, 93, 383 - 441." type="journal article" year="1987" yearSuffix="a">Pace 1987a</bibRefCitation>
) elytra are illustrated as being shorter than pronotum, but in
<bibRefCitation id="1E3FA4D96E34C40D8C5685A4FE07FD2C" author="Pace" box="[358,490,644,670]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace 1991</bibRefCitation>
this species is placed in the subgenus
<taxonomicName id="BDAEA2AB6E34C40D8EC885A4FB57FD2F" box="[1016,1210,644,669]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8EC885A4FB57FD2F" box="[1016,1210,644,669]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
</taxonomicName>
and not
<taxonomicName id="BDAEA2AB6E34C40D8C38858CFE64FD77" box="[264,393,684,709]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8C38858CFE64FD77" box="[264,393,684,709]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
. Some species assigned by Pace to
<taxonomicName id="BDAEA2AB6E34C40D8E07858CFC55FD77" box="[823,952,684,709]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8E07858CFC55FD77" box="[823,952,684,709]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
lack both diagnostic characters of
<taxonomicName id="BDAEA2AB6E34C40D8C1885F4FE44FD5F" box="[296,425,724,749]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8C1885F4FE44FD5F" box="[296,425,724,749]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
. For example, in
<taxonomicName id="BDAEA2AB6E34C40D8F4E85F4FC5FFD5C" authority="Pace, 1998" authorityName="Pace" authorityYear="1998" box="[638,946,724,750]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="eremita" subGenus="G.">
<emphasis id="48DA053A6E34C40D8F4E85F4FD7DFD5F" box="[638,656,724,749]" italics="true" pageId="10" pageNumber="11">P.</emphasis>
(
<emphasis id="48DA053A6E34C40D8F9485F4FD2DFD5F" box="[676,704,724,749]" italics="true" pageId="10" pageNumber="11">G.</emphasis>
)
<emphasis id="48DA053A6E34C40D8FE185F4FCC4FD5F" box="[721,809,724,749]" italics="true" pageId="10" pageNumber="11">eremita</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8E0185F4FC5FFD5C" author="Pace" box="[817,946,724,750]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">Pace, 1998</bibRefCitation>
</taxonomicName>
elytra are longer than pronotum (Fig.
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<bibRefCitation id="1E3FA4D96E34C40D8CA985DCFDF9FCA4" author="Pace" box="[409,532,764,790]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">Pace 1998</bibRefCitation>
) and spermatheca is not S­shaped (Fig.
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<bibRefCitation id="1E3FA4D96E34C40D890A85DCFB55FCA4" author="Pace" box="[1082,1208,764,790]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">Pace 1998</bibRefCitation>
).
</paragraph>
<paragraph id="7A11D9286E34C40D8C088404FBFBFBCC" blockId="10.[264,1324,284,1190]" pageId="10" pageNumber="11">
Examination of Pace’s description of
<taxonomicName id="BDAEA2AB6E34C40D8E358404FBB2FC8C" authority="Pace, 1984" authorityName="Pace" authorityYear="1984" box="[773,1119,804,830]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="himalayiensis">
<emphasis id="48DA053A6E34C40D8E358404FC22FC8F" box="[773,975,804,829]" italics="true" pageId="10" pageNumber="11">G. himalayiensis</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8EEA8404FBB2FC8C" author="Pace" box="[986,1119,804,830]" pageId="10" pageNumber="23" refString="Pace, R. (1984) Aleocharinae dell'Himalya [sic!]. LI. Contributo alla conoscenza delle Aleocharinae (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France (Nouvelle Serie), 20 (3), 309 - 339." type="journal article" year="1984">Pace, 1984</bibRefCitation>
</taxonomicName>
(
<typeStatus id="A515678A6E34C40D89438404FB4AFC8C" box="[1139,1191,804,830]" pageId="10" pageNumber="11">type</typeStatus>
species of
<taxonomicName id="BDAEA2AB6E34C40D8C38846CFE64FCD7" box="[264,393,844,869]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8C38846CFE64FCD7" box="[264,393,844,869]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
) and description of additional species included by Pace in
<taxonomicName id="BDAEA2AB6E34C40D896E846CFB32FCD7" box="[1118,1247,844,869]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D896E846CFB32FCD7" box="[1118,1247,844,869]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
(
