269 lines
32 KiB
XML
269 lines
32 KiB
XML
<document id="E065C1DFADD738E1C71BE306B72048D1" ID-DOI="10.5281/zenodo.183097" ID-GBIF-Dataset="df4cf84e-9a79-465b-87f2-c0fd4a7a98ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="183097" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459890880288" checkinUser="plazi" docAuthor="Brancucci, Michel & Hendrich, Lars" docDate="2008" docId="03FBE052FFF0FF9FFF38FCEA7129FC76" docLanguage="en" docName="zt01825p058.pdf" docOrigin="Zootaxa 1825" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Agabus joachimschmidti Brancucci & Hendrich, 2008, sp. n." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FFC2982AFFF1FF9CFFAFFFDF745EFF8C" masterDocTitle="5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae)" masterLastPageNumber="58" masterPageNumber="51" pageNumber="52" updateTime="1698228027061" updateUser="plazi">
|
||
<mods:mods id="6341734C0F88F92CDCE1C955D3B60440" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B1AA43376452436C79AFDD615683A60F">
|
||
<mods:title id="64F3551E59E49E642844BA44DF26A6C4">5100 m above sea level: Agabus joachimschmidti sp. n. and notes on other high altitude diving beetles from Tibet and Bhutan (Coleoptera, Dytiscidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6D48423110B456DA2B509C5AF33664E6" type="personal">
|
||
<mods:role id="05C4AF243A28F52C2F0D7E246BD6B7E2">
|
||
<mods:roleTerm id="B5148A077389A5ED33DC823847972D63">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F4DCE92C167C71FE939EE257B9DEF1A9">Brancucci, Michel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8321D47862DB8CB1C0C2B03970623196" type="personal">
|
||
<mods:role id="B2665AA8B92770E5833B109ECE881B83">
|
||
<mods:roleTerm id="4642345E97F2003D04E73DBD4DA3820A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8B9FDF0931F159C8F4138A2048728A62">Hendrich, Lars</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="64E489FFBD98A7D0601AD176374EBDEC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F3B86851127AB69CB95F4101C9EB4D18" type="host">
|
||
<mods:titleInfo id="6333FA2EC3C2C05A8B17259A5A9C4FCB">
|
||
<mods:title id="1688A8F3AA827C811A957974EE8155CE">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="299FA124BDB62B621529401B550DA80C">
|
||
<mods:date id="C2172DF4292608B6DAAC5B5600C3319F">2008</mods:date>
|
||
<mods:detail id="28ABFA25CF9F4A55C05567E5E22725ED" type="volume">
|
||
<mods:number id="03F8824EC811927CDC755D4CE5351F58">1825</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="5715B4E7F09E87368922201BB0E5A03F" unit="page">
|
||
<mods:start id="177F349A65A412D9D10BF264EF80965D">51</mods:start>
|
||
<mods:end id="4750F1BC8645ABB166ECB259B5DEBEB8">58</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="81C0D7FA330BE33D9D55233896C98EB1">journal article</mods:classification>
|
||
<mods:identifier id="D1561D811E8C679B11D311018F1994EE" type="DOI">10.5281/zenodo.183097</mods:identifier>
|
||
