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<document id="B87B19DFBF7A6FCFDE29394942BFD5D2" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. &amp; Herbert, D. G." docDate="2013" docId="03BC1F3CE52CD910070EFE384226B483" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Acroptychia bathiei : Fischer-Piette &amp; Bedoucha 1965" docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="5" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="275AE46996E2D2BE6045CEBF8A5723A9">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
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<mods:namePart id="BCEC3DED293A9BF86C6705B3BB7D6010">Griffiths, O. L.</mods:namePart>
<mods:affiliation id="D14E0F728F7D7AC17E232F2751BFE3DF">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
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<mods:namePart id="6415D16CBF647F7FBC5A58CBA68319DB">Herbert, D. G.</mods:namePart>
<mods:affiliation id="0770D4F03EE712E561184F2859EF7161">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
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<treatment id="03BC1F3CE52CD910070EFE384226B483" ID-DOI="http://doi.org/10.5281/zenodo.7670242" ID-GBIF-Taxon="209341508" ID-Zenodo-Dep="7670242" LSID="urn:lsid:plazi:treatment:03BC1F3CE52CD910070EFE384226B483" httpUri="http://treatment.plazi.org/id/03BC1F3CE52CD910070EFE384226B483" lastPageId="7" lastPageNumber="7" pageId="5" pageNumber="5">
<subSubSection id="C30FFDA1E52CD912070EFE3845BDB686" box="[323,924,427,454]" pageId="5" pageNumber="5" type="nomenclature">
<paragraph id="8BAAAE2AE52CD912070EFE3845BDB686" blockId="5.[323,924,427,454]" box="[323,924,427,454]" pageId="5" pageNumber="5">
<heading id="D0E21946E52CD912070EFE3845BDB686" box="[323,924,427,454]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="5" reason="3">
<taxonomicName id="4C15D5A9E52CD912070EFE3845BDB686" ID-CoL="64NBV" ID-ENA="928778" authority="Fischer-Piette &amp; Bedoucha, 1965" authorityName=": Fischer-Piette &amp; Bedoucha" authorityYear="1965" box="[323,924,427,454]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="5" phylum="Mollusca" rank="species" species="bathiei">
<emphasis id="B9617238E52CD912070EFE384401B685" box="[323,544,430,454]" italics="true" pageId="5" pageNumber="5">Acroptychia bathiei</emphasis>
Fischer-Piette &amp; Bedoucha, 1965
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E52CD912046CFE4044E1B6B3" box="[545,704,470,496]" pageId="5" pageNumber="5" type="description">
<paragraph id="8BAAAE2AE52CD912046CFE4044E1B6B3" blockId="5.[545,704,470,496]" box="[545,704,470,496]" pageId="5" pageNumber="5">
<figureCitation id="132EB2AFE52CD912046CFE404449B6B3" box="[545,616,470,496]" captionStart="Fig" captionStartId="5.[142,175,1582,1603]" captionTargetBox="[149,1091,895,1560]" captionTargetId="figure-253@5.[143,1103,890,1564]" captionTargetPageId="5" captionText="Fig. 2. Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965: (AC) Tsingy Beanka, stn 18/06, max. diameter 23.7 mm, NMSA L7174; (DF)Antsingimavo, stn 04/06,NMSA L7091:(D) typical shell, max. diameter 23.0; (E) pale yellow specimen with a less elevated spire, max. diameter 21.7 mm; (F) juvenile specimen, max. diameter 14.9 mm." figureDoi="http://doi.org/10.5281/zenodo.7670104" httpUri="https://zenodo.org/record/7670104/files/figure.png" pageId="5" pageNumber="5">Figs 2</figureCitation>
,
<figureCitation id="132EB2AFE52CD9120439FE4044A2B6B3" box="[628,643,470,496]" captionStart="Fig" captionStartId="6.[181,214,556,577]" captionTargetBox="[190,1134,204,538]" captionTargetId="figure-411@6.[189,1135,204,538]" captionTargetPageId="6" captionText="Fig. 3. Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, radula, NMSA L7039: (A) entire width of radula; (B) oblique view of left half row. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7670106" httpUri="https://zenodo.org/record/7670106/files/figure.png" pageId="5" pageNumber="5">3</figureCitation>
,
<figureCitation id="132EB2AFE52CD91204DDFE4044E1B6B3" box="[656,704,470,496]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, stn 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., stn 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette &amp; Bedoucha, 1965, stn 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, stn 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,stn 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, stn 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, stn 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., stn 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="5" pageNumber="5">25A</figureCitation>
</paragraph>
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<subSubSection id="C30FFDA1E52CD91206DDFE684402B501" pageId="5" pageNumber="5" type="reference_group">
<paragraph id="8BAAAE2AE52CD91206DDFE684402B501" blockId="5.[144,1104,509,859]" pageId="5" pageNumber="5">
<taxonomicName id="4C15D5A9E52CD91206DDFE684489B551" ID-CoL="64NBV" authority=": Fischer-Piette &amp; Bedoucha 1965: 61" authorityName=": Fischer-Piette &amp; Bedoucha" authorityPageNumber="61" authorityYear="1965" box="[144,680,509,530]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="5" phylum="Mollusca" rank="species" species="bathiei">
<emphasis id="B9617238E52CD91206DDFE684765B552" box="[144,324,510,529]" italics="true" pageId="5" pageNumber="5">Acroptychia bathiei</emphasis>
: Fischer-Piette &amp; Bedoucha 1965: 61
</taxonomicName>
, fig. 12, pl. 1, figs 911; Fischer-Piette
<emphasis id="B9617238E52CD912026DFE68426BB552" box="[1056,1098,510,529]" italics="true" pageId="5" pageNumber="5">et al</emphasis>
. 1993: 47, pl. 1, figs 1921.
<typeStatus id="54AE1088E52CD91207A1FD834438B569" box="[492,537,533,554]" pageId="5" pageNumber="5">Type</typeStatus>
loc.: près de la rivière Andranomavo (Ambongo) [Perrier de la Bâthie leg.], NW Madagascar.
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E52CD91106DDFDC74256B40E" lastPageId="6" lastPageNumber="6" pageId="5" pageNumber="5" type="description">
<paragraph id="8BAAAE2AE52CD91206DDFDC747A0B528" blockId="5.[144,1104,509,859]" box="[144,385,593,619]" pageId="5" pageNumber="5">Morphological notes:</paragraph>
<paragraph id="8BAAAE2AE52CD91206DDFDEE46CBB5B3" blockId="5.[144,1104,509,859]" pageId="5" pageNumber="5">
<emphasis id="B9617238E52CD91206DDFDEE476FB5D3" box="[144,334,632,656]" italics="true" pageId="5" pageNumber="5">External features</emphasis>
(
<figureCitation id="132EB2AFE52CD9120711FDE047E2B5D3" box="[348,451,630,656]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, stn 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., stn 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette &amp; Bedoucha, 1965, stn 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, stn 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,stn 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, stn 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, stn 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., stn 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="5" pageNumber="5">Fig. 25A</figureCitation>
): Head-foot mostly greyish, irregularly mottled with darker spots and blotches; eyestalks pale, but tentacles more or less uniformly dark; forehead and snout brown; tip of snout shallowly indented in mid-line; skin texture relatively smooth.
</paragraph>
<paragraph id="8BAAAE2AE52CD91206DDFD684516B456" blockId="5.[144,1104,509,859]" box="[144,823,763,790]" pageId="5" pageNumber="5">
<emphasis id="B9617238E52CD91206DDFD68472CB455" box="[144,269,766,790]" italics="true" pageId="5" pageNumber="5">Operculum</emphasis>
: Corneous, oligospiral with an eccentric nucleus.
</paragraph>
<paragraph id="8BAAAE2AE52CD91106DDFCB54269B5BA" blockId="5.[144,1104,509,859]" lastBlockId="6.[181,1143,639,1446]" lastPageId="6" lastPageNumber="6" pageId="5" pageNumber="5">
<emphasis id="B9617238E52CD91206DDFCB546C0B478" box="[144,225,803,827]" italics="true" pageId="5" pageNumber="5">Radula</emphasis>
(
<figureCitation id="132EB2AFE52CD91206BDFCB74715B478" box="[240,308,801,827]" captionStart="Fig" captionStartId="6.[181,214,556,577]" captionTargetBox="[190,1134,204,538]" captionTargetId="figure-411@6.[189,1135,204,538]" captionTargetPageId="6" captionText="Fig. 3. Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, radula, NMSA L7039: (A) entire width of radula; (B) oblique view of left half row. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7670106" httpUri="https://zenodo.org/record/7670106/files/figure.png" pageId="5" pageNumber="5">Fig. 3</figureCitation>
): Formula 1+2+1+2+1; length
<quantity id="4CED03CFE52CD91204DDFCB744C0B478" box="[656,737,801,827]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="5" unit="mm" value="12.0">12 mm</quantity>
, with
<emphasis id="B9617238E52CD9120568FCB5451EB478" box="[805,831,803,827]" italics="true" pageId="5" pageNumber="5">ca</emphasis>
140 tooth rows [
<emphasis id="B9617238E52CD91205B3FCB54239B478" box="[1022,1048,803,827]" italics="true" pageId="5" pageNumber="5">ca</emphasis>
11.5 rows/mm]; teeth robust. Rachidian tricuspid with a rounded central cusp and two smaller lateral ones; inner and outer laterals similar, each with four cusps, of which the second is consistently the largest; marginals bicuspid, the outer cusp larger. Very similar to that of
<taxonomicName id="4C15D5A9E52FD91106F8FD5744E5B59A" authority="Fischer-Piette &amp; Bedoucha, 1965" authorityName="Fischer-Piette &amp; Bedoucha" authorityYear="1965" box="[181,708,703,729]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="6" phylum="Mollusca" rank="species" species="culminans">
<emphasis id="B9617238E52FD91106F8FD574766B59A" box="[181,327,705,729]" italics="true" pageId="6" pageNumber="6">A. culminans</emphasis>
Fischer-Piette &amp; Bedoucha, 1965
</taxonomicName>
, as described and illustrated by
<bibRefCitation id="EF84D3DBE52FD911026AFD2947A9B5BA" author="PIETTE, E. &amp; BLANC, F. &amp; SALVAT, F." pageId="6" pageNumber="6" pagination="109 - 144" refId="ref18370" refString="FISCHER- PIETTE, E., BLANC, F. &amp; SALVAT, F. 1969. Complement aux mollusques opercules terrestres de Madagascar. Memoires du Museum National d'Histoire Naturelle, Serie A 55 (2): 109 - 144, pls 1, 2." type="book chapter" year="1969">
FisherPiette
<emphasis id="B9617238E52FD91106B1FD77470CB5BA" box="[252,301,737,761]" italics="true" pageId="6" pageNumber="6">et al</emphasis>
. (1969)
</bibRefCitation>
, except that the outer lateral is mostly tricuspid in that species.
</paragraph>
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<paragraph id="8BAAAE2AE52CD91206C3F9B844A1B1C8" blockId="5.[142,1104,1582,1675]" pageId="5" pageNumber="5">
Fig. 2.
<taxonomicName id="4C15D5A9E52CD912069BF9A644EEB100" authority="Fischer-Piette &amp; Bedoucha, 1965" authorityName=": Fischer-Piette &amp; Bedoucha" authorityYear="1965" box="[214,719,1582,1603]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="5" phylum="Mollusca" rank="species" species="bathiei">
<emphasis id="B9617238E52CD912069BF9A647AAB100" box="[214,395,1584,1603]" italics="true" pageId="5" pageNumber="5">Acroptychia bathiei</emphasis>
Fischer-Piette &amp; Bedoucha, 1965
</taxonomicName>
: (AC)
<taxonomicName id="4C15D5A9E52CD9120550F9B8458BB100" authorityName=", Antsingimavo" authorityYear="2013" box="[797,938,1582,1603]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, stn 18/06, max. diameter 23.7 mm, NMSA L7174; (DF) Antsingimavo, stn 04/06, NMSA L7091: (D) typical shell, max. diameter 23.0; (E) pale yellow specimen with a less elevated spire, max. diameter 21.7 mm; (F) juvenile specimen, max. diameter 14.9 mm.
</paragraph>
</caption>
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<paragraph id="8BAAAE2AE52FD91106F8FDBA450CB51A" blockId="6.[181,1142,556,601]" pageId="6" pageNumber="6">
Fig. 3.
<taxonomicName id="4C15D5A9E52FD91106B5FDB844C6B502" authority="Fischer-Piette &amp; Bedoucha, 1965" authorityName=": Fischer-Piette &amp; Bedoucha" authorityYear="1965" box="[248,743,556,577]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="6" phylum="Mollusca" rank="species" species="bathiei">
<emphasis id="B9617238E52FD91106B5FDB8478DB502" box="[248,428,558,577]" italics="true" pageId="6" pageNumber="6">Acroptychia bathiei</emphasis>
Fischer-Piette &amp; Bedoucha, 1965
</taxonomicName>
, radula, NMSA L7039: (A) entire width of radula; (B) oblique view of left half row. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8BAAAE2AE52FD91106F8FC9E4256B40E" blockId="6.[181,1143,639,1446]" pageId="6" pageNumber="6">
Locality data:
<emphasis id="B9617238E52FD911077EFC9C47B0B45E" box="[307,401,778,797]" italics="true" pageId="6" pageNumber="6">Namoroka</emphasis>
: stns 930/97, 932/97.
<emphasis id="B9617238E52FD9110415FC9C44F2B45E" box="[600,723,778,797]" italics="true" pageId="6" pageNumber="6">Antsingimavo</emphasis>
: stns 04/06, 06/06, 08/06.
<taxonomicName id="4C15D5A9E52FD911058CFC9C4260B45E" authorityName=", Antsingimavo" authorityYear="2013" box="[961,1089,778,797]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="6" phylum="Mollusca" rank="species" species="beanka">
<emphasis id="B9617238E52FD911058CFC9C4260B45E" box="[961,1089,778,797]" italics="true" pageId="6" pageNumber="6">Tsingy Beanka</emphasis>
</taxonomicName>
: stns 03/06, 11/0613/06, 16/0618/06, 01/09, 02/09, 06/0909/09, 11/09, 01/10, 03/10, 05/1008/10, 10/10.
<emphasis id="B9617238E52FD9110270FCB4470BB40E" italics="true" pageId="6" pageNumber="6">Tsingy de Bemaraha</emphasis>
: stns 07/95, 14/95, 18/95, 09/96, 10/96, 12/96, 14/96.
<emphasis id="B9617238E52FD9110548FCAC45D9B40E" box="[773,1016,826,845]" italics="true" pageId="6" pageNumber="6">South bank of Tsiribihina R.</emphasis>
: stn 02/99LP.
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E52FD91106F8FCCA4274B4D4" pageId="6" pageNumber="6" type="distribution">
<paragraph id="8BAAAE2AE52FD91106F8FCCA4274B4D4" blockId="6.[181,1143,639,1446]" pageId="6" pageNumber="6">
Distribution: Restricted to central W
<collectingCountry id="F302EEBAE52FD9110402FCCA44FBB435" box="[591,730,860,886]" name="Madagascar" pageId="6" pageNumber="6">Madagascar</collectingCountry>
; from the Tsiribihina R. and southern Bemaraha region through
<taxonomicName id="4C15D5A9E52FD9110797FCEA44A0B4D5" authorityName=", Antsingimavo" authorityYear="2013" box="[474,641,892,918]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
and Antsingimavo to the Namoroka area.
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E52FD91106F8FC324789B35C" pageId="6" pageNumber="6" type="biology_ecology">
<paragraph id="8BAAAE2AE52FD91106F8FC324789B35C" blockId="6.[181,1143,639,1446]" pageId="6" pageNumber="6">
Habitat: Dry mixed deciduous-evergreen forest growing on limestone; living in leaflitter, under limestone rocks and in soil pockets on karst boulders; patchily common at both
<taxonomicName id="4C15D5A9E52FD91106A2FC7247B7B4BD" authorityName=", Antsingimavo" authorityYear="2013" box="[239,406,996,1022]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
and Antsingimavo. Also common throughout most of the Tsingy de Bemaraha reserve.
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E52FD91006F8FBBA4226B483" lastPageId="7" lastPageNumber="7" pageId="6" pageNumber="6" type="discussion">
<paragraph id="8BAAAE2AE52FD91106F8FBBA45A0B385" blockId="6.[181,1143,639,1446]" pageId="6" pageNumber="6">
Remarks: A moderately sized, thick-shelled species with a single, well-developed, terminal, peristomal varix. Diameter rarely more than
<quantity id="4CED03CFE52FD91104ADFBDA4511B324" box="[736,816,1100,1127]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="6" unit="mm" value="24.0">24 mm</quantity>
. Shell essentially smooth save for fine axial pliculae on the early teleoconch whorls and fine, close-set growth-lines on later whorls. Umbilicus relatively narrow and deep. Juveniles have a much thinner shell that is frequently weakly angled at the periphery (
<figureCitation id="132EB2AFE52FD9110552FB3A4555B385" box="[799,884,1196,1222]" captionStart="Fig" captionStartId="5.[142,175,1582,1603]" captionTargetBox="[149,1091,895,1560]" captionTargetId="figure-253@5.[143,1103,890,1564]" captionTargetPageId="5" captionText="Fig. 2. Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965: (AC) Tsingy Beanka, stn 18/06, max. diameter 23.7 mm, NMSA L7174; (DF)Antsingimavo, stn 04/06,NMSA L7091:(D) typical shell, max. diameter 23.0; (E) pale yellow specimen with a less elevated spire, max. diameter 21.7 mm; (F) juvenile specimen, max. diameter 14.9 mm." figureDoi="http://doi.org/10.5281/zenodo.7670104" httpUri="https://zenodo.org/record/7670104/files/figure.png" pageId="6" pageNumber="6">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="8BAAAE2AE52FD911069DFB5A4795B2E5" blockId="6.[181,1143,639,1446]" pageId="6" pageNumber="6">The ground colour ranges from a pale yellowish white to a deeper pinkish or orangebrown; in fresh specimens this is overlain by a fine, irregular whitish mottling, which is in turn overlain by a thin, rather glossy, corneous periostracum. However, the latter is commonly eroded in parts, after which the whitish mottling is quickly worn off and the shell becomes more uniformly coloured. A darker spiral line is frequently present just below the periphery, occasionally another just above it and sometimes a third in the middle of the base.</paragraph>
<paragraph id="8BAAAE2AE52FD91107ABFA5A4541B2A5" blockId="6.[486,864,1484,1511]" box="[486,864,1484,1511]" pageId="6" pageNumber="6">
Family
<taxonomicName id="4C15D5A9E52FD9110471FA5A4541B2A5" authority="Newton, 1891" authorityName="Newton" authorityYear="1891" box="[572,864,1484,1511]" class="Gastropoda" family="Pomatiidae" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="6" phylum="Mollusca" rank="family">Pomatiidae Newton, 1891</taxonomicName>
</paragraph>
<paragraph id="8BAAAE2AE52FD9110787FA7A4540B145" blockId="6.[458,865,1516,1543]" box="[458,865,1516,1543]" pageId="6" pageNumber="6">
Genus
<taxonomicName id="4C15D5A9E52FD9110455FA794540B145" authority="Troschel, 1847" authorityName="Troschel" authorityYear="1847" box="[536,865,1516,1543]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="B9617238E52FD9110455FA794495B144" box="[536,692,1519,1543]" italics="true" pageId="6" pageNumber="6">Tropidophora</emphasis>
Troschel, 1847
</taxonomicName>
</paragraph>
<paragraph id="8BAAAE2AE52FD91107E9F99945A6B165" blockId="6.[420,903,1548,1575]" box="[420,903,1548,1575]" pageId="6" pageNumber="6">
<taxonomicName id="4C15D5A9E52FD91107E9F99945A6B165" authority="Fischer-Piette, 1949" authorityName=": Fischer-Piette" authorityYear="1949" box="[420,903,1548,1575]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="6" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E52FD91107E9F99944BEB164" box="[420,671,1551,1575]" italics="true" pageId="6" pageNumber="6">Tropidophora chavani</emphasis>
Fischer-Piette, 1949
</taxonomicName>
</paragraph>
<paragraph id="8BAAAE2AE52FD9110411F9A144EEB112" blockId="6.[604,719,1591,1617]" box="[604,719,1591,1617]" pageId="6" pageNumber="6">
<figureCitation id="132EB2AFE52FD9110411F9A14485B112" box="[604,676,1591,1617]" captionStart="Fig" captionStartId="7.[142,175,488,509]" captionTargetBox="[144,1103,211,461]" captionTargetId="figure-370@7.[143,1103,201,473]" captionTargetPageId="7" captionText="Fig. 4. Tropidophora chavani Fischer-Piette, 1949: Tsingy Beanka, stn 15/06, max. diameter 27.0 mm, NMSA L7129." figureDoi="http://doi.org/10.5281/zenodo.7670108" httpUri="https://zenodo.org/record/7670108/files/figure.png" pageId="6" pageNumber="6">
Figs
<geoCoordinate id="EE21C8EDE52FD91104D9F9A14485B112" box="[660,676,1591,1617]" degrees="4, 9" direction="east" orientation="longitude" pageId="6" pageNumber="6" precision="5555" value="4.9">4</geoCoordinate>
</figureCitation>
,
<figureCitation id="132EB2AFE52FD91104FDF9A144EEB112" box="[688,719,1591,1617]" captionStart="Fig" captionStartId="13.[144,177,1606,1627]" captionTargetBox="[153,1096,661,1587]" captionTargetId="figure-180@13.[151,1097,654,1588]" captionTargetPageId="13" captionText="Fig.9. Tropidophora sericea sp.n.and T. chavani: (AC) T.sericea, holotype,Tsingy Beanka,max.diameter 37.3mm, height18.9 mm,AMS C.474166;(D, E) adult teleoconch sculpture of T. sericea (holotype) and T. chavani (NMSA L8447) respectively, scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7670122" httpUri="https://zenodo.org/record/7670122/files/figure.png" pageId="6" pageNumber="6">9E</figureCitation>
</paragraph>
<paragraph id="8BAAAE2AE52FD91106F8F9C94257B1C9" blockId="6.[181,1142,1629,1674]" pageId="6" pageNumber="6">
<taxonomicName id="4C15D5A9E52FD91106F8F9C94477B131" authority=": Fischer-Piette 1949: 15" authorityName=": Fischer-Piette" authorityPageNumber="15" authorityYear="1949" box="[181,598,1629,1650]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="6" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E52FD91106F8F9C9475CB131" box="[181,381,1631,1650]" italics="true" pageId="6" pageNumber="6">Tropidophora chavani</emphasis>
: Fischer-Piette 1949: 15
</taxonomicName>
, pl. 1, figs 46; Fischer-Piette &amp; Bedoucha 1965: 73;
<bibRefCitation id="EF84D3DBE52FD9110261F9CB47F9B1C9" author="PIETTE, E. &amp; BLANC, C. P. &amp; BLANC, F. &amp; SALVAT, F." pageId="6" pageNumber="6" pagination="1 - 281" refId="ref18289" refString="FISCHER- PIETTE, E., BLANC, C. P., BLANC, F. &amp; SALVAT, F. 1993. Gasteropodes terrestres prosobranches. In: Faune de Madagascar. Vol. 80. Paris: Museum National d'Histoire Naturelle, pp. 1 - 281." type="book chapter" year="1993">
FischerPiette
<emphasis id="B9617238E52FD9110700F9E14753B1C9" box="[333,370,1655,1674]" italics="true" pageId="6" pageNumber="6">et al</emphasis>
. 1993: 102
</bibRefCitation>
, pl. 5, figs 68.
<typeStatus id="54AE1088E52FD911042FF9E344AEB1C9" box="[610,655,1653,1674]" pageId="6" pageNumber="6">Type</typeStatus>
loc.: gorges de Salapanga (Bemaraka) [= Bemaraha].
</paragraph>
<caption id="DF6AFEA2E52ED91006C3FE7E4779B556" ID-DOI="http://doi.org/10.5281/zenodo.7670108" ID-Zenodo-Dep="7670108" httpUri="https://zenodo.org/record/7670108/files/figure.png" pageId="7" pageNumber="7" startId="7.[142,175,488,509]" targetBox="[144,1103,211,461]" targetPageId="7">
<paragraph id="8BAAAE2AE52ED91006C3FE7E4779B556" blockId="7.[142,1104,488,533]" pageId="7" pageNumber="7">
Fig. 4.
<taxonomicName id="4C15D5A9E52ED910069EFE7C4442B6BE" authority="Fischer-Piette, 1949" authorityName=": Fischer-Piette" authorityYear="1949" box="[211,611,488,509]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="7" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E52ED910069EFE7C4780B6BE" box="[211,417,490,509]" italics="true" pageId="7" pageNumber="7">Tropidophora chavani</emphasis>
Fischer-Piette, 1949
</taxonomicName>
:
<taxonomicName id="4C15D5A9E52ED9100420FE7E44D9B6BE" authorityName=", Antsingimavo" authorityYear="2013" box="[621,760,488,509]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, stn 15/06, max. diameter 27.0 mm, NMSA L7129.
</paragraph>
</caption>
<paragraph id="8BAAAE2AE52ED91006DDFDAF45D6B525" blockId="7.[144,1104,569,614]" pageId="7" pageNumber="7">
Locality data:
<taxonomicName id="4C15D5A9E52ED9100759FDAD47B8B50D" authorityName=", Antsingimavo" authorityYear="2013" box="[276,409,571,590]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="7" phylum="Mollusca" rank="species" species="beanka">
<emphasis id="B9617238E52ED9100759FDAD47B8B50D" box="[276,409,571,590]" italics="true" pageId="7" pageNumber="7">Tsingy Beanka</emphasis>
</taxonomicName>
: stns 12/06, 14/06, 15/06, 01/09, 02/09, 06/09, 07/09, 08/09, 09/09, 01/10, 05/10, 07/10, 08/10, 09/10, 10/10.
<emphasis id="B9617238E52ED9100787FDC544A5B525" box="[458,644,595,614]" italics="true" pageId="7" pageNumber="7">Tsingy de Bemaraha</emphasis>
: stns 18/95, 04/96, 09/96, 10/96, 12/96.
</paragraph>
<paragraph id="8BAAAE2AE52ED91006DDFDE04787B5F3" blockId="7.[144,1104,630,960]" pageId="7" pageNumber="7">
Distribution: A narrow-range endemic; currently recorded only from the Bemaraha region and
<taxonomicName id="4C15D5A9E52ED91006BAFD004783B5F3" authorityName=", Antsingimavo" authorityYear="2013" box="[247,418,662,688]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
.
</paragraph>
<paragraph id="8BAAAE2AE52ED91006DDFD2846D9B5BB" blockId="7.[144,1104,630,960]" pageId="7" pageNumber="7">Habitat: Dry forest growing on limestone; found in leaf-litter and between limestone boulders.</paragraph>
<paragraph id="8BAAAE2AE52ED91006DDFC904226B483" blockId="7.[144,1104,630,960]" pageId="7" pageNumber="7">
Remarks:A moderately common species in the central
<taxonomicName id="4C15D5A9E52ED91004BDFC9045B9B463" authorityName=", Antsingimavo" authorityYear="2013" box="[752,920,774,800]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, but usually present at rather low density. It is characterised by its low spire and uniformly fine spiral sculpture. Specimens with a similarly dense spiral sculpture have been collected at Andranavory in the
<collectingRegion id="49D160C8E52ED910072BFCF04792B4C3" box="[358,435,870,896]" country="Madagascar" name="Toliara" pageId="7" pageNumber="7">Toliara</collectingRegion>
[Tuléar] region, but these are more elevated and have a flaring white lip, which is broadly reflected in the columella region, half covering the umbilicus. They are closer to
<taxonomicName id="4C15D5A9E52ED910072DFC3E44F0B483" authority="(Pfeiffer, 1847)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1847" box="[352,721,934,960]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="7" phylum="Mollusca" rank="species" species="semidecussata">
<emphasis id="B9617238E52ED910072DFC3E443CB483" box="[352,541,936,960]" italics="true" pageId="7" pageNumber="7">T. semidecussata</emphasis>
(Pfeiffer, 1847)
</taxonomicName>
than they are to
<taxonomicName id="4C15D5A9E52ED91005C1FC3E4220B483" authorityName=": Fischer-Piette" authorityYear="1949" box="[908,1025,936,960]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="7" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E52ED91005C1FC3E4220B483" box="[908,1025,936,960]" italics="true" pageId="7" pageNumber="7">T. chavani</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>