<bibRefCitation id="1E3FA4D96E34C40D89C1846CFEA8FC3C" author="Pace" pageId="10" pageNumber="23" refString="Pace, R. (1984) Aleocharinae dell'Himalya [sic!]. LI. Contributo alla conoscenza delle Aleocharinae (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France (Nouvelle Serie), 20 (3), 309 - 339." type="journal article" year="1984">Pace 1984</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E34C40D8C698454FE4EFC3C" author="Pace" box="[345,419,884,910]" pageId="10" pageNumber="24" refString="Pace, R. (1985 a) Aleocharinae riportate dall'Himalaya dal Prof. Franz. Parte I (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (Nouvelle Serie), 2 (1), 91 - 105." type="journal article" year="1985" yearSuffix="a">1985a</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E34C40D8C878454FE19FC3C" author="Pace" box="[439,500,884,910]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">1991</bibRefCitation>
,
<bibRefCitation id="1E3FA4D96E34C40D8F388454FDA9FC3C" author="Pace" box="[520,580,884,910]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">1998</bibRefCitation>
) demonstrates that
<taxonomicName id="BDAEA2AB6E34C40D8E708454FC2DFC3F" box="[832,960,884,909]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8E708454FC2DFC3F" box="[832,960,884,909]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
is an artificial group which includes those species of
<taxonomicName id="BDAEA2AB6E34C40D8F0C84BCFD10FC07" box="[572,765,924,949]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8F0C84BCFD10FC07" box="[572,765,924,949]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
</taxonomicName>
(
<taxonomicName id="BDAEA2AB6E34C40D8E2084BCFB8AFC04" authority="Pace, 1984" authorityName="Pace" authorityYear="1984" box="[784,1127,924,950]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="himalayiensis">
<emphasis id="48DA053A6E34C40D8E2084BCFC35FC07" box="[784,984,924,949]" italics="true" pageId="10" pageNumber="11">G. himalayiensis</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8ED184BCFB8AFC04" author="Pace" box="[993,1127,924,950]" pageId="10" pageNumber="23" refString="Pace, R. (1984) Aleocharinae dell'Himalya [sic!]. LI. Contributo alla conoscenza delle Aleocharinae (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France (Nouvelle Serie), 20 (3), 309 - 339." type="journal article" year="1984">Pace, 1984</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="BDAEA2AB6E34C40D894684BCFEABFC6C" authority="Pace, 1984" authorityName="Pace" authorityYear="1984" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="major">
<emphasis id="48DA053A6E34C40D894684BCFB0FFC07" box="[1142,1250,924,949]" italics="true" pageId="10" pageNumber="11">G. major</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D89DC84BCFEABFC6C" author="Pace" pageId="10" pageNumber="23" refString="Pace, R. (1984) Aleocharinae dell'Himalya [sic!]. LI. Contributo alla conoscenza delle Aleocharinae (Coleoptera, Staphylinidae). Annales de la Societe entomologique de France (Nouvelle Serie), 20 (3), 309 - 339." type="journal article" year="1984">Pace, 1984</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="BDAEA2AB6E34C40D8C6284E4FD40FC6C" authority="Pace, 1985" authorityName="Pace" authorityYear="1985" box="[338,685,964,990]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="annapurnensis">
<emphasis id="48DA053A6E34C40D8C6284E4FDC9FC6F" box="[338,548,964,989]" italics="true" pageId="10" pageNumber="11">G. annapurnensis</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8F1A84E4FD40FC6C" author="Pace" box="[554,685,964,990]" pageId="10" pageNumber="24" refString="Pace, R. (1985 a) Aleocharinae riportate dall'Himalaya dal Prof. Franz. Parte I (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (Nouvelle Serie), 2 (1), 91 - 105." type="journal article" year="1985" yearSuffix="a">Pace, 1985</bibRefCitation>
</taxonomicName>
a) which have short elytra, and some unrelated species (which should be reassigned to other genera) with the same
<typeStatus id="A515678A6E34C40D8EEC84CCFBE2FBB4" box="[988,1039,1004,1030]" pageId="10" pageNumber="11">type</typeStatus>
of pronotal pubescence (
<taxonomicName id="BDAEA2AB6E34C40D8C228334FDF9FB9C" authority="Pace, 1998" authorityName="Pace" authorityYear="1998" box="[274,532,1044,1070]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lii" subGenus="G.">
<emphasis id="48DA053A6E34C40D8C228334FEC8FB9F" box="[274,293,1044,1069]" italics="true" pageId="10" pageNumber="11">P.</emphasis>
(
<emphasis id="48DA053A6E34C40D8C0B8334FEBAFB9F" box="[315,343,1044,1069]" italics="true" pageId="10" pageNumber="11">G.</emphasis>
)
