333 lines
44 KiB
XML
333 lines
44 KiB
XML
<document id="AEDB8193CD86272117356A6D441DB7F6" ID-CLB-Dataset="8444" ID-DOI="10.11646/zootaxa.4853.4.6" ID-GBIF-Dataset="1d17a387-d55f-4a0d-8a98-e66b817981d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="4411149" ID-ZooBank="2F03212F-E526-422F-A7B7-C83CD57F70DB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609605990093" checkinUser="plazi" docAuthor="Gao, Zhizhong, Zhang, Feng & Chen, Huiming" docDate="2020" docId="0874CA41FFB0FF9FFF61F9DE113686CC" docLanguage="en" docName="zootaxa.4853.4.6.pdf" docOrigin="Zootaxa 4853 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrannochthonius zhai Gao & Zhang & Chen 2020, sp. nov." docType="treatment" docUuid="DF1220A7-37B6-4608-A4BB-C01A5215B70D" docUuidSource="ZooBank" docVersion="9" lastPageNumber="579" masterDocId="F44DB239FFB4FF98FFF6FF8414138209" masterDocTitle="Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China" masterLastPageNumber="580" masterPageNumber="572" pageNumber="576" updateTime="1701999495881" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="11B4461F12553803C8B4B89459CAEC8B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9327763A453AB0A0398F63552711295B">
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<mods:title id="BBC0C4E7079E8ABFDFA3D57BA8D6269F">Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China</mods:title>
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<mods:name id="13E31A5682D28C0C0ED8DF541C70803D" type="personal">
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<mods:roleTerm id="498094661F787AA56210FBEAEDDA3E92">Author</mods:roleTerm>
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<mods:namePart id="8A6D75266C5141DE6D6706BD7A6255C0">Gao, Zhizhong</mods:namePart>
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<mods:nameIdentifier id="085B831028D067F95080A41D5A4EA86E" type="ORCID">0000-0002-6666-8746</mods:nameIdentifier>
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<mods:affiliation id="96BAF5368CF150115A9FDABCAA7FEF58">Department of Biology, Wutai Mountain Institute of Resource and Environment, Xinzhou Teachers University, Xinzhou 034000, Shanxi Province, P. R. China. gaozhizhong 1987 @ 126. com; https: // orcid. org / 0000 - 0002 - 6666 - 8746</mods:affiliation>
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</mods:name>
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<mods:name id="3F4B34F01FFDD1C5F8CAC262A0FE1398" type="personal">
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<mods:role id="DC062CAA51215CBA5F50376BB86E526F">
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<mods:roleTerm id="4336C9FFFDD61EED5450F03FB09D7472">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1BBDAA97AD851F9656EBB15BDA5E2408">Zhang, Feng</mods:namePart>
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<mods:nameIdentifier id="74C0A76DF150F6A877ED9CB8001B4956" type="ORCID">0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="A937F67EA5F607223A09A1A93FBC33FB">The Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China. dudu 06042001 @ 163. com; https: // orcid. org / 0000 - 0002 - 3347 - 1031</mods:affiliation>
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<mods:roleTerm id="1E99AB2E5982F81073F286D502635B17">Author</mods:roleTerm>
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<mods:namePart id="F612E4909BC8A5F76B582A7408533E16">Chen, Huiming</mods:namePart>
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<mods:affiliation id="971B802AFF51D7F470A0DA67535D09E0">Institute of Biology, Guizhou Academy of Sciences, Guiyang, P. R. China.</mods:affiliation>
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<mods:title id="372ECEEAB80C695941E30307B158C90A">Zootaxa</mods:title>
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<mods:part id="DEFC5403FE07DEDD0998178F45B07A3B">
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<mods:date id="E6C0913187A0FE8F7797A52192162D1C">2020</mods:date>
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<mods:number id="A8AA26CC806D1CE61F078E590028C44A">2020-09-24</mods:number>
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<mods:number id="43547256E3ADAA03548BE6740FBF3D64">4853</mods:number>
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<mods:detail id="76BD32FAB40CA4A7C36AA8AD095DAE02" type="issue">
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<mods:number id="32819CC3FE9E9775D14921EABE6CB3C2">4</mods:number>
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<treatment id="0874CA41FFB0FF9FFF61F9DE113686CC" ID-DOI="http://doi.org/10.5281/zenodo.4411056" ID-GBIF-Taxon="177212024" ID-Zenodo-Dep="4411056" LSID="urn:lsid:zoobank.org:act:DF1220A7-37B6-4608-A4BB-C01A5215B70D" httpUri="http://treatment.plazi.org/id/0874CA41FFB0FF9FFF61F9DE113686CC" lastPageId="7" lastPageNumber="579" pageId="4" pageNumber="576">
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<heading id="DB2ACC3BFFB0FF9CFF61F9DE1605847C" bold="true" box="[151,534,1626,1653]" fontSize="11" level="1" pageId="4" pageNumber="576" reason="1">
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<emphasis id="B2A9A745FFB0FF9CFF61F9DE1605847C" bold="true" box="[151,534,1626,1653]" pageId="4" pageNumber="576">