<mods:identifier id="786B594C5B4E94116399FB9F3E4B1B22" type="GBIF-Dataset">df4cf84e-9a79-465b-87f2-c0fd4a7a98ca</mods:identifier>
|
||
<mods:identifier id="0DC5E64FCD37A7C75B3C1A82CE9F2FF7" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="F07941A0C851BC205E29F5D187152699" type="Zenodo-Dep">183097</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03FBE052FFF0FF9FFF38FCEA7129FC76" ID-DOI="http://doi.org/10.5281/zenodo.6235827" ID-GBIF-Taxon="119371980" ID-Zenodo-Dep="6235827" LSID="urn:lsid:plazi:treatment:03FBE052FFF0FF9FFF38FCEA7129FC76" httpUri="http://treatment.plazi.org/id/03FBE052FFF0FF9FFF38FCEA7129FC76" lastPageId="3" lastPageNumber="54" pageId="1" pageNumber="52">
|
||
<subSubSection id="C34802CFFFF0FF9DFF38FCEA755AFCFE" pageId="1" pageNumber="52" type="nomenclature">
|
||
<paragraph id="8BED5144FFF0FF9DFF38FCEA7655FCC3" blockId="1.[151,523,821,882]" box="[151,523,821,847]" pageId="1" pageNumber="52">
|
||
<heading id="D0A5E628FFF0FF9DFF38FCEA7655FCC3" bold="true" box="[151,523,821,847]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
|
||
<emphasis id="B9268D56FFF0FF9DFF38FCEA7655FCC3" bold="true" box="[151,523,821,847]" pageId="1" pageNumber="52">
|
||
<taxonomicName id="4C522AC7FFF0FF9DFF38FCEA759FFCC3" ID-CoL="7QDNW" box="[151,449,821,847]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="joachimschmidti" status="sp. nov.">
|
||
<emphasis id="B9268D56FFF0FF9DFF38FCEA759FFCC3" bold="true" box="[151,449,821,847]" italics="true" pageId="1" pageNumber="52">Agabus joachimschmidti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A215302DFFF0FF9DFE67FCEA7655FCC3" box="[456,523,821,847]" pageId="1" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF0FF9DFF31FC87755AFCFE" blockId="1.[151,523,821,882]" box="[158,260,856,882]" pageId="1" pageNumber="52">
|
||
<figureCitation id="13694DC1FFF0FF9DFF31FC87755AFCFE" box="[158,260,856,882]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="1" pageNumber="52">Figs 1–5</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34802CFFFF0FF9DFF38FC78751AFB06" pageId="1" pageNumber="52" type="materials_examined">
|
||
<paragraph id="8BED5144FFF0FF9DFF38FC787000FC4E" blockId="1.[151,1436,935,2002]" box="[151,1118,935,962]" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF38FC787562FC4D" bold="true" box="[151,316,935,961]" pageId="1" pageNumber="52">
|
||
<typeStatus id="54E9EFE6FFF0FF9DFF38FC78748AFC4D" box="[151,212,935,961]" pageId="1" pageNumber="52">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Small spring, Budha Valley N of Yangpachem, Tibet South Central.
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF0FF9DFF6AFC10751AFB06" blockId="1.[151,1436,935,2002]" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF6AFC107580FC65" bold="true" box="[197,478,975,1001]" pageId="1" pageNumber="52">
|
||
<typeStatus id="54E9EFE6FFF0FF9DFF6AFC10755CFC65" box="[197,258,975,1001]" pageId="1" pageNumber="52">Type</typeStatus>
|
||
material studied.
|
||
</emphasis>
|
||
<typeStatus id="54E9EFE6FFF0FF9DFE49FC0F760DFC66" box="[486,595,976,1002]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
|
||
male (
|
||
<collectionCode id="ED43C981FFF0FF9DFD0BFC0F76B3FC66" box="[676,749,976,1002]" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="1" pageNumber="52">NMB</collectionCode>
|
||
): “
|
||
<collectionCode id="ED43C981FFF0FF9DFCA4FC0F773FFC66" box="[779,865,976,1002]" pageId="1" pageNumber="52">TIBET</collectionCode>
|
||
South Central, Budha Valley N of Yangpachem, about 30°10'38
|
||
<geoCoordinate id="EE663783FFF0FF9DFEE7FC2775E8FB9E" box="[328,438,1016,1042]" direction="north" orientation="latitude" pageId="1" pageNumber="52" precision="925" value="90.5">N 90°30'</geoCoordinate>
|
||
18E,
|
||
<date id="FFEC7784FFF0FF9DFE5BFC277622FB9E" box="[500,636,1016,1042]" pageId="1" pageNumber="52" value="2007-06-20">20.VI.2007</date>
|
||
,
|
||
<quantity id="4CAAFCA1FFF0FF9DFD25FC277681FB9E" box="[650,735,1016,1042]" metricMagnitude="3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="52" unit="m" value="5100.0">5100m</quantity>
|
||
” [white printed label]. “
|
||
<typeStatus id="54E9EFE6FFF0FF9DFBAFFC27702EFB9E" box="[1024,1136,1016,1042]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
|
||
<taxonomicName id="4C522AC7FFF0FF9DFBD6FC2771C2FB9D" box="[1145,1436,1016,1041]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="joachimschmidti" status="sp. nov.">
|
||
<emphasis id="B9268D56FFF0FF9DFBD6FC2771C2FB9D" box="[1145,1436,1016,1041]" italics="true" pageId="1" pageNumber="52">Agabus joachimschmidti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A215302DFFF0FF9DFF38FBFF7490FBB6" box="[151,206,1056,1082]" pageId="1" pageNumber="52" rank="species">sp.n.</taxonomicNameLabel>
|
||
Brancucci & Hendrich des. 2008” [red printed label].
|
||
<typeStatus id="54E9EFE6FFF0FF9DFCE6FBFF77E1FBB6" box="[841,959,1056,1082]" pageId="1" pageNumber="52" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D549ACDFFF0FF9DFC65FBFF7007FBB6" box="[970,1113,1056,1082]" pageId="1" pageNumber="52" type="generic">9 specimens</specimenCount>
|
||
with same data as
|
||
<typeStatus id="54E9EFE6FFF0FF9DFA99FBFF71C2FBB6" box="[1334,1436,1056,1082]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCount id="9D549ACDFFF0FF9DFF0FFB9774A0FBEE" box="[160,254,1096,1122]" pageId="1" pageNumber="52" type="male">3 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D549ACDFFF0FF9DFE91FB9775ECFBEE" box="[318,434,1096,1122]" pageId="1" pageNumber="52" type="female">4 females</specimenCount>
|
||
<collectionCode id="ED43C981FFF0FF9DFE13FB97765CFBEE" box="[444,514,1096,1122]" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="1" pageNumber="52">NMB</collectionCode>
|
||
,
|
||
<specimenCount id="9D549ACDFFF0FF9DFDBEFB97763AFBEE" box="[529,612,1096,1122]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
|
||
and
|
||
<specimenCount id="9D549ACDFFF0FF9DFD0CFB977752FBEE" box="[675,780,1096,1122]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
|
||
<collectionCode id="ED43C981FFF0FF9DFCB8FB977709FBEE" box="[791,855,1096,1122]" pageId="1" pageNumber="52">CLH</collectionCode>
|
||
). All specimens are provided with a red printed
|
||
<typeStatus id="54E9EFE6FFF0FF9DFF38FBAF74A2FB06" box="[151,252,1136,1162]" pageId="1" pageNumber="52" type="paratype">paratype</typeStatus>
|
||
label.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34802CFFFF0FF9FFF6AFB487066FEA6" lastPageId="3" lastPageNumber="54" pageId="1" pageNumber="52" type="description">
|
||
<paragraph id="8BED5144FFF0FF9DFF6AFB487141FA8E" blockId="1.[151,1436,935,2002]" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF6AFB487678FB3D" bold="true" box="[197,550,1175,1201]" pageId="1" pageNumber="52">Description. Measurements:</emphasis>
|
||
<typeStatus id="54E9EFE6FFF0FF9DFD82FB4776C1FB3E" box="[557,671,1176,1202]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
|
||
: total length =
|
||
<quantity id="4CAAFCA1FFF0FF9DFCE2FB4777E6FB3E" box="[845,952,1176,1202]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="52" unit="mm" value="6.2">6.20 mm</quantity>
|
||
, total length-head =
|
||
<quantity id="4CAAFCA1FFF0FF9DFB08FB47714DFB3E" box="[1191,1299,1176,1202]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="52" unit="mm" value="5.6">5.60 mm</quantity>
|
||
, total width =
|
||
<quantity id="4CAAFCA1FFF0FF9DFF00FB1F7542FB56" box="[175,284,1216,1242]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="52" unit="mm" value="3.2">3.20 mm</quantity>
|
||
.
|
||
<typeStatus id="54E9EFE6FFF0FF9DFE88FB1F75C7FB56" box="[295,409,1216,1242]" pageId="1" pageNumber="52" type="paratype">Paratypes</typeStatus>
|
||
(male and female do not differ in size): total length =
|
||
<quantity id="4CAAFCA1FFF0FF9DFBB5FB1F709BFB56" box="[1050,1221,1216,1242]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.1" metricValueMin="5.6" pageId="1" pageNumber="52" unit="mm" value="5.85" valueMax="6.1" valueMin="5.6">5.60–6.10 mm</quantity>
|
||
(
|
||
<quantity id="4CAAFCA1FFF0FF9DFB7BFB1F711CFB56" box="[1236,1346,1216,1242]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" pageId="1" pageNumber="52" unit="mm" value="5.85">5.85 mm</quantity>
|
||
, n = 7), total length-head =
|
||
<quantity id="4CAAFCA1FFF0FF9DFED3FB377678FA8E" box="[380,550,1256,1282]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="1" pageNumber="52" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.20–5.50 mm</quantity>
|
||
(
|
||
<quantity id="4CAAFCA1FFF0FF9DFD9AFB3776FDFA8E" box="[565,675,1256,1282]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" pageId="1" pageNumber="52" unit="mm" value="5.35">5.35 mm</quantity>
|
||
, n = 7), total width =
|
||
<quantity id="4CAAFCA1FFF0FF9DFC0FFB377014FA8E" box="[928,1098,1256,1282]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="1" pageNumber="52" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.90–3.20 mm</quantity>
|
||
(
|
||
<quantity id="4CAAFCA1FFF0FF9DFBF6FB377099FA8E" box="[1113,1223,1256,1282]" metricMagnitude="-3" metricUnit="m" metricValue="3.08" pageId="1" pageNumber="52" unit="mm" value="3.08">3.08 mm</quantity>
|
||
, n = 4).
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF0FF9DFF6AFAD074E8FADE" blockId="1.[151,1436,935,2002]" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF6AFAD0756CFAA5" bold="true" box="[197,306,1295,1321]" pageId="1" pageNumber="52">Habitus:</emphasis>
|
||
Body short-oval, distinctly convex, black on head and pronotum and dark brown on elytra (
|
||
<figureCitation id="13694DC1FFF0FF9DFAC3FACF74FBFADE" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF0FF9DFF6AFA8077EDF996" blockId="1.[151,1436,935,2002]" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF6AFA80754FFAF5" bold="true" box="[197,273,1375,1401]" pageId="1" pageNumber="52">Head:</emphasis>
|
||
Black, shining, labrum testaceous. Antennae testaceous at base, becoming darker brown on distal part from the 4th joint on. Reticulation consisting of medium-sized and well-impressed polygonal meshes, irregular in size, with a small puncture on the inner side of some meshes. Row of punctures alongside eyes and clypeal grooves well-impressed. A few punctures beside eyes well-impressed but not confluent. Antennae short, joints almost globose, the 5th only 1.2 times as long as broad.
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF0FF9DFF6AF9F8767EF8BE" blockId="1.[151,1436,935,2002]" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF6AF9F87510F9CD" bold="true" box="[197,334,1575,1601]" pageId="1" pageNumber="52">Pronotum:</emphasis>
|
||
Black, shining. Reticulation consisting of polygonal meshes, irregular in size and often incomplete particularly on disc; some meshes with 1 very minute puncture, seldom more, on their inner surfaces. Latero-basal part with few deep wrinkles. Anterior row of punctures complete; punctures very large, confluent and in groups, leaving small spaces free. Posterior row broadly interrupted at middle; punctures large, irregularly distributed, confluent at sides. Lateral margin distinctly bordered with punctures alongside the grooves. Sides of pronotum rather straight, forming a distinct pronoto-elytral angle, base of pronotum narrower than base of elytra (
|
||
<figureCitation id="13694DC1FFF0FF9DFE65F8C77651F8BE" box="[458,527,1816,1842]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF0FF9EFF6AF8E07711FF56" blockId="1.[151,1436,935,2002]" lastBlockId="2.[151,1436,152,218]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
|
||
<emphasis id="B9268D56FFF0FF9DFF6AF8E07541F8D5" bold="true" box="[197,287,1855,1881]" pageId="1" pageNumber="52">Elytra:</emphasis>
|
||
Brownish-black with a distinct bronze lustre. Epipleura brown. Reticulation consisting of wellimpressed polygonal meshes, with one minute puncture on the inner side of some meshes. Sutural row of punctures restricted to apical two-thirds and consisting of some medium-sized well-spaced punctures. Discal, sublateral and lateral rows not reaching base; punctures well-spaced, here and there in groups. Punctures present in interspaces between discal and sublateral rows. Epipleura broad at base, strongly narrowed by first sternite and evenly but slightly tapered as far as apical part.
|
||
</paragraph>
|
||
<caption id="DF2D01CCFFF3FF9EFF38F9F170D3F9E4" httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="2" pageNumber="53" targetBox="[176,1400,280,1555]" targetPageId="2">
|
||
<paragraph id="8BED5144FFF3FF9EFF38F9F170D3F9E4" blockId="2.[151,1436,1582,1640]" pageId="2" pageNumber="53">
|
||
<emphasis id="B9268D56FFF3FF9EFF38F9F1751DF9CA" bold="true" box="[151,323,1582,1606]" pageId="2" pageNumber="53">FIGURES 1–5.</emphasis>
|
||
Habitus of: 1)
|
||
<taxonomicName id="4C522AC7FFF3FF9EFE47F9F176AFF9C9" box="[488,753,1582,1605]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="joachimschmidti" status="sp. nov.">
|
||
<emphasis id="B9268D56FFF3FF9EFE47F9F176AFF9C9" box="[488,753,1582,1605]" italics="true" pageId="2" pageNumber="53">Agabus joachimschmidti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9268D56FFF3FF9EFD58F9F1776AF9CA" bold="true" box="[759,820,1582,1606]" pageId="2" pageNumber="53">
|
||
<taxonomicNameLabel id="A215302DFFF3FF9EFD58F9F1776AF9CA" box="[759,820,1582,1606]" pageId="2" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 2)
|
||
<taxonomicName id="4C522AC7FFF3FF9EFCCFF9F177BEF9C9" box="[864,992,1582,1605]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF3FF9EFCCFF9F177BEF9C9" box="[864,992,1582,1605]" italics="true" pageId="2" pageNumber="53">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
; median lobe of aedeagus in lateral view of: 3)
|
||
<taxonomicName id="4C522AC7FFF3FF9EFF77F98E75FBF9E4" box="[216,421,1617,1640]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="joachimschmidti" status="sp. nov.">
|
||
<emphasis id="B9268D56FFF3FF9EFF77F98E75FBF9E4" box="[216,421,1617,1640]" italics="true" pageId="2" pageNumber="53">A. joachimschmidti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9268D56FFF3FF9EFE03F98F75B6F9E4" bold="true" box="[428,488,1616,1640]" pageId="2" pageNumber="53">
|
||
<taxonomicNameLabel id="A215302DFFF3FF9EFE03F98F75B6F9E4" box="[428,488,1616,1640]" pageId="2" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, 4)
|
||
<taxonomicName id="4C522AC7FFF3FF9EFDBCF98E76CCF9E4" box="[531,658,1617,1640]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF3FF9EFDBCF98E76CCF9E4" box="[531,658,1617,1640]" italics="true" pageId="2" pageNumber="53">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
; 5) right paramere of
|
||
<taxonomicName id="4C522AC7FFF3FF9EFCD3F98E7014F9E4" box="[892,1098,1617,1640]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="joachimschmidti" status="sp. nov.">
|
||
<emphasis id="B9268D56FFF3FF9EFCD3F98E7014F9E4" box="[892,1098,1617,1640]" italics="true" pageId="2" pageNumber="53">A. joachimschmidti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A215302DFFF3FF9EFBFEF98F70D3F9E4" box="[1105,1165,1616,1640]" pageId="2" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BED5144FFF3FF9EFF6AF94476FDF8D9" blockId="2.[151,1437,1691,2037]" pageId="2" pageNumber="53">
|
||
<emphasis id="B9268D56FFF3FF9EFF6AF944757DF939" bold="true" box="[197,291,1691,1717]" pageId="2" pageNumber="53">Venter:</emphasis>
|
||
Dark ferrugineous-brown, legs testaceous, femora dark brown. Prosternal process lanceolate, very narrow and strongly convex. Metacoxal line complete. Metafemora with a short row of setae, slightly transversely reticulated. Metatibiae with two rows of punctures, one along outer margin and consisting of 7–9 larger punctures each with a strong but short seta, other along inner margin, shortened distally with 4–5 large punctures; ground-surface distinctly shining.
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF3FF9FFF6AF8BC7066FEA6" blockId="2.[151,1437,1691,2037]" lastBlockId="3.[151,1437,152,1018]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
|
||
<emphasis id="B9268D56FFF3FF9EFF6AF8BC759BF8F1" bold="true" box="[197,453,1891,1917]" pageId="2" pageNumber="53">Sexual dimorphism:</emphasis>
|
||
Male with protarsi and mesotarsi distinctly dilated, with numerous small rounded pads. Protarsal and mesotarsal claws almost as long as protarsomere 5 and mesotarsomere 5, respectively. Anal sternite microreticulate on anterior third, somewhat wrinkled and punctured on posterior half. Posterior margin distinctly flattened, finely bordered. Median lobe of aedeagus, in lateral view, regularly curved, bidentate in apical part (
|
||
<figureCitation id="13694DC1FFF2FF9FFED6FF47759CFF3E" box="[377,450,152,178]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="3" pageNumber="54">Fig. 3</figureCitation>
|
||
); in dorsal view, suddenly tapered and prolonged into a sharp point at apex. Basal part of parameres short but narrow (
|
||
<figureCitation id="13694DC1FFF2FF9FFD90FF1F76D6FF56" box="[575,648,192,218]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="3" pageNumber="54">Fig. 5</figureCitation>
|
||
). Female similar to male. Anal sternite distinctly reticulate on posterior half, with coarse and irregularly distributed punctures on both sides of middle. Protarsal and mesotarsal claws only 1.4 and 1.3 times shorter than protarsomere 5 and mesotarsomere 5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34802CFFFF2FF9FFF6AFEE87664FCD6" pageId="3" pageNumber="54" type="etymology">
|
||
<paragraph id="8BED5144FFF2FF9FFF6AFEE8758CFEF6" blockId="3.[151,1437,152,1018]" pageId="3" pageNumber="54">
|
||
<emphasis id="B9268D56FFF2FF9FFF6AFEE875F2FEDD" bold="true" box="[197,428,311,337]" pageId="3" pageNumber="54">Derivatio nominis.</emphasis>
|
||
Dedicated to Joachim Schmidt (Admannshagen,
|
||
<collectingCountry id="F34511D4FFF2FF9FFC43FEE77005FEDE" box="[1004,1115,312,338]" name="Germany" pageId="3" pageNumber="54">Germany</collectingCountry>
|
||
) a well-known specialist of Himalayan ground beetles.
|
||
</paragraph>
|
||
<paragraph id="8BED5144FFF2FF9FFF6AFE587664FCD6" blockId="3.[151,1437,152,1018]" pageId="3" pageNumber="54">
|
||
<emphasis id="B9268D56FFF2FF9FFF6AFE587587FE2D" bold="true" box="[197,473,391,417]" pageId="3" pageNumber="54">Differential diagnosis.</emphasis>
|
||
This species clearly belongs to the
|
||
<taxonomicName id="4C522AC7FFF2FF9FFCD0FE577782FE2D" box="[895,988,392,417]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="confinis">
|
||
<emphasis id="B9268D56FFF2FF9FFCD0FE577782FE2D" box="[895,988,392,417]" italics="true" pageId="3" pageNumber="54">confinis</emphasis>
|
||
</taxonomicName>
|
||
-group as delimited by
|
||
<bibRefCitation id="EFC32CB5FFF2FF9FFB41FE5771C9FE2E" author="Nilsson" box="[1262,1431,392,418]" pageId="3" pageNumber="58" refString="Nilsson, A. (1990) Revisional notes on selected East Palaearctic species of Agabus Leach (Coleoptera, Dytiscidae). Entomologisk Tidskrift, 111, 149 - 161." type="journal article" year="1990">Nilsson (1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC32CB5FFF2FF9FFF38FE6F7488FE46" author="Nilsson" box="[151,214,432,458]" pageId="3" pageNumber="58" refString="Nilsson, A. (2001) World Catalogue of Insects, Volume 3. Dytiscidae (Coleoptera). Apollo Books, Stenstrup, 395 pp." type="book" year="2001">2001</bibRefCitation>
|
||
), mainly characterized by a bifid median lobe and a typical habitus of the species. It is related to
|
||
<taxonomicName id="4C522AC7FFF2FF9FFA2CFE6F755DFE7D" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF2FF9FFA2CFE6F755DFE7D" italics="true" pageId="3" pageNumber="54">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
, but can be easily distinguished by its smaller size. The total length is
|
||
<quantity id="4CAAFCA1FFF2FF9FFB95FE0770FCFE7E" box="[1082,1186,472,498]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" pageId="3" pageNumber="54" unit="mm" value="5.85">5.85 mm</quantity>
|
||
as the mean between all
|
||
<typeStatus id="54E9EFE6FFF2FF9FFF14FDDF7570FD96" box="[187,302,512,538]" pageId="3" pageNumber="54" type="paratype">paratypes</typeStatus>
|
||
, whereas it is
|
||
<quantity id="4CAAFCA1FFF2FF9FFE7BFDDF7670FD96" box="[468,558,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="54" unit="mm" value="6.8">6.8 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C522AC7FFF2FF9FFDFBFDDF7681FD95" box="[596,735,512,537]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF2FF9FFDFBFDDF7681FD95" box="[596,735,512,537]" italics="true" pageId="3" pageNumber="54">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
. The sides of the pronotum are rather straight and not regularly rounded, forming a distinct pronoto-elytral angle, the base of the pronotum being narrower than base of elytra (
|
||
<figureCitation id="13694DC1FFF2FF9FFF42FD8F756AFDE6" box="[237,308,592,618]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="3" pageNumber="54">Fig. 1</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C522AC7FFF2FF9FFEDDFD8F765EFDE5" box="[370,512,592,617]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF2FF9FFEDDFD8F765EFDE5" box="[370,512,592,617]" italics="true" pageId="3" pageNumber="54">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
, the base of pronotum and the base of elytra are approximately of the same width (
|
||
<figureCitation id="13694DC1FFF2FF9FFF43FDA7756EFD1E" box="[236,304,632,658]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="3" pageNumber="54">Fig 2</figureCitation>
|
||
). The posterior enlarged part of the metacoxal processes is provided laterally just with a few larger punctures, whereas it is distinctly bordered in
|
||
<taxonomicName id="4C522AC7FFF2FF9FFD56FD7F77DAFD35" box="[761,900,672,697]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF2FF9FFD56FD7F77DAFD35" box="[761,900,672,697]" italics="true" pageId="3" pageNumber="54">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
. The sutural rows of the elytra are incomplete and just marked by a few larger and sparse punctures, but are complete and very distinct in
|
||
<taxonomicName id="4C522AC7FFF2FF9FFAA3FD1771C5FD6D" box="[1292,1435,712,737]" class="Insecta" family="Dytiscidae" genus="Agabus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="tibetanus">
|
||
<emphasis id="B9268D56FFF2FF9FFAA3FD1771C5FD6D" box="[1292,1435,712,737]" italics="true" pageId="3" pageNumber="54">A. tibetanus</emphasis>
|
||
</taxonomicName>
|
||
according to the re-description by
|
||
<bibRefCitation id="EFC32CB5FFF2FF9FFD94FD2F76ACFC86" author="Nilsson" box="[571,754,752,778]" pageId="3" pageNumber="58" refString="Nilsson, A. (1990) Revisional notes on selected East Palaearctic species of Agabus Leach (Coleoptera, Dytiscidae). Entomologisk Tidskrift, 111, 149 - 161." type="journal article" year="1990">Nilsson (1990)</bibRefCitation>
|
||
. Finally, the anal sternite is less roughly puncturate and wrinkled. The median lobe of aedeagus does not allow a clear identification to be made, but the subapical tooth is more elongate (
|
||
<figureCitation id="13694DC1FFF2FF9FFE04FC9F7673FCD6" box="[427,557,832,858]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,269,1582,1606]" captionTargetBox="[176,1400,280,1555]" captionTargetId="figure@2.[152,1434,277,1555]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: 1) Agabus joachimschmidti sp. n., 2) A. tibetanus; median lobe of aedeagus in lateral view of: 3) A. joachimschmidti sp. n., 4) A. tibetanus; 5) right paramere of A. joachimschmidti sp. n." httpUri="https://zenodo.org/record/183098/files/figure.png" pageId="3" pageNumber="54">Figs 3 & 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34802CFFFF2FF9FFF6AFCB87133FC5E" pageId="3" pageNumber="54" type="description">
|
||
<paragraph id="8BED5144FFF2FF9FFF6AFCB87133FC5E" blockId="3.[151,1437,152,1018]" pageId="3" pageNumber="54">
|
||
<emphasis id="B9268D56FFF2FF9FFF6AFCB87573FC0D" bold="true" box="[197,301,871,897]" pageId="3" pageNumber="54">Habitat.</emphasis>
|
||
These specimens were collected at an altitude of
|
||
<quantity id="4CAAFCA1FFF2FF9FFC2CFCB777BFFC0E" box="[899,993,872,898]" metricMagnitude="3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="54" unit="m" value="5100.0">5100 m</quantity>
|
||
, which may be an altitude record for dytiscids. The habitat was a flat area of debris with water trickling over it, with the source exposed to the south. This appears to be a temporary habitat as the ice-free period at this altitude seems to be quite short.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34802CFFFF2FF9FFF6AFC007129FC76" box="[197,1399,991,1018]" pageId="3" pageNumber="54" type="distribution">
|
||
<paragraph id="8BED5144FFF2FF9FFF6AFC007129FC76" blockId="3.[151,1437,152,1018]" box="[197,1399,991,1018]" pageId="3" pageNumber="54">
|
||
<emphasis id="B9268D56FFF2FF9FFF6AFC007538FC75" bold="true" box="[197,358,991,1017]" pageId="3" pageNumber="54">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54E9EFE6FFF2FF9FFDDEFC3F76FDFC76" box="[625,675,992,1018]" pageId="3" pageNumber="54">type</typeStatus>
|
||
locality but probably more widespread in south central Tibet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |