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<document id="BABB710BBD0B8484B48DF5AD13B888D8" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. &amp; Herbert, D. G." docDate="2013" docId="03BC1F3CE524D9180783FEF84220B666" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Tropidophora sericea Griffiths &amp; Herbert 2013, sp. n." docType="treatment" docUuid="D3E7C8E8-D480-4510-A1A3-3E9748DB1BA0" docUuidSource="ZooBank" docVersion="8" lastPageNumber="14" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="13" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="92A7B3D8A4B23CEF0BF1BBC159CF8F7C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1D6188B604E1960092C829311E9E5F96">
<mods:title id="1828DF9CCC96AB517409963B0B957DA7">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
</mods:titleInfo>
<mods:name id="24A446CF7E23C6CFEF59E6ACC8B072EF" type="personal">
<mods:role id="AABECAF9462ADE8D716C6C3EACBAC62A">
<mods:roleTerm id="ACD625806B7E11F7D67906E8DB58FD69">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20602FD90EB2E9C146F0CDAB22C5F928">Griffiths, O. L.</mods:namePart>
<mods:affiliation id="DB32D4087869614D40D79F7B776E2FD9">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
<mods:nameIdentifier id="2BF0DBA145C32B5D8A607954BFEC3073" type="email">owen@bcm.intnet.mu</mods:nameIdentifier>
</mods:name>
<mods:name id="C26B0FDD9A653754FCB153ADBFF959D1" type="personal">
<mods:role id="A2F416A5200637564FE896C5556DE3EA">
<mods:roleTerm id="7DD359AD0380CB0A53D1EA79E6622F2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA7D0DF04FBEFDD14AE643B1D3F0CD86">Herbert, D. G.</mods:namePart>
<mods:affiliation id="B4332331FF1FFAAF6F7496900140DFE1">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
<mods:nameIdentifier id="9C247C5ADC113344052081E102E43224" type="email">dherbert@nmsa.org.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B33D52CCD20ABA82BB0C9D835698F6FB">text</mods:typeOfResource>
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<mods:titleInfo id="94A93A08F6BEE468C6F04AF802A635CD">
<mods:title id="DA0648F4EAB7356D3E650D98375B93BA">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="1DF15BE34EFF8D936BE95CF0242EE05C">
<mods:date id="9349CE72F825A2157F593D052673458A">2013</mods:date>
<mods:detail id="4BB59FEC23B3471230A35626F74A2FF2" type="pubDate">
<mods:number id="73C2772D3E5728069E84917FB90FFE81">2013-02-13</mods:number>
</mods:detail>
<mods:detail id="B9BFCF3D0BFCEE5F320A638387C3E0F4" type="volume">
<mods:number id="99EC821EC5133BFC277AB512C2FC7FFB">54</mods:number>
</mods:detail>
<mods:detail id="58AE7840149DF2DA0F7CC1A2E3E810DF" type="issue">
<mods:number id="09DABC9402D97E4455AB0DDB967E2591">1</mods:number>
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<mods:start id="B7369298F78879068452BC0658CE3ACB">1</mods:start>
<mods:end id="BD5B9A16BC1CE5F854FA3AC281CA9665">48</mods:end>
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<mods:classification id="4F6646AB7DC7C805BCCA6065B59A5D86">journal article</mods:classification>
<mods:identifier id="4273AFFC5AB1079AE0A7F441B1A42169" type="CLB-Dataset">54828</mods:identifier>
<mods:identifier id="14D4CDD40C33EED3C449F8A3BA948208" type="DOI">10.5733/afin.054.0101</mods:identifier>
<mods:identifier id="91F5611239A47A121354D71982B65399" type="GBIF-Dataset">753510a6-cffa-4637-a1bd-e9ae7d5d1660</mods:identifier>
<mods:identifier id="5C5B0CFF5F55627A49F85DFBE3F66DD1" type="ISSN">2305-2562</mods:identifier>
<mods:identifier id="ED0CEC199FC3B3F13E5831F5D2BC2735" type="Zenodo-Dep">7670100</mods:identifier>
<mods:identifier id="7DD374BCC5588B90167B3392A316BE17" type="ZooBank">3795B466-1227-4BED-AD8A-DC88CA3E14E1</mods:identifier>
</mods:mods>
<treatment id="03BC1F3CE524D9180783FEF84220B666" ID-DOI="http://doi.org/10.5281/zenodo.7649545" ID-GBIF-Taxon="209341512" ID-Zenodo-Dep="7649545" LSID="urn:lsid:zoobank.org:act:D3E7C8E8-D480-4510-A1A3-3E9748DB1BA0" httpUri="http://treatment.plazi.org/id/03BC1F3CE524D9180783FEF84220B666" lastPageId="15" lastPageNumber="14" pageId="13" pageNumber="13">
<subSubSection id="C30FFDA1E524D91A0783FEF84530B6CA" box="[462,785,366,393]" pageId="13" pageNumber="13" type="nomenclature">
<paragraph id="8BAAAE2AE524D91A0783FEF84530B6CA" blockId="13.[462,785,366,393]" box="[462,785,366,393]" pageId="13" pageNumber="13">
<heading id="D0E21946E524D91A0783FEF84530B6CA" box="[462,785,366,393]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="13" reason="2">
<taxonomicName id="4C15D5A9E524D91A0783FEF844EFB6CB" ID-CoL="59C9D" authority="Griffiths &amp; Herbert, 2013" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[462,718,366,392]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="13" phylum="Mollusca" rank="species" species="sericea" status="sp. nov.">
<emphasis id="B9617238E524D91A0783FEF844EFB6CB" bold="true" box="[462,718,366,392]" pageId="13" pageNumber="13">Tropidophora sericea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252CF43E524D91A0498FEF94530B6CA" box="[725,785,367,393]" pageId="13" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E524D9180402FE0F4220B666" lastPageId="15" lastPageNumber="15" pageId="13" pageNumber="13" type="description">
<paragraph id="8BAAAE2AE524D91A0402FE0F44B1B6F0" blockId="13.[591,656,409,435]" box="[591,656,409,435]" pageId="13" pageNumber="13">
<figureCitation id="132EB2AFE524D91A0402FE0F44B1B6F0" box="[591,656,409,435]" captionStart="Fig" captionStartId="13.[144,177,1606,1627]" captionTargetBox="[153,1096,661,1587]" captionTargetId="figure-180@13.[151,1097,654,1588]" captionTargetPageId="13" captionText="Fig.9. Tropidophora sericea sp.n.and T. chavani: (A–C) T.sericea, holotype,Tsingy Beanka,max.diameter 37.3mm, height18.9 mm,AMS C.474166;(D, E) adult teleoconch sculpture of T. sericea (holotype) and T. chavani (NMSA L8447) respectively, scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7670122" httpUri="https://zenodo.org/record/7670122/files/figure.png" pageId="13" pageNumber="13">Fig. 9</figureCitation>
</paragraph>
<paragraph id="8BAAAE2AE524D91A06DDFE554262B69E" blockId="13.[144,1104,451,614]" box="[144,1091,451,478]" pageId="13" pageNumber="13">
Etymology: From Latin
<emphasis id="B9617238E524D91A07ECFE5047DCB69D" box="[417,509,454,478]" italics="true" pageId="13" pageNumber="13">sericeus</emphasis>
(silky); in reference to the silky texture of the shell.
</paragraph>
<paragraph id="8BAAAE2AE524D91A06DDFE7D4219B526" blockId="13.[144,1104,451,614]" pageId="13" pageNumber="13">Diagnosis: Spire very low, shell almost planorboid, body whorl not conspicuously tumescent, umbilicus very wide; columella lip weakly reflected, but not broadly expanded over umbilicus; sculpture finely decussate, comprising low, close-set spiral ridges crossed by similar axial pliculae; yellowish white with traces of fine orange-brown spiral lines.</paragraph>
<caption id="DF6AFEA2E524D91A06DDF9D04505B1C8" ID-DOI="http://doi.org/10.5281/zenodo.7670122" ID-Zenodo-Dep="7670122" httpUri="https://zenodo.org/record/7670122/files/figure.png" pageId="13" pageNumber="13" startId="13.[144,177,1606,1627]" targetBox="[153,1096,661,1587]" targetPageId="13">
<paragraph id="8BAAAE2AE524D91A06DDF9D04505B1C8" blockId="13.[144,1104,1606,1675]" pageId="13" pageNumber="13">
Fig. 9.
<taxonomicName id="4C15D5A9E524D91A0681F9DE47AFB118" authority="Griffiths &amp; Herbert, 2013" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[204,398,1608,1627]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="13" phylum="Mollusca" rank="species" species="sericea" status="sp. nov.">
<emphasis id="B9617238E524D91A0681F9DE47AFB118" box="[204,398,1608,1627]" italics="true" pageId="13" pageNumber="13">Tropidophora sericea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A252CF43E524D91A07DCF9D047E0B118" box="[401,449,1606,1627]" pageId="13" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="4C15D5A9E524D91A07A5F9DE4465B118" authorityName=": Fischer-Piette" authorityYear="1949" box="[488,580,1608,1627]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="13" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E524D91A07A5F9DE4465B118" box="[488,580,1608,1627]" italics="true" pageId="13" pageNumber="13">T. chavani</emphasis>
</taxonomicName>
: (A–C)
<taxonomicName id="4C15D5A9E524D91A04C0F9DE44C3B118" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[653,738,1608,1627]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="13" phylum="Mollusca" rank="species" species="sericea">
<emphasis id="B9617238E524D91A04C0F9DE44C3B118" box="[653,738,1608,1627]" italics="true" pageId="13" pageNumber="13">T.sericea</emphasis>
</taxonomicName>
, holotype,
<taxonomicName id="4C15D5A9E524D91A050EF9D045EBB118" authorityName=", Antsingimavo" authorityYear="2013" box="[835,970,1606,1627]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, max. diameter 37.3 mm, height 18.9 mm,AMS C.474166; (D, E) adult teleoconch sculpture of
<taxonomicName id="4C15D5A9E524D91A05DAF9F645CCB130" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[919,1005,1632,1651]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="13" phylum="Mollusca" rank="species" species="sericea">
<emphasis id="B9617238E524D91A05DAF9F645CCB130" box="[919,1005,1632,1651]" italics="true" pageId="13" pageNumber="13">T. sericea</emphasis>
</taxonomicName>
(holotype) and
<taxonomicName id="4C15D5A9E524D91A06BBF9EE4775B1C8" authorityName=": Fischer-Piette" authorityYear="1949" box="[246,340,1656,1675]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="13" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E524D91A06BBF9EE4775B1C8" box="[246,340,1656,1675]" italics="true" pageId="13" pageNumber="13">T. chavani</emphasis>
</taxonomicName>
(NMSA L8447) respectively, scale bars = 2.5 mm.
</paragraph>
</caption>
<paragraph id="8BAAAE2AE527D91906F8FF5D4760B7A6" blockId="14.[181,1144,203,1674]" box="[181,321,203,229]" pageId="14" pageNumber="14">Description:</paragraph>
<paragraph id="8BAAAE2AE527D91906F8FF60441EB58E" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">
<emphasis id="B9617238E527D91906F8FF6046CCB64D" box="[181,237,246,270]" italics="true" pageId="14" pageNumber="14">Shell</emphasis>
: Medium to large, depressed-discoidal to planorboid, spire very low with little other than embryonic whorls projecting in apertural view; final part of last adult whorl descending gently prior to aperture; whorls more or less evenly rounded, suture indented; umbilicus very wide, its margin evenly rounded; underside of embryonic whorls clearly visible. Protoconch of 1¼–1½ whorls, essentially smooth, but microscopically shagreened. Teleoconch of a further 2¾–3 whorls; first whorl initially with 6 or 7 low spiral ridges increasing to double this by end of whorl; ridges crossed by numerous, close-set axial pliculae producing a finely decussate sculpture; intervals between ridges 2–3 times wider than those between pliculae; subsequent whorls with additional spiral ridges arising by intercalation, but sculpture remaining finely decussate more or less throughout, including base; spiral ridges more close-set on later whorls and sculptural interstices less axially elongate; ridges with minute granules where crossed by axial pliculae. Aperture subcircular, strongly oblique to vertical axis of shell; peristome virtually complete, somewhat angled in parietal region; aperture expanding slightly prior to lip and lip also weakly reflected.
</paragraph>
<paragraph id="8BAAAE2AE527D919069DFD45477BB40D" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">Ground colour dirty white to yellowish buff; the freshest specimens retaining evidence of a pattern of fine orange-brown spiral lines above the periphery, that at periphery strongest; similar lines just below periphery but umbilicus and most of base lacking colour pattern.</paragraph>
<paragraph id="8BAAAE2AE527D919069DFCC347E9B4D3" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">
Dimensions:
<typeStatus id="54AE1088E527D9190728FCC347F1B42C" box="[357,464,853,879]" pageId="14" pageNumber="14" type="holotype">Holotype</typeStatus>
, max. diameter
<quantity id="4CED03CFE527D91904CFFCC344C9B433" box="[642,744,853,880]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="14" pageNumber="14" unit="mm" value="37.3">37.3 mm</quantity>
, height
<quantity id="4CED03CFE527D919050CFCC34589B433" box="[833,936,853,880]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="14" pageNumber="14" unit="mm" value="18.9">18.9 mm</quantity>
; largest specimen, max. diameter
<quantity id="4CED03CFE527D9190710FCE347E5B4D3" box="[349,452,885,912]" metricMagnitude="-2" metricUnit="m" metricValue="4.14" pageId="14" pageNumber="14" unit="mm" value="41.4">41.4 mm</quantity>
.
</paragraph>
<paragraph id="8BAAAE2AE527D91906F8FC0D4490B373" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">
<materialsCitation id="3B7DA477E527D91906F8FC0D4253B4A3" collectingDate="2010-09-22" collectionCode="AMS" collectorName="O. Griffiths &amp; R. Randalana" country="Madagascar" elevation="215" latitude="-17.90517" location="Belitsaka" longLatPrecision="1" longitude="44.4824" pageId="14" pageNumber="14" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AE1088E527D91906F8FC0D472CB4F3" box="[181,269,923,944]" pageId="14" pageNumber="14" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F302EEBAE527D919075BFC0D479AB4F3" box="[278,443,923,944]" name="Madagascar" pageId="14" pageNumber="14">MADAGASCAR</collectingCountry>
:
<collectorName id="26E0CBFCE527D9190789FC0D44B6B4F3" box="[452,663,923,944]" pageId="14" pageNumber="14">Central W Madagascar</collectorName>
,
<emphasis id="B9617238E527D91904EDFC0B4494B4F3" box="[672,693,925,944]" italics="true" pageId="14" pageNumber="14">ca</emphasis>
<locationDeviation id="9A18B708E527D91904F1FC0D45B1B4F3" box="[700,912,923,944]" location="Maintirano" pageId="14" pageNumber="14">
<quantity id="4CED03CFE527D91904F1FC0D44D7B4F3" box="[700,758,923,944]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="14" unit="km" value="60.0">60 km</quantity>
E of Maintirano
</locationDeviation>
, NE of
<location id="8ECAF8F1E527D9190595FC0D420EB4F3" LSID="urn:lsid:plazi:treatment:03BC1F3CE524D9180783FEF84220B666:8ECAF8F1E527D9190595FC0D420EB4F3" box="[984,1071,923,944]" country="Madagascar" latitude="-17.90517" longLatPrecision="1" longitude="44.4824" name="Belitsaka" pageId="14" pageNumber="14">Belitsaka</location>
, central part of
<taxonomicName id="4C15D5A9E527D91906B8FC25475FB48B" authorityName=", Antsingimavo" authorityYear="2013" box="[245,382,947,968]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, in slots and small caves above
<collectorName id="26E0CBFCE527D91904D7FC25452DB48B" box="[666,780,947,968]" pageId="14" pageNumber="14">Bokarano R.</collectorName>
gorge, next to low dry deciduous forest,
<geoCoordinate id="EE21C8EDE527D91906F8FC5D4701B4A3" box="[181,288,971,992]" degrees="17.90517" direction="south" orientation="latitude" pageId="14" pageNumber="14" precision="1" value="-17.90517">17.90517°S</geoCoordinate>
<geoCoordinate id="EE21C8EDE527D919076BFC5D47B5B4A3" box="[294,404,971,992]" degrees="44.48240" direction="east" orientation="longitude" pageId="14" pageNumber="14" precision="1" value="44.4824">44.48240°E</geoCoordinate>
,
<emphasis id="B9617238E527D91907D0FC5B4793B4A3" box="[413,434,973,992]" italics="true" pageId="14" pageNumber="14">ca</emphasis>
<quantity id="4CED03CFE527D91907FAFC5D47D2B4A3" box="[439,499,971,992]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="14" pageNumber="14" unit="m" value="215.0">
<elevation id="00384919E527D91907FAFC5D47D2B4A3" box="[439,499,971,992]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="14" pageNumber="14" unit="m" value="215.0">215 m</elevation>
</quantity>
,
<date id="FFAB88EAE527D91907B6FC5D447FB4A3" box="[507,606,971,992]" pageId="14" pageNumber="14" value="2010-09-22">
<collectingDate id="EFEF7102E527D91907B6FC5D447FB4A3" box="[507,606,971,992]" pageId="14" pageNumber="14" value="2010-09-22">22.ix.2010</collectingDate>
</date>
,
<collectorName id="26E0CBFCE527D919042AFC5D44EEB4A3" box="[615,719,971,992]" pageId="14" pageNumber="14">O. Griffiths</collectorName>
&amp;
<collectorName id="26E0CBFCE527D91904A1FC5D4549B4A3" box="[748,872,971,992]" pageId="14" pageNumber="14">R. Randalana</collectorName>
, st’n 09/10 (
<collectionCode id="ED0436EFE527D9190595FC5D422BB4A3" box="[984,1034,971,992]" pageId="14" pageNumber="14">AMS</collectionCode>
C.474166)
</materialsCitation>
.
<materialsCitation id="3B7DA477E527D91906F8FC7D448CB373" collectingDate="2010-09-22" collectionCode="NMSA, AMS, TMAM, MHNP" collectorName="O. Griffiths &amp; R. Randalana" country="Madagascar" elevation="215" latitude="-17.90517" location="Belitsaka" longLatPrecision="1" longitude="44.4824" pageId="14" pageNumber="14" specimenCode="L8524, T161" specimenCount="20" specimenCount-adult="17" typeStatus="paratype">
<typeStatus id="54AE1088E527D91906F8FC7D4730B343" box="[181,273,1003,1024]" pageId="14" pageNumber="14" type="paratype">Paratypes</typeStatus>
: Same data as holotype (
<collectionCode id="ED0436EFE527D91907A6FC7D440CB343" box="[491,557,1003,1024]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="14" pageNumber="14" type="Museum">NMSA</collectionCode>
<specimenCode id="DBB30651E527D919047CFC7D444FB343" box="[561,622,1003,1024]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="14" pageNumber="14" type="Museum">L8524</specimenCode>
/T2979,
<specimenCount id="9D1365A3E527D91904F5FC7D450AB343" box="[696,811,1003,1024]" pageId="14" pageNumber="14" type="generic">3 specimens</specimenCount>
); st’n 06/09 (
<collectionCode id="ED0436EFE527D91905D3FC7D45F1B343" box="[926,976,1003,1024]" pageId="14" pageNumber="14">AMS</collectionCode>
C.469585,
<specimenCount id="9D1365A3E527D9190275FC7D4254B343" box="[1080,1141,1003,1024]" pageId="14" pageNumber="14" type="adult">3 adult</specimenCount>
specimens); st’n 07/09 (
<collectionCode id="ED0436EFE527D91907C3FB9547F7B35B" box="[398,470,1027,1048]" pageId="14" pageNumber="14">TMAM</collectionCode>
<specimenCode id="DBB30651E527D9190796FB95442DB35B" box="[475,524,1027,1048]" collectionCode="TMAM" pageId="14" pageNumber="14">T161</specimenCode>
,
<specimenCount id="9D1365A3E527D919045BFB954475B35B" box="[534,596,1027,1048]" pageId="14" pageNumber="14" type="adult">1 adult</specimenCount>
specimen); st’n 08/09 (
<collectionCode id="ED0436EFE527D9190567FB95457DB35B" box="[810,860,1027,1048]" pageId="14" pageNumber="14">AMS</collectionCode>
C.469586,
<specimenCount id="9D1365A3E527D919058BFB954222B35B" box="[966,1027,1027,1048]" pageId="14" pageNumber="14" type="adult">9 adult</specimenCount>
specimens); st’n 08/10 (
<collectionCode id="ED0436EFE527D9190751FB8D477FB373" box="[284,350,1051,1072]" country="France" name="Museum d'Histoire Naturelle Perpignan" pageId="14" pageNumber="14">MHNP</collectionCode>
IM-2010-20083,
<specimenCount id="9D1365A3E527D919044DFB8D441FB373" box="[512,574,1051,1072]" pageId="14" pageNumber="14" type="adult">4 adult</specimenCount>
specimens)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAAAE2AE527D91906F8FBAD4531B313" blockId="14.[181,1144,203,1674]" box="[181,784,1083,1104]" pageId="14" pageNumber="14">
Additional locality data:
<taxonomicName id="4C15D5A9E527D91907DAFBAB443AB313" authorityName=", Antsingimavo" authorityYear="2013" box="[407,539,1085,1104]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="beanka">
<emphasis id="B9617238E527D91907DAFBAB443AB313" box="[407,539,1085,1104]" italics="true" pageId="14" pageNumber="14">Tsingy Beanka</emphasis>
</taxonomicName>
: st’ns 06/10, 07/10, 09/12.
</paragraph>
<paragraph id="8BAAAE2AE527D91906F8FBC9478AB3DA" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">
Distribution: A narrow-range endemic; currently known only from the central region of the
<taxonomicName id="4C15D5A9E527D91906B0FBE94786B3DA" authorityName=", Antsingimavo" authorityYear="2013" box="[253,423,1151,1177]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
.
</paragraph>
<paragraph id="8BAAAE2AE527D91906F8FB314783B3A2" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">Habitat: Found only in subfossil form in deep slots and crevices within the tsingy, in dry deciduous forest.</paragraph>
<paragraph id="8BAAAE2AE527D91906F8FB794572B16A" blockId="14.[181,1144,203,1674]" pageId="14" pageNumber="14">
Remarks: Like
<taxonomicName id="4C15D5A9E527D9190729FB6447FBB249" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[356,474,1266,1290]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="humbug">
<emphasis id="B9617238E527D9190729FB6447FBB249" box="[356,474,1266,1290]" italics="true" pageId="14" pageNumber="14">T. humbug</emphasis>
</taxonomicName>
, this species is referable to the ‘groupe d’espèces du
<taxonomicName id="4C15D5A9E527D9190278FB6444A0B26A" authority="(Petit de la Saussaye, 1844)" baseAuthorityName="Petit de la Saussaye" baseAuthorityYear="1844" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="deshayesiana">
<emphasis id="B9617238E527D9190278FB644710B269" italics="true" pageId="14" pageNumber="14">T. deshayesiana</emphasis>
(Petit de la Saussaye, 1844)
</taxonomicName>
’ of Fischer-Piette
<emphasis id="B9617238E527D919051BFA8445ABB269" box="[854,906,1298,1322]" italics="true" pageId="14" pageNumber="14">et al</emphasis>
. (1993). Within this group, it is closest to
<taxonomicName id="4C15D5A9E527D91907E8FAA44520B20A" authority="Fischer-Piette, 1949" authorityName=": Fischer-Piette" authorityYear="1949" box="[421,769,1327,1354]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E527D91907E8FAA4443BB209" box="[421,538,1330,1354]" italics="true" pageId="14" pageNumber="14">T. chavani</emphasis>
Fischer-Piette, 1949
</taxonomicName>
(see above) in having a very low spire, relatively fine spiral sculpture and a colour pattern of narrow, brownish, spiral bands. It is, however, considerably larger than
<taxonomicName id="4C15D5A9E527D91904FEFAE44507B2C9" authorityName=": Fischer-Piette" authorityYear="1949" box="[691,806,1394,1418]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E527D91904FEFAE44507B2C9" box="[691,806,1394,1418]" italics="true" pageId="14" pageNumber="14">T. chavani</emphasis>
</taxonomicName>
, attaining as much as
<quantity id="4CED03CFE527D919025BFAF94257B2C9" box="[1046,1142,1391,1418]" metricMagnitude="-2" metricUnit="m" metricValue="4.14" pageId="14" pageNumber="14" unit="mm" value="41.4">41.4mm</quantity>
in max. diameter (compared to
<quantity id="4CED03CFE527D919046DFA194450B2E9" box="[544,625,1423,1450]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="14" unit="mm" value="32.0">32 mm</quantity>
for
<taxonomicName id="4C15D5A9E527D91904EEFA04453BB2E9" authorityName=": Fischer-Piette" authorityYear="1949" box="[675,794,1426,1450]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E527D91904EEFA04453BB2E9" box="[675,794,1426,1450]" italics="true" pageId="14" pageNumber="14">T. chavani</emphasis>
</taxonomicName>
). Furthermore, its sculpture is considerably finer than that of
<taxonomicName id="4C15D5A9E527D919044BFA244458B289" authorityName=": Fischer-Piette" authorityYear="1949" box="[518,633,1458,1482]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E527D919044BFA244458B289" box="[518,633,1458,1482]" italics="true" pageId="14" pageNumber="14">T. chavani</emphasis>
</taxonomicName>
, with subequal spiral and axial elements. In
<taxonomicName id="4C15D5A9E527D919022CFA24472FB2A9" authorityName=": Fischer-Piette" authorityYear="1949" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E527D919022CFA24472FB2A9" italics="true" pageId="14" pageNumber="14">T. chavani</emphasis>
</taxonomicName>
the axial pliculae are crisp, and much finer and more close-set than the spiral cords (compare
<figureCitation id="132EB2AFE527D919072BFA794432B14A" box="[358,531,1519,1545]" captionStart="Fig" captionStartId="13.[144,177,1606,1627]" captionTargetBox="[153,1096,661,1587]" captionTargetId="figure-180@13.[151,1097,654,1588]" captionTargetPageId="13" captionText="Fig.9. Tropidophora sericea sp.n.and T. chavani: (A–C) T.sericea, holotype,Tsingy Beanka,max.diameter 37.3mm, height18.9 mm,AMS C.474166;(D, E) adult teleoconch sculpture of T. sericea (holotype) and T. chavani (NMSA L8447) respectively, scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7670122" httpUri="https://zenodo.org/record/7670122/files/figure.png" pageId="14" pageNumber="14">Figs 9D and 9E</figureCitation>
).At some localities,
<taxonomicName id="4C15D5A9E527D91904BBFA644549B149" authorityName=": Fischer-Piette" authorityYear="1949" box="[758,872,1522,1546]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="chavani">
<emphasis id="B9617238E527D91904BBFA644549B149" box="[758,872,1522,1546]" italics="true" pageId="14" pageNumber="14">T. chavani</emphasis>
</taxonomicName>
occurs in subfossil form together with
<taxonomicName id="4C15D5A9E527D919071CF984479CB169" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[337,445,1554,1578]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="sericea">
<emphasis id="B9617238E527D919071CF984479CB169" box="[337,445,1554,1578]" italics="true" pageId="14" pageNumber="14">T. sericea</emphasis>
</taxonomicName>
, but the two remain clearly distinct.
</paragraph>
<paragraph id="8BAAAE2AE527D918069DF9A44220B666" blockId="14.[181,1144,203,1674]" lastBlockId="15.[144,1104,203,294]" lastPageId="15" lastPageNumber="15" pageId="14" pageNumber="14">
<taxonomicName id="4C15D5A9E527D919069DF9A4440AB10A" authority="(Grateloup, 1840)" baseAuthorityName="Grateloup" baseAuthorityYear="1840" box="[208,555,1583,1610]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="moulinsii">
<emphasis id="B9617238E527D919069DF9A44777B109" box="[208,342,1586,1610]" italics="true" pageId="14" pageNumber="14">T. moulinsii</emphasis>
(Grateloup, 1840)
</taxonomicName>
(
<figureCitation id="132EB2AFE527D9190470F9B944EBB10A" box="[573,714,1583,1609]" captionStart="Fig" captionStartId="37.[144,176,1510,1531]" captionTargetBox="[157,1095,235,1476]" captionTargetId="figure-89@37.[147,1108,223,1487]" captionTargetPageId="37" captionText="Fig.26.Speciesillustrated for comparative purposes:(A–C) Tropidophora moulinsii, holotype of Cyclostoma moulinsii Grateloup, 1840, ‘Madagascar’, max. diameter 33.1 mm, NHMUK 1907.11.22.12; (D–F) Tropidophora thesauri Fischer-Piette, 1949, Orangea, Diego Suarez [= Antsiranana] district, max. diameter 38.0 mm, NMSA L8438; (G–I) Tropidophora reticulata, one of three syntypes of Cyclostoma reticulatum (Adams &amp; Reeve, 1850), ‘Madagascar’, max.diameter 15.1 mm, NHMUK 1874.12.11.231; (J–L) Kalidos hova, syntype of Nanina hova Odhner, 1919, Catsèpe, Madagascar, max. diameter 24.4 mm (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.7670163" httpUri="https://zenodo.org/record/7670163/files/figure.png" pageId="14" pageNumber="14">Fig. 26A–C</figureCitation>
,
<typeStatus id="54AE1088E527D9190495F9B9451CB10A" box="[728,829,1583,1609]" pageId="14" pageNumber="14" type="holotype">holotype</typeStatus>
, NHMUK) and
<taxonomicName id="4C15D5A9E527D91905B4F9A447B6B12A" authority="Fischer-Piette, 1949" authorityName="Fischer-Piette" authorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="thesauri">
<emphasis id="B9617238E527D91905B4F9A44228B109" box="[1017,1033,1586,1610]" italics="true" pageId="14" pageNumber="14">T</emphasis>
.
<emphasis id="B9617238E527D9190257F9A44257B109" box="[1050,1142,1586,1610]" italics="true" pageId="14" pageNumber="14">thesauri</emphasis>
Fischer-Piette, 1949
</taxonomicName>
(
<figureCitation id="132EB2AFE527D91907EBF9D9440FB12A" box="[422,558,1615,1641]" captionStart="Fig" captionStartId="37.[144,176,1510,1531]" captionTargetBox="[157,1095,235,1476]" captionTargetId="figure-89@37.[147,1108,223,1487]" captionTargetPageId="37" captionText="Fig.26.Speciesillustrated for comparative purposes:(A–C) Tropidophora moulinsii, holotype of Cyclostoma moulinsii Grateloup, 1840, ‘Madagascar’, max. diameter 33.1 mm, NHMUK 1907.11.22.12; (D–F) Tropidophora thesauri Fischer-Piette, 1949, Orangea, Diego Suarez [= Antsiranana] district, max. diameter 38.0 mm, NMSA L8438; (G–I) Tropidophora reticulata, one of three syntypes of Cyclostoma reticulatum (Adams &amp; Reeve, 1850), ‘Madagascar’, max.diameter 15.1 mm, NHMUK 1874.12.11.231; (J–L) Kalidos hova, syntype of Nanina hova Odhner, 1919, Catsèpe, Madagascar, max. diameter 24.4 mm (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.7670163" httpUri="https://zenodo.org/record/7670163/files/figure.png" pageId="14" pageNumber="14">Fig. 26D–F</figureCitation>
), both from the north-eastern tip of
<collectingCountry id="F302EEBAE527D91905F2F9D94269B12A" box="[959,1096,1615,1641]" name="Madagascar" pageId="14" pageNumber="14">Madagascar</collectingCountry>
, are of a more similar size (attaining
<quantity id="4CED03CFE527D919046DF9F9444FB1C9" box="[544,622,1647,1674]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="14" unit="mm" value="38.0">38 mm</quantity>
or more in max. diameter), but
<taxonomicName id="4C15D5A9E527D919059DF9E4426BB1C9" authorityName="Fischer-Piette" authorityYear="1949" box="[976,1098,1648,1674]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="14" phylum="Mollusca" rank="species" species="thesauri">
<emphasis id="B9617238E527D919059DF9E445C1B1C9" box="[976,992,1650,1674]" italics="true" pageId="14" pageNumber="14">T</emphasis>
.
<emphasis id="B9617238E527D91905A3F9E4426BB1C9" box="[1006,1098,1650,1674]" italics="true" pageId="14" pageNumber="14">thesauri</emphasis>
</taxonomicName>
has much deeper whorls, a higher spire, narrower umbilicus, irregular scale-like subsutural pliculae and lacks the finely decussate microsculpture of
<taxonomicName id="4C15D5A9E526D9180553FF7845ADB645" authorityName="Griffiths &amp; Herbert" authorityYear="2013" box="[798,908,238,262]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="15" phylum="Mollusca" rank="species" species="sericea">
<emphasis id="B9617238E526D9180553FF7845ADB645" box="[798,908,238,262]" italics="true" pageId="15" pageNumber="15">T. sericea</emphasis>
</taxonomicName>
.
<taxonomicName id="4C15D5A9E526D91805D1FF784200B645" baseAuthorityName="Grateloup" baseAuthorityYear="1840" box="[924,1057,238,262]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="15" pageNumber="15" phylum="Mollusca" rank="species" species="moulinsii">
<emphasis id="B9617238E526D91805D1FF784200B645" box="[924,1057,238,262]" italics="true" pageId="15" pageNumber="15">T. moulinsii</emphasis>
</taxonomicName>
has stronger, cord-like, spiral sculpture, more tumescent whorls and a higher spire.
</paragraph>
</subSubSection>
</treatment>
</document>
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