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<taxonomicName id="47DD00D4FFB0FF9CFF61F9DE15A2847C" authority="Gao & Zhang & Chen, 2020" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[151,433,1626,1653]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="576" phylum="Arthropoda" rank="species" species="zhai" status="sp. nov.">
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<emphasis id="B2A9A745FFB0FF9CFF61F9DE15A2847C" bold="true" box="[151,433,1626,1653]" italics="true" pageId="4" pageNumber="576">Tyrannochthonius zhai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A99A1A3EFFB0FF9CFE4EF9DF1605847C" box="[440,534,1627,1653]" pageId="4" pageNumber="576" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C8C728DCFFB0FF9CFF61F9FB17DE84B4" pageId="4" pageNumber="576" type="description">
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<paragraph id="80627B57FFB0FF9CFF61F9FB14EC8490" blockId="4.[151,973,1626,1725]" box="[151,255,1663,1689]" pageId="4" pageNumber="576">
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<figureCitation id="18E667D2FFB0FF9CFF61F9FB14EC8490" box="[151,255,1663,1689]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,1089,1113]" captionStartId-1="5.[151,250,1960,1984]" captionStartId-2="6.[151,250,1499,1523]" captionTargetBox-0="[227,1361,181,1066]" captionTargetBox-1="[217,1369,1138,1937]" captionTargetBox-2="[171,1414,162,1460]" captionTargetId-0="figure-25@5.[227,1361,181,1066]" captionTargetId-1="figure-43@5.[217,1369,1138,1937]" captionTargetId-2="figure-19@6.[151,1437,151,1481]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 4. Tyrannochthonius zhai sp. nov., holotype female, habitus, dorsal view." captionText-1="FIGURE 5. Tyrannochthonius zhai sp. nov., holotype female: A. Chelal fingers (lateral view); B. Carapace (dorsal view); C. Left chelicerae (dorsal view); D. Genital (ventral view). Scale bars: 0.1 mm." captionText-2="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi-0="http://doi.org/10.5281/zenodo.4411159" figureDoi-1="http://doi.org/10.5281/zenodo.4411161" figureDoi-2="http://doi.org/10.5281/zenodo.4411163" httpUri-0="https://zenodo.org/record/4411159/files/figure.png" httpUri-1="https://zenodo.org/record/4411161/files/figure.png" httpUri-2="https://zenodo.org/record/4411163/files/figure.png" pageId="4" pageNumber="576">Figs. 4–6</figureCitation>
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</paragraph>
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<paragraph id="80627B57FFB0FF9CFF61F92617DE84B4" blockId="4.[151,973,1626,1725]" box="[151,973,1698,1725]" pageId="4" pageNumber="576">
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<uri id="F44C7755FFB0FF9CFF61F92617DE84B4" box="[151,973,1698,1725]" pageId="4" pageNumber="576">
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urn:lsid:zoobank.org:act:
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<uuid id="F47B4182FFB0FF9CFE5CF92617DE84B4" box="[426,973,1698,1725]" pageId="4" pageNumber="576">DF1220A7-37B6-4608-A4BB-C01A5215B70D</uuid>
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<subSubSection id="C8C728DCFFB0FF9CFF61F96F10448520" pageId="4" pageNumber="576" type="materials_examined">
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<paragraph id="80627B57FFB0FF9CFF61F96F10448520" blockId="4.[151,1436,1770,2013]" pageId="4" pageNumber="576">
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<emphasis id="B2A9A745FFB0FF9CFF61F96F1557850D" bold="true" box="[151,324,1770,1797]" pageId="4" pageNumber="576">Type material.</emphasis>
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<materialsCitation id="30B5710AFFB0FF9CFEBFF96F10448520" ID-GBIF-Occurrence="3029870303" collectingDate="2011-06-07" collectionCode="MHBU-GZ" collectorName="Zuwei Zha" country="China" county="Xingyi City" location="Jiangjia Cave" longitude="105.601944" municipality="Dushan Town" pageId="4" pageNumber="576" specimenCode="MHBU-GZ11060701" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
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<typeStatus id="5F66C5F5FFB0FF9CFEBFF96F15A2850C" box="[329,433,1771,1797]" pageId="4" pageNumber="576" type="holotype">Holotype</typeStatus>
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female (Ps.-
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<specimenCode id="D07BD32CFFB0FF9CFDC1F96F1726850F" box="[567,821,1771,1798]" collectionCode="MHBU-GZ" pageId="4" pageNumber="576">MHBU-GZ11060701</specimenCode>
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):
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<collectingCountry id="F8CA3BC7FFB0FF9CFCB0F96F179F850C" box="[838,908,1771,1797]" name="China" pageId="4" pageNumber="576">China</collectingCountry>
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,
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<collectingRegion id="4219B5B5FFB0FF9CFC60F96F1071850C" box="[918,1122,1771,1797]" country="China" name="Guizhou" pageId="4" pageNumber="576">Guizhou Province</collectingRegion>
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,
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<collectingCounty id="690303DBFFB0FF9CFB9AF96F10E2850C" box="[1132,1265,1771,1797]" pageId="4" pageNumber="576">Xingyi City</collectingCounty>
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,
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<collectingMunicipality id="6006E12DFFB0FF9CFB0DF96F118B850C" box="[1275,1432,1771,1797]" pageId="4" pageNumber="576">Dushan Town</collectingMunicipality>
