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<document id="E65A0DBDF6F9E45695234156786F7BBE" ID-DOI="10.5281/zenodo.157855" ID-GBIF-Dataset="2fcc9637-0666-42af-a7ea-47adc189b600" ID-ISSN="1175­5326" ID-Zenodo-Dep="157855" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461015877901" checkinUser="plazi" docAuthor="Pesic, Vladimir, Saboori, Alireza, Asadi, Mahdieh &amp; Vafaei, Reza" docDate="2004" docId="03FB7D20DD61FF83FEAFFED04ED10034" docLanguage="en" docName="zt00495.pdf" docOrigin="Zootaxa 495" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Atractides nikooae Pesic, sp. nov." docType="treatment" docVersion="11" lastPageNumber="24" masterDocId="FFC20558DD74FF94FFA7FFCC4A040462" masterDocTitle="Studies on water mites of the family Hygrobatidae (Acari, Hydrachnidia) from Iran, I. The water mite genus Atractides Koch, with the description of five new species" masterLastPageNumber="40" masterPageNumber="1" pageNumber="22" updateTime="1698568796051" updateUser="plazi">
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<mods:title id="2E2495117F40F9683D92B6492328F5BA">Studies on water mites of the family Hygrobatidae (Acari, Hydrachnidia) from Iran, I. The water mite genus Atractides Koch, with the description of five new species</mods:title>
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<mods:namePart id="2B1E6213F3E5FCAF44B496768156F52A">Pesic, Vladimir</mods:namePart>
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<mods:namePart id="0D48EC972E0517946513910DCA150078">Saboori, Alireza</mods:namePart>
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<mods:namePart id="7C5B1BFF2143CCF6FDAB09A7B8EAE4B8">Asadi, Mahdieh</mods:namePart>
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<mods:namePart id="9722084CB8F23DFA48D97DBA3E07264C">Vafaei, Reza</mods:namePart>
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<mods:date id="54A2A5CD45A1FC60382250E87A03A3D5">2004</mods:date>
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<treatment id="03FB7D20DD61FF83FEAFFED04ED10034" ID-DOI="http://doi.org/10.5281/zenodo.5689548" ID-GBIF-Taxon="119601534" ID-Zenodo-Dep="5689548" LSID="urn:lsid:plazi:treatment:03FB7D20DD61FF83FEAFFED04ED10034" httpUri="http://treatment.plazi.org/id/03FB7D20DD61FF83FEAFFED04ED10034" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="C3489FBDDD61FF81FEAFFED04BA2053B" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BEDCC36DD61FF81FEAFFED048910557" blockId="21.[264,661,283,345]" box="[264,661,283,310]" pageId="21" pageNumber="22">
<heading id="D0A57B5ADD61FF81FEAFFED048910557" bold="true" box="[264,661,283,310]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="B9261024DD61FF81FEAFFED048910557" bold="true" box="[264,661,283,310]" pageId="21" pageNumber="22">
<taxonomicName id="4C52B7B5DD61FF81FEAFFED048340557" authority="Pesic" authorityName="Pesic" box="[264,560,283,310]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD61FF81FEAFFED04BED0554" bold="true" box="[264,489,284,310]" italics="true" pageId="21" pageNumber="22">Atractides nikooae</emphasis>
Pesic
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD61FF81FD9FFED748910557" box="[568,661,283,309]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BEDCC36DD61FF81FEAFFEF34BA2053B" blockId="21.[264,661,283,345]" box="[264,422,319,345]" pageId="21" pageNumber="22">
(
<figureCitation id="1369D0B3DD61FF81FEB6FEF34B9A053B" box="[273,414,319,345]" captionStart-0="FIGURES 34 37" captionStart-1="FIGURES 38 40" captionStartId-0="20.[264,382,1799,1823]" captionStartId-1="22.[264,382,1461,1485]" captionTargetBox-0="[296,1288,362,1752]" captionTargetBox-1="[293,1259,634,1426]" captionTargetId-0="figure@20.[274,1314,329,1778]" captionTargetId-1="figure@22.[287,1300,609,1434]" captionTargetPageId-0="20" captionTargetPageId-1="22" captionText-0="FIGURES 34 37. Atractides nikooae sp. nov., male: 34 = idiosoma, ventral view; 35 = genital field; 36 = palp; 37 = I­L­ 5 / 6. Scale bars = 0.1 mm." captionText-1="FIGURES 38 40. Atractides nikooae sp. nov., female: 38 = I­L­ 5 / 6; 39 = palp; 40 = genital field. Scale bars = 0.1 mm." httpUri-0="https://zenodo.org/record/157864/files/figure.png" httpUri-1="https://zenodo.org/record/157865/files/figure.png" pageId="21" pageNumber="22">Figs. 3440</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD61FF81FEAFFE4348E40643" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BEDCC36DD61FF81FEAFFE4348E40643" blockId="21.[264,1325,399,1905]" pageId="21" pageNumber="22">
<emphasis id="B9261024DD61FF81FEAFFE434BB605CA" box="[264,434,399,424]" italics="true" pageId="21" pageNumber="22">
<typeStatus id="54E97294DD61FF81FEAFFE434B3B05CA" box="[264,319,399,424]" pageId="21" pageNumber="22">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="54E97294DD61FF81FE6DFE43483805CB" box="[458,572,399,425]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
: male, dissected and mounted in Hoyer's fluid;
<collectingCountry id="F3458CA6DD61FF81FB0EFE434ED905CB" box="[1193,1245,399,425]" name="Iran" pageId="21" pageNumber="22">Iran</collectingCountry>
: IR60 Markazi Province, Ghareh Chai stream
<quantity id="4CAA61D3DD61FF81FCA4FE7B495305B3" box="[771,855,439,465]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="22" unit="km" value="30.0">30 km</quantity>
after Tafresh city (
<quantity id="4CAA61D3DD61FF81FBF1FE7B4EAF05B3" box="[1110,1195,439,465]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="21" pageNumber="22" unit="km" value="90.0">90 km</quantity>
N Arak),
<date id="FFECEAF6DD61FF81FEAFFE134B8A059B" box="[264,398,479,505]" pageId="21" pageNumber="22" value="2003-08-24">24.08.2003</date>
, leg. Pesic.
<typeStatus id="54E97294DD61FF81FDBAFE134890059B" box="[541,660,479,505]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5407BFDD61FF81FD07FE134911059B" box="[672,789,479,505]" pageId="21" pageNumber="22" type="female">7 females</specimenCount>
, same data as
<typeStatus id="54E97294DD61FF81FC66FE134E2F059B" box="[961,1067,479,505]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
, one of these females dissected and mounted on Hoyer's fluid.
</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD61FF81FE9FFDE34E370773" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BEDCC36DD61FF81FE9FFDE34E370773" blockId="21.[264,1325,399,1905]" pageId="21" pageNumber="22">
<emphasis id="B9261024DD61FF81FE9FFDE34BAB062A" box="[312,431,559,584]" italics="true" pageId="21" pageNumber="22">Diagnosis</emphasis>
: dorsal integument: striated; muscle attachments: unsclerotized. I­L: I­L­5 S­1­2 distanced, S­1 slender, blunt, S­2 basally thickened, more pointed; I­L­6 slender, curved, with maximum H proximally. Genital field: Ac small, in a curved line; excretory pore: smooth; Vgl­1: not fused to Vgl­2. Palp: no sexual dimorphism, ventral margin P­2 straight, with slightly curved distal edge, P­3 straight, ventral margin of P­4 divided by hair insertions in sections 1:1:2, sword seta near the distoventral hair.
</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD61FF83FE9FFCD34B47053C" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
<paragraph id="8BEDCC36DD61FF81FE9FFCD34BC6075A" blockId="21.[264,1325,399,1905]" box="[312,450,799,824]" pageId="21" pageNumber="22">
<emphasis id="B9261024DD61FF81FE9FFCD34BC6075A" box="[312,450,799,824]" italics="true" pageId="21" pageNumber="22">Description</emphasis>
</paragraph>
<paragraph id="8BEDCC36DD61FF81FE9FFC8B499E0123" blockId="21.[264,1325,399,1905]" pageId="21" pageNumber="22">
<emphasis id="B9261024DD61FF81FE9FFC8B4B700702" box="[312,372,839,864]" italics="true" pageId="21" pageNumber="22">Male</emphasis>
: Length of idiosoma 514, width 357. Dorsally the integument is striated; muscle attachments: unsclerotized. Coxal field (
<figureCitation id="1369D0B3DD61FF81FD4BFCA3494307EB" box="[748,839,879,905]" captionStart="FIGURES 34 37" captionStartId="20.[264,382,1799,1823]" captionTargetBox="[296,1288,362,1752]" captionTargetId="figure@20.[274,1314,329,1778]" captionTargetPageId="20" captionText="FIGURES 34 37. Atractides nikooae sp. nov., male: 34 = idiosoma, ventral view; 35 = genital field; 36 = palp; 37 = I­L­ 5 / 6. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157864/files/figure.png" pageId="21" pageNumber="22">Fig. 34</figureCitation>
): mediocaudal margin of Cx­1+2 with a large, slightly concave area between the laterally directed apodemes of Cx­2 (
<figureCitation id="1369D0B3DD61FF81FB66FC5B4F1807D3" box="[1217,1308,919,945]" captionStart="FIGURES 34 37" captionStartId="20.[264,382,1799,1823]" captionTargetBox="[296,1288,362,1752]" captionTargetId="figure@20.[274,1314,329,1778]" captionTargetPageId="20" captionText="FIGURES 34 37. Atractides nikooae sp. nov., male: 34 = idiosoma, ventral view; 35 = genital field; 36 = palp; 37 = I­L­ 5 / 6. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157864/files/figure.png" pageId="21" pageNumber="22">Fig. 34</figureCitation>
); length between anterior end of first coxae and posterior end of fourth coxae 339; Cx­3 width 385, Cx­1+2 width 304; Cx­1+2 medial suture
<quantity id="4CAA61D3DD61FF81FC25FC2B49C90063" box="[898,973,999,1025]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="22" unit="in" value="100.0">100 in</quantity>
length; distance from lateralmost tips of Cx­2 to the medioposterior edge of Cx­1+2 212. Measurements of mouthparts: cheliceral claw
<quantity id="4CAA61D3DD61FF81FDACFBFB48430033" box="[523,583,1079,1105]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="21" pageNumber="22" unit="in" value="56.0">56 in</quantity>
length; palp total length 318, dorsal length/height and relative length (% total length, in parentheses) of single segments: P­
<date id="FFECEAF6DD61FF81FBA5FB934E5F001B" box="[1026,1115,1119,1145]" pageId="21" pageNumber="22" value="1929-01-31">1 31/29</date>
(9.7), P­2 71/54 (22.3), P­3 75/46 (23.6), P­4 104/39 (32.7), P­
<date id="FFECEAF6DD61FF81FC8AFB4B498100C3" box="[813,901,1159,1185]" pageId="21" pageNumber="22" value="1937-05-12">5 37/12</date>
(11.6); length ratio P­2/P­4 is 0.68; palp (
<figureCitation id="1369D0B3DD61FF81FEEDFB634BA600AB" box="[330,418,1199,1225]" captionStart="FIGURES 34 37" captionStartId="20.[264,382,1799,1823]" captionTargetBox="[296,1288,362,1752]" captionTargetId="figure@20.[274,1314,329,1778]" captionTargetPageId="20" captionText="FIGURES 34 37. Atractides nikooae sp. nov., male: 34 = idiosoma, ventral view; 35 = genital field; 36 = palp; 37 = I­L­ 5 / 6. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157864/files/figure.png" pageId="21" pageNumber="22">Fig. 36</figureCitation>
): ventral margin P­2 straight, with slightly curved distal edge, P­3 straight, P­ 4 1:1:2, sword seta near the distoventral hair. Genital field (
<figureCitation id="1369D0B3DD61FF81FC79FB1B4E3D0093" box="[990,1081,1239,1265]" captionStart="FIGURES 34 37" captionStartId="20.[264,382,1799,1823]" captionTargetBox="[296,1288,362,1752]" captionTargetId="figure@20.[274,1314,329,1778]" captionTargetPageId="20" captionText="FIGURES 34 37. Atractides nikooae sp. nov., male: 34 = idiosoma, ventral view; 35 = genital field; 36 = palp; 37 = I­L­ 5 / 6. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157864/files/figure.png" pageId="21" pageNumber="22">Fig. 35</figureCitation>
): apple­shaped with nearly straight, irregular anterior margin;
<quantity id="4CAA61D3DD61FF81FD50FB334930017B" box="[759,820,1279,1305]" metricMagnitude="0" metricUnit="m" metricValue="2.3876" pageId="21" pageNumber="22" unit="in" value="94.0">94 in</quantity>
length,
<quantity id="4CAA61D3DD61FF81FC31FB3349D7017B" box="[918,979,1279,1305]" metricMagnitude="0" metricUnit="m" metricValue="2.4384" pageId="21" pageNumber="22" unit="in" value="96.0">96 in</quantity>
width, Ac­1­3 length 21­23­ 27; excretory pore smooth; Vgl­1: separated from Vgl­2
</paragraph>
<paragraph id="8BEDCC36DD61FF81FE9FFA834EE50253" blockId="21.[264,1325,399,1905]" pageId="21" pageNumber="22">
I­L (
<figureCitation id="1369D0B3DD61FF81FEC9FA834BC2010B" box="[366,454,1359,1385]" captionStart="FIGURES 34 37" captionStartId="20.[264,382,1799,1823]" captionTargetBox="[296,1288,362,1752]" captionTargetId="figure@20.[274,1314,329,1778]" captionTargetPageId="20" captionText="FIGURES 34 37. Atractides nikooae sp. nov., male: 34 = idiosoma, ventral view; 35 = genital field; 36 = palp; 37 = I­L­ 5 / 6. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157864/files/figure.png" pageId="21" pageNumber="22">Fig. 37</figureCitation>
): S­1/2heteromorphic and distanced, S­1 long, S­2 shortened, enlarged; I­ L­6 long and slender; I­L­5 dorsal length 196, I­L­5 ventral length 115, ratio dorsal length I­L­5 /ventral length 1.7, I­L­5 central height 50, ratio dorsal length I­L­5/central height 3.92, S­1 length 100, ratio length S­1/width 11.6, S­2 length 71, ratio length S­2/ width 6.2, distance between sword setae of I­L­5 30; ratio length S­1/2 1.41; I­L­6 length 152, I­ L­6 central height 18, ratio length I­L­6/central height 8.3; ratio length I­L­5/6 1.23.
</paragraph>
<paragraph id="8BEDCC36DD61FF82FE9FF9F34B19059C" blockId="21.[264,1325,399,1905]" lastBlockId="22.[264,1324,284,510]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B9261024DD61FF81FE9FF9F34B94023A" box="[312,400,1599,1624]" italics="true" pageId="21" pageNumber="22">Female</emphasis>
: Length of idiosoma 892, width 650. Dorsally the integument is striated; muscle attachments: unsclerotized. Coxal field: mediocaudal margin of Cx­1+2 with a large, slightly concave area between the laterally directed apodemes of Cx­2; length between anterior end of first coxae and posterior end of fourth coxae 400, Cx­3 width 511, Cx­1+2 width 365 Cx­1+2 medial suture
<quantity id="4CAA61D3DD61FF81FD32F91348E5029B" box="[661,737,1759,1785]" metricMagnitude="0" metricUnit="m" metricValue="3.429" pageId="21" pageNumber="22" unit="in" value="135.0">135 in</quantity>
length; distance from lateralmost tips of Cx­2 to the medioposterior edge of Cx­1+2 260. Measurements of mouthparts: chelicera
<quantity id="4CAA61D3DD61FF81FB7AF8CB4F2F0343" box="[1245,1323,1799,1825]" metricMagnitude="0" metricUnit="m" metricValue="6.35" pageId="21" pageNumber="22" unit="in" value="250.0">250 in</quantity>
length, claw
<quantity id="4CAA61D3DD61FF81FE39F8E34BDD032B" box="[414,473,1839,1865]" metricMagnitude="0" metricUnit="m" metricValue="1.905" pageId="21" pageNumber="22" unit="in" value="75.0">75 in</quantity>
length; palp total length 408, dorsal length/height and relative length (% total length, in parentheses) of single segments: P­1 40/40 (9.8), P­2 91/69 (22.3), P­3 112/ 52 (27.5), P­4 125/34 (30.6), P­
<date id="FFECEAF6DD62FF82FD25FED048DE0554" box="[642,730,284,310]" pageId="22" pageNumber="23" value="1940-05-15">5 40/15</date>
(9.8); length ratio P­2/P­4 is 0.73; palp (
<figureCitation id="1369D0B3DD62FF82FB64FED04F190554" box="[1219,1309,284,310]" captionStart="FIGURES 38 40" captionStartId="22.[264,382,1461,1485]" captionTargetBox="[293,1259,634,1426]" captionTargetId="figure@22.[287,1300,609,1434]" captionTargetPageId="22" captionText="FIGURES 38 40. Atractides nikooae sp. nov., female: 38 = I­L­ 5 / 6; 39 = palp; 40 = genital field. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157865/files/figure.png" pageId="22" pageNumber="23">Fig. 39</figureCitation>
): ventral margin P­2 straight, with slightly curved distal edge, P­4 more slender than in male, 1:1:2, sword seta near the distoventral hair. Genital field (
<figureCitation id="1369D0B3DD62FF82FBAFFEA04E6605E4" box="[1032,1122,364,390]" captionStart="FIGURES 38 40" captionStartId="22.[264,382,1461,1485]" captionTargetBox="[293,1259,634,1426]" captionTargetId="figure@22.[287,1300,609,1434]" captionTargetPageId="22" captionText="FIGURES 38 40. Atractides nikooae sp. nov., female: 38 = I­L­ 5 / 6; 39 = palp; 40 = genital field. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157865/files/figure.png" pageId="22" pageNumber="23">Fig. 40</figureCitation>
): Ac in a curved line, genital plates with weak indentations at borders Ac­1/2 and Ac­2/3;
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length,
<quantity id="4CAA61D3DD62FF82FEAFFE704B5705B4" box="[264,339,444,470]" metricMagnitude="0" metricUnit="m" metricValue="4.4958" pageId="22" pageNumber="23" unit="in" value="177.0">177 in</quantity>
width, Ac­1­3 length 25­29­36; excretory pore smooth; Vgl­1: separated from Vgl­ 2.
</paragraph>
<caption id="DF2D9CBEDD62FF82FEAFFA794BEE018E" httpUri="https://zenodo.org/record/157865/files/figure.png" pageId="22" pageNumber="23" targetBox="[293,1259,634,1426]" targetPageId="22">
<paragraph id="8BEDCC36DD62FF82FEAFFA794BEE018E" blockId="22.[264,1323,1461,1516]" pageId="22" pageNumber="23">
<emphasis id="B9261024DD62FF82FEAFFA794BCB01AF" bold="true" box="[264,463,1461,1485]" pageId="22" pageNumber="23">FIGURES 3840.</emphasis>
<taxonomicName id="4C52B7B5DD62FF82FE71FA79489B01AE" box="[470,671,1461,1484]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD62FF82FE71FA79489B01AE" box="[470,671,1461,1484]" italics="true" pageId="22" pageNumber="23">Atractides nikooae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD62FF82FD01FA7948FF01AE" box="[678,763,1461,1484]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, female: 38 = I­L­5/6; 39 = palp; 40 = genital field. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEDCC36DD62FF83FE9FF9A14B47053C" blockId="22.[264,1324,1645,1871]" lastBlockId="23.[264,1323,284,1110]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
I­L (
<figureCitation id="1369D0B3DD62FF82FECAF9A14BC102E5" box="[365,453,1645,1671]" captionStart="FIGURES 38 40" captionStartId="22.[264,382,1461,1485]" captionTargetBox="[293,1259,634,1426]" captionTargetId="figure@22.[287,1300,609,1434]" captionTargetPageId="22" captionText="FIGURES 38 40. Atractides nikooae sp. nov., female: 38 = I­L­ 5 / 6; 39 = palp; 40 = genital field. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/157865/files/figure.png" pageId="22" pageNumber="23">Fig. 38</figureCitation>
): I­L­5 both S­1/2 not enlarged distally, S­2 strongly thickened in the basal third part, distance S­1­2 large (&gt; 50 µm); I­L­6 curved, maximum H basally, strongly narrowed in the centre (HB); I­L­5 dorsal length 252, I­L­5 ventral length 135, ratio dorsal length I­L­5 /ventral length 1.89, I­L­5 central height 77, ratio dorsal length I­L­5/central height 3.3, S­1 length 125, ratio length S­1/width 10.2, S­2 length 83, ratio length S­2/ width 4.4, distance between sword setae of I­L­5 57; ratio length S­1/2 1.5; I­L­6 length 198, I­L­6 central height 19, ratio length I­L­6/central height 10.3; ratio length I­L­5/6 1.27.
</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD63FF83FE9FFEA04B4B0064" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BEDCC36DD63FF83FE9FFEA04B8606A4" blockId="23.[264,1323,284,1110]" pageId="23" pageNumber="24">
<emphasis id="B9261024DD63FF83FE9FFEA04BBC05E7" box="[312,440,364,389]" italics="true" pageId="23" pageNumber="24">Discussion</emphasis>
: Among the species with relatively small Ac, excretory pore smooth, I­L­ 5/6 slender,
<taxonomicName id="4C52B7B5DD63FF83FE3FFE58481205CF" box="[408,534,404,429]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD63FF83FE3FFE58481205CF" box="[408,534,404,429]" italics="true" pageId="23" pageNumber="24">A. nikooae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD63FF83FDBAFE58487C05CC" box="[541,632,404,430]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C52B7B5DD63FF83FCF4FE584E9505CC" authority="Szalay, 1935" authorityName="Szalay" authorityYear="1935" box="[851,1169,404,430]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="diastema">
<emphasis id="B9261024DD63FF83FCF4FE5849DA05CF" box="[851,990,404,429]" italics="true" pageId="23" pageNumber="24">A. diastema</emphasis>
(
<bibRefCitation id="EFC3B1C7DD63FF83FC49FE584E8C05CC" author="Szalay" box="[1006,1160,404,430]" pageId="23" pageNumber="40" refString="Szalay, L. (1935) Eine neue Hydracarine aus der Gattung Megapus Neuman und das Weibchen von Megapus (M.) nodipalpis var. fluviatilis Szalay und Megapus (M.) acutirostris Motas. Zoologischer Anzeiger, 111 (11 - 12), 309 - 314." type="journal article" year="1935">Szalay, 1935</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C52B7B5DD63FF83FB39FE58480505B4" authority="Lundblad, 1942" authorityName="Lundblad" authorityYear="1942" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="algeriensis">
<emphasis id="B9261024DD63FF83FB39FE584B2305B7" italics="true" pageId="23" pageNumber="24">A. algeriensis</emphasis>
(
<bibRefCitation id="EFC3B1C7DD63FF83FE9FFE704BFD05B4" author="Lundblad" box="[312,505,444,470]" pageId="23" pageNumber="39" refString="Lundblad, O. (1942) Afrikanische Hydracarinen. Entomologisk Tidskrift, 63 (3 - 4), 155 - 209." type="journal article" year="1942">Lundblad, 1942</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C52B7B5DD63FF83FDE6FE70499905B4" authority="K.Viets, 1930" authorityName="K.Viets" authorityYear="1930" box="[577,925,444,470]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cultellatus">
<emphasis id="B9261024DD63FF83FDE6FE7048D905B7" box="[577,733,444,469]" italics="true" pageId="23" pageNumber="24">A. cultellatus</emphasis>
(
<bibRefCitation id="EFC3B1C7DD63FF83FD48FE70499105B4" author="Viets" box="[751,917,444,470]" pageId="23" pageNumber="40" refString="Viets, K. (1930) Zur Kenntnis der Hydracarinen-Fauna von Spanien. Archiv fur Hydrobiologie, 21 (2), 175 - 240, 21 (3), 359 - 446." type="journal article" year="1930">K.Viets, 1930</bibRefCitation>
)
</taxonomicName>
due to absence of muscle attachment sclerites, striated integument and the Vgl­1 not fused to Vgl­2. From the latter two species,
<taxonomicName id="4C52B7B5DD63FF83FECBFDC04BEF0647" box="[364,491,524,549]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD63FF83FECBFDC04BEF0647" box="[364,491,524,549]" italics="true" pageId="23" pageNumber="24">A. nikooae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD63FF83FE54FDC0484B0644" box="[499,591,524,550]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
is easily distinguished by the larger distance S­1­2, while the ventral margin of P­2 is straight, not distally protruding. Furthermore, from
<taxonomicName id="4C52B7B5DD63FF83FB2FFDF84F27062F" box="[1160,1315,564,589]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cultellatus">
<emphasis id="B9261024DD63FF83FB2FFDF84F27062F" box="[1160,1315,564,589]" italics="true" pageId="23" pageNumber="24">A. cultellatus</emphasis>
</taxonomicName>
, a species known only in the female sex,
<taxonomicName id="4C52B7B5DD63FF83FD5FFD90497D0617" box="[760,889,604,629]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD63FF83FD5FFD90497D0617" box="[760,889,604,629]" italics="true" pageId="23" pageNumber="24">A. nikooae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD63FF83FC25FD9049E40614" box="[898,992,604,630]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
differs, in a longer median suture Cx­1+2 (in
<taxonomicName id="4C52B7B5DD63FF83FE4EFD48488306FF" box="[489,647,644,669]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cultellatus">
<emphasis id="B9261024DD63FF83FE4EFD48488306FF" box="[489,647,644,669]" italics="true" pageId="23" pageNumber="24">A. cultellatus</emphasis>
</taxonomicName>
L = 90 µm) and the sword seta on P­4 not particularly thickened.
</paragraph>
<paragraph id="8BEDCC36DD63FF83FE9FFD1849B307D4" blockId="23.[264,1323,284,1110]" pageId="23" pageNumber="24">
<taxonomicName id="4C52B7B5DD63FF83FE9FFD184BB1068F" box="[312,437,724,749]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD63FF83FE9FFD184BB1068F" box="[312,437,724,749]" italics="true" pageId="23" pageNumber="24">A. nikooae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD63FF83FE1AFD18481C068C" box="[445,536,724,750]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C52B7B5DD63FF83FD15FD184939068F" box="[690,829,724,749]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="diastema">
<emphasis id="B9261024DD63FF83FD15FD184939068F" box="[690,829,724,749]" italics="true" pageId="23" pageNumber="24">A. diastema</emphasis>
</taxonomicName>
(in parentheses data from
<bibRefCitation id="EFC3B1C7DD63FF83FBD2FD184F19068C" author="Gerecke" box="[1141,1309,724,750]" pageId="23" pageNumber="39" refString="Gerecke, R. (2003) Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378." type="journal article" year="2003">Gerecke 2003</bibRefCitation>
), a weakly defined species from
<collectingCountry id="F3458CA6DD63FF83FDD0FD3048DA0774" box="[631,734,764,790]" name="Hungary" pageId="23" pageNumber="24">Hungary</collectingCountry>
and
<collectingCountry id="F3458CA6DD63FF83FCBFFD30496D0774" box="[792,873,764,790]" name="Poland" pageId="23" pageNumber="24">Poland</collectingCountry>
known only in the female sex, by the following features: the genital plates are not elongated, ventral margin of P­2 is straight (ventral marginof P­2 slightly protruding), ventral margin of P­4 divided by hair insertions in sections 1:1:2 (2:2:
<quantity id="4CAA61D3DD63FF83FDB6FCB8483A07EC" box="[529,574,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="23" pageNumber="24" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C52B7B5DD63FF83FDEFFCB848D107EF" box="[584,725,884,909]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="diastema">
<emphasis id="B9261024DD63FF83FDEFFCB848D107EF" box="[584,725,884,909]" italics="true" pageId="23" pageNumber="24">A. diastema</emphasis>
</taxonomicName>
), more slender palp segments (L/H P­3 2.77, P­4 4.20), and a more heteromorphic setae S­1/2 (L S­1/2 1.3).
</paragraph>
<paragraph id="8BEDCC36DD63FF83FE9FFC084B4B0064" blockId="23.[264,1323,284,1110]" pageId="23" pageNumber="24">Etymology: The species is named after Nikoo Saboori, a daughter of the second author (A.S).</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD63FF83FE9FFBD84E96004C" box="[312,1170,1044,1070]" pageId="23" pageNumber="24" type="biology_ecology">
<paragraph id="8BEDCC36DD63FF83FE9FFBD84E96004C" blockId="23.[264,1323,284,1110]" box="[312,1170,1044,1070]" pageId="23" pageNumber="24">
<emphasis id="B9261024DD63FF83FE9FFBD84B97004F" box="[312,403,1044,1069]" italics="true" pageId="23" pageNumber="24">Biology</emphasis>
:
<taxonomicName id="4C52B7B5DD63FF83FE05FBD8487B004F" box="[418,639,1044,1069]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nikooae" status="sp. nov.">
<emphasis id="B9261024DD63FF83FE05FBD8487B004F" box="[418,639,1044,1069]" italics="true" pageId="23" pageNumber="24">Atractides nikooae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215AD5FDD63FF83FD21FBD848E5004C" box="[646,737,1044,1070]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
is very likely a rhitrobiontic species.
</paragraph>
</subSubSection>
<subSubSection id="C3489FBDDD63FF83FE9FFBF04ED10034" box="[312,1237,1084,1110]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BEDCC36DD63FF83FE9FFBF04ED10034" blockId="23.[264,1323,284,1110]" box="[312,1237,1084,1110]" pageId="23" pageNumber="24">
<emphasis id="B9261024DD63FF83FE9FFBF04BD50037" box="[312,465,1084,1109]" italics="true" pageId="23" pageNumber="24">Distribution:</emphasis>
<collectingCountry id="F3458CA6DD63FF83FE7FFBF0480E0034" box="[472,522,1084,1110]" name="Iran" pageId="23" pageNumber="24">Iran</collectingCountry>
, only known from the
<emphasis id="B9261024DD63FF83FCB3FBF049A80037" box="[788,940,1084,1109]" italics="true" pageId="23" pageNumber="24">locus typicus</emphasis>
in the Markazi Province.
</paragraph>
</subSubSection>
</treatment>
</document>