<emphasis id="48DA053A6E34C40D8C5B8334FE69FB9F" box="[363,388,1044,1069]" italics="true" pageId="10" pageNumber="11">lii</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8CBD8334FDF9FB9C" author="Pace" box="[397,532,1044,1070]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">Pace, 1998</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="BDAEA2AB6E34C40D8F128334FC47FB9C" authority="Pace, 1998" authorityName="Pace" authorityYear="1998" box="[546,938,1044,1070]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kowloonensis" subGenus="G.">
<emphasis id="48DA053A6E34C40D8F128334FDD8FB9F" box="[546,565,1044,1069]" italics="true" pageId="10" pageNumber="11">P.</emphasis>
(
<emphasis id="48DA053A6E34C40D8F7B8334FD85FB9F" box="[587,616,1044,1069]" italics="true" pageId="10" pageNumber="11">G.</emphasis>
)
<emphasis id="48DA053A6E34C40D8F4B8334FCF4FB9F" box="[635,793,1044,1069]" italics="true" pageId="10" pageNumber="11">kowloonensis</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8E138334FC47FB9C" author="Pace" box="[803,938,1044,1070]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">Pace, 1998</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="BDAEA2AB6E34C40D8E888334FB1AFB9C" authority="Pace, 1998" authorityName="Pace" authorityYear="1998" box="[952,1271,1044,1070]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="eremita" subGenus="G.">
<emphasis id="48DA053A6E34C40D8E888334FC26FB9F" box="[952,971,1044,1069]" italics="true" pageId="10" pageNumber="11">P.</emphasis>
(
<emphasis id="48DA053A6E34C40D8ED18334FC13FB9F" box="[993,1022,1044,1069]" italics="true" pageId="10" pageNumber="11">G.</emphasis>
)
<emphasis id="48DA053A6E34C40D89218334FB87FB9F" box="[1041,1130,1044,1069]" italics="true" pageId="10" pageNumber="11">eremita</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D89438334FB1AFB9C" author="Pace" box="[1139,1271,1044,1070]" pageId="10" pageNumber="24" refString="Pace, R. (1998) Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (4), 911 - 982." type="journal article" year="1998">Pace, 1998</bibRefCitation>
</taxonomicName>
and probably
<taxonomicName id="BDAEA2AB6E34C40D8C4E831CFCFEFBE4" authority="Pace, 1991" authorityName="Pace" authorityYear="1991" box="[382,787,1084,1110]" class="Insecta" family="Staphylinidae" genus="Pelioptera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="problematica" subGenus="G.">
<emphasis id="48DA053A6E34C40D8C4E831CFE7CFBE7" box="[382,401,1084,1109]" italics="true" pageId="10" pageNumber="11">P.</emphasis>
(
<emphasis id="48DA053A6E34C40D8C9B831CFE2AFBE7" box="[427,455,1084,1109]" italics="true" pageId="10" pageNumber="11">G.</emphasis>
)
<emphasis id="48DA053A6E34C40D8CEF831CFD97FBE7" box="[479,634,1084,1109]" italics="true" pageId="10" pageNumber="11">problematica</emphasis>
<bibRefCitation id="1E3FA4D96E34C40D8FB8831CFCFEFBE4" author="Pace" box="[648,787,1084,1110]" pageId="10" pageNumber="24" refString="Pace, R. (1991) Aleocharinae nepalesi del Museo di Ginevra. Parte V: Athetini (conclusione) e Thamiaraeini (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 98 (4), 803 - 863." type="journal article" year="1991">Pace, 1991</bibRefCitation>
</taxonomicName>
). A revision of the
<typeStatus id="A515678A6E34C40D8913831CFBBBFBE4" box="[1059,1110,1084,1110]" pageId="10" pageNumber="11">type</typeStatus>
of
<taxonomicName id="BDAEA2AB6E34C40D89B9831CFAE7FBE7" box="[1161,1290,1084,1109]" class="Insecta" family="Staphylinidae" genus="Geostibida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D89B9831CFAE7FBE7" box="[1161,1290,1084,1109]" italics="true" pageId="10" pageNumber="11">Geostibida</emphasis>
</taxonomicName>
is necessary to formally synonymize the name with
<taxonomicName id="BDAEA2AB6E34C40D8E7E8344FBE2FBCF" box="[846,1039,1124,1149]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8E7E8344FBE2FBCF" box="[846,1039,1124,1149]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7A11D9286E34C40D8C0883ACFAE0FB14" blockId="10.[264,1324,284,1190]" box="[312,1293,1164,1190]" pageId="10" pageNumber="11">
The genus
<taxonomicName id="BDAEA2AB6E34C40D8C8883ACFD94FB17" box="[440,633,1164,1189]" class="Insecta" family="Staphylinidae" genus="Tropimenelytron" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="48DA053A6E34C40D8C8883ACFD94FB17" box="[440,633,1164,1189]" italics="true" pageId="10" pageNumber="11">Tropimenelytron</emphasis>
</taxonomicName>
is being reported from North
<collectingCountry id="02B999B86E34C40D8EEA83ACFBD2FB14" box="[986,1087,1164,1190]" name="United States of America" pageId="10" pageNumber="11">America</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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