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,
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<location id="85022D8CFFB0FF9CFF61F88B15228520" LSID="urn:lsid:plazi:treatment:0874CA41FFB0FF9FFF61F9DE113686CC:85022D8CFFB0FF9CFF61F88B15228520" box="[151,305,1807,1833]" country="China" county="Xingyi City" longitude="105.601944" municipality="Dushan Town" name="Jiangjia Cave" pageId="4" pageNumber="576" stateProvince="Guizhou">Jiangjia Cave</location>
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, dark zone, [
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<geoCoordinate id="E5E91D90FFB0FF9CFE49F88A16448520" box="[447,599,1806,1833]" pageId="4" pageNumber="576">25°18'096”N</geoCoordinate>
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,
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<geoCoordinate id="E5E91D90FFB0FF9CFD95F88A17168520" box="[611,773,1806,1833]" degrees="105" direction="east" minutes="36" orientation="longitude" pageId="4" pageNumber="576" precision="15" seconds="007" value="105.601944">105°36'007”E</geoCoordinate>
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],
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<date id="F4635D97FFB0FF9CFCEEF88B17B28520" box="[792,929,1807,1833]" pageId="4" pageNumber="576" value="2011-06-07">
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<collectingDate id="E427A47FFFB0FF9CFCEEF88B17B28520" box="[792,929,1807,1833]" pageId="4" pageNumber="576" value="2011-06-07">7 June 2011</collectingDate>
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</date>
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,
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<collectorName id="2D281E81FFB0FF9CFC5AF88B10348520" box="[940,1063,1807,1833]" pageId="4" pageNumber="576">Zuwei Zha</collectorName>
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leg.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C8C728DCFFB0FF9CFF31F8B711438544" box="[199,1360,1843,1869]" pageId="4" pageNumber="576" type="etymology">
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<paragraph id="80627B57FFB0FF9CFF31F8B711438544" blockId="4.[151,1436,1770,2013]" box="[199,1360,1843,1869]" pageId="4" pageNumber="576">
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<emphasis id="B2A9A745FFB0FF9CFF31F8B7155D8544" bold="true" box="[199,334,1843,1869]" pageId="4" pageNumber="576">Etymology.</emphasis>
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The specific epithet is a patronym in honor of Mr. Zuwei Zha, who collected the specimen.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8C728DCFFB0FF9CFF31F8D215A885D4" pageId="4" pageNumber="576" type="diagnosis">
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<paragraph id="80627B57FFB0FF9CFF31F8D215A885D4" blockId="4.[151,1436,1770,2013]" pageId="4" pageNumber="576">
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<emphasis id="B2A9A745FFB0FF9CFF31F8D215518579" bold="true" box="[199,322,1878,1904]" pageId="4" pageNumber="576">Diagnosis.</emphasis>
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Small sized troglomorphic species (Body length 1.40); carapace without eyes or eyespots, both anterior and posterior margin with 4 setae respectively, without epistome; tergites I–III with 4 setae; chelal fixed finger with intercalary teeth, movable finger with numerous pointed retrorse and contiguous teeth; no chemosensory setae on dorsum of chelal hand.
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</paragraph>
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</subSubSection>
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<caption id="D4A22BDFFFB1FF9DFF61FBC517F58653" ID-DOI="http://doi.org/10.5281/zenodo.4411159" ID-Zenodo-Dep="4411159" box="[151,998,1089,1114]" httpUri="https://zenodo.org/record/4411159/files/figure.png" pageId="5" pageNumber="577" startId="5.[151,250,1089,1113]" targetBox="[227,1361,181,1066]" targetPageId="5">
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<paragraph id="80627B57FFB1FF9DFF61FBC517F58653" blockId="5.[151,998,1089,1114]" box="[151,998,1089,1114]" pageId="5" pageNumber="577">
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<emphasis id="B2A9A745FFB1FF9DFF61FBC515008650" bold="true" box="[151,275,1089,1113]" pageId="5" pageNumber="577">FIGURE 4.</emphasis>
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<taxonomicName id="47DD00D4FFB1FF9DFEEFFBC616138653" authority="Gao & Zhang & Chen, 2020" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[281,512,1090,1114]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="zhai" status="sp. nov.">
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<emphasis id="B2A9A745FFB1FF9DFEEFFBC616138653" box="[281,512,1090,1114]" italics="true" pageId="5" pageNumber="577">Tyrannochthonius zhai</emphasis>
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</taxonomicName>
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<emphasis id="B2A9A745FFB1FF9DFDF1FBC616448653" bold="true" box="[519,599,1090,1114]" pageId="5" pageNumber="577">
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<taxonomicNameLabel id="A99A1A3EFFB1FF9DFDF1FBC616448653" box="[519,599,1090,1114]" pageId="5" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype female, habitus, dorsal view.
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</paragraph>
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</caption>
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<caption id="D4A22BDFFFB1FF9DFF61F82C178E85ED" ID-DOI="http://doi.org/10.5281/zenodo.4411161" ID-Zenodo-Dep="4411161" httpUri="https://zenodo.org/record/4411161/files/figure.png" pageId="5" pageNumber="577" startId="5.[151,250,1960,1984]" targetBox="[217,1369,1138,1937]" targetPageId="5">
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<paragraph id="80627B57FFB1FF9DFF61F82C178E85ED" blockId="5.[151,1437,1960,2020]" pageId="5" pageNumber="577">
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<emphasis id="B2A9A745FFB1FF9DFF61F82C150785C9" bold="true" box="[151,276,1960,1984]" pageId="5" pageNumber="577">FIGURE 5.</emphasis>
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<taxonomicName id="47DD00D4FFB1FF9DFEEDF82C161085C9" authority="Gao & Zhang & Chen, 2020" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[283,515,1960,1984]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="577" phylum="Arthropoda" rank="species" species="zhai" status="sp. nov.">
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<emphasis id="B2A9A745FFB1FF9DFEEDF82C161085C9" box="[283,515,1960,1984]" italics="true" pageId="5" pageNumber="577">Tyrannochthonius zhai</emphasis>
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</taxonomicName>
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<emphasis id="B2A9A745FFB1FF9DFDFDF82C164F85C9" bold="true" box="[523,604,1960,1984]" pageId="5" pageNumber="577">
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<taxonomicNameLabel id="A99A1A3EFFB1FF9DFDFDF82C164F85C9" box="[523,604,1960,1984]" pageId="5" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
|
||
, holotype female: A. Chelal fingers (lateral view); B. Carapace (dorsal view); C. Left chelicerae (dorsal view); D. Genital (ventral view). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D4A22BDFFFB2FF9EFF61FA5F16CB8456" ID-DOI="http://doi.org/10.5281/zenodo.4411163" ID-Zenodo-Dep="4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="6" pageNumber="578" startId="6.[151,250,1499,1523]" targetBox="[171,1414,162,1460]" targetPageId="6">
|
||
<paragraph id="80627B57FFB2FF9EFF61FA5F16CB8456" blockId="6.[151,1437,1499,1631]" pageId="6" pageNumber="578">
|
||
<emphasis id="B2A9A745FFB2FF9EFF61FA5F150087FA" bold="true" box="[151,275,1499,1523]" pageId="6" pageNumber="578">FIGURE 6.</emphasis>
|
||
<taxonomicName id="47DD00D4FFB2FF9EFEEDFA5F161187FA" authority="Gao & Zhang & Chen, 2020" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[283,514,1499,1523]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="578" phylum="Arthropoda" rank="species" species="zhai" status="sp. nov.">
|
||
<emphasis id="B2A9A745FFB2FF9EFEEDFA5F161187FA" box="[283,514,1499,1523]" italics="true" pageId="6" pageNumber="578">Tyrannochthonius zhai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B2A9A745FFB2FF9EFDFFFA5F164987FA" bold="true" box="[521,602,1499,1523]" pageId="6" pageNumber="578">
|
||
<taxonomicNameLabel id="A99A1A3EFFB2FF9EFDFFFA5F164987FA" box="[521,602,1499,1523]" pageId="6" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C8C728DCFFB2FF9FFF31F90915408188" lastPageId="7" lastPageNumber="579" pageId="6" pageNumber="578" type="description">
|
||
<paragraph id="80627B57FFB2FF9EFF31F909164084A1" blockId="6.[151,1437,1677,2028]" box="[199,595,1677,1704]" pageId="6" pageNumber="578">
|
||
<emphasis id="B2A9A745FFB2FF9EFF31F909154A84AE" bold="true" box="[199,345,1677,1703]" pageId="6" pageNumber="578">Description.</emphasis>
|
||
Adult female (
|
||
<figureCitation id="18E667D2FFB2FF9EFDF6F909165584A1" box="[512,582,1677,1704]" captionStart="FIGURE 4" captionStartId="5.[151,250,1089,1113]" captionTargetBox="[227,1361,181,1066]" captionTargetId="figure-25@5.[227,1361,181,1066]" captionTargetPageId="5" captionText="FIGURE 4. Tyrannochthonius zhai sp. nov., holotype female, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4411159" httpUri="https://zenodo.org/record/4411159/files/figure.png" pageId="6" pageNumber="578">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="80627B57FFB2FF9EFF31F935114A84C2" blockId="6.[151,1437,1677,2028]" box="[199,1369,1713,1740]" pageId="6" pageNumber="578">Colour: Generally pale yellow, carapace, chelicera, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
|
||
<paragraph id="80627B57FFB2FF9EFF31F95117C6853E" blockId="6.[151,1437,1677,2028]" pageId="6" pageNumber="578">
|
||
Chelicera (
|
||
<figureCitation id="18E667D2FFB2FF9EFEB5F95115B384F9" box="[323,416,1749,1776]" captionStart="FIGURE 5" captionStartId="5.[151,250,1960,1984]" captionTargetBox="[217,1369,1138,1937]" captionTargetId="figure-43@5.[217,1369,1138,1937]" captionTargetPageId="5" captionText="FIGURE 5. Tyrannochthonius zhai sp. nov., holotype female: A. Chelal fingers (lateral view); B. Carapace (dorsal view); C. Left chelicerae (dorsal view); D. Genital (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4411161" httpUri="https://zenodo.org/record/4411161/files/figure.png" pageId="6" pageNumber="578">Figs 5C</figureCitation>
|
||
,
|
||
<figureCitation id="18E667D2FFB2FF9EFE5BF95115C284F9" box="[429,465,1749,1776]" captionStart="FIGURE 6" captionStartId="6.[151,250,1499,1523]" captionTargetBox="[171,1414,162,1460]" captionTargetId="figure-19@6.[151,1437,151,1481]" captionTargetPageId="6" captionText="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi="http://doi.org/10.5281/zenodo.4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="6" pageNumber="578">6B</figureCitation>
|
||
): Five setae on hand, all setae acuminate; movable finger with one medial seta; fixed finger and movable finger with about 10 teeth; galea represented by a very slight bump on movable finger; rallum (
|
||
<figureCitation id="18E667D2FFB2FF9EFF56F89914E58531" box="[160,246,1821,1848]" captionStart="FIGURE 6" captionStartId="6.[151,250,1499,1523]" captionTargetBox="[171,1414,162,1460]" captionTargetId="figure-19@6.[151,1437,151,1481]" captionTargetPageId="6" captionText="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi="http://doi.org/10.5281/zenodo.4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="6" pageNumber="578">Fig. 6C</figureCitation>
|
||
) with 6 blades, distal one longer than others, all blades bipinnate.
|
||
</paragraph>
|
||
<paragraph id="80627B57FFB2FF9FFF31F8C5112D8348" blockId="6.[151,1437,1677,2028]" lastBlockId="7.[151,1437,151,1221]" lastPageId="7" lastPageNumber="579" pageId="6" pageNumber="578">
|
||
Pedipalp (
|
||
<figureCitation id="18E667D2FFB2FF9EFECCF8C5158A8555" box="[314,409,1857,1884]" captionStart="FIGURE 5" captionStartId="5.[151,250,1960,1984]" captionTargetBox="[217,1369,1138,1937]" captionTargetId="figure-43@5.[217,1369,1138,1937]" captionTargetPageId="5" captionText="FIGURE 5. Tyrannochthonius zhai sp. nov., holotype female: A. Chelal fingers (lateral view); B. Carapace (dorsal view); C. Left chelicerae (dorsal view); D. Genital (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4411161" httpUri="https://zenodo.org/record/4411161/files/figure.png" pageId="6" pageNumber="578">Figs 5A</figureCitation>
|
||
,
|
||
<figureCitation id="18E667D2FFB2FF9EFE50F8C515A78555" box="[422,436,1857,1884]" captionStart="FIGURE 6" captionStartId="6.[151,250,1499,1523]" captionTargetBox="[171,1414,162,1460]" captionTargetId="figure-19@6.[151,1437,151,1481]" captionTargetPageId="6" captionText="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi="http://doi.org/10.5281/zenodo.4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="6" pageNumber="578">6</figureCitation>
|
||
E–H): All setae acuminate; femur 6.00, patella 2.20, chela 6.37, hand 1.89 times longer than deep; movable finger 2.14 times longer than hand, without large basal apodeme, only slightly sclerotized section is present. Femur without tactile setae. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria:
|
||
<emphasis id="B2A9A745FFB2FF9EFE72F829158985CE" box="[388,410,1965,1991]" italics="true" pageId="6" pageNumber="578">ib</emphasis>
|
||
and
|
||
<emphasis id="B2A9A745FFB2FF9EFE26F82915E285CE" box="[464,497,1965,1991]" italics="true" pageId="6" pageNumber="578">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis id="B2A9A745FFB2FF9EFB4EF82910C185CE" box="[1208,1234,1965,1991]" italics="true" pageId="6" pageNumber="578">eb</emphasis>
|
||
,
|
||
<emphasis id="B2A9A745FFB2FF9EFB16F829111685CE" box="[1248,1285,1965,1991]" italics="true" pageId="6" pageNumber="578">esb</emphasis>
|
||
and
|
||
<emphasis id="B2A9A745FFB2FF9EFACDF82A114685CE" box="[1339,1365,1966,1991]" italics="true" pageId="6" pageNumber="578">ist</emphasis>
|
||
forming a straight oblique row at base of fixed chelal finger;
|
||
<emphasis id="B2A9A745FFB2FF9EFD09F856171C85E2" box="[767,783,2002,2027]" italics="true" pageId="6" pageNumber="578">it</emphasis>
|
||
slightly distal to
|
||
<emphasis id="B2A9A745FFB2FF9EFC3BF85617FF85E2" box="[973,1004,2002,2027]" italics="true" pageId="6" pageNumber="578">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis id="B2A9A745FFB2FF9EFB2BF85610E285E2" box="[1245,1265,2002,2027]" italics="true" pageId="6" pageNumber="578">et</emphasis>
|
||
slightly near to tip of fixed finger, very close to chelal teeth;
|
||
<emphasis id="B2A9A745FFB3FF9FFD7EFF13168C82B9" box="[648,671,151,176]" italics="true" pageId="7" pageNumber="579">xs</emphasis>
|
||
situated distal to
|
||
<emphasis id="B2A9A745FFB3FF9FFC94FF13176582B9" box="[866,886,151,176]" italics="true" pageId="7" pageNumber="579">et</emphasis>
|
||
, microsetae (chemosensory setae) absent on both palpal fingers;
|
||
<emphasis id="B2A9A745FFB3FF9FFEB7FF3F154982DC" box="[321,346,187,213]" italics="true" pageId="7" pageNumber="579">sb</emphasis>
|
||
situated slightly closer to
|
||
<emphasis id="B2A9A745FFB3FF9FFD72FF3F168182DC" box="[644,658,187,213]" italics="true" pageId="7" pageNumber="579">b</emphasis>
|
||
than to
|
||
<emphasis id="B2A9A745FFB3FF9FFD19FF3F171182DD" box="[751,770,187,212]" italics="true" pageId="7" pageNumber="579">st</emphasis>
|
||
;
|
||
<emphasis id="B2A9A745FFB3FF9FFCE4FF3F173382DC" box="[786,800,187,213]" italics="true" pageId="7" pageNumber="579">b</emphasis>
|
||
and
|
||
<emphasis id="B2A9A745FFB3FF9FFCAEFF3F177382DD" box="[856,864,187,212]" italics="true" pageId="7" pageNumber="579">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis id="B2A9A745FFB3FF9FFBBAFF3F104782DD" box="[1100,1108,187,212]" italics="true" pageId="7" pageNumber="579">t</emphasis>
|
||
situated at same level as
|
||
<emphasis id="B2A9A745FFB3FF9FFA83FF3F118782DD" box="[1397,1428,187,212]" italics="true" pageId="7" pageNumber="579">est</emphasis>
|
||
;
|
||
<emphasis id="B2A9A745FFB3FF9FFF61FF5B14B682F0" box="[151,165,223,249]" italics="true" pageId="7" pageNumber="579">b</emphasis>
|
||
situated basal to
|
||
<emphasis id="B2A9A745FFB3FF9FFE9EFF5B159482F1" box="[360,391,223,248]" italics="true" pageId="7" pageNumber="579">est</emphasis>
|
||
; microsetae (chemosensory setae) absent on both palpal fingers. Sensilla absent. Chelal teeth (
|
||
<figureCitation id="18E667D2FFB3FF9FFF56FE8714E98317" box="[160,250,259,286]" captionStart="FIGURE 6" captionStartId="6.[151,250,1499,1523]" captionTargetBox="[171,1414,162,1460]" captionTargetId="figure-19@6.[151,1437,151,1481]" captionTargetPageId="6" captionText="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi="http://doi.org/10.5281/zenodo.4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="7" pageNumber="579">Fig. 6H</figureCitation>
|
||
) heterodentate: fixed finger with 22 large, slightly retrorse, well-spaced teeth, plus 15 small intercalary teeth; movable finger with 26 middle size retrorse, pointed and continuous teeth, without intercalary teeth.
|
||
</paragraph>
|
||
<paragraph id="80627B57FFB3FF9FFF31FECF165283FC" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="579">
|
||
Cephalothorax (
|
||
<figureCitation id="18E667D2FFB3FF9FFE8AFECF15CB836C" box="[380,472,330,357]" captionStart="FIGURE 5" captionStartId="5.[151,250,1960,1984]" captionTargetBox="[217,1369,1138,1937]" captionTargetId="figure-43@5.[217,1369,1138,1937]" captionTargetPageId="5" captionText="FIGURE 5. Tyrannochthonius zhai sp. nov., holotype female: A. Chelal fingers (lateral view); B. Carapace (dorsal view); C. Left chelicerae (dorsal view); D. Genital (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4411161" httpUri="https://zenodo.org/record/4411161/files/figure.png" pageId="7" pageNumber="579">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="18E667D2FFB3FF9FFE13FECF161F836F" box="[485,524,331,358]" captionStart="FIGURE 6" captionStartId="6.[151,250,1499,1523]" captionTargetBox="[171,1414,162,1460]" captionTargetId="figure-19@6.[151,1437,151,1481]" captionTargetPageId="6" captionText="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi="http://doi.org/10.5281/zenodo.4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="7" pageNumber="579">6A</figureCitation>
|
||
): Carapace as broad as long; anterior margin smooth; lateral margins constricted posteriorly; without any traces of eyes; without epistome; with 16 setae arranged 4: 4: 4: 2: 2; without furrows. Chaetotaxy of coxae: 4: 3: 4: 4; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; coxae II with 5–6 terminally indented coxal spines on each side, set in oblique row; intercoxal tubercle absent; without sub-oral seta.
|
||
</paragraph>
|
||
<paragraph id="80627B57FFB3FF9FFF31FE7B17C48068" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="579">Abdomen: Pleural membrane papillostriate. Tergites and sternites undivided; setae uniseriate and acuminate. Each tergite with four setae except last one which has two only. Sternal chaetotaxy V–XI: 8: 8: 10: 9: 10: 10: 8. Anterior genital operculum with 10 setae plus 8 setae on posterior margin.</paragraph>
|
||
<paragraph id="80627B57FFB3FF9FFF31FDEF162680A0" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="579">
|
||
Legs (
|
||
<figureCitation id="18E667D2FFB3FF9FFEFBFDEF1547808F" box="[269,340,619,646]" captionStart="FIGURE 6" captionStartId="6.[151,250,1499,1523]" captionTargetBox="[171,1414,162,1460]" captionTargetId="figure-19@6.[151,1437,151,1481]" captionTargetPageId="6" captionText="FIGURE 6. Tyrannochthonius zhai sp. nov., holotype female: A. Carapace (dorsal view); B. Left chelicerae (dorsal view); C. Rallum; D. Coxal spines on coxae II (ventral view); E. Finger tips of chela (lateral view); F. Left pedipalp (minus chela, dorsal view); G. Left chela (lateral view); H. Chelal fingers (lateral view, showing the teeth); I. Leg I (lateral view); J. Leg IV (lateral view). Scale bars: 0.10 mm (C–E); 0.25 mm (A–B, F–J)." figureDoi="http://doi.org/10.5281/zenodo.4411163" httpUri="https://zenodo.org/record/4411163/files/figure.png" pageId="7" pageNumber="579">Figs 6</figureCitation>
|
||
I–J): Typical. Femur + patella of leg IV 2.90 times longer than deep; arolium slightly shorter than the claws, not divided; claws simple.
|
||
</paragraph>
|
||
<paragraph id="80627B57FFB3FF9FFF31FD3715408188" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="579">
|
||
Measurements: (length/breadth or depth in mm, ratios in parentheses). Female
|
||
<typeStatus id="5F66C5F5FFB3FF9FFBAFFD3710AC80C4" box="[1113,1215,691,717]" pageId="7" pageNumber="579" type="holotype">holotype</typeStatus>
|
||
: Body length 1.40. Pedipalps: trochanter 0.24/0.14 (1.71), femur 0.84/0.14 (6.00), patella 0.33/0.15 (2.20), chela 1.21/0.19 (6.37), hand length 0.36/0.16 (1.89), movable finger length 0.77. Carapace 0.48/0.48 (1.00). Leg I: trochanter 0.15/0.11 (1.36), femur 0.44/0.07 (6.29), patella 0.24/0.06 (4.00), tibia 0.22/0.06 (3.67), tarsus 0.46/0.05 (9.20). Leg IV: trochanter 0.20/0.11 (1.82), femur + patella 0.61/0.21 (2.90), tibia 0.45/0.08 (5.63), basitarsus 0.19/0.06 (3.17), telotarsus 0.46/0.05 (9.20).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8C728DCFFB3FF9FFF31FC0F113686CC" pageId="7" pageNumber="579" type="discussion">
|
||
<paragraph id="80627B57FFB3FF9FFF31FC0F113686CC" blockId="7.[151,1437,151,1221]" pageId="7" pageNumber="579">
|
||
<emphasis id="B2A9A745FFB3FF9FFF31FC0F152881AC" bold="true" box="[199,315,907,933]" pageId="7" pageNumber="579">Remarks.</emphasis>
|
||
Of the Chinese species,
|
||
<taxonomicName id="47DD00D4FFB3FF9FFDA4FC0F168E81AC" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[594,669,907,933]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="B2A9A745FFB3FF9FFDA4FC0F168E81AC" box="[594,669,907,933]" italics="true" pageId="7" pageNumber="579">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
has the same tergal chaetotaxy on tergites I–II as
|
||
<emphasis id="B2A9A745FFB3FF9FFB27FC0F14D781C1" italics="true" pageId="7" pageNumber="579">
|
||
<taxonomicName id="47DD00D4FFB3FF9FFB27FC0F115681AC" authorityName="Mahnert" authorityYear="2009" box="[1233,1349,907,933]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="akaelus">T. akaelus</taxonomicName>
|
||
,
|
||
<taxonomicName id="47DD00D4FFB3FF9FFAA7FC0F14D781C1" authorityName="Gao & Zhang & Chen" authorityYear="2020" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="harveyi">T. harveyi</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="47DD00D4FFB3FF9FFEF6FC2B15D181C0" authorityName="Mahnert" authorityYear="2009" box="[256,450,943,969]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="ganshuanensis">
|
||
<emphasis id="B2A9A745FFB3FF9FFEF6FC2B15D181C0" box="[256,450,943,969]" italics="true" pageId="7" pageNumber="579">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
(only 2 setae), but can be distinguished from
|
||
<taxonomicName id="47DD00D4FFB3FF9FFC2CFC2B105E81C0" authorityName="Mahnert" authorityYear="2009" box="[986,1101,943,969]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="akaelus">
|
||
<emphasis id="B2A9A745FFB3FF9FFC2CFC2B105E81C0" box="[986,1101,943,969]" italics="true" pageId="7" pageNumber="579">T. akaelus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47DD00D4FFB3FF9FFB7EFC2B115881C0" authorityName="Mahnert" authorityYear="2009" box="[1160,1355,943,969]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="ganshuanensis">
|
||
<emphasis id="B2A9A745FFB3FF9FFB7EFC2B115881C0" box="[1160,1355,943,969]" italics="true" pageId="7" pageNumber="579">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
by the number of setae on the anterior margin of carapace (
|
||
<taxonomicName id="47DD00D4FFB3FF9FFD2DFC57173681E4" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[731,805,979,1005]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="B2A9A745FFB3FF9FFD2DFC57173681E4" box="[731,805,979,1005]" italics="true" pageId="7" pageNumber="579">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
with 4,
|
||
<taxonomicName id="47DD00D4FFB3FF9FFC88FC5717FC81E4" authorityName="Mahnert" authorityYear="2009" box="[894,1007,979,1005]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="akaelus">
|
||
<emphasis id="B2A9A745FFB3FF9FFC88FC5717FC81E4" box="[894,1007,979,1005]" italics="true" pageId="7" pageNumber="579">T. akaelus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="47DD00D4FFB3FF9FFBD5FC5710F081E4" authorityName="Mahnert" authorityYear="2009" box="[1059,1251,979,1005]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="ganshuanensis">
|
||
<emphasis id="B2A9A745FFB3FF9FFBD5FC5710F081E4" box="[1059,1251,979,1005]" italics="true" pageId="7" pageNumber="579">T. ganshuanensis</emphasis>
|
||
</taxonomicName>
|
||
with 6;
|
||
<bibRefCitation id="E44C06A6FFB3FF9FFACBFC5714F88618" author="Mahnert, V." pageId="7" pageNumber="579" pagination="185 - 201" refId="ref4227" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116, 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert (2009))</bibRefCitation>
|
||
and by the stouter pedipalps (e.g. femur 6.00 times vs. 6.60 times;
|
||
<bibRefCitation id="E44C06A6FFB3FF9FFC16FC73108D8618" author="Mahnert, V." box="[992,1182,1015,1041]" pageId="7" pageNumber="579" pagination="185 - 201" refId="ref4227" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116, 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert (2009))</bibRefCitation>
|
||
.
|
||
<taxonomicName id="47DD00D4FFB3FF9FFB5EFC7310E08618" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[1192,1267,1015,1041]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="B2A9A745FFB3FF9FFB5EFC7310E08618" box="[1192,1267,1015,1041]" italics="true" pageId="7" pageNumber="579">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="47DD00D4FFB3FF9FFA7EFC7314FE863C" authorityName="Mahnert" authorityYear="2009" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="akaelus">
|
||
<emphasis id="B2A9A745FFB3FF9FFA7EFC7314FE863C" italics="true" pageId="7" pageNumber="579">T. akaelus</emphasis>
|
||
</taxonomicName>
|
||
by the number of chelal teeth fixed finger with 22 large, slightly retrorse, well-spaced teeth, plus 15 small intercalary teeth in
|
||
<taxonomicName id="47DD00D4FFB3FF9FFE84FBBB15AE8650" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[370,445,1087,1113]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="B2A9A745FFB3FF9FFE84FBBB15AE8650" box="[370,445,1087,1113]" italics="true" pageId="7" pageNumber="579">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
but 43 pointed teeth of equal length in
|
||
<taxonomicName id="47DD00D4FFB3FF9FFC8DFBBB17FE8650" authorityName="Mahnert" authorityYear="2009" box="[891,1005,1087,1113]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="akaelus">
|
||
<emphasis id="B2A9A745FFB3FF9FFC8DFBBB17FE8650" box="[891,1005,1087,1113]" italics="true" pageId="7" pageNumber="579">T. akaelus</emphasis>
|
||
</taxonomicName>
|
||
; movable finger with 26 middle-sized retrorse, pointed and continuous teeth in
|
||
<taxonomicName id="47DD00D4FFB3FF9FFD95FBE716BE8674" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[611,685,1123,1149]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="B2A9A745FFB3FF9FFD95FBE716BE8674" box="[611,685,1123,1149]" italics="true" pageId="7" pageNumber="579">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
but with 37 marginal teeth and
|
||
<specimenCount id="96DBB0DEFFB3FF9FFBE5FBE710B28674" box="[1043,1185,1123,1149]" pageId="7" pageNumber="579" type="intercalary">8 intercalary</specimenCount>
|
||
teeth in
|
||
<taxonomicName id="47DD00D4FFB3FF9FFAF4FBE711678674" authorityName="Mahnert" authorityYear="2009" box="[1282,1396,1123,1149]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="akaelus">
|
||
<emphasis id="B2A9A745FFB3FF9FFAF4FBE711678674" box="[1282,1396,1123,1149]" italics="true" pageId="7" pageNumber="579">T. akaelus</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName id="47DD00D4FFB3FF9FFED5FB03158286A8" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[291,401,1159,1185]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B2A9A745FFB3FF9FFED5FB03158286A8" box="[291,401,1159,1185]" italics="true" pageId="7" pageNumber="579">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
by having only 2 setae on the posterior margin of the carapace (
|
||
<quantity id="4725D6B2FFB3FF9FFBA0FB03109286A8" box="[1110,1153,1159,1185]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="579" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="47DD00D4FFB3FF9FFB71FB0310E686A8" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[1159,1269,1159,1185]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B2A9A745FFB3FF9FFB71FB0310E686A8" box="[1159,1269,1159,1185]" italics="true" pageId="7" pageNumber="579">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
), and the fixed chelal finger with 15 small intercalary teeth in
|
||
<taxonomicName id="47DD00D4FFB3FF9FFD55FB2F16FD86CC" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[675,750,1195,1221]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="zhai">
|
||
<emphasis id="B2A9A745FFB3FF9FFD55FB2F16FD86CC" box="[675,750,1195,1221]" italics="true" pageId="7" pageNumber="579">T. zhai</emphasis>
|
||
</taxonomicName>
|
||
, whereas intercalary teeth are absent in
|
||
<taxonomicName id="47DD00D4FFB3FF9FFB59FB2F110D86CC" authorityName="Gao & Zhang & Chen" authorityYear="2020" box="[1199,1310,1195,1221]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="579" phylum="Arthropoda" rank="species" species="harveyi">
|
||
<emphasis id="B2A9A745FFB3FF9FFB59FB2F110D86CC" box="[1199,1310,1195,1221]" italics="true" pageId="7" pageNumber="579">T. harveyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |